OF ILLINOIS LIBRARY AT URBANA-CHAMPA1GN BIOLOGY FIEL Botany NEW SERIES, NO. 27 PTERIDOPHYTA OF PERU Part IV 17. Dryopteridaceae Rolla M. Tryon Robert G. Stolze With the collaboration of: John T. Mickel Robbin C. Moran "Mr 21 February 28, 1991 Publication 1424 PUBLISHED BY FIELD MUSEUM OF NATURAL HISTORY Information for Contributors to Fieldiana General: / warily a journal for Field Museum staff members and research associates, although nsidered as space permits. i65.00 per printed page or fraction thereof. Payment of at least 50% of page which reduces the publication time. Contributions from staff, research authors will be considered for publication regardless of ability to pay page charges, however, the full i'.ed authors of unsolicited manuscripts. 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FIELDIANA Botany NEW SERIES, NO. 27 PTERIDOPHYTA OF PERU Part IV 17. Dryopteridaceae Rolla M. Tryon Harvard University Herbaria 22 Divinity Avenue Cambridge, Massachusetts 02138 and Department of Biology University of South Florida Tampa, Florida 33620-5150 Robert G. Stolze Associate Curator Department of Botany Field Museum of Natural History Roosevelt Road at Lake Shore Drive Chicago, Illinois 60605-2496 With the collaboration of: John T. Mickel, New York Botanical Garden and Robbin C. Moran, Missouri Botanical Garden Accepted August 10, 1990 Published February 28, 1991 Publication 1424 PUBLISHED BY FIELD MUSEUM OF NATURAL HISTORY 1 99 1 Field Museum of Natural History Library of Congress Catalog Card Number: 90-85755 ISSN 00 15-0746 PRINTED IN THE UNITED STATES OF AMERICA Table of Contents List of Illustrations 1 1. INTRODUCTION Ctenitis: C. submarginalis; C. ampla . 6 ACKNOWLEDGMENTS . 1 2. Megalastrum biseriale 12 17. DRYOPTERIDACEAE 2 3. Triplophyllum funestum var. funestum 20 Ctenitis . 5 4. Tectaria incisa var. incisa 22 Megalastrum 11 5. Cyclopeltis semicordata 28 Triplophyllum 19 6. Rumohra adiantiformis 30 Tectaria 21 7. Lastreopsis: L. effusa; L. killipii 32 Cyclopeltis 29 8. Dryopteris: D. paleacea; D. denticulata 34 Rumohra 31 9. Cyrtomium dubium 39 Lastreopsis 31 10. Didymochlaena truncatula 41 Dryopteris 35 1 1 . Stigmatopteris: S. heterophlebia; S. pel- Cyrtomium 38 lucidopunctata 43 Didymochlaena 40 12. Cyclodium: C. trianae; C. meniscioides Stigmatopteris 42 var. meniscioides 46 Cyclodium 47 1 3. Polystichum: P. pycnolepis; P. platyphyl- Polystichum 49 lum; P. montevidense var. montevi- Olfersia 55 dense 50 Polybotrya 57 14. Olfersia cervina 56 Diplazium 65 15. Polybotrya: P. fractiserialis; P. osmun- Athyrium 88 dacea; P. polybotryoides 58 Hemidictyum 90 16. Diplazium: D. cristatum; D. roemeri- Cystopteris 92 anum; D. striatum 66 Woodsia 94 1 7. Athyrium dombeyi 89 Oleandra 96 18. Hemidictyum marginatum 91 Bolbitis 98 19. Cystopteris fragilis 93 Lomariopsis 105 20. Woodsia montevidensis 95 Lomagramma 109 21. Oleandra: O. articulata; O. lehmannii . 97 Elaphoglossum Ill 22. Bolbitis: B. serrata; B. serratifolia; B. Peltapteris 167 lindigii 99 MAP OF PERU 171 23. Lomariopsis: L. fendleri; L. japurensis; INDEX TO NAMES . .172 L. nigropaleata 104 24. Lomagramma guianensis 108 25. Elaphoglossum: E. paleaceum; E. eri- naceum; E. piloselloides; E. squamipes ..110 26. Peltapteris peltata f. peltata 168 111 PTERIDOPHYTA OF PERU Part IV 17. Dryopteridaceae Introduction intergradation and its geographic basis are more fully understood. This fourth part of the "Pteridophyta of Peru" is Type collections from Peru are mentioned in devoted exclusively to the family Dryopterida- the nomenclature but are not repeated in the spec- ceae. The general style, typography, form of ci- imen citations. They are, however, included in the tations, and so forth follow the previously pub- Peru range and ecology. The nomenclature of the lished parts I and II. These matters are adequately genera and species is not intended to be complete. dealt with in part I (Fieldiana: Botany, n.s. No. It includes all names based on Peru material and 20, 1989), and it is not necessary to repeat them other names that are considered useful to mention. here. C. V. Morton (Amer. Fern J. 62: 57-64. 1972) Stigmatopteris, Olfersia, and Polybotrya have has argued that the Cryptogamae Vasculares Qui- been contributed by Robbin C. Moran, and Elaph- tenses of Sodiro was published over a period of oglossum and Peltapteris have been contributed years, most of it in the Anales Univ. Quito, 1890- by John T. Mickel. Other genera have been a joint 1895, and a small part in the Anales Univ. Central effort of Rolla M. Tryon and Robert G. Stolze, Rep. Ecuador, 1897. He proposed that the book each critically reviewing the treatments prepared was not published until 1897. This conclusion may by the other. be correct, although Luis Mille, who was certainly The Dryopteridaceae are the largest family of in a position to know, indicated (Nov. Rec. Crypt. Pteridophyta in Peru, with 26 genera and about Vase. Ecuador, page 1 and page 5, 1927) that the 200 species. Many of the genera are taxonomically book was published in 1893. We accept that date difficult because of considerable variation and the and consequently only one of the names cited for lack of recent critical work on the Andean ele- the Pteridophyta was published earlier in the An- = ments that would provide guidance. Among these ales: Acrostichum setigerum Sodiro, 1890 may be mentioned Ctenitis, Megalastrum, Tec- Elaphoglossum setigerum (Sodiro) Diels. tana, Polystichum, Diplazium, and Elaphoglos- Abbreviations of periodicals generally follow the sum. In these genera there is clearly a maximum system of Botanico-Periodicum-Huntianum of variation in the Andes from Bolivia north to (1 968), while those of books and authors generally Colombia, and also extensive intergradation be- follow the system of Taxonomic Literature (TL- tween many of the variations of a species. The 2, 1976 et seq.) The acronyms for herbaria follow taxonomy is often less complicated to the east and Index Herbariorum and are also provided below. north, especially in Mexico and Central America. Accordingly, a number of commonly recognized species are placed as synonyms for Peru or they are maintained with some doubt. Although the treatments for Peru are based on all materials from Acknowledgments South America, or from tropical America, there is need for monographic study of many genera, so The authors are extremely grateful to Dr. John that the species characters are better known and T. Mickel (Elaphoglossum, Peltapteris) and Dr. TRYON & STOLZE: PTERIDOPHYTA OF PERU.