THE UNIVERSITY OF KANSAS PALEONTOLOGICAL CONTRIBUTIONS Serial Number 46 ECHE\ ODERMATA ARTICLE 9 Pages 1-86, Figures 1-6, Plates 1-28 CLASSIFICATION AND NOMENCLATURE OF FOSSIL CRINOIDS BASED ON STUDIES OF DISSOCIATED PARTS OF THEIR COLUMNS RAYMOND C. MOORE and RUSSELL M. JEFFORDS The University of Kansas Paleontological Institute HAROLD NORMAN FISK MEMORIAL PAPERS Humble Oil & Refining Company THE UNIVERSITY OF KANSAS PUBLICATIONS JANUARY 26, 1968 THE UNIVERSITY OF KANSAS PALEONTOLOGICAL CONTRIBUTIONS Serial Number 46—Echinodermata, Article 9, Pages 1-86, Figures 1-6, Plates 1-28 CLASSIFICATION AND NOMENCLATURE OF FOSSIL CRINOIDS BASED ON STUDIES OF DISSOCIATED PARTS OF THEIR COLUMNS RAYMOND C. MooRE 1 and RUSSELL M. JEFFORDS 2 CONTENTS [Asterisk (*) indicates type species of genera] PAGE PAGE ABSTRACT 7 Order Cladida Moore & Laudon, 1943 33 INTRODUCTION 7 Suborder Dendrocrinina Bather, 1899 33 Nature and Occurrence of Fossil Crinoid Frag- I Group l'entamer' I 33 ments Family Proctothylacocrinidae Kier, 1952 34 Dissociation of Skeletal Remains 9 Genus Proctothylacocrinus Kier, 1952 34 Materials for Study 9 Proctothylacocrinus *Iongus Kier, 1952? 34 Scope of Article 9 Proctothylacocrinus esseri Kesling, 1965 35 Acknowledgments 1 1 Genus Decacrinus Yeltysheva, 1957 35 ARTICULATED SKELETONS OF CRINOIDS l i Decacrinus *pcnnatus Yeltysheva, 1957 35 Main Morphological Features Il Decacrinus tenuicrenulatus, new 35 Theca Genus Kuzbassocrinus Yeltyshcva, 1957 35 Pelma Kuzbassocrinus *bystrotvi Yeltyshcva, 1957 .. 36 Main Taxonomic Groups 14 Subclass Camerata Wachsmuth & Springer, 1885 .. 36 Occurrence 14 Order Diplobathrida Moore & Laudon, 1943 36 Stratigraphie Significance 16 I Group Pentameri I 36 Pztleoecologic Significance 17 Family Lampterocrinidae Bather, 1899 36 DISARTICULATED SK LETAL REMAINS OF CRINOIDS . 18 Genus Lampterocrinus Roerner, 1860 36 Nature and Usefulness of Remains 18 Lampterocrinus *tennesseensis Roemer, 1860 36 Methods of Study 19 Lampterocrinus sp. A 37 Previous Work 20 Lampterocrinus sp. B 37 Classification and Nomenclature of Crinoids I Group Cyclici 37 Based on Studies of Disarticulated Stem Parts 24 Family Rhodocrinitidae Bassler, 1938 (1854) 37 Objectives and Principles 24 Genus Atactocrinus Weller, 1916 37 Unacceptable Taxonomie Groups 25 Atactocrinus? sp. 37 Acceptable Taxonomie Groups 26 Genus Gilbertsocrinus Phillips, 1836 38 FUTURE STUDIES 28 Gilbertsocrinus vetulus, new 38 SYSTEMATIC DESCRIPTIONS 31 Gilbertsocrinus aequalis, new 39 Subclass Inadunata Wachsmuth & Springer, 1885 .. 31 Gilbertsocrinus cassiope, new 39 Order Disparida Moore & Laudon, 1943 31 Gilbertsocrinus concinnus, new 39 IGroup Pentameri I 31 Genus Elytroclon, new 39 Family locrinidae Moore & Laudon, 1943 31 Elytroclon *elimatus, new 40 Genus locrinus Hall, 1866 32 Genus Ilematerisma, new lotrinus *subcrassus (Meek & Worthen), Ilematerisma *enamma, new 40 1865 32 Order Monobathrida Moore & Laudon, 1943 40 Family Myelodactylidae S. A. Miller, 1883 32 I Group Elliptici I 40 Genus Myelodactylus Hall, 1852 '2 Family Platycrinitidae Bassler, 1938 (1842) ... 40 Myelodactylus ammonis (Bather), 1893 32 Genus Platycrinites Miller, 1821 41 Genus Hyperexochus, new 33 Platycrinites? irroratus, new 41 Hyperexochus *immodicus, new 33 Genus Platystela, new 41 Plat ystela *proiecta, new 42 1 The University of Kansas, and Gin- Paleontohigical Institute, Genus Eucladocrinus Meek, 1872 42 sultant, Esso Production Research Company; 'Esso Poiduction Research Company, Houston, Texas Eucladocrinus? springeri, new 42 PACE PAGE Eucladocrinus? kentuckiensis, new 42 Pentacauliscus *nodosus?, new 52 Genus Platyparallelus, new 43 Genus Pentagonostipes, new 52 Platyparallelus *parilis, new 43 Pentagonstipes *petaloides, new 52 Genus Platyclonus, new 43 Genus Wanakastaurus, new 53 Platyclonus *dispar, new 44 Wanakastaurus *delicatus, new 53 Genus Platyplateium, new 44 Genus Petalerisma, new 53 Platyplateium *texanum, new 44 Petalerisma *eriense, new 53 Platyplateium sp. 45 Genus Pentagonomischus, new 53 Platyplateium pro videncense, new 45 Pentagonomischus *plebeius, new 53 Genus Platycion, new 46 Pentagonomischus *plebeius?, new 54 Platycion *mingusensis, new 46 Genus Asteromischus, new 54 [Group Cyclici I 46 Asteromischus *stellatus, new 54 Family Eucalyptocrinitidae Bassler, 1938 (1854) 46 Asteromischus *stellatus?, new 54 Genus Eucalyptocrinites Goldfuss, 1831 46 Genus Pentagonostaurus, new 54 Eucalyptocrinites? shelbyensis, new 46 Pentagonostaurus *leptus, new 54 Family Dolatocrinidae S. A. Miller, 1890 47 Genus Pentaridica, new 54 Genus Dolatocrinus Lyon, 1857 47 Pentaridica *rothi, new 55 Dolatocrinus exculptus, new 47 Pentaridica simplicis, new 55 Dolatocrinus avis, new 47 Pentaridica pentagonalis, new 55 [Group Varii I 47 Genus Pentagonopternix, new 55 Family Dichocrinidae S. A. Miller, 1889 48 Pentagonopternix *insculptus, new 55 Genus Cam ptocrinus Wachsmuth & Springer, Family Pentamerostelidae, new 55 1897 48 Genus Pentamerostela, new 56 Cam ptocrinus beaveri, new 48 Pentamerostela *delicatula, new 56 Subclass Articulata Miller, 1821 48 Pentamerostela minuta, new 56 Order Isocrinida Sieverts-Doreck in Ubaghs, 1953 48 Genus Floripila, new 56 [Group Pentameril 48 Floripila *florealis, new 57 Family Isocrinidae Gislén, 1924 48 [Group Cycylici I 57 Genus Isselicrinus Rovereto, 1914 48 Family Cyclopagodidae, new 57 lsselicrinus berm udezi, new 49 Genus Cyclo pagoda, new 57 Genus Austinocrinus de Loriol, 1889 49 Cyclopagoda *alternata, new 57 Austinocrinus mexicanus (Springer), 1922 .. 49 Cyclo pagoda costata, new 57 Order Millericrinida Sieverts-Doreck in Ubaghs, Genus Cyclomonile, new 57 1953 49 Cyclomonile *monde, new 58 Suborder Millericrinina Sieverts-Doreck in Genus Apiastrum, new 58 Ubaghs, 1953 49 Apiastrum *candidum, new 58 Suborder Bourgueticrinina Sieverts-Doreck in Genus Euloncherostigma, new 58 Ubaghs, 1953 49 Euloncherostigma *impunitum, new 58 [Group Ellipticil 49 Family Cyclomischidae, new 58 Family Bourgueticrinidae de Loriol, 1882 49 Genus Cyclomischus, new 58 Genus Bourgueticrinus d'Orbigny, 1840 50 Cyclomischus *shelbyensis, new 59 Bourgueticrinus alabamensis de Loriol, 1882 .. 50 Cyclomischus alternatus, new 59 Genus Dunnicrinus Moore, 1967 50 Cyclomischus tennesseensis, new 59 Dunnicrinus *mississippiensis Moore, 1967 .. 50 Genus Hattinanteris, new 59 Subclass and Order Uncertain 50 Hattinanteris *indianensis, new 59 I Group Pentameri I 50 Hattinanteris regularis, new 59 Family Dianthicoelomatidae, new 50 Genus Amsdenanteris, new 60 Genus Dianthicoeloma, new 50 Amsdenantey-is *tennesseensis, new 60 Dianthicoeloma *insuetum, new 50 Genus Rhysocamax, new 60 Dianthicoeloma monile (Eichwald), 1856 .... 51 Rhysocamax *cristata, new 60 Dianthicoeloma pentaporus (Eichwald), 1840 51 Rhysocamax grandis, new 60 Family Pentacauliscidae, new 51 Rhysocamax tuberculata, new 60 Genus Pentacauliscus, new 52 Genus Lomalegnum, new 61 Pentacauliscus *nodosus, new 52 Lomalegnum *hormidium, new 61 PAGEAGE Genus Stiberostaurus, nev, 61 Family Exaesiodiseidae, new 73 Stiberostau rus *aestimatus, new 61 Genus Exaesiodiscus, new 73 Genus Graphosterigma, new 61 Exaesiodiscus *acutus, new 73 Graphosterigma *scriptum, new 62 Exaesiodiscus truncatus, new 73 Graphosterigma synthetes, new 62 Exaesiodiscus minutus, new 74 Graphosterigma gram modes, new 62 Genus Scelidiopternix, new 74 Genus Cylindrocauliscus, new 62 Scelidiopternix *norops, new 74 Cylindrocaultscus *fiski, new 62 Genus Clematidiscus, new 74 Genus Cyphostelechus, new 62 Clematidiscus *denotatus, new 74 Cyphostelechus *claudus, new 63 Genus Exedrodiscus, new 74 Genus Catholworhachis, new 63 Exedrodiscus *excussus, new 74 Catholicorhachis *multifaria, new 63 Genus Ampholenium, new 75 Genus Blothronagma, new 63 Ampholenium *apolegma, new 75 Blothronagma *cinctutum, new 63 Genus Goniostathmus, new 75 Genus Baryschyr, new 64 Goniostathmus *annexus, new 75 Baryschyr *anosus, new 64 Genus Goniocion, new 75 Genus Cyclocaudex, new 65 Goniocion *gonimus, new 75 Cyclocaudex *typicus, new 65 Goniocion turgidus, new 75 Cyclocaudex congregalis, new 65 Genus Cyclostelechus, new 75 Cyclocaudex aptus, new 66 Cyclostelechus *turritus, new 76 Cyclocaudex plenus, new 66 Family Cyclocharacidae, new 76 Cyclocaudcx jucundus, new 66 Genus Cyclocharax, new 76 Cyclocaudex insaturatus, new 66 Cyclocharax *fasciatus, new 76 Cyclocaudex costatus, new 66 Cyclocharax modestus, new 76 Genus Mooreanteris Miller, new 66 Genus Cionerisma, new 76 Mooreanteris *waylandensis Miller, new 67 Cionerisma *exile, new 76 Mooreanteris pert oratus, new 67 Family Floricyclidae, new 76 Mooreantcris sp. 68 Genus Floricyclus, new 76 Family Euracidae, new 68 Floricyclus *hebes, new 77 Genus Furax, new 68 Floricyclus tvelleri, new 77 Furax *ethas, new 68 Floricyclus angustimargo, new 77 Eurax eu genes, new 68 Floricyclus kansasensis, new 77 Genus Pandocrinus StukalMa, 1965 69 Floricyclus pulcher, new 77 Pandocrinus stoloniferus (Hall), 1859 69 Floricyclus granulosus, new 77 Genus Avicantus, new 70 Genus Cyclocion, new 78 Avicantus *dunbari, new 70 Cyclocion *distinctus, new 78 Genus Desidiamphidia, new 70 Genus Plummeranteris, new 78 Desidiam ph idea *frondea, new 70 Plummeranteris *sansaba, new 78 Family Fluetieharacidae, new 70 Genus Lam prosterigma , new 78 Genus Flucticharax, new 70 Lamprosterigma *mirificum, new 79 Flucticharax *undatus, new 71 Lamprosterigma erathense, new 79 Genus Fabalium, new 71 Family Leptocarphiidae, new 79 Fabalium *fabale, new 71 Genus Leptocarphium, new 79 Genus Crenatames, new 71 Leptocarphium *gracile, new 79