Evolutionary trends in Oxfordian and Kimmeridgian Subbetic Aspidoceratinae (Southern Spain). A proposition of Null Hypotheses about the evolutionary course in n highly significant group of tethydian Upper Jurassit ammonites Departamenro de Gcologla y de Paiesntologia Unrve~s~ci~~cidr Granada K!:)' \V01</1\' - Ai~~t~orrrtt~r,hledrf~rrrl~t~on, Upper Jlrrurrrc tlfpprr Qxfnrrlrrlrr Kirrtr~/rrtiigBlrr),ii~altrtlon. .-111.Y'l Rrl(:?' - rErt ca\tenslve strrtEy of a large drtzount DJ il~pzd~~t~rusu rrtaterrirl from the st~bbericIJpp~xr furirsssc has been car- ~tr.J~r~t 011r p,ii~~zp,~li~~ttt hdr bi,c,~r EU ritahi~~h~~ZIC t'v~luf~~ndr~ contormatlons izus~~tion tht* rttodr)irrlt~onsof erlrlter tharuc. /crt tri~iltile rrrtorpor,rtro,i Q/ rh~~rgc.srind r~znov~itrorrsThrs r,rerbodology hrts allowed trs to propose st~vera! hgpotElrses strggestrng pr~,\il>!~sn"/rE(tolf$ cji~/~~g ~ht-sc basic rvoluttonary coniorrndtrons ,ind the ejtabfrshrtry uf rxuolfdrfoifary~rrrdmczes 1 h&, /orrnzll'airun of th1.b~ jmprtlll~c.~t 1~~~0~lwrr~r,r,rparlrllue cfiir.t[u~tronp~,rmrts iir to present dedrrctrvc IIZOJL~Stt~5zch represetzf tbr oplnrorr of the .lrrlho/~ IC#~//I )t'jildrc/ /er [he L'UO~IILION',I~ /rLrr~ti'~o~~X'O/ I~C g)r,ttp ~\~/IlltiL'rll'hcsc pdftrrcls ~ICJC* fir 1hcp flld! h~lpolhesc., which rurJl br /c.~l~.rl111 fbr t8rx.f ~arrrhrrrgpbilrrs - spcSrres trrafmcrrl ~'rflwr(oral und/or niotloyrapblr i%fdrrtover,tbev rt~/plgnuut rt*la- trorrj dir~totr~~~pecic~ tJ they arc ~otzic~uedIN ti trrltIrtlo~tiI ~JI~W.The contltlsrorr~reached rlrr the /rr,i' bypothrsts on strattgraphtcul6y c-oi!c ( t~'i/r/~~,iirr~,rran~wn n n~prdocerus u. Kl.~Ii\IIi"\"lC) -- ['l'cn~lcn~ccvolutive neglr H :irlslridocerds R dcl1Y)xfurdidno c Krnlmerrdyrdno subtxitcu (L&>J~I~Jtilerrdro- ~r.~lct 1'ttrprrst.i tl~a llull hypothrscs D sur nrndelli evolt~tivitl~ tin importanre gruppo di ammoniti del Grur.issrco s~il~"r8u~re"j- LO J<OPO </L'![fiijef~lt! lclunro e [a fortiuiuzloar /It trrtr~ a rtrrll hvpothc.rrs # sul con~purlnrtzmto evoErtl~uodrgit c Asprritrter~s*, duiu- r/lt'tt/irfrt rlrillcl ii~sti~~zo~~~n~u~OSSIIL* La rrcercir P srata cumprufa rt~riirlr~zdorm'r collrzron~dr crtca 1.000 tndivrdrtr i umpmndtr Jrlglr rluforr rrellu Lorw S'r~hb~atzca(Spdyna merr,fiavaalr). Tutto 11 ~n/rtrttulrC? rluto sottoposto ad trn precrso co~rrrollostrirrrgrdfilo, e rr trorad rirpos~t~topre~~o zI Dtpart~~t~eniodi P~leontologia deb$'Utzrurtstt& dt Grgnoda (Spagrrai. I)opr, 'irJcprrrpplrt uto te~nrcbestutisi~cbr al rurutfert mrsrtrtrbzll, sono sfate riconoscritrr quenr che nor abbrdtrti-, brlttcxir~to u ~M.Z~L~'vojt~t~~mdry confoltnstions n, stablllre tra l'rnsierne dtr cdrirtterr rredlrarr Per 1i1 loro rappresenlazione graJiin svtru sldtr rrr~!r::,rt, a block-d~ugrrirt~su trrrlr~~~enrior~ull,tzer qrtali l'allezzu corrlspontlc ulla Jtsfrrbuzrorie strulrgrufica uccertata 1 ra I rirurrsl Jragrar~zrrirr rrrerenft le basic rvolurionary conformarions D sr P tercato dt trovure iporebt di relarioni eucriutrrle, Jr /t rtc~rt,rtrrlo qrielle L he L~VI~C*~ZI~(Y~NO/P YP!J:IOI~I p11; drrettc, L 10; le I~O~PSZprri erono~ni~Aescientifi~atnenfe. La decrstrsn~ rEr ~nrbrrrrtrrek rttterprctuzrorrt o le rpotesr prri cornptrssr potrebbe scr~zbrarr rl r~strltafodi una certu greferentu per I modi"II~ grr- rlrriilrrtrtr, tr~ir rron i. yut!sto ;l c-aso, conszderuto che ?ran P statcz valzitata la contlnuttri dei processi euolutrvt. In reoltd In rrostrd pr~~tc~'~in ttrrtit i/~t<lfrt</I ro~rtirlerureyu,~lunyrrc rntr,rpretazrone L-Z~O~UIII~~Idr I/NC#JIO grfippo IN uw contrsto, inirrale, in act ordo con lir prc~~zllrht!~*~t~tbrlttd J~*El'rlrtlhzi'~te ecoI~gico, e 112 cancorriett~rzcot1 t lrttrdtr iit fol&.rdnra presupposti per guesta firms: turto yae- sto Z rldlo ronsrdcrdlo uccettubric rn quej/u fuse dell,/ nostra ricerca. 1 diversi gruppt di S basic evolutionary conformations w, relo- zrc~~utttru lc~ro,e dtstrrrtr srrfernatfcanrente a livello generico, ci dunno Ir B ntrfl /r'lypothesrj M do cornprorare, studrandde nlano- ~~i((iid~l.t'fll~nel ff~versr Jettort plrkeogengrafict, o nteglro, in base ad un nrodeilo generule che d permetta dt rntegrare qzrestr stud; !.it t~etodoioglosegrrtta hu farto errzergere tenderize a cantbiantenti morfolog~crncl tempo, ell'mterno dell'insieme degli r Aspr- docsrus medtterranet dell'0xfordiano e Kirnrneridgiano subbetrco I rrsulfatr debbono essere considerafi come derzuatr du untr [rlfrrra direttn della dacunzentazzone fossile, e le conclusioni derivanti possnno cos? riussumersi. 1) Necessitri di creme nlrove con- nersront interspecrfiche se si conridera Ea sistematica tradrzionaae appiicafa zn questo grsrppo 2) Duramte il Kimmeridgtano e Z'Ox- ff~rdtdnosuperrore, oue ha origme, il genere Aspidoceras svduppa mu Itma euohtiua in cul persirte fa brtube~ularioneduranre l'ontogenesr, IN utta secondu lrnea si mantfesta la tendenza alta inuncmra piO a ttieno tnrdrva della sene di rubercoli esterni. 5) I'se~iciounayen~os~rlrrppu Irr itiu rrror~ologrupri, euolutir Jcl K/l,r~f/c~rrr/,~~i/~rorr/cr/to e ritpcrrorc fra /'msremr dt /lortur irrf ~r.wpin omhclito c irrpgnlizrt,rerrtr hrtt~hrrrcrlcri~,rhe orzgrnilrJa pli l-Iyhi-incrt~ccrasdrl yrrtppn prcssulum NEIINIIIY~4) I'hysoclocctas r2 Qtthns- p~docer.ia qlllPlQONO irl Ioro IISNIEII?;~ruiiuppo durante I! Krrritr?o.rllgrorro ttr/crrorr Le droi'rsrfirirzron~sr pmd~icotro a pcrrtrre dol- !'t~trt.nrr~rrtotit~llc drrrtcrrrrotrr r tit[ nti??fcvo der frrhenolr, cd irrrt!3cs prr ~'irii~rrrrrrronedr serlnfrr ptir ~.rrnlorrr/ofcNt!l~ (1~1er1 dr r)rtrrmrc n/icrtortc p~ir fnr~t~t3 n1.r gtrr r~rjlrtmirrnrr lir ilrrxc rsrfrctizrolrc ric't~i//rr,per&, la dit~rrbtrzrnrrc rfrntlgrrr~itai. rrrr'oftd 5 ~Vc*i' ~'O~/r~rcf~~rno~err~trno/'r~ c K~t?~rr/crr(f~runortztzr'd~~ izhbrdrrro trttrf~//(~/ortttL, qt<jhr~<(',ttr ctt~~<Ivz~I/:<~ sr~t(~~~/</ftcd rtr~1uf7e (t~rt<~rrr r C rlri (1) (;r rii~ii!/r/r.n/i, trc rrrrjrcr stoc t/t c n~/tgtrrd;ror?c prrm!/ir !.a/,firti;11 K!f?ittitF~~~RIIIII~I trlfr't for<' 7) I rt<lllr rli dfc't I 5ljrc it:~ol?f' deglr c. ,/lrjacJoitar,rr 8, -it prcrerzrd ncl Krmnrerrdgtano irt/crsore, rrr.ilc :rtric. a \,rr,/rrhcrli cz I)~cr\rtrrr X) Ucllc ict ltrrt-c r-t~idrrtrirc*chc trrprriznn rl h'r?tfmrrrriiqrir~t,r~nfr~rrorr, drre sr orrgtnano nellii zonu ~r \trt~lnht'r-kieiI 11 rcrtr) trcsil* ZONII (I U~I~IJIIIN9) IN ~t~ficIc IINL,L~ 1 rntremcnto drlld @'rrttd~'2;ci tr rnan6jerta per Iff prtma uolla rfcblki:orrii rl \rrornhtrt kr, yricilo jerzolrieno ctrlfr si rrpl'lr IIC~~/~l~ftei. I'VOIN~~~Y*rnolto cfrr>cr~rfi<nlc10) ltle~tr Aspidoccras. Psrud~>\\~aagcn~av l'hysodoccras, !C{ /c*udenza irll'i~rtmenlo d~/l'~J?lJrt nrnnrJt star rn trcti'e /e Irrrec crrcrif~trtre.f l) iVel E;rnrnrertd.q~anunrprrrore 11 ~er?creAS~IL~OC~T,I\ ~1'1Ir1ppiz l~brrco/f p~*rtr~~!//~~'~~tri[i EJ LW ronfi~urnztone& m' an3p1a base e orrrntaziotfe rridrclie o retrcltgersn RlJ,\tliMEN - El ohlettvo de cste trcrblrto ha srdo [a arjiunrrntucrrj?~ dc nrrrn * hzprjtests iz~li~B r(>hre el i~ottcportrirrtrettro evol~trtto rie Ios a Asptdocerasia medtterraineos basada en eE registlo I'rjr~I Lu tnz~rrilgcrrzrjnSF ha rmlr:adi, sobre rrnzl rtrrrcstrn de ~rt~rout??riidnt?rctrt~fOOfl tndrotdtios, retofccfutfcrr por (0s at/torcr, c'rr [ir /oft# luh/iL:l:iCi(f-rp~ctii? ~ncrrtlrf~trii/) i cr~tritrmEo titd tnirfcrrrr! llir 11110 rjhlrnriltr) bnjo tin erfrrrfn cofltrot' rrtfutrgriilrto, v iCz c~nirirntnrrir.poii(crtlr> <,rr (.I Drprr/ftrr/.r~n/orlrx I'Nl'r.m~~fdnfiin dr. la Iittrrtc>rrdi/<lrlc, Clrtinrcda 0:spoAu) Trrrs lo irpl,rtrcl<ir~ili. r,ri/rrmt~rttr,r r qtedir/rcor 11 lor rrirai /rrc3r mencrtrtihlec (I' llrils re tr?nrrtitlo IN+ rlrlrri clcrror~1rvtrrilr1~Estrticturscr~nes evolutrvas hdkicas rcgrr~nrf,re(fir erttre tonlrt~rtr~~(/L* <,itrrt[csrc t htrc.c/rtti~\crr ~i/ctt (I(;FZ '1 /'l3 II?NUVUlIfJWCi ~fl'(~'f~ildd~ I'cfrrr trr rc~pvrrc~nlarriitrgrifira re hnn rcfr[irtrdo hlcfrfrrr,r /r~i~trtr~nrro?r,i~r*serr [or r{rrc. 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