ANNOTATED CHECKLIST of the VASCULAR PLANTS of SANTA CRUZ COUNTY, CALIFORNIA SECOND EDITION Dylan Neubauer Artwork by Tim Hyland & Maps by Ben Pease CALIFORNIA NATIVE PLANT SOCIETY, SANTA CRUZ COUNTY CHAPTER Copyright © 2013 by Dylan Neubauer All rights reserved. No part of this publication may be reproduced without written permission from the author. Design & Production by Dylan Neubauer Artwork by Tim Hyland Maps by Ben Pease, Pease Press Cartography (peasepress.com) Cover photos (Eschscholzia californica & Big Willow Gulch, Swanton) by Dylan Neubauer California Native Plant Society Santa Cruz County Chapter P.O. Box 1622 Santa Cruz, CA 95061 To order, please go to www.cruzcps.org For other correspondence, write to Dylan Neubauer [email protected] ISBN: 978-0-615-85493-9 Printed on recycled paper by Community Printers, Santa Cruz, CA For Tim Forsell, who appreciates the tiny ones ... Nobody sees a flower, really— it is so small— we haven’t time, and to see takes time, like to have a friend takes time. —GEORGIA O’KEEFFE CONTENTS ~ u Acknowledgments / 1 u Santa Cruz County Map / 2–3 u Introduction / 4 u Checklist Conventions / 8 u Floristic Regions Map / 12 u Checklist Format, Checklist Symbols, & Region Codes / 13 u Checklist Lycophytes / 14 Ferns / 14 Gymnosperms / 15 Nymphaeales / 16 Magnoliids / 16 Ceratophyllales / 16 Eudicots / 16 Monocots / 61 u Appendices 1. Listed Taxa / 76 2. Endemic Taxa / 78 3. Taxa Extirpated in County / 79 4. Taxa Not Currently Recognized / 80 5. Undescribed Taxa / 82 6. Most Invasive Non-native Taxa / 83 7. Rejected Taxa / 84 8. Notes / 86 u References / 152 u Index to Families & Genera / 154 u Floristic Regions Map with USGS Quad Overlay / 166 “True science teaches, above all, to doubt and be ignorant.” —MIGUEL DE UNAMUNO 1 ~ACKNOWLEDGMENTS ~ ANY THANKS TO THE GENEROUS DONORS without whom this publication would not M have been possible—and to the numerous individuals, organizations, insti- tutions, and agencies that so willingly gave of their time and expertise. D O N O R S LUPINE MANZANITA LIVE OAK (UNDER $100) ($100 OR MORE) ($1,000 OR MORE) Avis Boutell Anonymous Linda Brodman Valerie Haley Julie Anne Hopkins REDWOOD Nicole Nedeff Dr. and Mrs. Brian Keelan ($10,000 OR MORE) Peter Slattery Ken Moore Ellen Uhler Heath Bartosh Ken Moore Linda Brodman Randall Morgan Kevin Bryant Dr. John B. Nelson Debbie Bulger Dr. Guy Nesom Eva Buxton Dr. Kevin Nixon Toni Corelli Ben Pease Taylor Crow Scott Peden Dr. Ellen Dean John Rawlings Dr. Richard Dodd Dr. Matt Ritter Deanna Giuliano Austin Robey Valerie Haley Dr. Andrew Sanders Brett Hall Suzanne Schettler Dr. Grey Hayes Kayla Schoos Tim Hyland Aaron Sims Tim Kask Daniel Slakey Dr. Brian Keelan Dr. David Styer Dr. David Keil Dr. Dean W. Taylor Al Keuter Ellen Uhler Neal Kramer James A. West Dr. Job Kujit Linda Willis Chris Lay Ray Wootten Stephen McCabe Vern Yadon Dr. Abigail Moore Dr. Jenn Yost Calflora California Academy of Sciences California Invasive Plant Council (Cal-IPC) California Native Plant Society (CNPS), Santa Cruz County Chapter California Natural Diversity Database (CNDDB) Calphotos CNPS Rare Plant Program CNPS Santa Cruz County Chapter Flora Committee Consortium of California Herbaria (CCH) Jepson Herbarium UCSC Arboretum; UCSC Museum of Natural History U.S. Fish and Wildlife Service 2 Santa Cruz County, CA Pe sca 35 85 der o Saratoga Saratoga Gap SKYLINE Creek 9 Butano Y T N 9 17 BL U S Creek Castle Rock VD O A C State Park S N Castle A S A N M A T E O N Los Gatos Rock T T Creek San A A Gazos 9 Big Basin 236 C CHINA GRADE 35 Lexington Redwoods Kings L A Reservoir State Park Lorenzo R A Creek CHALKS ek e r SU C M M Chalk IT Little Mtn Fork R Basin D C Eagle Rock Creek Creek R Pine Mtn CHALKS U East EMPIRE JAMISON Z CREEK RD Boulder S Bear U GRADE Creek M Newell M IT See Detail River RD Waddell Loch D C O R Lomond U N Scott E T Creek T Y N YA A DR Z . D Mill Creek E O 17 Rancho O Ben Lomond W N del Oso Big RD M E RD L G Soquel Demonstration O SW Greyhound Rock A Creek N Fall Creek San State Forest U T Creek O FLAT HILL RD State Park VINE N N T 1 Vicente RD PINEBonny Doon A Ecological 9 I San Bonny Creek EMPIRE N Doon Reserve GLEN RDLAUREL Creek S Felton Lorenzo Molino Scotts Scott Creek Beach D R Creek Valley RD Coast Dairies (BLM) E E SMITH S GRADE D no public access T N O R R J O O Henry Cowell O P F D I H N A GRANITE CREEK RD CREEK GRANITE Y Redwoods C C S Coast Dairies L A N Creek N Soquel A - U N GRADE State Park R L State Beach O B G l E C B l River k U e e O d Q d a e E I i un r UCSC D O L De Laveaga S g O C Davenport a Creek F L Park R Pogonip I nW ilder Ranch Soquel wi C d State Park 1 Majors l a Sand Hill Bluff B Arana Moore Capitola O Santa Cruz Greenbelt New Creek Brighton C Preserve E State A Beach N Lighthouse Twin Lakes Twin Bridges State Beach State Beach Field State Beach R E N T E Y Parks M O B A Y Other Public Lands Sandhills 0 5 Miles 0 5 Kilometers 3 9 San Jose Detail Loch Lomond Boulder San EMPIRE Recreation Area Creek Loch ANTE RD Y Lomond 85 9 Lorenzo Lompico E. ZA GRADE 17 S ALBA RD A Quail Hollow Ranch Ben Lomond N RD County Park T Los Gatos Creek A Fall Creek Olympia FLAT South State Park Watershed 17 Ridge limited access Vicente TO FEL N Lexington PINE EM P Reservoir Bonny IRE Scotts San Bonny Doon RD Doon Ecological Valley EMPIRE Reserve Felton River Camp Evers (site) ON O D SMITH GRADE Y RD 1 Mile C N Henry Cowell 0 N R O Redwoods U B GRADE State Park 0 1 Kilometer Z S U Marshall M Field M IT RD 9 De Laveaga C O Wilder Ranch UCSC Park U N Loma T Y Prieta 1 State Park Pogonip 1 H Arana IG RD HL M Moore Greenbelt AN D O Creek Soquel Demonstration W State Forest AY U Santa Cruz N 101 Twin Lakes Forest of T EUREKA A State Beach Nisene Marks BUZZARD I State Park N GLEN RDLAUREL Creek S CANYON k e L e A r G C O E O S D N O R RD J k HA H e Z N RD Corallitos E e L A C r S RD. L D Soquel - C E U L L L G Aptos Mt. Madonna E R U 152 O D E Q ALLEY D O V RD Gilroy O S R ALLEY V Valencia RD. Soquel HAMES BROWNS BLVD GREEN Aptos OM D E Capitola E New R F F R E Brighton E D Pinto Lake 152 O State M College PAJARO HILLS Beach B Lake Seacliff LAR LV Creek SAN KIN V D AL State Beach LE Twin Lakes Y State Beach TRABING RDRD Kelly 1 DR Lake ANDREAS AIRPORTRD Lake Tynan LAKE AVE Manresa BUENA VISTA MAIN ST E R E Watsonville 129 Soda N T Y State Beach M O B Watsonville RIVERSIDE DR Lake A Y Wetlands M O N T E R RD. E Y Harkins Sl C O U Sl N T e Y River ill v RD n Pajaro o ts Sunset a State Beach W BEACH PEASE PRESS CARTOGRAPHY www.peasepress.com 4 INTRODUCTION “There are no fixtures in nature. The universe is fluid and volatile.”—RALPH WALDO EMERSON OR THOSE WHO GRUMBLE AND GROAN about the recent changes in botanical nomencla- Fture, it is good to be reminded that this has been an ongoing trend. Indeed, a quick perusal of naturalist Dr. Charles Lewis Anderson’s 1892 “Catalogue of Flowering Plants and Ferns of Santa Cruz County” and “List and Notes of Native and Other Grasses Found Growing Wild in Santa Cruz County” yields many strange names for familiar plants. Those lists constitute the very first inventory of our local flora. The second major treatment was John Hunter Thomas’sFlora of the Santa Cruz Mountains of California, published in 1961. His work—which includes San Francisco, San Mateo, and western Santa Clara counties, as well as Santa Cruz County—has remained the principal botanical reference for the area. The first edition of this Checklist was published in 2005 by the Santa Cruz County Chapter of the California Native Plant Society (CNPS)—the result of six years of data-gathering by an ad hoc Flora Committee headed by Randall Morgan. The main impetus for the creation of this new edition was the 2012 publication of The Jepson Manual: Vascular Plants of California, Second Edition (TJM2)—a massive revision of the information presented in The Jepson Manual: Higher Plants of Cali- fornia (TJM1), published in 1993. With some exceptions, this Checklist follows the nomenclature of TJM2 (and of its online, periodically updated version, the Jepson eFlora) and is to be used in tandem with the Manual, along with the Jepson Online Interchange for California Floristics. The Checklist is targeted toward an audience already familiar with botanical concepts and terminology. Its purpose is to provide the current nomenclature, distribution, and rarity information for all taxa (= species, subspecies, and variet- ies) known to occur in Santa Cruz County, including those that have been more recently documented and those that still remain unpublished.