<p>ESM S3: Significant clustered and dispersed region on each linkage group in the female and male map.</p><p>Region with clustered markers </p><p>Group λij Nij ∑ N.j P Sij Eij Iij ∑ Iij Li Li % Ni Ni% I 0.89 5 11 0.002 ** 104.3 109.7 5.4 9.8 240 4.1 40 27.5 0.73 6 0.000 ** 176.3 180.7 4.4 II 0.18 3 10 0.001 ** 8.1 9.2 1.1 8.7 155.7 5.6 27 37.0 1.26 7 0.000 ** 114.3 121.9 7.6 IV 0.38 6 6 0.000 ** 48.0 50.3 2.3 2.3 139.1 1.7 22 27.3 V 1.06 5 5 0.005 ** 0.0 6.4 6.4 6.4 73.4 8.7 11 45.5 VI 0.36 3 6 0.006 ** 26.0 28.2 2.2 5.7 182.3 3.1 29 20.7 0.58 3 0.021 * 59.7 63.2 3.5 VIII 0.65 3 8 0.028 * 13.7 17.6 3.9 7.2 106.1 6.8 23 34.8 0.55 5 0.000 ** 102.7 106.0 3.3 e l</p><p> a IX 0.00 2 13 0.000 ** 44.4 44.4 0 27.9 85.8 32.5 25 52.0 m</p><p> e 4.61 11 0.008 ** 57.6 85.5 27.9 F X 2.35 8 8 0.003 ** 0.0 14.2 14.2 14.2 135.7 10.5 19 42.1 XI 0.98 4 4 0.018 * 12.3 18.2 5.9 5.9 80.6 7.3 11 36.4 XII 3.31 9 9 0.007 ** 37.0 57.0 20 20 74.4 26.9 17 52.9 XIII 0.31 2 6 0.039 * 36.0 37.9 1.9 5.3 137.6 3.9 24 25.0 0.56 4 0.003 ** 80.9 84.3 3.4 XIV 0.56 3 3 0.019 * 110.2 113.6 3.4 3.4 113.9 3.0 14 21.4 XVI 5.31 15 15 0.000 ** 41.2 73.3 32.1 32.1 86.1 37.3 22 68.2 XVII 0.33 5 7 0.000 ** 29.5 31.5 2 3.8 82 4.6 20 35.0 0.30 2 0.037 * 80.0 81.8 1.8 XVIII 0.89 5 5 0.002 ** 48.1 53.5 5.4 5.4 81.5 6.6 14 35.7 Ic 0.25 3 9 0.002 ** 0.0 2.1 2.1 17.8 103.6 17.2 18 50.0 1.86 6 0.012 * 76.3 92.0 15.7 IIa 1.21 6 6 0.002 ** 91.0 101.2 10.2 10.2 121.6 8.4 15 40.0 III 0.89 5 5 0.002 ** 147.4 154.9 7.5 7.5 155 4.8 19 26.3 Va 1.27 7 12 0.000 ** 20.9 31.6 10.7 24.2 114.9 21.1 19 63.2 1.60 5 0.024 * 48.1 61.6 13.5 VI 1.00 4 4 0.019 * 40.1 48.5 8.4 8.4 173.9 4.8 17 23.5 VII 0.71 3 3 0.035 * 58.0 64.0 6 6 64.1 9.4 12 25.0 VIII 0.12 2 2 0.007 ** 20.1 21.1 1 1 152.2 0.7 15 13.3 e l</p><p> a X 3.23 7 7 0.046 * 0.0 27.2 27.2 27.2 125.6 21.7 21 33.3 M XI 0.84 4 4 0.011 * 59.3 66.4 7.1 7.1 122.1 5.8 15 26.7 XIII 0.26 3 10 0.002 ** 28.7 30.9 2.2 10.3 107.8 9.6 20 50.0 0.91 5 0.002 ** 53.7 61.4 7.7 0.05 2 0.001 ** 91.8 92.2 0.4 XIV 0.40 3 3 0.008 ** 50.6 54.0 3.4 3.4 105.3 3.2 12 25.0 XV 1.01 4 4 0.020 * 0.0 8.5 8.5 8.5 82 10.4 9 44.4 XVI 2.56 9 9 0.001 ** 40.4 62.0 21.6 21.6 92.8 23.3 18 50.0 XVII 1.48 5 5 0.018 * 48.1 60.6 12.5 12.5 127.6 9.8 17 29.4 XIX 0.31 2 2 0.039 * 26.0 28.6 2.6 2.6 81.1 3.2 10 20.0 Region with dispersed markers </p><p>Group λij Nij ∑ N.j P Sij Eij Iij ∑ Iij Li Li % Ni Ni% III 11.16 4 4 0.014 * 49.7 117.2 67.5 67.5 155.7 43.4 18 22.2</p><p> e V 7.94 2 2 0.014 * 25.3 73.3 48 48 73.4 65.4 11 18.2 l a</p><p> m VI 7.84 3 3 0.047 * 127.6 175.0 47.4 47.4 182.3 26.0 29 10.3 e F X 8.82 2 2 0.007 ** 82.3 135.6 53.3 53.3 135.7 39.3 19 10.5 XIV 14.93 7 7 0.019 * 0.0 90.3 90.3 90.3 113.9 79.3 14 50.0 e</p><p> l IV 10.64 4 4 0.019 * 6.8 96.5 89.7 89.7 123.4 72.7 11 36.4 a</p><p>M VIII 6.51 2 2 0.043 * 51.2 106.1 54.9 54.9 152.2 36.1 15 13.3</p><p>** significant at α=0.01; * significant at α=0.05; λij is the expected number of markers in a distance of Iij; Nij is the observed number of markers in a distance of Iij; ∑ N.j is the sum of Nij on linkage group i P is the Poisson one-tailed P-value of Nij ≤ λij or Nij ≥ λij; Ni is the total number of markers on linkage group i; Ni % is the percentage of markers clustered or in lower density than expected; Sij is the start position (cM) of Iij on linkage group i; Eij is the end position (cM) of Iij on linkage group i; Iij is the j distance (cM) formed by continuous intervals whose size is larger or smaller than average on linkage group i; ∑Iij the sum (cM) of Iij on linkage group i; Li is the map distance (cM) between terminal markers of linkage group i; Li % is the percentage of regions with clustered or dispersed markers on linkage group i; </p>