Hantaviridae 0.7 Filoviridae 0.6 Picornaviridae 0.3 Wenling red spikefish hantavirus Rhinovirus C Ahab virus * Possum enterovirus * Aronnax virus * * Wenling minipizza batfish hantavirus Wenling filefish filovirus Norway rat hunnivirus * Wenling yellow goosefish hantavirus Starbuck virus * * Porcine teschovirus European mole nova virus Human Marburg marburgvirus Mosavirus Asturias virus * * * Tortoise picornavirus Egyptian fruit bat Marburg marburgvirus Banded bullfrog picornavirus * Spanish mole uluguru virus Human Sudan ebolavirus * Black spectacled toad picornavirus * Kilimanjaro virus * * * Crab-eating macaque reston ebolavirus Equine rhinitis A virus Imjin virus * Foot and mouth disease virus Dode virus * Angolan free-tailed bat bombali ebolavirus * * Human cosavirus E Seoul orthohantavirus Little free-tailed bat bombali ebolavirus * African bat icavirus A Tigray hantavirus Human Zaire ebolavirus * Saffold virus * Human choclo virus *Little collared fruit bat ebolavirus Peleg virus * Eastern red scorpionfish picornavirus * Reed vole hantavirus Human bundibugyo ebolavirus * * Isla vista hantavirus * Seal picornavirus Human Tai forest ebolavirus Chicken orivirus Paramyxoviridae 0.4 * Duck picornavirus Hepadnaviridae 0.4 Bildad virus Ned virus Tiger rockfish hepatitis B virus Western African lungfish picornavirus * Pacific spadenose shark paramyxovirus * European eel hepatitis B virus Bluegill picornavirus Nemo virus * Carp picornavirus * African cichlid hepatitis B virus Triplecross lizardfish paramyxovirus * * Fathead minnow picornavirus Avian avulavirus-1 Queequeg virus Pasivirus A1 Teratoscincus roborowskii picornavirus * Avian avulavirus-3 * Sockeye salmon hepatitis B virus * * Mapple virus Bat paramyxovirus Stubb virus Human parechovirus * Human mumps virus Yellow drum hepatitis B virus * Ljungan virus Salmon aquaparamyxovirus * Tashtego virus Boomer virus Gabriel virus * Human respirovirus-3 Australasian snapper hepatitis B virus * * Fleece virus Bovine respirovirus-3 * White sucker hepatitis B virus Yancheng anchovy picornavirus *Swine parainfluenza virus-3 * * Coho salmon hepatitis B virus Beihai pentapodus picornavirus Canine morbillivirus Duck hepatitis B virus Eel picornavirus * * Eastern sea garfish picornavirus Rinderpest virus * Parrot hepatitis B virus * Bat paramyxovirus * * Skink hepatitis B virus Reoviridae 0.3 * Rodent paramyxovirus * * Mount mabu lophuromys virus-1 Spiny lizard hepatitis B virus Aquareovirus C * * Feline paramyxovirus Tibetan frog hepatitis B virus American grass carp reovirus Tailam virus Daggoo virus Human orthoreovirus-2 * Crocodile icefish hepatitis B virus Bat Nelson Bay orthoreovirus 0.5 * * Arenaviridae * Eastern sea garfish hepatitis B virus Avian orthoreovirus Haartman Institute snake virus * Bluegill hepatitis B virus * Mahlapitsi orthoreovirus * Conseil virus * Bush viper reovirus * Eastern bent wing bat hepatitis B virus * * Wenling frogfish arenavirus-1 * Ground squirrel hepatitis B virus Reptilian orthoreovirus * Moby virus * Orangutan hepatitis B virus Guangxi changeable lizard reovirus * Pacific spadenose shark reovirus-1 Wenling frogfish arenavirus-2 * Chimpanzee hepatitis B virus * Boa constrictor kaltenbach virus * * Largemouth bass reovirus Human hepatitis B virus * Human lujo mammarenavirus Ariel virus * Chinook salmon orthoreovirus Mouse Latino mammarenavirus Rhabdoviridae 0.4 * * *Rainbow trout orthoreovirus-3 * Human Argentinian mammarenavirus Paddle-tail newt Fujian dimarhabdovirus *Coho salmon orthoreovirus-3 * Woodrat Real de Catorce virus * Pip virus * * Brown trout orthoreovirus * Bear canyon mammarenavirus Lagos bat lyssavirus * Rat Tamiami mammarenavirus Rabies lyssavirus Astroviridae 0.3 Walkabout Creek virus Wenling blackspotted gurnard astrovirus Flaviviridae 0.7 Keuraliba virus * * Ursula virus Yellow fever virus * Le Dantec virus Ngaingan hapavirus * Spotted paddle tail newt astrovirus * Zika virus * * * * Bovine ephemeral fever virus Zhejiang gunthers frog astrovirus Dengue virus-1 * Beatrice Hill virus * Nanhai ghost shark astrovirus * Lumpfish flavivirus Turbot flatfish rhabdovirus * Wenling banjofish astrovirus * Tamana bat virus * * Perch perhabdovirus Hainan black spectacled toad astrovirus Eastern red scorpionfish flavivirus * Lake trout rhabdovirus * Wenzhou shark flavivirus * Turkey astrovirus * Vesicular stomatitis New Jersey virus * Chicken astrovirus Nanhai dogfish shark pesti-like virus * American bat vesiculovirus * *Xiamen fanray pesti-like virus * Isfahan virus Mouse astrovirus Oriental leaf toed gecko hepacivirus * Carp sprivivirus * Porcine astrovirus * GB virus-B *Grass carp virus * Human astrovirus * *Pike fry sprivivirus * Feline astrovirus * Rodent hepacivirus * Hepatitis C virus *Tench rhabdovirus * Dongbei arctic lamprey astrovirus * Simian pegivirus * Wenling longspine snipefish astrovirus * Matonaviridae 0.3 Western African lungfish hepacivirus * Eastern sea garfish astrovirus Chinese water snake rubivirus * * Ishmael virus * Fedallah virus Yancheng grey stingfish astrovirus Wenling shark hepacivirus * Largetooth flounder astrovirus * * Human rubella virus Guangxi houndshark hepacivirus *Human rubella virus * Wenling rattails astrovirus Xiamen sepia stingray hepacivirus *Human rubella virus Triton virus * Xiamen guitarfish hepacivirus Human rubella virus * Yellow striped sandperch astrovirus * Human rubella virus * Squarepants virus fish mammals Human rubella virus Beihai fish astrovirus Human rubella virus birds, reptiles, vector-borne Human rubella virus Wenling righteye flounder astrovirus amphibians Patrick star virus Eastern red scorpionfish astrovirus novel viruses new in (Geoghegan et al. 2018 Virus Evolution).