DEVELOPMENT AND EVALUATION OF THE DIPLOTAXIS MURALIS (L.) DC (muD CYTOPLASMIC MALE STERILITY (CMS) SYSTEM IN SUMMER RAPE (ERASSICA NAPUS L.) A Thesis Submitted to the Faculty of Graduate Studies by TERESIO CAESAR RIUNGU in Partial Fulfilment of the Requirements for the Degree of Doctor of Philosophy Department of Plant Science December 1995 N,{onarLibrav Bibliothèque nationale I*I du Canada Acquisitions and Direction des acquisitions et Bibliographic Services Branch des services bibliographiques 395 Wellington Street 395, rue Wellington Otiawa, Ontario Ottawa (Ontario) K1A ON4 K1A ON4 Yout lile Votrc Élércnce Ou lile Nolrc rélércnce The author has granted an L'auteur a accordé une licence irrevocable non-exclus¡ve licence irrévocable et non exclus¡ve allowing the National Library of permettant à la Bibliothèque Canada to reproduce, loan, nationale du Canada de distribute or sell cop¡es of reproduire, prêter, distribuer ou his/her thesis by any means and vendre des copies de sa thèse in any form or format, making de quelque manière et sous this thesis available to interested quelque forme que ce soit pour persons. mettre des exemplaires de cette thèse à la disposition des person nes i ntéressées. 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Wffi}A UMI SUUECTIERM SUBJECT CODE Subiect Cotegories THE HUITT'\NIIIES SOCIAL SCIENCES (oilmuiltüiloNs AilD IHE ARTS 'TND PHII,OSOPHY, RII.IGION AND Anc¡61 ...............................0579 Archilecture ........ .........-.... -...-..-O7n ïHt0L0GY Mediwol ............................ 0581 Art Hisþrv................................0377 Philosophv.......... ...................... 0Á22 G¡¡erno ..'.......,....,............,......,. OSOO Relioioir ' Donce ......................................0378 ëe¡¡erol ..............................031 Fine Ärb .................................. O35/ .........................053¿ I Biblicdl stud;æ.................... 0321 lnbrmolion Scier¡c d..... -. -...,..,.... 07 23 ............................. 03¿0 Cl"rsy Joumolism................................ 0391 --............................ 03t 9 librory Scime ......................... 0399 Historv ot............................ 0320 F*rilæåphv of ......................0322 Ms Communicqlion¡.............., 0708 Thælosy .f ...i............................ Music.......................................O41 3 o¡ó9 Speecl¡ Communicofion .............0459 Vocotioml......... .......-............... o7Á7 so(lAl, s0tNcs Tl¡eots............... ... -. -..-..- -....-....Oß5 Amsicon Studiæ...................... 0323 IANGUAGT, UIERAÏURT AND TDUGÏIOII Anthrcælow uNGUtSil(s Aröo"p%s/ .......................0324 Lonorcæ Culturol .............. .... -...........O32ó ëeËrol ............ .................. 0679 Phwicql .............. ................0327 0616 Ancient............................... 0289 Busine'ss Administrolion ...........0617 Limuistics ...........................0290 Generol ..............................031 0 081¿ oÁ52 .........onz lvl¡ìÍlem .............................. 029¡ Accountinq ......................... 027 2 [ilso]ure Bonlring ..L.......... -...............o77o ..........oó88 Gsñ€rol ..............................o¿ol Mqnoq€m€nl....,,....-............0454 --........o275 Clqs¡col .....-...07?f - -. - --. - ^ -...........O8Á Mqrkãinq ...........................0338 ..........0518 Comp<¡rqtive ..........,............ Oæ5 Conodion Stüdiæ .....................0385 Mediwol ........... -. -.. -...........o?F7 Ecoæmic -..-...-..osu jt'lodem .............................. 0æ8 Gene¡ol .............................. 0501 Alricon ............................... 03l ó Agricu1turo1......................... 0503 Americon............................O591 CommerceBusiness .............0505 As¡'on .................................0305 Fimnce ..............................0508 Cqnodiqn {Enqlishl .............. 0352 His¡orv................................ 0509 Cqnodion {Frõrhl............... 0355 l¡bor'................................. 05l O Enq|ish ..............:................ 0593 Thærv................................051 I Gõmonic ...........................O31 I Folklore ..'.................................. 0358 Lclin Ànericon ........,.........,. 031 2 Gæqroohv............................... 03óó Middle Easlem .................... 031 5 Gero.-nlciloc¡v ............................. O35l Romonce ............................ 031 3 History Slwic ond Ecrsl Ph¡iccil..i................................. 0523 Europeon.....O3l4 Gámrql .............................. o5l8 IHE SGTENGES AND ENGINEERING Btotoct(Ât sffilGs Sp*h Pothobgy................0¿ó0 Engineering Aoriculture- Toxicoloqy .......................... 0383 Gelgol Ger¡sol ..........- -. -. -.--. ---.--....oo3 Home Ecmìíis ...................... 038ó .............0285 PHY5IGI SCTNGS ............................0¿2ó Pure S<þnces .............................04t I Clrmistry ............0359 Genríql .............................. o¿ss .......--..o478 Aqriolturol......... .-.-......-.....o7 Á9 .....-......oÁ79 AñoMicol ...........................04t¡ó .............0¿l 5 Biodímistrv .......................0487 lrcrqonic ..:......................... 0¿89 IIIAIÏI{ AND TNVIRONÍIITI{TAI Nucl6ø .............................. 0738 Orqonic...........................-.. 0¿90 sctltcs Phà'rmqcætico1.................... 049¡ Erwionmentol Sciencæ .............O7æ Phwicol .............. ...-............0494 Hælth Scio<æ Anqþmv Pofn¡r .............. ................O49 5 -....-...-. ---.-...--.-.....0287 Ge¡rerol .............................. 05óó Bioccdisirrca ......................... 0308 Ro8iotion ............ ..-. -...........O7 5Á Audiobcy........................... 0300 Mqthemqtie¡ ............................. 0¿05 8olonv................................ 0309 Cherptfrrqpv ................... ..'................. 0992 Phwie cdl .................o37e Denrisky ....:.:...................... 05ó7 ' Ecobcv ............. ..... -...........0329 Ger¡sol ..............................0óo5 Enþnì61ory....-.....-.-...-.--.... 0353 Educotián ...........................0350 AouCic ............................ 098ó l-lospilsl Monoqemsr¡!.,,.....,.07ó9 Ashomvond Gmt¡c .. - - - - - O%9 --.'.......... -. -....... Humon DevebÉnef,t ........... 0758 lí¡rurolocv .......... .. AsiophGics..................... OóOó -....--...-....0793 lñununoloqy ...'..................... 0982 M¡crot¡'ðfoqv ...................... O¿t O Arrrodpiic Science............ 0óO8 M€d¡ctm ãíd Surqsrv .........Osótr Alomi¿.................-...... -.......O7 Á8 lvlolerulor ::.............-.-........ 0307 Mgltol Hoolth .-..:...'............oU7 Neu¡osc¡ence ...................... Ele¿ronic¡ ond Electricity .....O&7 031 7 Nursim Geor¡oqrohy.................... 0¿l ó ..............................05ó9 Elemontol Põrticl€s onJ Nuhiño-n ............................. O5lO .......................... 0ó2t Phvs¡olõv :...:..................... o4¡3 Hiqh Enärqv..................... 0798 OËddrics nd Gynecoloqv ..0380 FluiJqnd Plãíma ................. 0759 nod¡qr¡ñ'............................ 0821 0 622 ôccuætimol Heíilh oJ' Mohculo¡ ...........................0ó09 ..................... -.... ldgimry Scienca...............O18 Tlxiropv ........................... ..........................oó20 |.............................0472 035¿ Nucleor .............................. 0ól 0 _. +"Þsr Oplrùol¡áobcv ................... 0381 .......................... 0 623 BþÐhEr6lrp|l>"id Optics ................................07 52 ..........................062l Pddþb<ry ...Ll.....................os2't Ródiotion ............ ................07 56 078ó Phormèôlogy .....................O¿l 9 ..........................0ó25 Medicol 0760 sol¡d sr"te ..........................0ól I ..........................0989 Phqrmqcy ................... -.... -..0 572 Srotistica ...................................0¿ó3 Phraicol theropv ........................... 03¿9 TÂRTH SqTH(ES ................. 0382 Publ¡c Heolh .].1................... 0573 Aoolied *ie¡res Bbqæch€mirtrv.. ...................... 0 Á25 Rod¡oloqy ........................... o5l4 Aååied Mæhonia ................... 034ó Geõdrenristry :..........-............ -..0n 6 RecrætËh .......................... o5l5 Cäinputer Science ..................... 098¿ DEVELOPHENT ÂND EVAI.IIATION OF l'trE DIPLOTAXIS Tfl'RAtIS (L. ) uc(¡nur) cyropr.Asurc HALE srtRrl,rly (cus) SYSIE}I IÌf SIIUHER R.APE (BRASSIcA xepus r.. ) BY IERESIO CAESAR RIIINGII A Thesis submitted to the Faculty of Graduate Studies of the University of Manitoba in partial fulfillment of the requirements of the degree of DOGÏOR OF PEILOSOPEY @ 7996 Perurission has been granted to the LIBRARY OF THE IINIYERSITY OF MANITOBA to lend or sell copies of this thesis, to the NATIONAL LIBRARY OF CANADA to microfilm this thesis and to lend or sell copies of the film, and LIBRARY MICROEILMS to publish an abstract of this thesis.