Annotated Checklist of the Vascular Plants of the Washington - Baltimore Area Part II Monocotyledons Stanwyn G. Shetler Sylvia Stone Orli Botany Section, Department of Systematic Biology National Museum of Natural History Smithsonian Institution, Washington, DC 20560-0166 MAP OF THE CHECKLIST AREA Annotated Checklist of the Vascular Plants of the Washington - Baltimore Area Part II Monocotyledons by Stanwyn G. Shetler and Sylvia Stone Orli Department of Systematic Biology Botany Section National Museum of Natural History 2002 Botany Section, Department of Systematic Biology National Museum of Natural History Smithsonian Institution, Washington, DC 20560-0166 Cover illustration of Canada or nodding wild rye (Elymus canadensis L.) from Manual of the Grasses of the United States by A. S. Hitchcock, revised by Agnes Chase (1951). iii PREFACE The first part of our Annotated Checklist, covering the 2001 species of Ferns, Fern Allies, Gymnosperms, and Dicotyledons native or naturalized in the Washington-Baltimore Area, was published in March 2000. Part II covers the Monocotyledons and completes the preliminary edition of the Checklist, which we hope will prove useful not only in itself but also as a first step toward a new manual for the identification of the Area’s flora. Such a manual is needed to replace the long- outdated and out-of-print Flora of the District of Columbia and Vicinity of Hitchcock and Standley, published in 1919. In the preparation of this part, as with Part I, Shetler has been responsible for the taxonomy and nomenclature and Orli for the database. As with the first part, we are distributing this second part in preliminary form, so that it can be used, criticized, and updated while the two parts are being readied for publication as a single volume. Corrections and comments are welcome. We hope that our now-completed Checklist stimulates much new fieldwork to check the current status of the local flora relative to what is being reported by us. The complete Checklist is available on our Web Site for the Flora of the Washington- Baltimore Area (www.nmnh.si.edu/botany/projects/dcflora) in pdf format and as a searchable database. The Herbarium of the Washington-Baltimore Area (“D.C. Herbarium”), consisting of nearly 60,000 preserved plant specimens collected over the past 125 years or so, is also accessible at this site. It has served as the basis for the Checklist. The Web database provides the label information for the specimens, distribution maps for the species, and, in many cases, a link to an image. We gratefully acknowledge the help of many persons. We are indebted in the first place to Carol Annable, ChristineBegle, MarcieBeyersdorfer, Laura Lehtonen, andSusan Wiser--all former assistants of Shetler--for their vital help in compiling the original working materials, including the initial list. In particular, Christine was largely responsible for compiling the working list from Hitchcock’s and Standley’s Flora (1919) and Hermann’s Checklist (1946). Help with the grasses was provided by Paul M. Peterson (especially, Agrostis, Bromus, Cenchrus, Cynodon, Muhlenbergia) and Robert (“Rob”) J. Soreng (esp. Festuca/Schedonorus, Hordeum, Poa). Rob reviewed the entire draft of the Poaceae and offered many useful comments. In preparing their recently published Atlas of Carex for Maryland and the District of Columbia (2002), Christopher T. Fryeand Christopher Lea examined all of ourCarex specimens and annotated many. Our treatment closely follows theirs, and we thank Chris Lea in particular for responding to many questions. Robert R. Haynes kindly annotated all of our Najadaceae and Potamogetonaceae, and his annotations guided our treatment of these families. Likewise, we were fortunate to have Larry J. Davenport annotate our Lemnaceae, providing the basis for our treatment. Mark Strong iv revised and annotated our collections of the species formerly treated broadly as Scirpus, and his treatment is followed. He also annotated other Cyperaceae and some Juncaceae. Dan H. Nicolson continued to give of his time and expertise to solve nomenclatural problems. Finally, we thank the BotanySection ofthe Department ofSystematic Biologyand especially W. John E. Kress, Head of the Section, for their continued encouragement and support of this project; and the Virginia E. Crouch Memorial Fund for Native Plant Conservation and Research, administered by Larry E. Morse of Natureserve, for partial support of the printing and distribution of Part II. Stanwyn G. Shetler Sylvia Stone Orli Department of Systematic Biology - Botany Section National Museum of Natural History Smithsonian Institution Washington, DC 20560-0166 [email protected] August 2002 v CONTENTS Preface........................................................................................................................................ iii Introduction................................................................................................................................ vii Geographic Coverage..................................................................................................... vii Taxonomic Considerations............................................................................................ viii Revision Procedures...................................................................................................... ix Primary Sources............................................................................................................. ix Principal References...................................................................................................... x Format of Species Entries.............................................................................................. xi Excluded Species........................................................................................................... xiii Statistics......................................................................................................................... xiv Web Site for Flora of Washington-Baltimore Area....................................................... xiv Annotated Checklist................................................................................................................... 1 Key to Abbreviations of Authors............................................................................................... 53 Literature Cited.......................................................................................................................... 73 Index of Common and Scientific Names................................................................................... 83 vi vii INTRODUCTION by Stanwyn G. Shetler A brief history of floristics in the Washington-Baltimore was presented in the Introduction to Part I. The procedures and format for the Checklist, given in Part I, are repeated here for the convenience of the user. There are three innovations in this part that should make it easier to use than the first part. Running heads indicate the family or families treated on the pages, common and scientific names are in the same index, and genera with more than 15 species, i.e., genera likely to run to a second page or more, are indexed to species. Geographic Coverage As indicated in the Preface, our Checklist is based in the first instance on the D.C. Herbarium. The limits of the D.C. Herbarium, as expanded in the 1930s and 1940s for the Conference on District Flora, are the limits of the “Washington-Baltimore Area” as defined by Hermann (1941, 1946) in his Checklist and also used here. This Area (see map inside front cover)is bounded on the north by the Maryland-Pennsylvania line, on the east by the Chesapeake Bay (western shore), on the south by the Rappahannock and Rapidan Rivers, and on the west by the east foot of the Blue Ridge Mountains. It includes the Bull Run and Catoctin mountains, but excludes Shenandoah National Park and the Eastern Shore. Encompassed are the District of Columbia and the following 23 counties and 5 independent cities in Maryland and Virginia: Maryland Counties: Anne Arundel, Baltimore, Calvert, Carroll, Charles, Frederick, Harford, Howard, Montgomery, Prince George’s, St. Mary’s. Cities: Baltimore Virginia Counties: Arlington, Culpeper, Fairfax, Fauquier, KingGeorge, Lancaster, Loudoun, Northumberland, Prince William, Richmond, Stafford, Westmoreland. Cities: Alexandria, Fairfax, Falls Church, Manassas So defined, the Washington-Baltimore Area represents a radius of coverage from the Capitol in Washington, D.C., of about 50 miles, more than three times the 15-mile radius of the previous Floras. In compiling the Atlas of the Virginia Flora, Harvill et al. (1992), included the independent cities within counties for the purpose of mapping specimen distributions. Thus, Alexandria viii specimens were mapped as though from Arlington Co., Fairfax and Falls Church specimens as though from Fairfax Co., and Manassas specimens as though from Prince William Co. It is important to keep this in mind when voucher herbarium specimens from these cities are checked against the Atlas. Taxonomic Considerations The Checklist includes all native and naturalized species of vascular plants known to occur or to have occurred within the defined geographic area. Naturalized species are alien (exotic) species introduced into the area that are now believed to be reproducing and spreading in the wild without cultivation if only on a limited scale. A few species are included that appear to spread only as clones but persist indefinitely