161 162 All-Russian Institute of Plant Protection RAAS Справочный список и определитель родов и видов ISSN 1815-3682 хищных мух Dolichopodidae (Diptera) Кавказа и ÂÅÑÒÍÈÊ Восточного Средиземноморья. Гричанов И.Я. Санкт- ÇÀÙÈÒÛ ÐÀÑÒÅÍÈÉ Петербург: ВИЗР РАСХН, 2007, 160 c. (Приложение к Приложение журналу «Вестник защиты растений»). A checklist and keys to Dolichopodidae (Diptera) of the Caucasus and East Mediterranean. Igor Ya. Grichanov. St.Petersburg: VIZR RAAS, 2007, 160 p. (Plant Protection News, Supplement). Supplement Составлен справочный список (518 видов) и определитель 52 родов и 512 видов хищных мух Dolichopodidae (Diptera), известных на Кавказе A checklist and keys to (Азербайджан, Армения, Грузия; Россия: Ростовская область, Краснодар- ский и Ставропольский края, Адыгея, Алания, Дагестан, Кабардино- Dolichopodidae (Diptera) Балкария, Карачаево-Черкессия) и в странах Восточного Средиземноморья (Греция, Египет, Израиль, Ирак, Кипр, Молдавия, Сирия, Турция, Украина). Для каждого вида даны оригинальные родовые комбинации, of the Caucasus and East основные синонимы, глобальное распространение. Во вводном разделе приведены сведения о систематическом положении, морфологии, Mediterranean экологии и практическом значении имаго мух-зеленушек. Работа будет полезна специалистам – энтомологам и экологам, интересующимся энтомофагами, студентам и аспирантам учебных и научных учреждений. Igor Ya. GRICHANOV Рецензент: канд. биол. наук И.В. Шамшев Работа выполнялась в рамках ОНТП Россельхозакадемии (2001-2005, 2006-2010). Рекомендовано к печати редакционной коллегией Всероссийского научно-исследовательского института защиты растений РАСХН 26 октяб- ря 2007 г. Copyright 2007© I.Ya. Grichanov ISSN 1815-3682 St.Petersburg 2007 2 Contents Introduction................................................................................................3 Systematic position of Dolichopodidae........................................................3 Morphology of Dolichopodidae..................................................................5 Ecological notes..........................................................................................9 Basic references........................................................................................10 Checklist of East Mediterranean Dolichopodidae......................................11 Key to East Mediterranean genera of Dolichopodidae...............................74 Keys to East Mediterranean species of Dolichopodidae.............................88 Achalcus & Australachalcus ................................................................88 Acropsilus............................................................................................88 A checklist and keys to Aphrosylus...........................................................................................88 Dolichopodidae (Diptera) of the Caucasus and East Argyra .................................................................................................90 Asyndetus.............................................................................................93 Mediterranean Campsicnemus.....................................................................................93 Chrysotimus (including Guzeriplia)......................................................96 Chrysotus.............................................................................................96 Igor Ya. GRICHANOV Diaphorus............................................................................................98 Dolichopus.........................................................................................101 Epithalassius......................................................................................115 Gymnopternus & Ethiromyia..............................................................115 Hercostomus, Ethiromyia & Poecilobothrus chrysozygos ...................117 All-Russian Institute of Plant Protection, Hydrophorus......................................................................................124 Lamprochromus.................................................................................125 Podbelskogo 3, St.Petersburg-Pushkin, 196608, Russia. Medetera ...........................................................................................126 Email: [email protected] Melanostolus......................................................................................131 Micromorphus ...................................................................................132 Nematoproctus...................................................................................132 Neurigona..........................................................................................132 Abstract Oncopygius........................................................................................134 Peloropeodes .....................................................................................135 Poecilobothrus...................................................................................135 A check list and keys to genera and species of Dolichopodidae Rhaphium ..........................................................................................136 of the Caucasus and East Mediterranean are compiled. 518 species Scellus ...............................................................................................139 and 52 genera known in this region are included in the list, belong- Sciapus ..............................................................................................140 ing to the nine subfamilies. Introductory notes concerning system- Sybistroma & Hercostomus caudatus .................................................144 atic position, ecology and morphology of Dolichopodidae and new Sympycnus .........................................................................................146 species records for some regions are given. New synonyms and a Syntormon..........................................................................................147 new name are proposed. 512 species are included into the keys. Systenus.............................................................................................151 Tachytrechus......................................................................................152 Key words: Diptera, Dolichopodidae, catalogue, keys, Cauca- Teuchophorus ....................................................................................154 Thinophilus........................................................................................154 sus, East Mediterranean. Thrypticus..........................................................................................156 Xanthochlorus....................................................................................157 Acknowledgements.................................................................................158 References..............................................................................................158 3 4 INTRODUCTION sockets)................................................................Cyclorrhapha-Schizophora – No lunule above antennae.............................................................................3 The Dolichopodidae fauna of the world is very large, with approximately 3. Tarsal empodium in the form of pulvilli, i.e., 3 subequal pads below claws..... 7000 described species and 240 genera (Grichanov, 2003-2007). These mostly .........................Stratiomyidae, Xylophagidae, Rhagionidae, Tabanidae et al. predatory flies are distributed throughout the world including the tropics and – Empodium bristle-like or not discernible, at most at most 2 well-developed high-latitude islands and territories. In North Europe adults and larvae of almost pads below claws........................................................................................4 all species of long-legged flies are predators inhabiting moist substrata. Species 4. Basal cells of wing long; vein CuA long, reaching wing margin near A , or of only one genus (Thrypticus) are known to be phytophages living inside stems 2 1 joining A1 close to wing margin (at less than a quarter of its length back of cereal grasses. Most of the numerous species of the cosmopolitan genus from wing margin) ....................................................................................... Medetera are associated with tree trunks, especially in boreal forests, where ................Asilidae, Syrphidae, Bombyliidae, Therevidae, Pipunculidae et al. their larvae are predacious mainly on bark-beetles (Coleoptera). – Second basal cell short, or even confluent with discal cell; anal cell short and East Mediterranean Dolichopodidae have never been generalised in one closed or absent; vein CuA2 absent, reduced, or joining A1 far from wing book. Now 518 species and 52 genera are known in this region, belonging to the margin........................................................................................................5 following nine subfamilies: Achalcinae, Diaphorinae, Dolichopodinae, Hydro- 5. Wing venation without any crossvein in median region of wing ..................... phorinae, Medeterinae, Neurigoninae, Rhaphiinae, Sciapodinae and Sympycni- .............................................................Lonchopteridae, Opetiidae, Phoridae nae. A number of new subfamilies and tribes have been proposed in the second – Venation with at least one crossvein in median region of wing......................6 half of the XX century that means that a revision