
<p><strong>Phytohormones </strong></p><p>The growth and development of a plant are influenced by: </p><p>•ꢀGeneꢀc factors •ꢀExternal environmental factors •ꢀChemical hormones inside the plant </p><p>Secondary messengers 1.ꢀ Involve in the transfer informaꢀon from sources to targets </p><p>2.ꢀ Amplify the signal produced by the phytohormone </p><p><strong>Phytohormones </strong></p><p>Plant hormones are organic compounds that are effecꢀve at very low concentraꢀon </p><p>(1g 20,000 tons<sup style="top: -0.45em;">-1</sup>) </p><p>They interact with specific target ꢀssues to cause physiological responses <br>•ꢀGrowth •ꢀFruit ripening </p><p><strong>Phytohormones </strong></p><p>•ꢀHormones sꢀmulate or inhibit plant growth Major groups of hormones: </p><p>1.ꢀ Auxins 2.ꢀ Gibberellins 3.ꢀ Ethylene 4.ꢀ Cytokinins 5.ꢀ Abscisic acid </p><p>6.ꢀ Brassinostereoids 7.ꢀ Salicylic acid 8.ꢀ Polyaminas 9.ꢀ Jasmonates 10.ꢀSystemin 11.ꢀNitric oxide </p><p><em>Arabidopsis thaliana </em></p><p><strong>Phytohormones </strong></p><p>EARLY EXPERIMENTS ON PHOTROPISM SHOWED THAT A STIMULUS (LIGHT) RELEASED CHEMICALS THAT INFLUENCED GROWTH </p><p><strong>Auxins </strong></p><p>Auxin causes several responses in plants: <br>* Phototropism * Geotropism * Promoꢀon of apical dominance * Flower formaꢀon * Fruit set and growth * Formaꢀon of advenꢀꢀous roots * Differenꢀaꢀon of vascular ꢀssues <br>(<em>de novo </em>or repairing existent vascular ꢀssue) </p><p><strong>Auxins </strong></p><p>Addiꢀon of auxins produce <strong>parthenocarpic </strong>fruit. </p><p><strong>Stenospermocarpy </strong></p><p>Biological mechanism that produces seedlessness in some fruits (many table grapes, watermelon) </p><p>diploid + tetraploid parent = triploid seeds </p><p><strong>vegetaꢀve parthenocarpy </strong></p><p>Plants that do not require pollinaꢀon or other sꢀmulaꢀon to produce parthenocarpic fruit (cucumber) </p><p><strong>Auxins </strong></p><p>Syntheꢀc auxins <br>Widely used in agriculture and horꢀculture <br>•ꢀ prevent leaf abscission •ꢀ prevent fruit drop •ꢀ promote flowering and fruiꢀng •ꢀ control weeds </p><p>Agent Orange - 1:1 raꢀo of 2,4-D and 2,4,5-T Dioxin usually contaminates 2,4,5-T, which is linked to miscarriages, birth defects, leukemia, and other types of cancer. </p><p><strong>Cytokinins </strong></p><p>In 1964, zeaꢀn was the first naturally occurring cytokinin isolated from corn. </p><p>Zeaꢀn and zeaꢀn riboside are found in coconut milk. </p><p>Natural </p><p>Syntheꢀc </p><p><strong>Cytokinins </strong></p><p><strong>Lateral bud development </strong><br>•ꢀ<strong>shoot iniꢀaꢀon </strong>•ꢀ<strong>roots </strong>•ꢀ<strong>seeds </strong><br><strong>Callus development <[auxin] = roots <[kineꢀn] = buds </strong><br><strong>Move non-polarly in xylem, phloem, and parenchyma cells. </strong></p><p>•ꢀ<strong>fruits </strong>•ꢀ<strong>young leaves </strong>•ꢀ<strong>chloroplast development </strong></p><p><strong>Ethylene </strong></p><p>•ꢀIs found only in the gaseous form •ꢀInduces ripening (e.g., green bananas). •ꢀEpinasty (causes leaves to droop in the fall ) •ꢀCauses abscission of fruits and flowers </p><p><strong>Ethylene </strong></p><p>Most produced organic compound 107 million metric tons in 2005 </p><p>Once fruit iniꢀated red coloraꢀon, ethylene release becomes elevated in a linear fashion (without diurnal fluctuaꢀons). </p><p><strong>Abscisic acid (ABA) </strong></p><p>Abscisin is made from carotenoids and moves non-polarly through plant ꢀssue. </p><p>Abscisic acid (ABA) is a general plant-growth inhibitor. </p><p>Induces dormancy Prevents seeds from germinaꢀng </p><p>Viviparous mutants Reduced sensiꢀvity for ABA </p><p><strong>Gibberellins </strong></p><p>In 1930's, Ewiꢀ Kurosawa and colleagues were studying plants suffering from bakanae, or "foolish seedling" disease in rice. <br>•ꢀSynthesized in apical porꢀons of stems and roots </p><p><strong>Thompson seedless grapes (Viꢀs vinifera) </strong></p><p>Can be readily obtained in large quanꢀꢀes from </p><p>fermentaꢀons of the fungus <strong>Gibberella fujikuroi </strong></p><p>•ꢀcell division and elongaꢀon •ꢀbreak seed dormancy •ꢀspeed germinaꢀon </p><p><strong>Gibberellins </strong></p><p><strong>bolꢀng </strong></p><p><strong>External factors and plant growth. Tropic responses </strong></p><p>Tropic responses Direcꢀonal movements by growth in response to a direcꢀonal sꢀmulus </p><p><strong>Phototropism </strong>is a </p><p>response to blue light </p><p><strong>Phytphormone: Auxin Second messenger: Calcium </strong></p><p><strong>Geotropism or gravitropism Phytphormones: cytokinin and auxin </strong></p><p><strong>External factors and plant growth. Thigmotropism </strong></p><p>Thigmotropism is direcꢀonal growth response to contact with an object. </p><p>Greek <em>thigma </em>= touch </p><p>Rapid response Diferenꢀaꢀon in the elongaꢀon of the cells </p><p>tendrils </p><p><strong>External factors and plant growth. Nasꢀc movements and heliotropism </strong></p><p>Nasꢀc movements are independent of the sꢀmulus' posiꢀon </p><p>Touch, mechanical, electrical, thermal or chemical </p><p>Venus Flytrap, <em>Dionaea muscipula. </em>Acid growth (cells that to elongate or expand quickly at low pH) and turgor </p><p><strong>External factors and plant growth. Heliotropism </strong></p><p>Heliotropism is the diurnal moꢀon of plant parts (flowers or leaves) in response to the direcꢀon of the sun. </p><p>During the night, the flowers may assume a random orientaꢀon Heliotropism is a response to blue light. </p><p><strong>External factors and plant growth. Circadian rhythms </strong></p><p>Roughly 24-hour cycle in the biochemical, physiological, or behavioral organisms’ processes </p><p>Hours of light </p><ul style="display: flex;"><li style="flex:1">Day </li><li style="flex:1">Night </li></ul><p></p><p>Endogenous and synchronized by the environment Biological clock </p><p>Photosynthesis Movement </p><p><strong>External factors and plant growth. Photoperiodism </strong></p><p>According to their photoperiods, plants can be classified into three groups: a)ꢀ Short day plants. Examples: <em>Nico8ana tabacum</em>, soybean, strawberry, and chrysanthemum. b) Long day plants. Examples: Onion, carrot and spinach. c)ꢀ Day neutral plants. Examples: Tomatoes, cucumbers, sunflower, dandelions and coꢁon. </p>
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