Zootaxa 3791 (1): 001–099 ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Monograph ZOOTAXA Copyright © 2014 Magnolia Press ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.3791.1.1 http://zoobank.org/urn:lsid:zoobank.org:pub:1C6E3E9C-E1F4-4A0E-9481-B59B817C8355 ZOOTAXA 3791 The Madagascan endemic myrmicine ants related to Eutetramorium (Hymenoptera: Formicidae): taxonomy of the genera Eutetramorium Emery, Malagidris nom. n., Myrmisaraka gen. n., Royidris gen. n., and Vitsika gen. n. BARRY BOLTON1 & BRIAN L. FISHER2 1 c/o Life Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, U.K. 2 Department of Entomology, California Academy of Sciences, Golden Gate Park, San Francisco, California 94118, U.S.A. Magnolia Press Auckland, New Zealand Accepted by J. Longino: 4 Feb. 2014; published: 24 Apr. 2014 Licensed under a Creative Commons Attribution License http://creativecommons.org/licenses/by/3.0 BARRY BOLTON & BRIAN L. FISHER The Madagascan endemic myrmicine ants related to Eutetramorium (Hymenoptera: Formicidae): taxonomy of the genera Eutetramorium Emery, Malagidris nom. n., Myrmisaraka gen. n., Royidris gen. n., and Vitsika gen. n. (Zootaxa 3791) 99 pp.; 30 cm. 24 Apr. 2014 ISBN 978-1-77557-374-6 (paperback) ISBN 978-1-77557-375-3 (Online edition) FIRST PUBLISHED IN 2014 BY Magnolia Press P.O. Box 41-383 Auckland 1346 New Zealand e-mail: [email protected] http://www.mapress.com/zootaxa/ © 2014 Magnolia Press ISSN 1175-5326 (Print edition) ISSN 1175-5334 (Online edition) 2 · Zootaxa 3791 (1) © 2014 Magnolia Press BOLTON & FISHER Table of Contents Abstract . 4 Introduction . 4 Abbreviations of depositories . 5 Standard measurements and indices . 5 Images . 6 The Eutetramorium genus group. 6 Diagnosis of worker. 6 Comments on Eutetramorium group worker characters . 7 Synopsis of genera of the Eutetramorium group. 8 Key to genera of the Eutetramorium group (workers and queens) . 8 Genera of the Eutetramorium group . 9 Genus Eutetramorium Emery . 9 Diagnosis of worker . 9 Diagnosis of queen (gyne). 9 Diagnosis of male . 9 Comments on Eutetramorium . 10 Synopsis of Eutetramorium species . 10 Key to Eutetramorium species (workers) . 10 Eutetramorium mocquerysi Emery . 10 Eutetramorium monticellii Emery. 12 Eutetramorium parvum Bolton & Fisher . 14 Genus Malagidris . 16 Malagidris Bolton & Fisher . 16 Diagnosis of worker . 16 Diagnosis of queen (gyne). 16 Diagnosis of male . 17 Comments on Malagidris . 17 Synopsis of Malagidris species. 18 Key to Malagidris species (workers) . 18 Provisional key to males of Malagidris species . 18 Malagidris alperti Bolton & Fisher . 19 Malagidris belti (Forel). 21 Malagidris dulcis Bolton & Fisher . 23 Malagidris galokoa Bolton & Fisher . 26 Malagidris jugum Bolton & Fisher . 28 Malagidris sofina Bolton & Fisher . 30 Genus Myrmisaraka . 33 Myrmisaraka Bolton & Fisher . 33 Diagnosis of worker . 33 Diagnosis of queen (gyne). 34 Diagnosis of male . 34 Comments on Myrmisaraka . ..