NOTES ON THE FAUNA OF THF: CH!POLA FORI\!AT!O:\ ~XXX ON THE PRESENCE OF EUDOL/UM tGALEODOL/UMt SUBFASCIATU,\1 SACCO I GASTROPODA· TON:\"lDA£1 F:~Ili.Y II . VOKES 1TL,\SF l.\1\'f.'RSlT'l

Tn the exposures of the Chipo lgenus hitherto unreported C\'l'l'. rt would appbu· tfwt thl'rc arc tv-:o in the fossil record of the Weste r n H t-n1ts­ clistinl'l linl·agl'S within the Eucfoliwn line. phere. although it is widespread in the Trut· EuTonnidae 11948. p. \()() l_ ohsern•d: \\·ith another r~Toup r t l'll .n·gl•d rug;:K at ""All species I in this fa mil~ · 1 appea r to hP l':\ the po: t(.•nor l"'lr t fonnmg drl ned notch ceedingl~· \'ari<.tble. not on ! ~- to sil.l'. but In the western Paclfrc here are also also in coloration. sculpture and gl'lll'r;:d species of Eudoliu 11 .s which are prE shape of the shell. .. Certain !~-. this is the sumed to have descended ·rom the Middlt: case in E. sul~f"nscintum in tlw Chipola For­ I\-'fiocene F:. lJiornatu;n 1Tatc, 1893J from mation. where we have som e two dozen Muddy· Creek, Victoria, AustraJJa. How­ examples from Sl.'\'l.'ll local ities. Thl' ever, tn the Earlv "!\'fiocene of New Zealand number and strength of thl· spi r al cords rs there is anoth~er species referable to differ·t_·nt. not only among d iffl' rent speci Galeodolium; indt·ed. E. aoteanum Be11 mens. but ;tlso in different growth stagl'S of 11970, p. 127. pl. :l. figs. 19~261 is so similar the same specimen. The stre ngth of tlw in appearance to E. tGuleodolitWI) mutt nodes on the ma_ior cords and thl' numbl'l" cum I Michelotti. 18()1 J from the Italian of nodecl cords also ,-aries . thPrl' being Oligocenl: that it must be a direct de!;cen­ from one to four such ormlnll'nll>cl spiral dant. Both of these latter species dif ·e, cords. although the usual numlwr is thrl'l'. from E. suQ{asciatum in having coarser In the R.ecent fauna of the w est<.·rn At dXial nodules and in lu ving the back of the !antic there are onl~· two sJwcies of outer lip less deeply excavated. Ewluli11111. both referabl(: to E 11(loliwn s.s. Occurring in the same area along Ten­ IE. ("I'USS('(II/11111. the t~'J)(' of the w·nus. dllci mile Creek is the superficially similar E. tlwmpsu11i McGint_\'. JD55 1. Both ;:1n.· in PhaUum rSemieussisl aldrichi IDall, 1890J, habitants of moderate!~· clt·t•p wakr and which also has a noded shoulder and a neither is pcu'ticularl~· simil<~r to the fossil pustulosl' inductura. However. the sharplv species. Sacco erected a subgenus Guleo­ recurved siphonal canal is an unmistaka­ clulill.m for E. sul~/(tsciuttlll! and the wnkr bk, identifier of the Phalium; furthermore, feels th;:tl. there is a valid need for thi there is but one row of nodes ahvays pre­ titxon. Dr. Alodo/iull! ( ;~ s '' -ell c.t:-:. about four times thl' size of that of P S

177 178 Tulane Studies in Geology and Paleontology Vol. 19

As noted above, the species of Eudolium Eudolimn DALL, 1889, Harvard Mus. Comp. s.s. live in relatively deep water (E . cross­ Zoo!., Bull., v. 18, p. 232 (new name for eanum from 40 to 450 fms [73 to 822 m], fide Doliopsis Monter·osato, 1872, not Vogt. 1852, Turner, 1948, pp. 179-180; E. thompsoni in nor Conrad, 1865). Type species: Dolium c1·osseanum Mon­ 75 fms [137 m], fide McGinty, 1955, p. 80; terosato, 1869, by monotypy. the Pacific species E. pyrifomw in 50-80 fms [91-146 m] and E. injlatum in 100 fms Subgenus GALEODOLIUM Sacco, 1891 [183 m], fide Kira, 1962, p. 59) but there IS Eudolium (Galeodolium) SACCO, 1891, Moll. no evidence that the beds along Tenmile Terr. Terz. Piemonte e Liguria, pt. 8, p. 4. Creek were deposited in water anywhere Type species: Cassidaria mutica Michelotti. near that deep. According to Scolaro (1968, 1861, by subsequent designation, Cossmann, pp. 46-47), the water depth at Tenmile 1903. Creek during Chipola time was about 20 Discussion: Sacco named Galeodolium meters, an interpretation that is corrobo­ as a "Sezione" of the genus Eudoliunt, in­ rated by the molluscan fauna in general. In cluding two nominal species: E. muticum view of the fact that the species occurs only (Michelotti), with three varieties; and E. along Tenmile Creek and has been taken subfasciatum, n. sp., with twelve varieties. nowhere else in the Chipola beds, possibly He did not designate a type species but bottom sediment type had a greater con­ Cossmann (1903, p. 139), in a confusing ac­ trol over the distribution of the animal than count of the family Doliidae (i.e., he places did water depth. The beds along Tenmile Sacco's Galeodolium, Tube'rculodolium, Creek are generally lower in carbonates and Simplicodolium in synonymy with and of finer grain size toward the western Dolium, but includes the species Sacco as­ end (TU localities 70 and 655); localities to­ signed to these groups in the subgenus ward the eastern end (TU 546, 830, and Eudolium), noted that in the Oligocene 998) are not different in any measurable there are two species of Eudolium.: "type parameter from those along the Chipola de Galeodolium et de Tuberculodolium: E. River (Furlong, 1980) and yet the species antiquum Sacco, Cassid. mutica Miche­ occurs at both ends of the Tenmile section lotti. " This is considered a valid cfesigna­ in approximately equal numbers (see Oc­ tion of the type species for both taxa, even cunence, below) but not on the Chipola though the order of the two names is re­ River. This problem has been encountered versed. in other molluscan species (see Vokes, 1977, p. 146, for a discussion), and so far, EUDOL!UM (GALEODOLIUM) no satisfactory answer has been achieved. SUBFASCJATUM Sacco Text figs. 1-3 ACKNOWLEDGMENTS

The writer wishes to acknowledge the EudoliHm snl~f'nscial1w! SACCO. 1890. Soc. contribution of the late Mrs. Shirley E. Geol. Ita!., Bol.. v. 9. p. 20\linclucling 12 val·­ Haerle, who originally recognized our ieties and subvarieties; all nude names. list species as being the same as the European onlyJ; 1890. Mus. Zoo!. e Anal. Comp. Torino. one. Alan Beu also made numerous sug­ Bol.. \'. 5. no. 86. p. 22 (including described gestions as to the improvement of the man­ \·arieties: supn1tuberc11liferu. uuilllbercttl.f­ em. percostatu: and nine undescribed var·­ uascript, and the fact that the writer does ietiesJ. not entirely agree with his generic concept E11dolium rGnleodoliHm) stlbfusciatum Sacco. of the group (while hastening to admit that SACCO. 1891. Moll. Ten·. Tcrz. Piemonte e he knows a whole lot more about the group Liguria. pt. 8. p. 6. pl. 1. fig. 4 ( f varieties than she does!), in no way diminishes her SHbtHberculnw. p. 7. pl. 1. fig. 5: HWfJIWPoiclea. gratitude to him. p. 7: coHicu. p. 7; Sllpmwberct~lifenL p. 8. pl. I. fig. 6: oblonwnn. p. 8. poutnnensis. p. 8: SYSTEMATIC PALEONTOLOGY !n!ituberclllifera. p. 8. pl. I. fig. 7: percostClla. p. 8. pl. I. fig. 8: "'subvariety" cmssilabiatn. p. Superfamily TONNACEA Suter, 1913 9: tubercHiifem. p. 9: and ill.f1nta. p. 9. all Family TONNIDEA Suter, 1913 from the Turin Hills; and tl·i!utelllberculntn. Genus EUDOLIUM Dall, 1889 p. 7. from Gnmd. Vienna Basin. Ausir·ial. No.4 Fauna q{the Chipoln Formation- XXX 179

Dolium fEHdolium) sub.faseia tltm tSaecol. The wide distribution of E. subfnsciatum COSSMANN. 1903. Essais Paleoconch. is not unexpected in \'iew of the fact that Comp.. v. 5. p. 138. pl. 5. fi g . 18. the type specie::; of Eudolium. E. cross­ Discussi011: This species occurs in the eanum. occurs from the Mediterranean. ''Helvetian'' of northern Ita ly and the Vi­ the south Atlantic and Indian Ocean to as enna Basin. and is here rep~rted from the far north as Natal. South Africa {Barnard. Burdigalian of northwestern Florida. 196:l. p. 81 and the western Atlantic. with Other than the Oligocene E. muticwn and records in the latter from 1'\ew Jersey to the New Zealand Mioce ne E. aotemwm Barbados !Turner. 1948. p. 1801. there are no othe r me m bers of Gal('o­ Occurrence in tl!e Chipula Fonrwtiou cloliton presently known an_ywhere. All of 1number of specimens in parentheses): all the other species (including those refl'ITecl localities on Tenfllile Creek. between bv Sacco to Tt~berculodo lium . which lack Florida Highway 70 and the base of the for­ the reflexed and denticu la ted outer lip. rnation - ron west to east - TU locality and Simplicodoliu.m. which lack both the numbers 7012 6551 l, 456121. 546151, 998121, outer lip and the nodulose spiral cords! arc 951 4). 830181 placed in Eu.dolium s.s .. the type of which has a reflected outer lip but no strong inner LITERATURF CITED lip. The true extent of gene ric vanatwn 1s BARr\AkD K. } Elll:~. CoPtribution~ tn the uncertain due to the fact that entire speci­ k wwleclgc ol South A1 io..;c.-. n; nne HHh _ mens are rare in the fossil record: because r ... t 1.1. .... o<.o. tr)J: do Pr lbr·mciJ;tc: of the extreme ly thin she lls rnost are bro­ Tclt_nlOI'il ~ Al1 s~ u1h Au· :\n ... \ ..li ken. l n the Chipola ma te rial \ V e ha\·e mu.n:-,' pt . t= 1 H!) _ xt ~ 1 :_r, more broken shells. or fragrnents. than en­ Bb ·\ C. 9'11). DE ·•pi 01 " ol new pl l:lt tire specimens. -.rl o1 on tax1 1~ 11V 1f I'\t v-: Zeal<1l d :\I)\-

3b

Text figure 1 (X 1 1 '4). US NM 404594. ht>1ghl :lG.8 1111. diametc>r ,26 I mm: lo~alit~· TU 998. Text figure 2 (X 1). USNM 404595: lw1ght 44.4 mm, chanwtt>r .l1.4 mm locahty TU 655. Text figure 3 (X 10). USNM 40459(): !wight 14.7 mm .. chanwler 8.9 mm. locahty TU 546. All localities on Tenmile Creek, Calhoun Count)-', F'lon(b 180 Ttdane Studies in Geology and Paleontology Vol. 19

lusca: Roy. Soc. New Zealand (Earth Sci­ Part 8. Galeodoliidae. Doliidae, Ficulidae and ences). Tr.ans .. v. 7. no. 8. p. 113-136. pls. 1-5. Naticidae: (ex R. Acca d . Sci. Torino. Mem., COSSMAKN. A E. M.. 1n03. Essais de Ser. 2, v. 41). p. 1-114. pis. 1. 2. pa\eoconchologie comparee. v. 5. Paris. 215 SCOLARO. R. J . 1968, P a leoecology of' the p .. n pis. Bryozoa of the Chipola Formation, FURLONG. W. J .. 1980. Depositional environ­ Clarksville area. Florida: Unpubl. disserta­ ment of the Chipola Formation. Calhoun tion. Tulane University. 251 p .. 20 pis.. 2 County. Florida: UnpubL thesis. Tulane Uni­ maps. ,·ersity. -10 p .. 7 text figs. TURNER. R. D .. 1948. T he family Tonnidae in KIRA. TETSUAKI. 1962. Shells of lhe western the western At.lantic: J oh nsonia. v. 2. no. 26. Pacific in color. v. I. OsMollusca' MCGINTY, T. L.. 1955. New marine mollusks B ivalvia) from lhe Chipola Formation. Cal­ from Florida: Acad. Nat. Sci. Philo .. Proc .. v. houn County, Florida: Tulane Stud . Geol. 108, p. 75-85. pis. I. 2, 1 text fig. Paleont., v. i3. no. 4. p. 143-189. 10 pis., ltext SACCO. FEDERICO. 18nl. I Molluschi dei ler­ !ig. . 6 tables. reni Terziarii del Piemonle e della Liguria. December 15. 1986

REVIEW

PALAEOFLORA O F SOUTHERN AF­ faunas, and a sequence of locality maps for RICA: P rodromus of South African each geological period , Devonian to Cre­ Megafloras, Devonian to Lower Cre­ taceous); 2) Localities, Assemblages, As­ taceous, by J ohn M. Anderson and sociations, Environment (with floristic ta­ Heidi M. Anderson. Published by A. A. bles, locality maps, a nd photos of localities Balkema, Rotterdam, The Nether­ for the eighteen successive megafloras rec­ lands, for the Botanical Research Insti­ ognized and documented); 3) Pre-Angios­ tute, 1985, 423 pp .. 8 1/2 x 12 inches, perm Megaflora! Succession (with the ill us., alias of 226 pis., index, $55.00 principal features of megaflora! evolution This study is concerne d with the floras outlined for South Africa, compared and a nd evolutionary tre nds preserved in the contrasted with patte rns for Gondwana fossil record of South Africa from the De­ and Pangaea); 4) Coll ectors (with biog­ vonian to the Lower Cretaceous (from the raphical sketches intended lo give a suc­ appearance of vascular land plants lo the cinct account of lhe history of megaplant rise of the flowering plants) which coi n­ fossil collecting in South Africa); 5) cides broadly with the Pangaeic phase of Taxonomic Revision (with systematic revi­ Earth history. Specimens of land plants sion undertaken for 91 genera and 263 have been coll ected actively since 1845 by species of South African megaflora! taxa); at least 35 ind ividuals; succinct biographies 6) P hotographic Cata logue (with 226 of these collectors a re included in the vol­ plates, 1888 individual photos, Ill genera , ume, as a record of the history of collect­ a nd 259 species); a nd, 7) B ib liography ing. ll is mainly a pictorial catalogue, bul is (nearly 500 references, classified and de­ divided into seven distinct sections, as fol­ tailed). A glossary and a n index are ap­ lows: I) Global & African Selling, in which pended. a global and continental fabric, against This comprehensive compilation is im­ which to visualise the successive South Af­ pressive and should be most useful for the r ican megafloras is developed (with maps specialized student of the megaOoras of of the drifting continents, phylogenetic South Africa. It appears so comprehensive charts depicting the evolution of vascular that there should be lillie need for future plants and the evolution of vertebrate attention lo this subject. --H .CS.