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Short Communication Assignment Of

Journal of Vertebrate Paleontology 29(1):295–302, March 2009 # 2009 by the Society of Vertebrate Paleontology SHORT COMMUNICATION ASSIGNMENT OF YAMACERATOPS DORNGOBIENSIS AND ASSOCIATED REDBEDS AT SHINE US KHUDAG (EASTERN GOBI, DORNGOBI PROVINCE, MONGOLIA) TO THE REDESCRIBED JAVKHLANT FORMATION (UPPER CRETACEOUS) DAVID A. EBERTH,*,1 YOSHITSUGU KOBAYASHI,2 YUONG-NAM LEE,3 OCTA´ VIO MATEUS,4 FRANC¸ OIS THERRIEN,1,5 DARLA K. ZELENITSKY,5 and MARK A. NORELL6; 1Royal Tyrrell Museum, Box 7500, Drumheller, Alberta, Canada, [email protected]; 2Hokkaido University, Hokkaido, Japan; 3Korean Institute of Geoscience & Mineral Resources, Daejeon, Korea; 4Museu da Lourinha˜ & Universidade Nova de Lisboa, Lourinha˜, Portugal; 5Department of Geoscience, University of Calgary, Calgary, Alberta, Canada; 6Division of Paleontology, American Museum of Natural History, New York City, USA Assigning Central Asian nonmarine Cretaceous vertebrate dorngobiensis, a taxon whose age, previously, had been fossils and their host rocks to lithostratigraphic units and estab- assessed provisionally as late Early Cretaceous (Makovicky lishing their sub-epochal ages can be challenging. Researchers and Norell, 2006:3) based on an erroneous inference that the frequently struggle with inconsistently applied locality and strat- host strata occur stratigraphically below the Baynshiree Forma- igraphic names, have to consider the impact of two distinct tion/Svita. Our review of GPS-derived locality data for the type stratigraphic philosophies (Benton 2001a; Gladenkov, 2007), and referred materials of Yamaceratops dorngobiensis confirms and often encounter additional challenges in working with that they were collected from the Shine Us Khudag redbeds manuscripts published in, or translated from, a variety of in a localized area of badlands called Khugenslavkhant languages (e.g., Benton, 2001b). In addition to these challenges, (Makovicky and Norell, 2006; Fig. 1). Because the Shine Us chronostratigraphic data are rare due both to a lack of discov- Khudag redbeds conformably overlie the richly fossiliferous ered rocks amenable to radiometric dating, and issues com- Upper Cretaceous Baynshiree Formation/Svita, and because plicating paleomagnetic sampling in the Cretaceous (e.g., geologists and biostratigraphers working in the Eastern “Cretaceous Quiet Zone,” Ogg et al., 2004). There are also Gobi have consistently shown the Baynshiree Formation/ inherent limitations in nonmarine biostratigraphic schemes and, Svita and the Shine Us Khudag redbeds to be Late Cretaceous in some cases, field areas may have complex stratigraphic archi- in age (Cenomanian-Campanian), we conclude that all tectures and depositional histories that are difficult to resolve known specimens of Yamaceratops dorngobiensis are also without the aid of subsurface data (e.g., Graham et al., 2001; Late Cretaceous in age, not Early Cretaceous as previously Johnson, 2004). Thus, although there have been significant advances in our knowledge about the ages and relationships of suggested. nonmarine Cretaceous vertebrate fossils and assemblages from In addition to Yamaceratops dorngobiensis, the KMIDP docu- southern Mongolia in the past 85 years, there remains much to mented the presence in these redbeds of a soft-shelled turtle, learn and clarify. and small nonavian theropod dinosaur skeletal remains and egg- During the 2007 field season, the Korea-Mongolia Interna- shell. Khand et al. (2000) noted the presence of ornithomimids. tional Dinosaur Project (KMIDP) worked two weeks in the Most importantly, however, the Mongolian-American Expedi- Eastern Gobi (Dorngobi Province, southwest of Sainshand) at tion (MAE) working in and around Khugenslavkhant in three Upper Cretaceous localities: Shine Us Khudag, Khar 1991, 2002-2004 and 2006-2007 collected hundreds of vertebrate Khutul, and Bayn Shiree (Fig. 1). Exposures of the 280 m thick, fossil specimens from the Shine Us Khudag redbeds, including buff-colored Baynshiree Formation/Svita at each locality pro- the type and referred materials of Yamaceratops dorngobiensis, duce a rich assemblage of Cenomanian-Santonian vertebrate and the partial skulls and postcrania of two new eutherian mam- and invertebrate fossils, including dinosaurs (Vasil’ev et al., mals (Giallombardo and Novacek, 2006; Giallombardo, 2007). 1959; Martinson et al., 1969; Barsbold, 1972; Sochava, 1975; In addition to these published remains, MAE also collected Martinson, 1982; Jerzykiewicz and Russell, 1991; Hicks et al., multiple skeletons and skulls of a hypsilophodont dinosaur, 1999; Ishii et al., 2000; Khand et al., 2000; Jerzykiewicz, 2001; several taxa of variously-sized theropod dinosaurs, turtles and Shuvalov, 2001; Graham et al., 2001). However, north of Shine lizards, additional specimens and taxa of mammals, and several Us Khudag and south-southwest of Khar Khutul, a 380 m thick types of dinosaur eggs and nests. One of the eggs includes the succession of fossiliferous alluvial redbeds (here called the embryo of a protoceratopsian dinosaur (probably Yamaceratops Shine Us Khudag redbeds) is broadly exposed (50 km2) and dornogobiensis; Balanoff and Norell, 1996), and many of conformably overlies the Baynshiree Formation/Svita (Fig. 2). the dinosaur specimens are juveniles. Given this rich and dis- The Shine Us Khudag redbeds have variously been assigned to tinctive fossil assemblage, we regard these Upper Cretaceous the Jibhalanta Suite, Javkhlant Formation, Baruungoyot Svita, redbeds as a critical source of new paleobiogeographic, Djadokhta Formation, and the “Barunbayan” (Table 1), and biostratigraphic, and paleoecologic data that will improve since 1969 have been regarded by most stratigraphers working our understanding of the Upper Cretaceous geologic history of in the area as having a Santonian-Campanian age (Martinson Central Asia. et al., 1969; Jerzykiewicz and Russell, 1991; Khand et al., 2000; Here, we review the details of the nomenclatural and age Jerzykiewicz 2001; Shuvalov, 2001). assignment history of the Shine Us Khudag redbeds, and present Locally abundant in the Shine Us Khudag redbeds are new lithostratigraphic and preliminary sedimentologic informa- remains of the basal protoceratopsian dinosaur, Yamaceratops tion that allows them to be formally assigned to the Javkhlant Formation. Our data more clearly establish and serve to redescribe the Javkhlant stratotype. We underscore that many of the basic sedimentologic descriptions and paleoenvironmental *Corresponding author. interpretations presented here are not substitutes for detailed 295 296 JOURNAL OF VERTEBRATE PALEONTOLOGY, VOL. 29, NO. 1, 2009 sedimentological and paleoenvironmental analyses of the Shine Us Khudag redbeds, which will be presented elsewhere. We follow the spellings provided in Benton (2001b) for Mongolian locations and stratigraphic units, except in those cases where we quote original literature. However, following the most recent revision to the Russian Stratigraphic Code (see Gladenkov, 2007) and contra to recent trends (e.g., Benton 2001a), we do not use “Svita” (a chronostratigraphic term) and “Formation” (a lithostratigraphic term) interchangeably. DESCRIPTION OF THE SHINE US KHUDAG REDBEDS The preserved thickness of the Shine Us Khudag redbeds is 380 m (Fig. 2). The base of the unit is well exposed in the southern portion of the field area, where it rests conformably on the Bayn Shiree Formation (Fig. 2I). At the top of the sec- tion, the Shine Us Khudag redbeds are truncated by an angular unconformity overlain by Quaternary Gobi sediments. Through- out the field area, the Shine Us Khudag redbeds and the under- lying Bayn Shiree Formation are structurally tilted in a variety of directions (Fig. 1), most likely as a result of the Late Mesozoic tectonic rift history of the area (cf. Graham et al. 2001; Meng et al., 2003; Johnson, 2004). Although the Shine Us Khudag redbeds consist overwhelm- ingly of rooted and mottled paleosols, massive sandy mudrocks, and sheet-shaped-to-lenticular sandstones and conglomerates, we recognize a three-fold stratigraphic subdivision of the unit that reflects an upsection increase in both grain size and the relative abundance of coarse-grained versus fine-grained sedi- ments and facies. We use this three-fold division to further orga- nize our description of the Shine Us Khudag redbeds. The lowest 110 meters of the Shine Us Khudag redbeds rest conformably on the light-tan colored sediments of the Bayn- shiree Formation. This lower interval is dominated by paleosols developed on mudstones comprising decimeter-thick couplets of deep red (10R 4/6; Rock-Color Chart Committee, 1995), clay- enriched horizons with slickensides and clay coatings, overlain by lighter red (10R 5/4-5YR 5/6), clay-impoverished and often well rooted horizons (Fig. 2G, H). Following Birkeland (1999), we tentatively interpret these mudstone couplets as ancient argi- lic (Bt) soil horizons overlain by ancient eluvial (E?) or surface (A) horizons. Interbedded with these paleosols are (1) decime- ter-thick sheets of grey, fine-grained sandstone exhibiting root traces, finger-size invertebrate burrow fills, and soft-sediment deformation structures, (2) decimeter-thick deposits of massive, light red, sandy mudstone with very sparse assemblages of cal- careous glaebules, interpreted here as immature caliches (Allen, 1986; Retallack, 1990), and (3) rare, 1-2 meter thick, lenticular sandstones with occasional granule to pebble strings, interpreted FIGURE 1. Locality map showing the distribution of

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