LIBRARY OF THE UNIVERSITY OF ILLINOIS AT URBANA-CHAMPAIGN 550.5 FI IS. V. 12 3z SEouoai this ^ '"re- ^do^i^cpy ft i THE STRATIGRAPHIC DISTRIBUTION OF THE PALEOZOIC EUMALACOSTRACA SOME MIDDLE PENNSYLVANIAN HOPLOCARIDA (CRUSTACEA) AND THEIR PHYLOGENETIC SIGNIFICANCE FREDERICK R. SCHRAM FIELDIANA: GEOLOGY VOLUME 12, NUMBERS 13 AND 14 ijrii«ersiiy of Illinois Published by FIELD MUSEUM OF NATURAL HISTORY JAN 28 1970 SEPTEMBER 30, 1969 tkiq GEOLOGY (JBRAinc TT^t^^ ^^,. FIELDIANA: GEOLOGY * A Continuation of the ^ <x \^ GEOLOGICAL SERIES oov^^'^S-^^*"^ of FIELD MUSEUM OF NATURAL HISTORY VOLUME 12 FIELD MUSEUM OF NATURAL HISTORY CHICAGO. U.S.A. INDEX Acanthotelson 9, 213 Crustaceans 9, 213-229, 235-286 Adelophthalmus mazonensis 85-100 cydophthalmus, Mazonipterus, n. sp. Aeschronectida, n. ord. 236 85, 101-103 aetherius, Anthrakcris, n. sp. 36-38 Amphibians 9 Deltaic deposition 10, 11 Amphineura 61 Dipnoans 9, 193-209 Annelida 9, 60 Anthracaris 213, 219 edesi, Conchopoma, n. sp. 195 Anthracomedusa, n. gen. 110 Essex Fauna 109, 119, 213 Anthracomedusa turnbidli, n. sp. 110- Eumalacostraca 213-229 112 Eurypterids 85-104 Anthracophausia 213 Anthrakoris, n. gen. 36 Fishes 9, 193-209 Anthrakoris aetherius, n. sp. 36-38 Fossil fauna 8-10 Anthrapalaemon 9, 219 Fossil flora 8-10 Arachnids 9 Fossils, abundance 9 Aratidedhes, n. gen. 251 Fossils, occurrence 4, 8, 9, 15, 121 Aratidedhes johnsoni, n. sp. 252-259 Fossils, preservation 109, 122-127, 137, Archaeostomatopodea, n. subord. 260 155 Arlhropleura 71-76 Francis Creek Shale 3, 10, 11 Arthropleura crislata, n. sp. 79-82 Arthropleurida 9, 71-76, 79-82 Genopterygidae, n. fam. 41-43 Aviculopecten mazonensis 65-66 Genopteryx 43 Glaphyrokoris, n. gen. Bandringa, n. gen. 158 Gl<iphyrokoris mirandus, n. sp. 38-39 Bandringa rayi, n. sp. 160-169 gregarium, Tullimonstrum 119-148 Bandringidae, n. fam. 157 Grundy County, Illinois 3-6 Belotelson 213 Blattaria 17 Helminthochiton concinnus, n. sp. 62- Braidwood, 111. 3, 5, 7 65 Braidwood Fauna 109, 119, 213 Holothurians 121 Hoplocarida 235-286 33 Cacurgidae Howellitubus, n. gen. 60 Caloneurodea 41 17, 18, Howellitubus whitfieldorum, n. sp. 60- Carbondale Formation 3 61 carpenteri, Lithoneura, n. sp. 24 Hughmilleriidae 86 Cephalopod 59 Channel-fill sandstone 3 inopinus, Rossites, n. sp. 44-45 Coal No. 2 3, 10 Insects, faunas 15-17 Coal City, 111. 3, 5, 6 Insects, immature 52-54 Conchopoma 193-195, 207 Insects, occurrence 15, 16 Conchopoma edesi, n. sp. 195-207 Insects, study 15, 16 concinnus, Helminthochiton, n. sp. 62- 65 n. 252-259 Concretions 3, 4, 8, 109, 121, 126-127 johnsoni, Aratidedhes, sp. Coprolites 9 Crinoids 151-156 Kallidedhes, n. gen. 237 cristata, Arthropleura, n. sp. 79-82 Kallidedhes richardsoni, n. sp. 238-251 Crossopterygians 9 Kallidecthidae, n. fam. 237 INDEX Langford, George 16 Protodonata, 17 Lithoneura 2U Protorthoptera 17, 33-41 Lithoneura carpenieri, n. sp. 24 Liverpool Cyclothem 3 Radula, polyplacophoran 64 linearis 61 Longitubus rayi, Bandringa, n. sp. 160-169 n. 20-24 luciae, Mcluckiepteron, sp. richardsoni, Kallidecthes, n. sp. 238- Lungfishes 9, 193-209 251 rossae, Nacekomia, n. sp. 34-36 219 Mamayocaris Rossites, n. gen. 44 Marine fossils 109-115 11, 59-66, Rossites inopinus, n. sp. 44-45 Mazon Creek 3-5, 8, 9, 15, 16 mazonensis, Adelophthalmus 85-100 Scorpions 171 mazonensis, Aviculopecten 65-66 Serpulidae 60 Mazonia woodiana 172-189 Sharks 59, 157 Mazonipterus, n. gen. 100-101 Spilapteridae 19-24 n. Mazonipterus cyclophthalmns, sp. Strip mines, specimens from 4, 16 101-103 85, Strip mining 3-7 Mcluckiepteron, n. gen. 20 Stomatopoda 260 Mcluckiepteron luciae, n. sp. 20-24 Stylonuridae 100 Medusae 109-115 Syntonopteridae 24-27 Megasecoptera 17, 18 mirabile, Teneopteron 47-52 Teneopteridae, n. fam. 46 mirandus, Glaphyrokoris, n. sp. 38-39 46 Mycterops 85, 103-104 Teneopteron 47-52 Myriapods 9 Teneopteron mirabile theridion, Tyrannophontes, n. sp. 261- 266 Nacekomia, n. gen. 33 Trilobite 59 Nacekomia rossae, n. sp. 34-36 79-82 Nymph, insect 52-54 Trilobitomorpha 71-76, Tullimonstrum gregarium 119-148 n. 110- Octomedusa, n. gen. 112 turnbulli, Anthracomedusa, sp. 112 Octomedusa pieckorum, n. sp. 113- 115 Turnbullia, n. gen. 27 Turnbullia n. 28-31 Opisterostomatopodea, n. subord. 266 priscillae, sp. Ostracodes 9 Tyrannophontes, n. gen. 260 Tyrannophontes theridion, n. sp. 261- 266 Palaeocaris 9, 213 Palaeodictyoptera 17-32 Unnamed crinoid 151 155-156 Unnamed insects 31, 32, 40, 41 Palaeoniscoids 9 Unnamed lungfish 208 Palaeostomatopoda 259 Paleoecology 10-12, 17, 59, 147, Waupecan Sandstone 3 Pelecypods 9, 65-66 whitfieldorum, Howellitubus, n. sp. 60- Perimecturidae 260 61 pieckorum, Octomedusa, n. sp. 113-115 Will County, 111. 3-6 Pleasantview Sandstone 3 woodiana, Mazonia 172-189 Polyplacophora 61 Prestwichianella 9 Xenacanths 9, 59 priscillae, Turnbullia, n. sp. 28-31 Xiphosures 9 TABLE OF CONTENTS PAGE 1. Pennsylvanian Invertebrates of the Mazon Creek Area, Illinois. Intro- duction. By Eugene S. Richardson, Jr 1 2. Pennsylvanian Invertebrates of the Mazon Creek Area, Illinois. Insects. By Eugene S. Richardson, Jr 13 3. Pennsylvanian Invertebrates of the Mazon Creek Area, Illinois. Marine Fauna. By Eugene S. Richardson, Jr 57 4. Pennsylvanian Invertebrates of the Mazon Creek Area, Illinois. Trilo- bitomorpha, Arthropleurida. By Eugene S. Richardson, Jr 69 5. Pennsylvanian Invertebrates of the Mazon Creek Area, Illinois. Trilo- bitomorpha, Arthropleurida, II. By Eugene S. Richardson, Jr 77 6. Pennsylvanian Invertebrates of the Mazon Creek Area, Illinois. Euryp- terida. By Erik N. Kjellesvig-Waering 83 7. Pennsylvanian Invertebrates of the Mazon Creek Area, Illinois. The Essex Fauna and Medusae. By Ralph Gordon Johnson and Eugene S. Richardson, Jr 107 8. Pennsylvanian Invertebrates of the Mazon Creek Area, Illinois. The Morphology and Affinities of Tullimonstrum. By Ralph Gordon Johnson and Eugene S. Richardson, Jr 119 9. A Crinoid from the Pennsylvanian Essex Fauna of Illinois. By N. Gary Lane 151 10. Bandringa rayi, a New Ctenacanthoid Shark from the Pennsylvanian Essex Fauna of Illinois. By Rainer Zangerl 157 11. Scorpionida: the Holotype of Mazonia woodiana Meek and Worthen, 1868. By Erik N. Kjellesvig-Waering 171 12. New Pennsylvanian Lungfishes from Illinois. By Robert H. Denison . 193 13. The Stratigraphic Distribution of the Paleozoic Eumalacostraca. By Frederick R. Schram 213 14. Some Middle Pennsylvanian Hoplocarida (Crustacea) and Their Phylo- genetic Significance. By Frederick R. Schram 235 THE STRATIGRAPHIC DISTRIBUTION OF THE PALEOZOIC EUMALACOSTRACA SOME MIDDLE PENNSYLVANIAN HOPLOCARIDA (CRUSTACEA) AND THEIR PHYLOGENETIC SIGNIFICANCE FREDERICK R. SCHRAM FIELDIANA: GEOLOGY VOLUME 12, NUMBERS 13, AND 14 Published by FIELD MUSEUM OF NATURAL HISTORY SEPTEMBER 30, 1969 CONTENTS PACE The Stratigraphic Distribution of the Paleozoic Eumalacostraca. by Frederick R. Schram 213 Some Middle Pennsylvanian Hoplocarida (Crustacea) and Their Phylogenetic Significance, by Frederick R. Schram 235 FIELDIANA . GEOLOGY Volume 12, No. 13 September 30, 1969 Publication 1079 The Stratigraphic Distribution of the Paleozoic Eumalacostraca Frederick R. Schram Walker Museum University of Chicago Department of Zoology, Eastern Illinois University INTRODUCTION Johnson and Richardson (1966) deHneated two Middle Pennsyl- vanian faunas within the Mazon Creek area in Grundy, Will, and Kankakee counties in northeastern Illinois —the Braidwood fauna and the Essex fauna. The fossils of these assemblages are found in the ironstone concretions of the Francis Creek Shale, Carbondale Formation, directly above the coal of the Liverpool Cyclothem. These beds are presently classified as being in the Kewanee Group of the Illinois Pennsylvanian (Kosanke et al., 1960). The Braidwood fauna is found along Mazon Creek and in the abandoned strip mines north of the towns of Coal City and Braidwood. The Essex fauna is found in the recently excavated strip mines near the town of Essex. These faunas are considered to represent a marginal deltaic regime, the Braidwood being the freshwater facies while the Essex is a ma- rine one. Crustacea are found in both of these faunas. Five species are de- scribed from the Braidwood localities: Acanthotelson stimpsoni Meek and Worthen, 1865; Palaeocaris typus Meek and Worthen, 1865; Belotelsoji magister Packard, 1886; Anthracaris gracilis (Meek and Worthen, 1865); and Anthracophausia strongi Brooks, 1962c. The Essex fauna contains some 20 recognizable forms, including the above. The proportions of occuiTence of species are quite different between the two faunas. The Braidwood Crustacea are mostly syncarids; the Essex Crustacea are mostly eocarids and hoplocarids. The study of this late Paleozoic fauna, comparable in preservation to that of the Burgess shale in the Middle Cambrian or the Solen- hofen limestone in the Upper Jurassic, is of great import toward un- derstanding the earlj'^ history of the Eumalacostraca. Until now the Library of Congress Catalog Card Number: 73-9291 7 213 214 FIELDIANA: GEOLOGY, VOLUME 12 higher Crustacea have had a record of isolated occurrences through- out the latter half of the Paleozoic. The Paleozoic Eumalacostraca are listed below (Table 1) along with their stratigraphic ranges. H. K. Brooks is at present revising the European forms. Considerable taxonomic confusion exists at the specific level in the European material. No attempt is made to for- mally express synonymies in this paper except in instances where the work is done and in press, where such has been indicated in private conversations