aua o ew eaa Ko te Aiaga eeke o Aoeaoa AUA AISOY GOU ACAE ESEAC EESEAIES . ema Mn Whn ndr rh dvrt & Cnrvtn Grp nln Sn Cntr .O. x 6, nln lnd .Κ. Cosy nd M. - C. aiièe Mn Whn ndr rh dvrt & Cnrvtn Grp Mnt Albrt rh Cntr rvt 20, Alnd lnd UIESIlES EESEAlE .M. Emeso prtnt f Entl .O. x 84, nln Unvrt lnd MUSEUMS EESEAIE M .. ama trl Envrnnt prtnt M f lnd .O. x 46, Wllntn lnd OESEAS EESEAE .. awece CSIO vn f Entl G..O. x 00, Cnbrr Ct Α.C.Τ. 260, Atrl SEIES EIO AUA O EW EAA M C.. ua Mn Whn ndr rh dvrt & Cnrvtn Grp Mnt Albrt rh Cntr rvt 20, Alnd, lnd Fauna of New Zealand K t Atn ēp Atr Number / Nama 35 Cyiae Acaosomaiae a eaomiae (Insecta: Heteroptera) : systematics, geographical distribution, and bioecology M.-C. Larivière Manaaki Whenua - Landcare Research Biodiversity and Conservation Group Mount Albert Research Centre Private Bag 92170, Auckland New Zealand Manaaki Whenua PRESS ico Caeuy ew eaa 1995 Coyig © I995 acae eseac ew eaa Νo a o is wok coee y coyig may e eouce o coie i ay om o y ay meas (gaic eecoic o mecaica icuig oocoyig ecoig aig iomaio eiea sysems o oewise wiou e wie emissio o e uise Caaoguig i uicaio LARIVIÈRE, M.-C. Cyiae Acaosomaiae a eaomiae (Iseca eeoea sysemaics geogaica isiuio a ioecoogy / M-C aiièe — ico Caeuy ew eaa Maaaki Weua ess 1995 (aua o ew eaa ISS 111-533 ; o 35 IS -7-931- I aiièe M-C (Maie-Caue II ie III Seies UC 59575(931 eae o uicaio y e seies eio a e auo usig comue-ase e ocessig ayou scaig a iig a acae eseac ew eaa M Ae eseac Cee iae ag 917 Aucka ew eaa Māoi e y aciic Ieaioa asaios Aucka ie y G i Weigo uise y Maaaki Weua ess acae eseac ew eaa O o ico Caeuy ew eaa Front cover: The insect depicted i s Hypsithocus hdn (Illustrator: D.W. Helmore) Aro mua Ko e ēeke nei ko Hpth hdn (Kai-wakāua W emoe OUA SUMMAY E WAKAOOGA ΜΑ E MAEA Cass / Kaāie Iseca Oe / Ōa Heteroptera Sueamiy/ Wāmee-ui eaomoiea amiies / Wāmee Cydnidae Acanthosomatidae, a Pentatomidae Iusaio / Wakāua pth hdn a zr vrdl (Iusao / Kai -wakāua W emoe Shield bugs Ko e wāau io ai o eaomoiea e ia oa au wāmeeāuaākai kei ēei roto a a i e gāaa e e sueamiy eaomoiea icues seea amiies āua gāaa momo oo a ae au ki e kāui maa oa o geeay sie-sae ue ugs wi a wo aua o e ia ei mao momo i e ao waui E ou ga wāau i seea ousas o secies ee amiies ae ee- oo i iu īei e ii oa e āuu uaga – ekau mā see i ew eaa wic as a eaiey sma aua wau momo aae i ga mea ka oo ii ki ga wāi ai —1 secies-comae o age o wame egios oe maaa āei e Cyiae ae usuay siig ack o ow ugs Ko e hua o e Cyiae e kii e magu e kāaa- uowig i e soi a eeig o oos sems o ae aa e gāaa aaui āei ka wiiaeeiroto e ki sees wo secies occu oy i ew eaa a wo oeoe kaikai aee ai i ga akiaka kakau ākau kākao occu aso esewee i Ausaasia e eemic secies aka āei E ma momo ka oo kei iu īei aae e ma Chrdn nrnt ies i eaee aias aa ka ua i ga akiwā o Aus traasia Ko ga momo ūuu suc as coasa sa ues ia oo ais a e- Choerocydnus nigrosignatus ka oo i ga wāi akuaku ee ussock gassas mo āou ēei i ga ueau oeū o e akuai moaa A ou secies o Acaosomaiae i ew eaa weua māia i oo au māia waiuke ā me ga ā ae eemic Secies o hplrph iai gasses kaaee oeoe kua aa au aee uses a seges i oe aias oeig seams o Ko ga momo e wā o Acaosomaiae uu ūuu mases oe a e oes ege om coasa o suaie kaoa o iu īei Ko ga momokaRhopalimorpha eiomes icuig ussock gassas Eggs ae ai oo i oo i e rai kaaee auau ākaikai e uae amog eeoig sees o o e uesie o eaes o aee ana ga wāi ooaga i ga aaaa awaawa as suc as cuy gass e ie immaue gow sages eoeo e ii ki ga aaaa gāeeee mai e akuai ae usuay se amog e sees o e os a wic moaaāuaākau ki ga eiei akiwā ei uugae aiao see as oo e sige secies o geus Onnt is ae am ki ga uuaga oeoe Ko ga ēki ka ukua ki ga ou oy sie ug iaiig aie oes kākaoākau e uu ake ana i ao āeiēei i ga raurau Eig secies o eaomiae ca e ou i ew i ga kaaee koikoi Ko ga wāaga ima e āa uu eaa wo ae eemic a wo oes ae aie o kugauga ake aa ka oo i ga kākao o e ākau e ew eaa u occu aso esewee i Ausaasia a eūaamai ei āou ei kaikai oki Ko e momo ei i au as o e Sou aciic e emaiig ou ae ee i e āai Onnt, ko ia aae e gāaa akeke ka iouce ee Oe o em e gee egeae ug oo awa i e wao zr vrdl, is a imoa a cosmooia es o e wau ga momo o eaomiae ka kiea i iu īei a wie age o ecoomicay imoa cos Mos r ka i koei e ua au ūuu ake o iu Τīei eoi (ntnd vrlf (ara hr tn eaomiae ee o a uices a ie aoe gou egai e kiea aee aa i ga akiwā o Ausaasia ā me o ei os as Some secies ae esucie o cui- ēaiwāici Ko o e e oega Sou momoaci e wā i ae as Oe gou o secies is eaceous a is āaaau mai ki koei uaweua Ko eai ko e g ew eaa memes ae imoa a eeicia ea- kākaiki eaa iiua e mea ia oo oa ake aiakoa os o esucie isecs ki ea kaiau gāaa wāui ki ga kaiuaweua ia ka I ew eaa ee is aaey oe geeaio e okoia Ko e uiga o eaomiae ka kai i e wai ākau yea i mos eaomi secies Wie is geeay asse ākauka oo i uga i e weuae aaaeekaiga ai kie y e au sage seeig i ceices o as a gass ei e āou Ko eai momo kaui e mai kio au awa i cums o ue ak ocks eaes a oe oecs e ga uu e wakaia aa Ko ēai a momo ko e ū maoiy o secies ae o osa-seciic u i some kaiauau aa ko a mema o ui īei kaui e ia e isaces e ie cyce is quie iimaey ike wi ce- oa gāaa kaikai ei aiga oki ei wiuga gāaa ai a geea eg Cpr o ttpr. au kio Maig geeay occus i sig a/o summe e- Ι oo i iu īei koai uuaga ka ua i ia aumo e eig o e secies ae-sae eggs ae ai i ig uiga o e momo eaomi Ko e oke e koe uaga cuses gue o e os a usuay o e uesie o gāaa akeke ka uuau i oo i ga kāii ākau ao i eaes Ae acig youg sie ugs oe say o- ga ū kaaee ao āei i e kii ākau e kwau auau gee cigig o e emy egg-ses ui ey mou oaoa ēā au mea Ko e uiga o ga momo kīai i ū i aa yms o e seco isa (gow sage e isese awa ki ēai akau aae eoi i ēai wā ko e wīa oa i seac o oo yms mou ie imes o ecome e oo aa ki ēai momo ākau a ēei i e Coprosma, aus Sie ugs ae a ume o eemies seea o Pittosporum āei wic ae iy wass o ies a aasiise e eggs Ka īmaa aee ēai momo i e maimai i e mauu oay e mos isicie sie ug i ew eaa i e aumai āei Ko ga ēki ie ki e āua kāo ei e is pth hdn (see iusaio Is eaiosi aga ka ukua aeeia i oo i e auau kikī ka ii aee o oe Soue emisee eaomiae is ucea ai e uiga i ao i ga au ākau I mui i e ēiga ko ga a ey ie is ecoe aou is ie isoy a ais gāaa kuga ka oo ai iiaiai ki ga wae ēki I is kow om oy ie ouaios isiue sea- oe kau ei e mea o oo ā aka oa e kii uaai Κo ga aey i suaie a aie eiomes o e Sou kuga uuaga uaua ka miaa aee ki e auau Isa Maes wee ukow ui quie ecey e kai e ima akaga ki o e kuga i mua au i e aega secies as imie aiiy o isese—oy maes ae ki e aumaa akeke Ko e gaaa kii akeke ei kaui og oewigs a o sees ack i wigs—a a o āou oa ii ko ēai ei e kai ii ei e ao occuies aias a ae cosiee o e a isk āei ka oo ei kai au aee i ga ēki Ko e mea ūoo o e ii gāaa kii akeke ka kiea i Coiuo Maie-Caue aiièe was o a eu- iu īei ko pth hudsonae (ioia e ikiia Ko cae i Quéec gauaig wi a i eomoogy a waaugaaga kia ēai au o eaomiae i e om McGi Uiesiy Motréa i 199 o wo yeas Τakiwā oga kāoe e māama awa aa ā e iiii oa ake se i osocoa wok o aiae a e Cee o kua ekoaaia m ā ikaga me e āua oo Ka a a ioogica esouces eseac o Agicuue mioia aae na ga ima momo ka oo wee aee i e Caaa Oawa Maie-Caue is ow eseac eae o wao ākau āua makaii ki e wao io ikeike me e e Ieeae Sysemaics ogamme o e ioiesiy keke āei i e aiao o e Waiouamu o aiaei aa a Coseaio gou o Maaaki Weua - acae a ka mioia e ame Ko e momo ei kīai i wai eseac Aucka Se as auoe oe 5 aes o wāaga awa ki e miaa aee oa – kia ga ame aae e sysemaics o aious amiies o ugs icuig ooa aa ga aiau o mua ko e ame me e uwa gaai aua a ecoogica asecs Se as a secia iees i kīai e aiau i mui – ka muu ka oo i ga wāi e aicaio o comue ecoogy o sysemaics akuaku kīai ei i io ai mo āou Kth: I wāau a Maie-Caue aiièe ka wai i aa emaics i oo i ga mai momo uga aiao ū Kai- māauaga i Queec ka ua i aa ougaaga ākua iakiaga o Μaāki Weua i Akaaa i Τāmaki ei oki auia gāaa i e Wae Wāaga o McGi i Mo trea i e Ko uia kua ua i ai ia maa au i e ima ekau ea e ā au 199 e ua au ia e mai aa i ga mai mai o mui i aa ki e momo āua o ēā au ēā au o ga wāau aa ākuaaga mo aiae i e Wāi agaau Weua gāaa ae au ki ga āua oo i e wao kie ikaga ka ā me e awa aiao Auweua i Kāaa Oawa Iaiei e ki e aiao Ko ēai o aa io ia kia waiwai i ēai kaiaaaki a Maie-Caue i e kee āai Ierteae Sys- āua wakamai miii ooiko ki ga āai ASAC ew eaas siee secies o eaomoiea—eeseig e amiies Cyiae Acaosomaiae a eaomiae—ae eise A eoye is esigae fr pth hdn ego a a ecoye is esigae o Chrdn nrnt F.B.