!"#$$%&'()*+!"#$%&'%(()%(#*!%+", -!%+",!./0123405652!52!789!:6;<9!=!>7?@A!! B77C!"(",="D,E!$F30/G&!H!B77C!%D"I=,+E,!$9/<0/2& 590'!JGGF'HH5KL590L93MH"+L""I+IH044/L%D"I=,+E,L1"#0%F%((=%(# !"#$%&'#(()*+',%+#* !"#$%&'(")*+',+-."+/'0#1)20+"0#"3$&+!"#$%&$%''(') '*+,'%-"(')4561232-27+ 8$9:';:'%%$#2"<=+>$-.+ description of the frst known juvenile specimen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loque de Chortís, diploglossid, distribución geográfca, Mesoamerica, morfología. A2:2()2%?&.2("7+ América Central, Bloco de Chortis, diploglossídeo, distribuição geográfca, (+&,+/+0G%9 S15&%).#* %)F* T./3+)* F#31&.U#F* ';#* /.V%&F* L&#(+)'%)#* S+.3'* %)F* ^+Z#&* S+)'%)#* T#'* !"#$%&$%''(')W*)+,$,'-('X)'./0'%1"(')WS15&%).#* M+&#3'*.)*"#$'+9*_+&+4*J+)F:&%34*&#$&#3#)'3*';#* %)F* T./3+)4* !==>X) W2Y:%(%'%D* ".$/+0/+33.F%#X* +)/8* (%'#&.%/* +,* ';.3* 3$#1.#3* #7#&* 1+//#1'#F* U%3#F* +)* 'Z+* %F:/'* ,#(%/#3* 1+//#1'#F* )#%&/8* %* WS15&%).#*<Q!RX9 F#1%F#* %$%&'9* [;#* $%&%'8$#4* 1+//#1'#F* .)* !=R?4* `)* !!* a:)#* <Q!P4* )#%&/8* <@* 8#%&3* %,'#&* ';#* .).'.%//8*Z%3*%33.0)#F*'+*+,$,'-(')2"3"--/-('4)U:'* 3$#1.#3* Z%3* /%3'* ])+Z)* '+* ;%7#* U##)* 1+//#1'#F4* %,'#&* %* 3#1+)F* 3$#1.(#)* W';#* ;+/+'8$#X* Z%3* Z#* ,+:)F* %* b:7#)./#* !"#$%&$%''(') './0'%1"(') 1+//#1'#F* .)* !==?4* ';#3#* 3$#1.(#)3* Z#&#* WM.0:&#*!X*.)*';#*/%'#*%,'#&)++)*+:'3.F#*%*;+:3#* F#31&.U#F* %3* %* )#Z* '%\+)* U%3#F* +)* 7#)'&%/* %U+7#*B/*c%$+'#4*)#%&*';#*3+:';Z#3'#&)*#F0#*+,* 1+/+&%'.+)* %)F* ';#* ):(U#&* +,* ,+:&';C'+#* %)F** L%&Y:#* H%1.+)%/* L.1+* L.b+/4* "#$'+9* _+&+4* third fnger lamellae (McCranie and Wilson J+)F:&%39* [;.3* /+1%/.'8* W!Od!QeQ=9QRee* H4* !==>X9* [;#* '8$#* 3#&.#34* 1+//#1'#F* ,&+(* 'Z+* RPd?Pe<>9<=ee* T4* Tf2R@* F%':(g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`';#&*&#$'./#3*,+:)F* .)* 38($%'&8* ';%'* ).0;'* .)1/:F#F* 60%$"'* Wh* 7%1%#'X) 8%1%,0'"'* WS15&%).#4* H.1;+/3+)* %)F* ij;/#&* <QQ!X4* 9,#-%:,"1/) ',#-,0-1"%0/$"'* Wi#)).1+''4*!RO=X4*;</:"0,$$/)="0=,$"0"*W-+#''0#&4* !R=RX4* %)F* >.,$%#%1(') <%0:(1,0'"'* S15&%).#4* <Q!R9* [;#* 3$#1.(#)* Z%3* F#$+3.'#F* .)* ';#* 5%&)#0.#*S:3#:(*+,*H%':&%/*J.3'+&8*2#1'.+)*+,* J#&$#'+/+08* 5+//#1'.+)4* L.''3U:&0;4* LA4* 62A* (CM 163365; feld ID CLP2542). CM 163365 is the frst juvenile and third ])+Z)*3$#1.(#)*+,*!"#$%&$%''(')'./0'%1"('?)N)* !"#$%&'()! "#$%&'(%!!"#$%&$%''(')'*+,'%-"('))*+!,-..-/0! '#&(3*+,*;#%F*%)F*U+F8*31%/%'.+)4*.'*.3*1+)3.3'#)'* '&!('1%2 Z.';* ';#* +&.0.)%/* F#31&.$'.+)* +,* S15&%).#* %)F* T./3+)*W!==>X9*2)+:'k7#)'*/#)0';*.3*?R*((*%)F* ';#*'%./*Z%3*@O*((*%'*1%$':&#*,+&*%*'+'%/*/#)0';* 5+/+&*.)*/.,#*.3*3;+Z)*.)*M.0:&#*!9*[;#*1+/+&* +,* R?* ((g* ;+Z#7#&4* <Q* ((* +,* ';#* '%./* '.$* Z%3* %,'#&* !?* (+)';3* .)* $&#3#&7%'.7#* .34* %3* &#(+7#F* ,+&* '.33:#* 3%($/#3* $&.+&* '+* ,+&(%/.)C ,+//+Z3D* (.FF+&3%/*3:&,%1#3*F%&]*0&%8.3;CU&+Z)4* fxing the specimen. There is a single prefrontal with a pair of well-defned, pale bluish gray ';%'*.3*U&+%F#&*';%)*/+)0g*'Z+*&+Z3*+,*.)'#&)%3%/34* dorsolateral stripes originating on the frst pair of Z.';* %)'#&.+&* $%.&* .)* 1+)'%1'* Z.';* ';#* )%3%/* %)F* .)'#&)%3%/3* %)F* #\'#)F.)0* +)'+* ';#* '%./4* Z;#&#* :$$#&* $+3')%3%/4* %)F* $+3'#&.+&* $%.&* .)* 1+)'%1'* ';#8* ,%F#* %)F* U/#)F* .)'+* ';#* 8#//+ZC+&%)0#* '%./* with the upper postnasal and frst two loreals; the coloration; upper 50–90% of most supralabials )%3%/* .3* .)* 1+)'%1'* Z.';* ';#* &+3'&%/g* ';#* )%&&+Z* F%&]*0&%8CU&+Z)4*.&&#0:/%&/8*U+&F#&#F*Z.';*$%/#* ,&+)'%/* .3* %$$&+\.(%'#/8* <9O* '.(#3* /+)0#&* ';%)* 1&#%(* '+Z%&F3* /+Z#&* #F0#34* Z.';* %* F%&]#&* 0&%8* Z.F#4* .)* 1+)'%1'* Z.';* 'Z+* (#F.%/* 3:$&%+1:/%&34* U+&F#&* %/+)0* $+3'#&.+&* #F0#3* +,* #%1;* 31%/#g* ';#* ,&+)'+$%&.#'%/34* %)F* ';#* .)'#&$%&.#'%/g* $%.&#F* 3:$&%/%U.%/3* P* %)F* R* %/(+3'* 1+($/#'#/8* 1&#%(* $%&.#'%/3* %&#* 3#$%&%'#F* U8* ';#* .)'#&$%&.#'%/* %)F* 1+/+&#F4*Z.';*F%&]*0&%8*3:&&+:)F.)0*.)'#&7#).)0* .)'#&+11.$.'%/g*';#*.)'#&$%&.#'%/*.3*%U+:'*';#*3%(#* 3:':&#3g*/%'#&%/*3:&,%1#3*F%&]*0&%8CU&+Z)4*3/.0;'/8* 3.V#* %3* ';#* $%&.#'%/4* 3/.0;'/8* /%&0#&* ';%)* .)'#C /.0;'#&* ';%)* (.FF+&3:(4* Z.';* 31%''#&#F* $%/#* &+11.$.'%/9*[;#&#*%&#*?C?*/+&#%/3g*OCO*3:$&%+1:/%&3g* U/:.3;C0&%8* 3$+'3* U#1+(.)0* (+&#* %U:)F%)'* !<C!!* 3:$&%/%U.%/34* Z.';* 3:$&%/%U.%/3* P* %)F* R* 7#)'&%//8*%)F*%)'#&.+&/84*%$$#%&.)0*(+3'*$&+,:3#* F.&#1'/8*U#/+Z*';#*+&U.'*+)*U+';*3.F#3g*%)F*!QC!Q* %)'#&.+&* +,* ,+&#/.(U3* %)F* %Fb%1#)'* '+* 7#)'&%/* .),&%/%U.%/39*[;#&#*%&#*>=*F+&3%/*31%/#3*%/+)0*';#* coloration; infralabials 25–50% dark gray on F+&3%/*(.F/.)#*,&+(*';#*.)'#&+11.$.'%/*'+*';#*/#7#/* Z;.'#* 0&+:)F* 1+/+&* %)F* F%&]* $.0(#)'* 1#)'#&#F* +,* $+3'#&.+&* #F0#* +,* ';.0;g* @>* 31%/#3* %/+)0* ';#* %&+:)F*3:':&#3*U#'Z##)*31%/#3g*7#)'&%/*1+/+&%'.+)* F+&3%/* (.F/.)#* ,&+(* /#7#/* +,* %\.//%* '+* /#7#/* +,* .((%1:/%'#* 1&#%(CZ;.'#g* /.(U3* U&+Z).3;C0&%8* 0&+.)g* %$$&+\.(%'#/8* >P* 7#)'&%/* 31%/#3* %/+)0* %/+)0*(+3'*F+&3%/*$+&'.+)34*(+''/#F*/%'#&%//84*%)F* midline from frst pair of chin shields to the 8#//+Z.3;CZ;.'#* 7#)'&%//8g* 7#)'&%/* 3:&,%1#3* +,* %)'#&.+&* #F0#* +,* 7#)'g* @<* 7#)'&%/* 31%/#3* %/+)0* ;.)F* /.(U3* 3+(#Z;%'* F%&]#&* 8#//+Z* ';%)* (.F/.)#* U#'Z##)* ';#* %\.//%* %)F* 0&+.)9* ".0.'%/* ,+&#/.(U34*.)'#&(#F.%'#*.)*1+/+&*'+*,+&#/.(U3*%)F* claws unsheathed; third fnger with 16-15 '%./g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his is the frst report of terrestrial activity for 1+)3.F#&#F* %&U+&#%/* (.0;'* ;%7#* %* '#)F#)18* this species, the specifc epithet of which, '+Z%&F3*'#&&#3'&.%/*;%U.'3*%3*b:7#)./#34*+&*$#&;%$3* l'./0'%1"('4m) (#%)3* l+,* 1/.(U.)0m* WS15&%).#* ';#* $%:1.'8* +,* +U3#&7%'.+)3* +,* ';#3#* &%&#/8* %)F*T./3+)*!==>D* <><X9*[+Z)3#)F*,-)/$?*W<QQOX* #)1+:)'#&#F*3$#1.#3*;%3*U.%3#F*+:&*:)F#&3'%)F.)0* &#$+&'#F* '#&&#3'&.%/* %1'.7.'8* .)* %* b:7#)./#* +,*';#.&*U#;%7.+&*WK%:0;%)*%)F*-&+Z)*<QQ@X9 !"#$$%&'()*+!"!#$%&'(!)*+*,-*.!&/#0 !"$ @%5A/00)#'*%/9 Debate continues about the classifcation of !",")"0&"% F.$/+0/+33.F3*.)*';#*Z%]#*+,*L8&+)*,-)/$?B'*W<Q!?X* fnding that the subfamily Diploglossinae is -+1;%'+)4*594*p9*-+.3'#/4*M9*5%3%0&%)F#4*29*f&+:%&F4*%)F*29* $%&%$;8/#'.14* %3* +,$,'-(') %)F* !"#$%&$%''(') %&#* -%./+)9*<Q!>9*A*,+33./*!"#$%&$%''(')W2Y:%(%'%D* A)0:.F%#X* /.V%&F* ,&+(* -%33#C[#&&#* %)F* f&%)#C[#&&#* N3/%)F3* $%&%$;8/#'.1*Z.';*&#3$#1'*'+*U+';*#%1;*+';#&*%)F* Wf:%F#/+:$#4* M&#)1;* T#3'* N)F.#3X9* Scientifc Reports ';#* 0#):3* C#<"%:,'?* A/';+:0;* ';.3* Z+&]* .3* ';#* EF* <R@PO9 (+3'* 1+($&#;#)3.7#* (+/#1:/%&* F%'%3#'* +,* ^+'V]%'4*294*A9*J#&'V4*%)F*f9*ij;/#&9*<Q!>9*A*)#Z*3$#1.#3*+,* F.$/+0/+33.F3* '+* F%'#4* ';#&#* Z#&#* 3#Y:#)1#* F%'%* +,$,'-(') W2Y:%(%'%D* A)0:.F%#X* ,&+(* Z#3'#&)* L%)%(%9* ,+&* +)/8* >* +,* ';#* O<* F#31&.U#F* 3$#1.#3* +,* G,'%/A,1"./0)@,1#,-%$%&8)HF* =><k=PO9 ".$/+0/+33.)%#)W6#'V*,-)/$?*<Q!=X?)-%3#F*+)*';.3* S15&%).#4* a9* p9* <Q!O9* A* 1;#1]/.3'* +,* ';#* %($;.U.%)3* %)F* fnding, McCranie (2015, 2018) considered &#$'./#3* +,* J+)F:&%34* Z.';* %FF.'.+)34* 1+((#)'3* +)* +,$,'-(') './0'%1"(') '+* U#* .)* ';#* ,%(./8* '%\+)+(84* 3+(#* &#1#)'* '%\+)+(.1* F#1.3.+)34* %)F* %&#%3* ".$/+0/+33.F%#* %)F* ';#* 0#):3* !"#$%&$%''('4) +,*,:&';#&*3':F.#3*)##F#F9*I%%-/J/)HKHLF* ?O<k?R>9 )+'.)0* ';%'* !"#$%&$%''(') .3* %)* +/F#&* %7%./%U/#* S15&%).#4*a9*p9*<Q!R9*[;#*/.V%&F34*1&+1+F./#34*%)F*':&'/#3*+,* )%(#9* S#%)Z;./#4* +';#&* Z+&]#&3* ;%7#* &#'%.)#F* J+)F:&%39* 283'#(%'.134* F.3'&.U:'.+)4* %)F* 1+)3#&7%'.+)9* ';#*:3#*+,*A)0:.F%#*%)F*+,$,'-(')W#9094*-+1;%'+)* M($$,-"0) %5) -<,) G(',(A) %5) +%A#/1/-"3,) I%%$%&8) ,-) /$?* <Q!>4* ^+'V]%'* ,-) /$?* <Q!>4* S#3+%(#&.1%)* LNF* !k!<=9 J#&$#'+/+08* [%\+)+(.1*