ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Jahrbuch der Geologischen Bundesanstalt Jahr/Year: 1989 Band/Volume: 132 Autor(en)/Author(s): Kozur Heinz, Mostler Helfried Artikel/Article: Echinoderm Remains from the Middle Permian (Wordian) from Sosio Valley (Western Sicily) 677-685 ©Geol. Bundesanstalt, Wien; download unter www.geologie.ac.at __ J_b_.G_eo_l_._B_.-_A_. IS_S_N_OO_1_6_-_7_8_00 B_a_nd_1_32 H_e_ft_4 S_._6_7_7_-6_8_5__ I~ien, Dezember 1989 Echinoderm Remains from the Middle Permian (Wordian) from Sosio Valley (Western Sicily) By HEINZ KOZUR & HELFRIEDMOSTLER*) With 1 Text-Figure and 2 Plates Italy Sicily Sosio Val/ey Paleontology Echinoderms Taxonomy Contents Zusammenfassung 677 Abstract 677 1. Introduction 677 2. Geological Setting 677 3. Paleontological Part 678 4. Stratigraphic Evaluation of the Fauna 682 Acknowledgments 682 References 685 Zusammenfassung cluded into Gondolella rosenkrantzi BENDER & STOPPELby 2 neue Familien, 5 neue Gattungen und 10 neue Arten von these authors. This species indicates (? Kubergandi- mikroskopisch kleinen Echinodermenresten, die zu den nian) Wordian age, like also the ammonoid faunas. Ophiocistioidea, Ophiuroidea, Holothuroidea und Echinoder- One of the authors (H. MOSTLER)sampled the Rupe men inc. sedis gehören, werden aus dem Mittelperm von del Passo di Burgio, Rocca di San Benedetto and Rupe dei Passo di Burgio (Sosio- Tal, West-Sizilien) beschrie- Pietra di Salomone sections in 1984. The other author, ben. Die Gattung Microantyx, die bisher zu den Holothuroidea gestellt wurde, wird den Ophiocistioidea SOLLASzugerechnet. H. KOZUR,sampled the same sections together with Dr. P. 01 STEFANO, Palermo, in 1987. Ammonoid-bearing sediments of the Rupe del Passo di Burgio section with Abstract Waagenoceras, the index genus of the Wordian, yielded 2 new families, 5 new genera and 10 new species of micro- rich microfaunas with the conodonts Mesogondella sicilien- scopic echinoderm remains belonging to the Ophiocistioidea, sis (KOZUR), Merrillina galeata (BENDER & STOPPEL), Hin- Ophiuroidea, Holothuroidea and echinoderm remains incertae deodus n. sp. A and Hindeodus n. sp. B that indicate sedis were desribed from the Middle Permian of Rupe del Passo di Burgio (Sosio Valley, Western Sicily). The genus Mi- likewise a Wordian age. croantyx, so far placed into the Holothuroidea is assigned to Both authors found also many echinoderm remains the Ophiocistioidea SOLLAS. (crinoids, echinids, ophiuroids, Ophiocistioidea, holothurian sclerites and echinoderms, inc. sedis) partly described in the present paper. Most of these echinoderm remains derived from the ammonoid-bear- 1. Introduction ing beds of Rupe del Passo di Burgio. The Middle Permian of the Sosio Valley has been in- vestigated for long time, but mainly macrofaunas (e. g. ammonoids, crinoids, sponges) were described. Except 2. Geological Setting fusulinids, microfossils were only described by BENDER & STOPPEL(1965), who found for the first time cono- The stratigraphy of the Permian of western Sicily was donts in the Sosio Valley. The stratigraphically most quite revised by CATALANO;01 STEFANO& KOZUR (1988 important species from the Rupe del Passo di Burgio and in press). According to these authors a continuous section, Mesogondolella siciliensis (KOZUR) was still in- deep-water sequence is known from the Kungurian (higher Lower Permian) up to the Late Permian. Beside of this deep-water sequence (e. g. Kungurian flysch, *) Authors' addresses: Dr. sc. HEINZ KOZUR, Reszü ut 83, H-1029 Budapest, Hungary; Univ.-Prof. Dr. HELFRIEDMOST- Kubergandinian olistostrome unit, Wordian to Ozhulfian LER, Institut für Geologie und Paläontologie, Universität soft, predominantly reddish clays) also slope sedi- Innsbruck, Innrain 52, A-6020 Innsbruck. ments are present, known from olistoliths in the olisto- 677 ©Geol. Bundesanstalt, Wien; download unter www.geologie.ac.at o 805io Valley, Rupe del Passo di Burgio Text-Fig. 1. o 50 l00Km Location of the sampling locality, Rupe del Passo di Burgio in the Sosio Valley, Western Sicily). strome unit and from big blocks (tectonic klippen) of group has to be elevated from the Lower Carbonifer- Middle Permian light-coloured limestones, which have ous (until now known youngest Ophiocistioidea) up yielded the famous Wordian faunas from the Sosio Val- to the top of the Permian. ley described since the end of the last century. Until now, Microantyx was placed into the holothurian Our investigated rocks were sampled in these blocks family Protocaudinidae OEFLANDRE-RIGAUD,1962. But of Middle Permian slope sediments. The here de- these holothurian sclerites are basically different by scribed echinoderm remains derived all from light-col- the absence of any hub. oured, ammonoid-bearing limestones of the locality Rupe del Passo di Burgio in the Sosio Valley (location see Fig. 1). Microantyx sosioensis n. sp. (PI. 1, Figs. 1,2) Oe r i vat ion 0 mi n is: According to the occurrence in the Sosio Valley. 3. Paleontological Part Hol 0 typ e: The specimen on pI. 1, fig. 1; rep.-no. KoMo 1988 1/1/IV-55a, lstituto Geologico, Palermo University. Class Ophiocistioidea SOLLAS, 1899 Locus typicus: Rupe del Passo di Burgio, Sosio Family Rotasaccidae Valley, Sicily HAUDE & LANGENSTRASSEN, 1976 Stratum typicum: Sample 120, light coloured, Wordian ammonoid-bearing limestone with Mesogon- Genus Microantyx KORNICKER & IMBRIE, dolella siciliensis (KOZUR). 1958 emend. MOSTLER, 1983 Material: 14 specimens. o i a g nos is: Wheel with moderately undulated rim and Ty pes pee i es: Microantyx permiana KORNICKER& IMBRIE, large, especially in the central part very high hub. 8 1958. spokes short, tapering towards the rim. Lower side Rem ark s: The wheels of Microantyx are very similar to with relatively small, but deep excavation. Arches the wheels of Rotasaccus dentifer HAUDE& LANGENSTRAS- narrow, with distinct median bar and 4 rays on its SEN (Ophiocistioidea SOLLAS). Only the symmetry of ends. The pores between the archs and the margin the archs on the lower sufaces of the wheels is diffe- of hub are smaller than the pores between the rent (4 in Microantyx and 3 in Rotasaccus). We place here spokes and the rim. One pair of opposite central also the genus Microantyx into the Ophiocistioidea pores is usually somewhat larger than the other one. SOLLAS.This is also supported by the fact that KOR- Occurrence: Middle Permian of Sosio Valley. Re- NICKER& IMBRIE(1958) published together with Micro- marks: Microantyx botoni GUTSCHICK, 1959, from the antyx permiana, the type species of Microantyx, wrinkle Lower Mississippian has cross-shaped, very broad teeth (goniodonts), further very characteristic skele- arches and small central pores of equal size. ton elements of the Ophiocistioidea. We have found Moreover, the rim is quite round or only indistinctly in our material wheels (Microantyx spp.). goniodonts undulated. and sieve plates, the most important skeleton ele- Microantyx tarazi MOSTlER & RAHIMI-YAZD, 1976, from ments of Ophiocistioidea. Some elements could be the highest Oorashamian (higher part of Lower rotulae. By our assignment of Microantyx into the Changxingian) has a strongly undulated outer rim. It Ophiocistioidea the known upper range of this fossil has obviously evolved from M. sosioensis n. sp. 678 ©Geol. Bundesanstalt, Wien; download unter www.geologie.ac.at Mieroantyx pseudobotoni n. sp. Pararotasaeeus permieus n. gen. n. sp. (PI. 1, Figs. 3,4) (PI. 1, Fig. 5) Der i vat ion 0 m i n is: According to the similary with Der i vat ion 0 m i n is: According to the occurrence in M. botoni GUTSCHICK,1959. the Permian. Holotype: The specimen on pI. 1, fig. 3; rep.-no. Holotype: The specimen on pI. 1, fig. 5; rep.-no. KoMo 1988/1/2/111-94 a. KoMo 1988 1-3/IV-55 Locus typicus: As for M. sosioensis. Locus typicus and stratum typicum: As for Stratum typicum: Sample S 9, Iight-coloured, am- Mieroantyx sosioensis n. sp. monoid-bearing limestone with Mesogondo/e//a siei/iensis Material: 5 specimens. (KOZUR), Wordian. Dia g nos is: Wheels circular to subcircular with con- Material: 23 specimens. stantly 6 spokes tapering toward the wheels. Hub Dia g nos is: Wheels with round or only indistinctly un- centrally higher than rim. Lower surface of hub in a dulated rim. Hub higher than rim, large. 8 short relatively small inner part deeply excavated. Arches spokes, tapering toward the rim. Lower surface basically triradiate. These 3 primary branches are deeply excavated. 4 arches thin, with median bar. bibranchiated shortly after the centre. Near to the Central pores generally large, but sometimes smaller margin some of the secondary branches are secon- than the outer pores. One pair of opposite central darily branched. pores may be a little smaller than the other one. Oe cur ren ce: Middle Permian (Wordian) of Sosio Val- Occurrence: Higher Mississippian to Late Permian. ley. Rem ark s: The holotype and the paratype of Mieroantyx Rem ark s: In Pararotasaeeus mudgei (GuTSCHICK;CANIS & botoni GUTSCHICK,1959, have archs in form of a broad BRill, 1967) the branchiating of the 3 primary cross and the central moderately undulated rim. It is branches of the arch is near to the margin. Mostly a transitional form between the conservative M. one, sometimes even two primary branches are un- pseudobotoni n. sp. and M. tarazi MOSTlER & RAHIMI- branchiated. No branching of the secondary YAZO, 1976, from the Late Permian. branches can be observed. Moreover, in this species an incomplete secondary outer pore ring is de- veloped. Genus Pararotasaccus n. gen. Class Ophiuroidea GRAY, 1840 Family Calclyridae FRIZZELL & EXLINE, 1955 Ty pes pee i es: Pararotasaeeus permieus n. gen. n. sp. Dia g nos is: Wheel circular to subcircular. Hub in the Genus Calclyra FRIZZELL & EXLINE, 1955 central portion rather strongly elevated. Lower sur- Typ e s pee i es: Prosynapta eise/iana SPANDEl, 1898. face in the central part deeply excavated, with 3 at least terminally dibranchiated archs. Remarks: MOSTlER (1971) could demonstrate that Ga/e/yra is not a holothurian sclerite, but belongs to Oe cur ren ce: Mississippian to Middle Permian.