"#$%&"'(%&")#)*+ 8/95%6"7 50 %12"34 /0 %,-. Aelia acuminata L. %&'(')*+,-'./0'(*$1234'$' 5/#4'(+6 !"#$ )+78#+941':#3;<'*2 (Hemiptera: Pentatomidae) E(*=-+8>0'?@A9';BCD 2:D% E4FG 1"#$%&")#)*+ %C ?7A%B 2"#:;<= Q95 %L50P0 MNJOG%KJ2"+ 95 05058%L I'"J2=%KJ2"+ H )F>GH %&2"?W%&"XY<% "$%D%=4 A%T?*A%&V%C" R7%C -3.%S< Aelia acuminata (L.) (Hemiptera: Pentatomidae) %R7 %^ ,D A%e3D%R7%RJ<%C" $bJY%IW"4%c%](%I#_3.% 3D%d#-*+%Y% 7"4 I?JS%C _`"aA %^ I7<%1,4%M2<F$%.%Y%Z,$%\2<FA%2=%]( %C"NkA<2"G%Y%"3;%Y%:;%C _`"aA%Y%C2Y( j3.%dA(%C )aA%2=%Z,$%C i2FA%S<%B A. acuminata %R7%C 12"c%d?;%dA"D%&<N:g%2hA%RJ,c %Ii"7 8P % C2;% 12Y=% Y % v us %t Ps % p?;% Icq2% Br#?-7% .2= 5o ±8% C"A= H% 1"n:J"AS(% 2=% 1,4="J% C" R7 %^ ,4% Z"l;<% ](% -mA,#`+ %{#+N+%c A. acuminata %R7% $2G%R7%xG%Y%yz+%C"3;Y%:;%]<2Y=%R#n;"#A %^ ,;,4%JOw+%Z,$%C ;<=%"c %Q EJ2"+%Ii"7 u%Y% J"4Y2 %Y%NA %v us %S<%~#c ^,A(%I7=%c%SY2 u0P/ ± 0s /|} %Y P0oP ± 0s 8uo %BP05u ± 0s 5/u %B9088 ± 0s 558 %B|09P ± 0s soP %B}08| ± 0s s5u %{#+N+% Hc %I4<,;%NnJ,EJ%"c%C2<= _A%&Y"+%1="A%Y%N;%&<N:g%C"3;Y%:;%]<2Y= %^ =c%ZY=%Y%6Y<%R#7%C" 12G%Y%yz+%dg<NA%c%cNA%N#A %6q%Y %Q yz+ H%dmA%,#`+%`"W%Y%`"W";%N;%B Qt8SY2 H%I#_3.%,42% J";%Y% +<%N;%C"NkA<2"G %^Q SY2 |/ 0ss ± s0u}%Y |/ 0/P ± s0u8 %^ ,4%=2Y(Nc Pu 099 ± 8o 0}|%Y s08P ± 0s s9| %B8s }0 ± o0u8 0s%B 9}| ± 0s 58s 0s%t%B s5P8 ± 0s s885 %{#+N+%c %Q SY2 H%d?;%eJ%C 12Y= ^,k?#;% "n:J"AS(%J<N4%2=% 42YNG% -D%2<N)k7<%c%2="X%" ](%D%,=% A%]":;%R7%RJ<%C 12"c%d?;%=<N'<% -mA%,#`+ % #_%=NE-3i "n:J"AS(%M2YNG%B&V%C" R7%B '<N$A= L(F>JK'(+6 )I3$ [email protected] %L 6?A%1,?J; 4%+%5,! %8%+,29:;<%2%.=81<%+! ./012 *+, !"#% ( ?@ # !" %,: "T#C% .M% #(a "% !1 |. ;N/:% H!!h %H%,29 G!1<%+! 4%+ C%<; Aelia %AC%+! 4 %N#% 2% }N% 2% +`1, sa% *!% 2% "% +!% 5:;:% + 5LN#/N %R L; 5LM;NO%,"M%P! %<;%+,!#8 %,:%M%LQ .%K8( %2%5:N" %H!d;%*c2,%1;%~M% %R T4;%=s%w;N Aelia acuminata (L.) 4%AC%1;%+! /%<;%.S1 %5N"w%+,29:;<%2%.=81<%+! ./012%G;N#O%.!M%T\8 %T#;% +;,;:% U# % 2% M% *!,"M% <;% .WN % ,:% M% T4; %#(a%2%1`a%. Nd%<; %R (Carey, 1993) %L19 .%T4: %! % * (Lodos, 1981; Burlaka, 2009) % T4;% +:!X= ; %T!W:%!4;%N %2%.4%+ 5012 +wN%c2,% %! 5:;:%1; (Panizzi %1L %+!#: %,:%4%1;%UM;NO%./:N=8/%:C2 %;N %;,%+Na !C%H!d;%*5:!%2%N%AC%2:%N%+! 5:;: Alexandrov Z% G;N1;% 2 et al ., 2000; Rider, 2006) A. %4% %P N% !%.4,N %,: %R (Chi, 1988) %:,29 . % % T#;% 2 % [ and Mirzayan, 1949; Radjabi, 1998 %T#C% %Ca%! %R L"%5L!"%H!d;%1; acuminata %.!/%!;%* (Panizzi et al. , 2000) %G9%+:!X= ;%M%T\8 %*,! % 1;2;% ,:% G;,zv!=8<% +! 4% L;% 2% 5LM;NO %2% ^L/% _,;`% ,:% G9% <;% .Ca% ! % +! T#C% H! 2; %T#C% T\8% % M % [ ^2:% 8 Z 5,! % +! 4% + !h %c,;`/ (Lodos, 1981; Javahery, 1995) %bN %.!/ %:;N ;%. !1<,;%U20O%1;%L %R T4;%Na ;N %L,;:%+N="# %+! M;Na%,:%G;N1;% d!%<;%+,!#8 %,:%4%1; %R T4; 5LQ %.Q,2NO%.M%#S"a%G!S;%,% %4%1;%+ 5,! %8 %+ 5N#a%<;%<N%2%.aN%+! %2%.;,<%G!!#/%+2,%*M RT4;%! G9%.=81<%+! ./012%TW!Q%2%.!v"1!<9 Radjabi, Z% :,W .% "g% % `# (Poaceae) % H +, $%&'%'()* R[ 2000 Z%! Z%! % 9% TQL % ^L/% _,;`% <;% 5!:;:NW% NW9% ,: A. % 4 Kustanai % + h% ,:% G!=8 ;` % i!Q% ,: *+ )(!( " #" %&' ! % .!Q% N .1!# ;NC% H!Xs" %4% 5;N% 2% 5: % +:!X= ;% T#;% +;,;: acuminata +* =) "89 ;&<"#" 5 ' 67 )6 ! "#"%&' (* ", ./"02"3 5 ' %+! 4% T#C % m kl % j1N Aelia siberica Reuter A. "L F&9I"A"L'<M;"L.5O F.GH"&;"9I "E 9>&'"?@A"B<".C %#42,%,: %R (Burov, 1962) %:;: .%#S"a%;, Aelia %AC ",T"L9U"F9VG>9BW"&'"T"M/"L&#W % acuminata %_,;`%+! 4%T#C%Lo,: %!an %*l A. acuminata %4 "L9' ]" ,9VG>9BW" \><./" &'" 9 A" >< " Z '<'" 5YG;" <& R(Burlaka, 2009) %: Samara %G!=4;%H ± "L&"F&#'"# "d c7 b!7 ",U_"aI 2&"3^5_A"%&' *( 6 %N="# A. acuminata %4%UM;NO%./:N=8/%:C2%! "fW"#"gMO"L <'"9I "E ,Y>&9;"a9A c"#",>9/#&"a9A 6! %i ;% 2% r2;% 2% U1;L #O% G;N#O% *G9% .4!Q ^ % H!!h ZMM/">h; Tischler, 1938; Z%T4;%=s% d!%,:%4%1;%T#C "n;" L9 CA'" 9I" \><./" &'" , <.O'" l>9BW" L<.I %R[ Burov, 1962; Benedek, 1971; Radjabi, 2000 "o#&'" 9O<M%" &2 I" #" a/<'.I" ,T" B<" <B#&" &2 I %<O!1:%+ 5L1LO%uXW%,:%.Ca% ! %+! .4,N %#t "9I"l>9BW "Z a .O"&<. "E .C ,C9A !"9'".@ ]"L.Cp"q#.r Hodek, Z% T4; = N/% H,o% 5!v"1!<9% ,:% 5N"w% 1; "FM9/p"h9T"9I"L.Cp"q#.r"T "Z M/"B9W"n ;"'M (+ %4 %1;%.!v"1!<9%.4!Q T81<%+ 5,! ,: %R[ 1971, 1979 *= " 9; 6* " 5I" 9 <'" T" FM/ u5v" gMO" L9 <'" B< " Z M/ %<;%yT4;%5LQ%G!# %.\!h%U# %2%M%.4!Q 5N"w% !%,: ""x> "Z M/"F'9 CA<">h;"L<.I"3MC/<'"&<. "fW"&'"a9A %1;% G;,zv!=8<% :;N ;% .{% L#a% +! ./012% .WN % C "B95"'&"fW"3oW"L 9'"&'"L< Up"9I"fW"L#9H",G>9BW (Javahery, %T4; = N/%,;N %.4,N %:,%5!v"1!<9%,:%4 "M>'B9I" q#.r" ><" B<" B#&" . " Z av9A ," F'&#W.I" <&" A R1967) 1 demography ( 6 +*"09A"3 *F&9/"3 M%"3,95O"y9 W"y9 5 z; "L< H."L9 I "Z MW"aA' I"B#& (c !) ± !)( "9I9"o<&#' "T"h9T"#"fW"3"<h"|>;"}"9;"a .O ,"y &{ "L9 I">.CG5I "Z aA< FMW 6 0#M%"&'"5"9I9"Cn"H<. "B<"MI "Z '/"a/<''9>"LB<M< aAp"#".5#"~."&9W"3q.r "oW">.CT"#"0#<"A"L9 F&p"I"I. "d c= ) "9I"L< H. "y&{ I" #" FM/" 5" 9 oW" u%" 9I" y<.GH" l><M5p ZMW"aA'"I "d * *) "9I"g#'"A"L9 F&p"I"I. "9 F'9"L>& n;"o<5"#">&9; "Z MC .O ,"&<. ""9I"a % 6"Y/"&'"5 (Sxj )"Cn"H<."L9 I"."L9 ,z "> <"I "Z M/"a/<'9>"<B#&"&2 I"F'9"#"."'<. <".5"#"~."# "L9 F&p"#"n;"H<."&'",T"&2 "ZI aA< FM/"F'<'"o9G "x5 <.O'"L9.C<&9p"'&#W.I"L<.I"A9"L9 F'<'"5;.; Z'<'"&",T".5#"~. "d c7 "3#'"#"x>"5A ZMW"<. A. acuminata "A ",>9"# "E rm]",;< "."L9.C<&9p"&95"L9@v"#"5V95 ",OMB" 0#M%" 0M" ^9A<" .I" FMW aA'" I" L9 F'<' "" M5;" 9v" # " E Ro]" 9v9" ." 3 Eλ]" a5%" M/& "# Chi and Liu (1985) "\A;"T",_%#'",A"L F># "F' <'"o9G *"0#M%"&' "E T]"_"x>"L F&#'"02"# "E GRR ] TWOSEX-MSChart "&< < g."9I"3FM/"5; Chi (1988) ZaA< FM/ ".C<&9p " Z a .O" &<. " L&9W" L >;" '&" 3 (Chi, 2012) "_"&'"a5%"l>< <",;< ".",>".C<&9p">.; b.#<"'9"^9A<".I "E rm]"a5%"l>< <",;< "." ω A. acuminata # " "."oW"B<"L#.5p"I"#"'I", "A"L F&9I ! rm ( x )1 "T" L&2 I" M/" '&#W.I " E 1 lxmxe ]" 9Y; "'v"T"'I "E )7 +c ]"x>"B<".CT"a5%"l>< <",9C x!0 Goodman, ]"aA< FM/"C .O"."&'". {"B#&"B<"F.GH"A ZaA<"F.GH"a5%"L <B#&"l9T".V95I SE 1982 &)$ "#&" >.2"B<"FM/'9>"L9.C<&9p "E ]"&95"L9@v "ZE (Huang 6 "&2 I" F.GH" x>" FU< 5" 9>" ,9VG>9BW" &#.p aA<"C .>hp"y &{"&<.Y;"&9I 67777 "9I .C_;I and Chi, 2012) ",C_>B" , 5T" #" ,T" L9 ,O>#"av9/" &" I" 0 "y<.GH"L9"G"o<&#'"02"L _>9 "Z SPSS t-test ".CO&I"^95 "&'"#"'v"L I"I"T"'.5O ,"y&{"oW ZM/"g9< "&< < g."9I"# "oBW"\A;"F'9"#". '!#%E "L9 9.I"g9<"L<.I",/&#.p"y<.GH"B<"F'9 CA<"I"M<; , "T" #" ,O&p" A" p" 3n;" L9" G" L F&#'" 02 "L 9@ " Z (Knipling, 1984) " M995I" a W" x>" 0.CT ">.; ,2 " Z aA< FM/" F'<'" o9G 6" 0#M%" &'" 9I9"o<&#' "A LF&9I"_"'<. <"L<.I"9 ,O>#"><"L ><&<"}".}9H "G"o<&#' "Z aA<"B#& 66)+ "9I"g#'"A"L F&p"I"I."F&#' "_"><"y<.GH"B<"F'9 CA<"T"'<'"o9G A. acuminata ± c)7 "# = !) ± c6)7 "5;.;"I"F'9"#"."y<.GH"L9"# '/Z ,",I9>B&<"5}",9VG>9BW",T"5YG;"L<.I t = ]"a/<M"L&<' ,"y#9 ;"T"MW"aA'"I"B#& = 77) "g#'" A A. acuminata " A" L9" #" G" ,A&.I" &' "."9T"y<.GH"."02 "ZE 0.298, df = 28, P = 0.77 "G"L F&#'">.; ,2",O&p"5A".>9A"I"aU _",O&p "B#& 6= 6+) ± 7!)= " # 6* *6) ± 7() " 9I"5;.;" I" F'9" # ",A" L H." .5#" ~." >.CG5I" # " E B#& 66)+ ]" 9" # t = -0.380, df = 28, P ]"a/<M""9I"L&<' ,"y#9 ; "&'" y" 0" &2 I" "a5U2"&' " Z a/<'" <& " Ed = ]6) ZE = 0.71 "I"oW"L FMT >h;"L9 F&p"F># I"3g#'"_"l><M5p"o9B "T" MM/" C/<hO" 'M 6= " 9; *" 5I" n;" L9 CA' "I" .>O9" #&" >B<" 3MA& ," <'" oM/ a A" L H.