THE RAFFLES BULLETIN OF ZOOLOGY 2000 Supplement No. 8: 209-232 © National University of Singapore

POLYCHAETOUS FROM THE

Hannelore Paxton Department ofBiological Sciences. Macquarie University. NS. W 2109,

L. M. Chou Department ofBiological Sciences, National University of Singapore, Kent Ridge 119260, Republic of Singapore

ABSTRACT. - A checklist of polychaetous anneIids from the South China Sea is generated from available literature. On record are 661 belonging to 54 families. Biodiversity distribution appears linked to collecting efforts rather than a reflection of the real situation. The potential for discovery of new species remains high.

INTRODUCTION

This check-list, based mainly on literature reports, lists 661 species ofpolychaetous annelids from the South China Sea. The apparently highest biodiversity ranked as mainland China, Vietnam, Hong Kong, Taiwan, Singapore, , Thailand, Malaysia, is due mainly to the collecting efforts of residing specialists or expeditions rather than true biodiversity.

The listed belong to 54 families. The families are grouped according to Hartman (1959), except for the Eunicidae and related families. We are emphasising that this list is primarily based on literature rather than personal knowledge and are presenting references for each record. This list is only a baseline; we expect it to change considerably with the removal of names as families are revised, and the addition of new species, as the areas of the South China Sea become studied more extensively. This is publication number 300 of the Commonwealth Key Centre for Biodiversity and Bioresources, Macquarie University.

NAME AUTHOR COUNTRY REFERENCE Aphroditidae Aphrodita talpa Quatrefages, 1865 China Yang & Sun 1988:133 Hermonia hystrix Hartman, 1959 Philippines Pillai 1965:112 China Yang & Sun 1988: 133 Pontogenia indica Grube, 1878 Philippines Grube 1878:19 Singapore Tan & Chou 1993:28 I Pontogenia lichaucoi Pillai, 1965 Philippines Pillai 1965:112

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Pontogenia nuda Horst, 1917 China Yang & Sun 1988:135

Polynoidae Eulagisca sp. Hong Kong Mak 1982:599 Gastrolepidia clavigera Sehmarda, 1861 China Wu et al. 1980:114 Halosydna brevisetosa Kinberg, 1855 Hong Kong Mak 1982:599 Harmothoe ampullijera (Grube, 1878) Philippines Pillai 1965: 117 Harmothoe dictyophora (Grube, 1878) S. Vietnam Gallardo 1968:47 China Yang & Sun 1988:148 Harmothoe extenuata (Grube, 1840) Philippines Natividad & Palpal-Latoe 1986:296 Harmothoe imbricata (Linnaeus, 1767) Hong Kong Shin 1982: 162 Harmothoe minuta (Potts, 1910) China Wu et al. 1980:114 Harmothoe nigricans Horst, 1915 China Wu et al. 1980:114 Harmothoe (Lagisca) waahli Kinberg, 1855 Philippines Natividad & Palpa1-Latoe 1986:294 Hermenia acantholepis (Grube, 1876) China Wu et al. 1980:114 Hololapidella commensalis Willey,1905 China Wu et al. 1980:114 Intoshella aphthalma Gallardo, 1968 S. Vietnam Gallardo 1968:47 ?Intoshella holothuricola (Izuka, 1912) S. Vietnam Gallardo 1968:48 Iphione muricata (Savigny, 1818) China Wu et al. 1980:114 Hong Kong Mak 1982:599 Singapore Tan & Chou 1993:281 carinulatus Grube, 1870 Philippines Pillai 1965:115 Lepidonotus cristatus (Grube, 1876) China Wu et al. 1980:114 Lepidonotus hedleyi Benham, 1915 Philippines Natividad & Palpal-Latoe 1986:290 Lepidonotus jacksoni Kinberg, 1855 Philippines Natividad & Palpal-Latoe 1986:286 (Linnaeus, 1767) Hong Kong Mak 1980:599 Lepidonotus tenuisetosus (Gravier, 1901) Hong Kong Shin 1982:162 Lepidasthenia izukai Imajima & Hartman, 1964 S. Vietnam Gallardo 1968:46 China Yang & Sun 1988:144 Lepidasthenia microlepis Potts, 1910 China Wu et al. 1980:114 Lucopia magnicirra Pillai, 1965 Philippines Pillai 1965: 117 Parahalosydnopsis hartmanae Pettibone, 1977 Hong Kong Shin 1982:162 Paralepidonotus ampullijerus (Grube, 1878) Singapore Tan & Chou 1993:281 Singapore Tan & Chou 1993:281 Scalisetosus jragilis (Claparede, 1868) China Wu & Wu 1987:79 Scalisetosus longicirrus (Sehmarda, 1861) China Wu et al. 1980:114 Thormora jukesii Baird, 1865 China Wu et al. 1980:114 Philippines Natividad & Palpa1-Latoe 1986:285

Polyodontidae Eupanthalis maculosa (Treadwell, 1931) China Wu et al. 1980:114 Neopolyodontes lepidus Shen & Wu, 1993 China Shen & Wu 1993:537 Neopanthalis muricatus Shen & Wu, 1993 China Shen & Wu 1993:537 Panthalis jogasimae Izuka, 1912 China Wu et al. 1980:114 Polyodontes gracilis Pflugfelder, 1932 S. Vietnam Gallardo 1968:49 Polyodontes maxillosus (Ranzani, 1817) China Wu et al. 1980:114 Polyodontes melanonotus (Grube, 1876) Hong Kong Shin 1982:162 China Yang & Sun 1988:151

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Sigalionidae Ehlersileanira incisa (Grube, 1877) China Yang & Sun 1988:154 Euleanira ehlersi Horst, 1916 S. Vietnam Gallardo 1968:50 Mayel/a sp. Taiwan Hsieh, pers. comm. Psammolyce fijiensis Mclntosh, 1885 S. Vietnam Gallardo 1968:50 Psammolyce malayana Horst, 1913 Philippines Natividad & Pa1pal-Latoc 1986:301 Sigalion sp. Taiwan Hsieh, pers. comm. Sthenelais boa (Johnston, 1839) Hong Kong Shin 1982: 162 Sthenelais mitsuii (Okuda, 1938) S. Vietnam Gallardo 1968:51 Sthenelais nami Gallardo, 1968 S. Vietnam Gallardo 1968:51 Augener, 1927 S. Vietnam Gallardo 1968:52 Sthenelais japonica (Mclntosh, 1885) S. Vietnam Gallardo 1968:52 Sthenolepis izuensis (Takahashi, 1938) China Wu et a1. 1980:115 Sthenolepis tentaculata (Horst, 1917) China Wu et a1. 1980: 115 Thalenessa oculata (Peters, 1854) China Wu & Wu 1987:79 Thalenessa cf. spinosa asiatica Uschakov & Wu, 1959 Taiwan Hsieh, pers. comm. Thalenessa stylolepis Willey, 1905 S. Vietnam Gallardo 1968:53 Thalenessa tropica Hartman, 1954 S. Vietnam Gallardo 1968:53

Eulepethidae Pareulepis malayana (Horst, 1913) S. Vietnam Gallardo 1968:54

Chrysopetalidae brevis Gallardo, 1968 S. Vietnam Gallardo 1968:54 Hong Kong Shin 1982: 162 Bhawania cryptocephala Gravier, 1901 China Yang & Sun 1988:158 Bhawania goodei Webster, 1884 Hong Kong Mak 1982:599 Philippines Natividad & Palpal-Latoc 1986:304 China Wu & Wu 1987:79 ehlersi Gravier, 1901 China Yang & Sun 1988:160 Paleanotus bel/is (Johnson, 1897) Hong Kong Mak 1982:599 Paleanotus chrysolepis Schmarda, 1861 China Yang & Sun 1988:161 Paleanotus debilis (Grube, 1855) China Wu et a1. 1980:115

Amphinomidae Amphinome pulchra Horst, 1912 China Wu et a1. 1980:115 Amphinome rostrata (Pallas, 1766) China Yang & Sun 1988:164 Philippines Natividad & Pa1pal-Latoc 1986:321 Chloeia flava (Pallas, 1766) S. Vietnam Gallardo 1968:56 Hong Kong Shin 1982:163 Philippines Natividad & Palpal-Latoc 1986:321 China Yang & Sun 1988:165 Singapore Tan & Chou 1993:281 Chloeia fusca Mclntosh, 1885 China Yang & Sun 1988:165 Chloeia entypa Chamberlin, 1919 Philippines Natividad & Palpal-Latoc 1986:321 Chloeia inermis Quatrefages, 1865 China Wu & Wu 1987:79 Chloeia parva Baird, 1870 S. Vietnam Gallardo 1968:56 China Yang & Sun 1988:165 Chloeia rosea Pous, 1909 Philippines Natividad & Pa1pa1-Latoc 1986:324

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Chloeia violacea Horst, 1910 S. Vietnam Gallardo 1968:56 China Wu et at. 1980:115 Eurythoe complanata (pallas, 1766) Philippines Pillai 1965:119 China Wu et at. 1980:115 Singapore Tan & Chou 1993:282 Eurythoe parvecarunculata Horst, 1912 China Wu et at. 1980:115 Unopherus hirsuta (Wesenberg-Lund, 1949) Hong Kong Shin 1982: 163 Notopygos gigas Horst, 1911 China Wu et at. 1980:115 Notopygos labiatus Fauvel, 1932 Philippines Natividad & Palpal-Latoe 1986:326 Notopygos ornata Grube, 1856 Philippines Natividad & Palpal-Latoe 1986:326 Notopygos sibogae Horst, 1911 China Wu et at. 1980:115 Notopygos supragigas Usehakov & Wu, 1962 China Wu et a.11980:115 Notopygos variabilis Grube, 1855 Philippines Pillai 1965: 120 Paramphinome indica Fauvel, 1932 China Wu et at. 1980:115 Pareurythoe borealis (Sars, 1862) China Yang & Sun 1988:170 Pherecardia striata (Kinberg, 1857) Philippines Pillai 1965:121 China Wu et at. 1980:115 Pseudeurythoe hirsuta Wesenberg-Lund, 1949 S. Vietnam Gallardo 1968:57 Pseudeurythoe oligobranchia Wu et aI, China Wu et at. 1980:115

Euphrosinidae Euphrosyne foliosa Audouin & M. Edw., 1833 China Sun et at. 1978:161 Euphrosine myrtosa Savigny, 1818 Hong Kong Mak 1982:599 China Yang & Sun 1988:172

Phyllodocidae Anaitides madeirensis (Langerhans, 1880) China Sun et at. 1978:135 as P.(Anait.)mad. Hong Kong Mak 1982:599 Singapore Tan & Chou 1993:282 Anaitides tenuissima (Grube, 1878) Philippi nes Grube 1878:95 Anaitides ?tenuissima S. Vietnam Gallardo 1968:57 Eteone (Mysta) siphodonta (Delle Chiaje, 1822) Singapore Tan & Chou 1993:282 Eteone (Mysta) tchangsii Usehakov & Wu, 1959 China Yang & Sun 1988:109 Eulalia (Pterocirrus) magalhaensis Kinberg, 1866 Singapore Tan & Chou 1993:282 Eulalia viridis (Linnaeus, 1767) Hong Kong Mak 1982:599 China Yang & Sun 1988: 105 Eumida sanguinea (Marenzeller, 1879) Hong Kong Mak 1982:599 Genetyllis castanea (Marenzeller, 1879) Hong Kong Mak 1982:599 China Yang & Sun 1988: 108 Philippines Natividad & Palpal-Latoe 1986:275 Genetyllis sp. Taiwan Hsieh, pers. eomm. Mysta ornata (Grube, 1878) S. Vietnam Gallardo 1968:58 China Yang & Sun 1988:109 Notophyllum splendens (Sehmarda, 1861) China Wu et at. 1980:115 Paranaitis zeylanica (Willey, 1905) S. Vietnam Gallardo 1968:58 Phyllodoce (Anaitides) dissotyla Willey, 1905 China Wu & Sun 1978:176 Phyllodoce (Anait.) madeirensis Langerhans, 1880 China Wu et at. 1980:115 Phyllodoce fristedti Bergstriim, 1914 China Wu et at. 1980:115

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Phyllodoce ?laminosa Savigny, 1818 S. Vietnam Gallardo 1968:58 Phyllodoce malmgreni Gravier, 1900 China Yang & Sun 1988:110 Pterocirrus macroceros (Grube, 1860) China Yang & Sun 1988:111

Alciopidae Alciopina parasitica Claparecte & Panceri, 1867 China Shen 1978:215 Krohnia lepidota (Krohn, 1845) China Sun & Wu 1979:64 Naiades cantrainii Delle Chiaje, 1830 China Shen 1978:211 Plotohelmis capitata (Greeff, 1876) China Shen 1978:214 Rhynchonerella petersii (Langerhans, 1880) China Sun & Wu 1979:63 Rhynchonerella gracilis Costa, 1864 China Wu et at. 1980:116 Rhynchonerella xishaensis Shen, 1978 China Shen 1978:214 Vanadis crystallina Greeff, 1876 China Shen 1978:212 Vanadis juscapunctata Treadwell, 1906 China Shen 1978:212 Vanadis minuta Treadwell, 1906 China Shen 1978:212 Singapore Tan & Chou 1993:282

Lopadorrhynchidae Lopadorrynchus brevis Grube, 1855 China Shen 1978:203 Lopadorrynchus krohni (Claparecte, 1870) China Shen 1978:205 Lopadorrhynchus tuberculus Shen, 1978 China Shen 1978:205 Lopadorrhynchus uncinatus Fauvel, 1915 China Shen 1978:203 Maupasia caeca Vignier, 1886 China Shen 1978:207 Pedinosoma curtum Reibisch, 1895 China Shen 1978:207 Pelagobia longicirrata Greeff, 1879 China Shen 1978:206

Lacydoniidae Paralacydonia paradoxa Fauvel, 1913 S. Vietnam Gal1ardo 1968:59 China Wu et at. 1980:117 Hong Kong Shin 1982: 163

Iospilidae Phalacrophorus pietus Greeff, 1879 China Shen 1978:208 Phalocrophorus uniformis Reibisch, 1895 China Shen 1978:209

Pontodoridae Pontodora pelagica Greef, 1879 China Shen 1978:209

Typhloscolecidae Sagitella kowalevskii Wagner, 1872 China Shen 1978:222 Travisiopsis dubia Stop-Bowitz, 1948 China Wu et at. 1980:117 Travisiopsis levinseni Southern, 1910 China Shen & Wu 1978:105 Travisiopsis lobifera Levinsen, 1885 China Shen 1978:223 Typhloscolex muelleri Busch, 1851 China Shen 1978:221

Tomopteridae Tomopteris apsteini Rosa, 1908 China Sun & Wu 1979:65 Tomopteris cavalii Rosa, 1908 China Shen 1978:218 Tomopteris duccii Rosa, 1908 China Shen 1978:220

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Tomopteris dunckeri Rosa, 1908 China Wu et al. 1980: 117 Tomopteris elegans Chun, 1888 China Shen 1978:217 Tomopteris ligulata Rosa, 1908 China Shen 1978:220 Tomopteris mariana Greef, 1885 China Wu et al. 1980:117 Tomopteris planktonis Apstein, 1900 China Shen 1978:218 Tomopteris rolasi Greef,1885 China Shen & Wu 1978:103 Tomopteris septentrionalis Steenstrup, 1849 China Sun & Wu 1979:67

Hesionidae Bonuania parva Pillai, 1965 Phillippines Pillai 1965: 123 Gyptis labatus (Hessle, 1925) China Yang & Sun 1988:80 Gyptis cf. maraunibinae Taiwan Hsieh, pers. eomm. Hesione genetta Grube, 1866 China Wu et al. 1980: 117 Hesione intertexta Grube, 1878 Philippines Grube 1878: China Wu et al. 1980:117 Hesione splendida Savigny, 1818 China Wu et al. 1980:117 Leocrates chinensis Kinberg, 1866 Hong Kong Mak 1982:599 China Yang & Sun 1988:82 Leocrates claparedii (Costa in C1aparede, 1868) Singapore Tan & Chou 1993:282 Ophiodromus berrisfordi Day, 1967 China Wu & Wu 1987:79 Ophiodromus pugettensis (Johnson, 1901) Hong Kong Shin 1982:164 Podarke latifrons (Grube 1878) Singapore Tan & Chou 1993:282 Podarke cf. obscura (Verrill, 1837) China Yang & Sun, 1988:84 Sinohesione genitaliphora Westheide et aI., 1994 Thailand Westheide et al. 1994:95

Pilargidae Ancistargis sp. Taiwan Hsieh, pers. eomm. Loandalia indica Thomas, 1963 S. Vietnam Gallardo 1968:60 Pilargis mohri Gallardo, 1968 S. Vietnam Gallardo 1968:60 Pilargis verrucosa pacific Usehakov, 1955 China Yang & Sun 1988:77 Sigambra cf. bassi Taiwan Hsieh, pers. eomm. Sigambra hanaokai (Kitamori, 1960) S. Vietnam Gallardo 1968:61 Sigambra tentaculata (Treadwell, 1941) Hong Kong Shin 1982:164 Synelmis albini (Langerhans, 1881) S. Vietnam Gallardo 1968:61 China Wu et al. 1980:117 Hong Kong Shin 1982: 164 Synelmis annamita Gallardo, 1968 S. Vietnam Gallardo 1968:62

Syllidae Autolytus robustisetus Wu & Sun, 1978 China Sun et al. 1978:137 Autolytus setoensis 1majima, 1966 China Yang & Sun 1988:55 Autolytus sp. Taiwan Hsieh, pers. comm. Ehlersia cornuta (Rathke, 1843) Hong Kong Shin 1982: 164 China Yang & Sun 1988:65 Ehlersia ef. rosea Imajima, 1966 China Yang & Sun 1988:66 Eusyllis edenticulata Pillai, 1965 Philippines Pillai 1965:125 Eusyllis ef. irregulata Imajima, 1966 China Yang & Sun, 1988:57

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Exogone verugera (Claparede, 1868) China Sun & Wu 1981:621 Haplosyllis spongicola (Grube, 1855) China Yang & Sun 1988:67 Haplosyllis spongicola tentaculata (Marion, 1879) China Yang & Sun 1988:67 Langerhansia cornuta (Rathke, 1843) S. Vietnam Gallardo 1968:62 Myrianida cf. pachycera (Augener, 1913) China Yang & Sun 1988:56 Odontosyllis gibba Claparede, 1863 China Sun et al. 1978:140 Odontosyllis rubrofasciata Grube, 1879 China Wu & Sun 1978:178 Parautolytus luzonensis Pillai, 1965 Philippines Pillai 1965:123 Sphaerosyllis hystrix Claparede, 1863 China Sun & Wu 1981:622 Streptosyllis baolingi Ding & Westheide, 1994 China Ding & Westheide 1994:304 Streptosyllis hainanensis Ding & Westheide, 1994 China Ding & Westheide 1994:308 Syllis amica Quatrefages, 1865 China Yang & Sun 1988:68 Syllis gracilis Grube, 1840 China Yang & Sun 1988:69 Singapore Tan & Chou 1993:283 Trypanosyllis taeniaformis (Haswell, 1886) Hong Kong Mak 1982:599 China Yang & Sun 1988:71 as T. (Tryp. )taen. Trypanosyllis zebra (Grube, 1860) China Wu et al. 1980:118 Typosyllis aciculata orientalis Imajima & Hartman, 1964 S. Vietnam Gallardo 1968:63 Hong Kong Mak 1982:599 Typosyllis alternata (Moore, 1908) China Wu & Sun 1978:179 Typosyllis armillaris (Muller, 1771) China Wu et al. 1980:118 T)posyllis exilis Gravier, 1900 Philippines Natividad & Palpal-Latoc 1986:310 Typosyllis maculata Imajima, 1966 China Yang & Sun 1988:74 Typosyllis magnipectinis (Storch, 1967) China Licher et al. 1995: 107 Typosyllis monilata Imajima, 1966 China Sun et al. 1978:141 Typosyllis prolifera (Krohn, 1852) China Wu et al. 1980:118 Typosyllis variegata (Grube, J860) S. Vietnam Gallardo 1968:63 China Sun et al. 1978: 140 Hong Kong Mak 1982:599

Nereididae Ceratonereis anchylochaeta (Hors!, 1924) China Wu et al. 1981:Table 1 Ceratonereis cf. burmensis Monro, 1937 China Sun et al. 1978: 148 Ceratonereis costae (Grube, 1840) China Sun et al. 1978:146 Hong Kong Mak 1982:599 Ceratocephale fauveli (Pillai, 1961) S. Vietnam Gallardo 1968:63 Ceratonereis erythraeensis Fauvel, 1918 China Wu et al. 1981:Table 1 Ceratonereis hircinicola (Eisig, 1870) China Wu et al. 1981:Table I Ceratonereis ?hircinicola Singapore Tan & Chou 1993:283 Ceratonereis japonica Imajima, 1972 China Sun et al. 1978:145 Ceratonereis marmorata Horst, 1924 China Wu et al. 1981:Table 1 Ceratonereis mirabilis Kinberg, 1866 China Sun et al. 1978:144 Hong Kong Mak 1982:599 Singapore Tan & Chou 1993:283 Ceratonereis pachychaeta Fauvel,1918 China Sun et al. 1978:147 Ceratonereis tripartita Horst, 1918 China Wu et al. 1980:118

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Ceratonereis sp. Taiwan Hsieh, pers. comm. Gymnonereis fauveli (Pillai, 1961) Thailand Hylleberg & Nateewathana 1988:8 Dendronereis pinnaticirris Grube, 1878 Philippines Pillai 1965:127 China Wu et al. 1981:Tb1e 1 Taiwan Hsieh, pers. comm. Leonnates decipiens Fauve1, 1929 China Wu et al. 1981 :Tab1e 1 Singapore Tan & Chou 1993:283 Leonnates decipiens manilensis Pillai, 1965 Philippines Pillai 1965:144 Leonnates jousseaumei Gravier, 1911 China Wu et al. 1981:Table 1 Singapore Tan & Chou 1993:283 Leonnates nipponicus 1majima, 1972 Hong Kong Mak 1982:599 Leonnates persica Wesenberg-Lund, 1949 S. Vietnam Gallardo 1968:64 China Wu et al. 1981 :Tab1e 1 Hong Kong Shin 1982: 165 Leonnates sp. Taiwan Hsieh, pers. comm. Namalycastis aibiuma (Muller in Grube, 1881) China Wu et al. 1981:Tab1e 1 Taiwan Hsieh, pers. comm. Namalycastis rigida Pillai, 1965 Philippines Pillai 1965:133 Neanthes arenaceodentata (Moore, 1903) S. Vietnam Gallardo 1968:65 Neanthes bongcoi Pillai, 1965 Philippines Pillai 1965:142 Neanthes diversicolor (Muller, 1776) Philippines Natividad & Pa1pa1-Latoc 1986:315 Neanthes glandicincta (Southern, 1921) China Wu et al. 198 I:Tab1e 1 Taiwan Hsieh, pers. comm. Neanthes maculata Wu et aI., 1981 China Wu et al. 1981:66 Neanthes manatensis Pillai, 1965 Philippines Pillai 1965:139 Neanthes nanhaiensis Wu et aI., 1981 China Wu et al. 1981:68 Neanthes unifilSciata (Willey, 1905) China Wu et al. 1980:119 Hong Kong Mak 1982:599 Nectoneanthes ijimai (Izuka, 1912 ) China Wu et al. 1981:Tab1e 1 Nectoneanthes multignatha Wu et aI., 1981 China Wu et al. 1981:73 Nectoneanthes oxypoda (Marenzeller, 1879) China Wu et al. 1981:Tab1e I Nectoneanthes cf. oxypoda Hong Kong Shin 1982: 164 Nereis coutieri Gravier, 1899 China Wu et al. 1981 :Table I Ne re is dayana Sun & Shen, 1978 China Sun et al. 1978:155 Nereis denhamensis Augener, 1913 China Sun et al. 1978:151 Nereis falcaria (Willey, 1905) China Wu et al. 1980:119 Nereis guangdongensis Wu et aI., 1981 China Wu et al. 1981:75 Nereis heteromorpha Horst, 1924 China Sun et al. 1978:157 Nereis jacksoni Kinberg, 1866 China Wu et al. 1981:Tab1e I Nereis multignatha Imajima & Hartman, 1964 China Wu et al. 1981 :Tab1e 1 Nereis nichollsi Kotl, 1951 China Wu et al. 1980:]]9 Nereis pers;c'a Fauve1, 1911 China Wu et al. 1981:Table 1 Hong Kong Shin 1982: 165 Nereis surugaense nanhaiensis Sun & Shen, 1978 China Sun et al. 1978:153 Nereis trifasciata Grube, 1878 China Wu et al. 1980:119 Nereis unifasciata Willey,1905 China Sun et al. 1978:154 Nereis zhongshaensis Shen & Sun, 1978 China Sun et al. 1978:158

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Nereis zonata Malmgren, 1867 China Wu et al. 1980:119 Nereis cf. zonata Taiwan Hsieh, pers. comm. Nicon japonica 1majima, 1972 China Wu & Sun 1979:104 Nicon maculata Kinberg, 1866 S. Vietnam Gallardo 1968:65 as N. maculatus China Wu & Sun 1979:101 Nicon sinica Wu & Sun, 1979 China Wu & Sun 1979:99 Paraleonnates uschakovi Chlebovitsch & Wu, 1962 China Wu et al. 1981:Table I Perinereis aibuhitensis Grube, 1878 China Wu et al. 1981:Table 1 Taiwan Hsieh, pers. comm. Perinereis barbara Momo, 1926 Singapore Tan & Chou 1993:283 Perinereis camiguinoides Augener, 1922 China Wu et al. 1981:Table 1 Perinereis cavifrons Ehlers, 1920 China Wu et al. 1981:Table I Perinereis cultrifera cultrifera (Grube, 1840) China Wu et al. 1981:Table I Hong Kong Mak 1982:599 Philippines Natividad & Palpal-Latoc 1986:313 Perinereis cultrifera jloridana Ehlers, 1868 China Wu et al. 1985:218 Perinereis cultrifera helleri Grube, 1878 China Wu et al. 1985:217 Perinereis cultrifera obfuscata Grube, 1878 China Wu et al. 1985:219 Perinereis cultrifera perspicillata Grube, 1878 Singapore Tan & Chou 1993:283 Perinereis cultrifera striolata Grube, 1878 Singapore Tan & Chou 1993:284 Perinereis nuntia (Savigny, 1818) China Wu et al. 1981:Table 1 Perinereis nuntia brevicirris (Grube, 1878) Singapore Tan & Chou 1993:284 China Wu et al. 1985:208 Perinereis nuntia majungaensis Fauvel, 1918 China Wu et al. 1985:213 Perinereis nuntia vallata (Grube, 1857) China Wu et al. 1985:210 Perinereis quatrefagesi (Grube, 1878) Thailand Hylleberg et al. 1986:5 Perinereis rhombodonta Wu et aI., 1981 China Wu et al. 1981:83 Perinereis singaporiensis (Grube, 1878) Singapore Grube 1878:84 Perinereis suluana Horst, 1924 China Sun et al. 1978: 150 Perinereis vancaurica (Ehlers, 1868) China Wu et al. 1981:Table 1 Singapore Tan & Chou 1993:284 Perinereis weijhouensis Wu et aI., 1981 China Wu et al. 1981:83 Perinereis cf. weijhouensis Taiwan Hsieh, pers. comm. Platynereis abnormis (Horst, 1924) China Wu et al. 1980:119 Platynereis bicanaliculata (Baird, 1863) China Wu et al. 1981:Table 1 Platynereis dumerilli (Audouin & M. Edw., 1833) China Wu et al. 1980:119 Platynereis pulchella Gravier, 1902 S. Vietnam Gallardo 1968:65 China Wu et al. 1981:Table 1 Platynereis sinica Sun & Shen, 1978 China Sun et al. 1978: 148 Pseudonereis anomala Gravier, 1902 China Wu et al. 1980:119 Pseudonereis gallapagensis Kinberg, 1866 China Wu et al. 198\ :Table 1 Pseudonereis variegata (Grube, 1857) China Wu et al. 1981:Table 1 Rullierinereis misakiensis (Imajima & Hayashi, 1972) China Wu & Sun 1979:106 Sinonereis heteropoda Wu & Sun, 1979 China Wu & Sun 1979:96 Tambalagamia fauveli Pillai, 1961 China Yang & Sun 1988:28 Taiwan Hsieh, pers. comm. Tylonereis bogoyawleskyi Fauvel, 1911 China Wu et al. 1981:Table 1

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Tylonereis heterochaeta Tan & Chou, 1994 Singapore Tan & Chou 1994:665 Tylorrhynchus heterochaetus (Quatrefages, 1865) China Wu et al. 1981:Tab1e 1

Nephtyidae Aglaophamus dibranchis (Gmbe, 1877) Singapore Tan & Chou 1993:284 Aglaophamus dicirroides Fauehald, 1968 S. Vietnam Faueha1d 1968: 10 Aglaophamus ef. jeffreysii (McIntosh, 1885) China Wu et al. 1980:120 Aglaophamus lyrochaeta (Fauve1, 1902) Hong Kong Shin 1982: 165 Aglaophamus munamaorii Gibbs, 1971 China Wu et al. 1980:120 Aglaophamus orientalis Faueha1d, 1968 S. Vietnam Faucha1d 1968:12 Aglaophamus sinensis (Fauve1, 1932) S. Vietnam Fauchald 1968:13 China Yang & Sun 1988:94 Aglaophamus vietnamensis Faucha1d, 1968 S. Vietnam Faucha1d 1968:13 Aglaophamus sp. Taiwan Hsieh, pers. comm. Inermonephtys gallardi Faueha1d, 1968 S. Vietnam Faucha1d 1968: 15 Inermonephtys inermis (EhIers, 1887) S. Vietnam FauchaId 1968:16 China Wu et al. 1980:120 Micronephthys sphaerocirrata (Wesenberg-Lund, 1949) S. Vietnam Fauchald 1968: 18 Singapore Tan & Chou 1993:284 Micronephthys sp. Taiwan Hsieh, pers. comm. Nephtys cf. califomiensis Taiwan Hsieh, pers. comm. Nephtys oligobranchia Southern, 1921 S. Vietnam Faucha1d 1968: 18 Nephtys polybranchia Southern, 1921 S. Vietnam FauchaId 1968:19 China Yang & Sun 1988:98

Sphaerodoridae Sphaerodorum sp. S. Vietnam Gallardo 1968:67

Glyceridae Glycera alba (Muller, 1788) S. Vietnam Gallardo 1968:68 Hong Kong Shin 1982:166 China Yang & Sun 1988:87 Glycera ?capitata Oersted, 1843 S. Vietnam Gallardo 1968:68 Glycera chirori lzuka, 1912 China Yang & Sun 1988:87 Glycera convoluta Keferstein, 1862 Singapore Tan & Chou 1993:284 Glycera cf. convoluta Taiwan Hsieh, pers. comm. Glycera lancadivae Schmarda, 1861 China Sun et al. 1978:136 Glycera nicobarica Grube, 1868 S. Vietnam Gallardo 1968:69 Glycera onomichiensis Izuka, 1912 S. Vietnam Gallardo 1968:69 China Wu et al. 1980:120 Glycera posterobranchia Hoagland, 1920 Philippines Hoag1and 1920:620 S. Vietnam Gallardo 1968:70 Glycera parashadi Fauve1, 1932 China Wu et al. 1980:120 Glycera rouxii Audouin & M. Edw., 1833 China Wu & Wu 1987:79 Singapore Tan & Chou 1993:285 Glycera cf. subaenea Taiwan Hsieh, pers. comm. Glyera cf. tenuis Taiwan Hsieh, pers. comm. Glycera tesselata Gmbe, 1863 S. Vietnam Gallardo 1968:70 China Wu et al. 1980:119

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Goniadidae Glyeinde oligodon Southern, 1921 S. Vietnam Gallardo 1968:71 Goniada annulata Moore, 1905 China Wu et al. 1980:119 Goniada emerita Audouin & M. Edw., 1833 China Wu et al. 1980:119 Goniada cf. emerita Taiwan Hsieh, pers. comm. Goniada cf. maeulata Oersted, 1843 China Wu et al. 1980:119 Singapore Tan & Chou 1993:285 Taiwan Hsieh, pers. comm. Goniada multidentata Arwidsson, 1899 S. Vietnam Gallardo 1968:72 Goniada paueidens Grube, 1878 Hong Kong Mak 1982:599 Goniada tridens Gallardo, 1968 S. Vietnam Gallardo 1968:73 Goniada uneinigera EhJers, 1901 S. Vietnam Gallardo 1968:73 Goniada cf. uneinigera Taiwan Hsieh, pers. comm. Goniadopsis sp. Taiwan Hsieh, pers. comm.

Dorvilleidae Dorvillea australiensis (McIntosh, 1885) China Wu et al. 1980:121 as Papilliod.aust. Sehistomeringos ineerta (Schmarda, 1861) S. Vietnam Gallardo 1968:91 as Dorvillea ine. Hong Kong Shin 1982: 167

Eunicidae Euniee afra Peters, 1854 China Wu & Wu 1987:79 Euniee antennata (Savigny, 1820) Philippines Pi1lai 1965: 148 China Wu et al. 1980:121 Hong Kong Mak 1982:599 Singapore Tan & Chou 1993:285 Euniee aphroditois (Pallas, 1788) China Wu & Wu 1987:79 Singapore Tan & Chou 1993:285 Euniee ?australis Quatrefages, 1865 S. Vietnam Gallardo 1968:77 Euniee australis Hong Kong Mak 1982:599 China Wu & Wu 1987:79 Euniee eineta (Kinberg, 1865) Hong Kong Mak 1982:599 Euniee eoeeinea Grube, 1878 China Wu et al. 1980:121 Euniee eoeeinioides Augener, 1922 Singapore Tan & Chou 1993:285 Euniee grubei Gravier, 1900 Singapore Tan & Chou 1993:286 Euniee hirsehi Fauchald, 1992 Singapore Tan & Chou 1993:286 Euniee indiea Kinberg, 1865 S. Vietnam Gallardo 1968:78 Philippines Gallardo 1968:78 China Wu et al. 1980:121 Euniee lueei Grube, 1856 Singapore Tan & Chou 1993:286 Euniee marenzelleri Gravier, 1900 China Wu et al. 1980:121 Euniee medicina Moore, 1903 S. Vietnam Gallardo 1968:78 Euniee nesiotes (Chamberlin, 1919) Singapore Tan & Chou 1993:286 Euniphysa aeuleata Wesenberg-Lund, 1949 China Shen & Wu 1991:131 Singapore Tan & Chou 1993:286 Euniphysa oeulata Wu & Sun in Wu et aI., 1979 China Wu et al. 1979:89 Euniphysa unieusa Shen & Wu 1991 China Shen & Wu 1991:131 Heterophysa tridontesa Shen & Wu, 1991 China Shen & Wu 1991:138

219 Paxton & Chou: Polychaetous annelids from the South China Sea

NAME AUTHOR COUNTRY REFERENCE

Lysidice collaris Grube, 1870 S. Vietnam Gallardo 1968:79 Philippines Gallardo 1968:79 China Sun et al. 1978: 162 as L.nin.collaris Hong Kong Mak 1982:599 Singapore Tan & Chou 1993:287 Lysidice ninetta Audouin & M. Edw., 1833 Hong Kong Mak 1982:599 China Yang & Sun 1988:184 Marphysa adenensis Gravier, 1900 Hong Kong Mak 1982:600 Marphysa bellii (Aud. & M. Edwards,1833) S. Vietnam Gallardo 1968:80 Thailand Gallardo 1968:80 Hong Kong Shin 1982: 166 Marphysa bifurcata Kott, 1951 S. Vietnam Gallardo 1968:80 Marphysa depressa (Schmarda, 1861) China Yang & Sun 1988:186 Marphysa disjuncta Hartman, 1961 Singapore Tan & Chou 1993:286 Marphysa gravelyi Southern, 1921 Philippines Pillai 1965: 150 Marphysa macintoshi Crossland, 1903 China Wu et al. 1980:121 Singapore Tan & Chou 1993:286 Marphysa mossambica (Peters, 1854) China Wu & Wu 1987:79 Singapore Tan & Chou 1993:287 Marphysa sanguinea (Montagu, 1815) Hong Kong Shin 1982: 166 China Yang & Sun 1988:186 Marphysa stragulum (Grube, 1878) China Wu et al. 1980:121 Nauphanta sp. Tan & Chou, in press Singapore Tan & Chou, in press Nematonereis unicornis (Grube, 1840) S. Vietnam Gallardo 1968:81 China Wu et al. 1980:121 Palola siciliensis (Grube, 1840) China Wu et al. 1980:121 Hong Kong Mak 1982:600 Paraeuniphysa falciseta Shen & Wu, 1991 China Shen & Wu 1991:134 Paraeuniphysa spinea (Miura, 1977) China Shen & Wu 1991:136 Paraeuniphysa taiwanensis Wu & He, 1988 Taiwan Shen & Wu 1991:134 Paramarphysa sp. Hong Kong Mak 1982:600

Lumbrineridae Lumbrineris acutiformis Gallardo, 1968 S. Vietnam Gallardo 1968:82 Lumbrineris ? amboinensis (Grube, 1877) S. Vietnam Gallardo 1968:82 Lumbrineris bifurcata (Mclntosh, 1885) China Wu et al. 1980:121 Lumbrineris caudaensis Gallardo, 1968 S. Vietnam Gallardo 1968:83 Lumbrineris cf. cruzensis Hartman, 1944 China Wu et al. 1980:121 Taiwan Hsieh, pers. comm. Lumbrineris hartmani (Day, 1953) Singapore Tan & Chou 1993:287 Lumbrineris heteropoda (Marenzeller, 1879) S. Vietnam Gal1ardo 1968:84 China Yang & Sun 1988:192 Lumbrineris inflata Moore, 1911 Hong Kong Mak 1982:600 Lumbrineris japonica (Marenzeller, 1879) S. Vietnam Gallardo 1968:84 Lumbrineris latreilli Audouin & M. Edw., 1834 China Wu et al. 1980:121 Singapore Tan & Chou 1993:287 Lumbrineris mucronata (Ehlers, 1908) S. Vietnam Gallardo 1968:85 Lumbrineris nagae Gallardo, 1968 S. Vietnam Gallardo 1968:85 China Yang & Sun 1988:191

220 THE RAFFLES BULLETIN OF ZOOLOGY 2000 Supplement No. 8

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Lumbrineris ?notocirrata (Fauvel, 1932) S. Vietnam Gallardo 1968:86 Lumbrineris pterignatha Gallardo, 1968 S. Vietnam Gallardo 1968:86 Lumbrineris shiinoi Gallardo, 1968 S. Vietnam Gallardo 1968:87 Hong Kong Shin 1982: 167 Lumbrineris simplex Southern, 1921 China Wuetal.1980:121 Lumbrineris sphaerocephala (Sehmarda, 1861) China Wu et al. 1980: 121 Ninoe bruuni Gallardo, 1968 S. Vietnam Gallardo 1968:88 Ninoe mando (Crossland, 1924) S. Vietnam Gallardo 1968:88

Oenonidae Arabella iricolor (Montague, 1804) Singapore Tan & Chou 1993:287 Arabella mu tans (Chamberlin, 1919) China Wu & Wu 1987:79 Arabella novecrinita Crossland, 1924 S. Vietnam Gallardo 1968:89 Drilonereis logani Crossland, 1924 S. Vietnam Gallardo 1968:89 China Wu et al. 1980:121 Drilonereis ef. logani Taiwan Hsieh, pers. eomm. Drilonereis ?paucidentata Treadwell, 1922 S. Vietnam Gallardo 1968:90 Notocirrus biaciculus Sehmarda, 1861 S. Vietnam Gallardo 1968:90 Notocirrus japonicus (Okuda, 1939) S. Vietnam Gallardo 1968:91 Oenone julgida (Savigny, 1818) Hong Kong Mak 1982:600 Singapore Tan & Chou 1993:287

Onuphidae Diopatra bulohensis Tan & Chou, 1996 Singapore Tan & Chou 1996:359 Diopatra neapolitana Delle Chiaje, 1841 Singapore Tan & Chou 1993:285 Diopatra sugokai lzuka, 1907 China Yang & Sun 1988:175 as D. chiliensis Taiwan Hsieh & Chang 1991 :331 as D. bilobata Diopatra variabilis Southern, 1921 S. Vietnam Gallardo 1968:74 China Wu et al. 1980:120 Hong Kong Shin 1982:166 Hyalinoecia tubicola (MUller, 1788) China Wu et al. 1980: 120 Onuphis punggolensis Tan & Chou, in press Singapore Tan & Chou, in press Onuphis sp. Taiwan Hsieh, pers. eomm. Nothria conchylega (Sars, 1835) S. Vietnam Gallardo 1968:75 Nothria holobranchiata (Marenzeller, 1879) S. Vietnam Gallardo 1968:75 Nothria nr. atlantisa Hartman, 1965 S. Vietnam Gallardo 1968:75 Onuphis eremita Audouin & M. Edw. 1833 S. Vietnam Gallardo 1968:76 China Wu et al. 1980:120 Onuphis pseudodibranchiata Gallardo, 1968 S. Vietnam Gallardo 1968:76 China Wu et al. 1980:120 Onuphis variolata Shen, 1987 China Shen 1987:222 Paradiopatra willemoesii (Mclntosh, 1885) China Wu et al. 1980:120

Orbiniidae Haploscoloplos elongatus (Johnson, 1901) S. Vietnam Gallardo 1968:92 China Yang & Sun 1988:204 Haploscoloplos ef. jragilis (Verrill, 1873) S. Vietnam Gallardo 1968:92

221 Paxton & Chou: Polychaetous annelids from the South China Sea

NAME AUTHOR COUNTRY REFERENCE

Leitoscoloplos elongatus Taiwan Hsieh, pers. eomm. Leitoscoloplos ef. kerguelensis (McIntosh, 1885) Hong Kong Shin 1982:167 Naineris hainanensis Q. WU China Wu & Wu 1987:79 Orbinia ef. dicrochaeta Taiwan Hsieh, pers. eomm. Orbinia vietnamensis Gallardo, 1968 S. Vietnam Gallardo 1968:93 China Yang & Sun 1988:205 Phylo kupfferi (Eh1ers, 1875) Hong Kong Shin 1982: 167 Phylo ornatus (Verrill, 1873) China Yang & Sun 1988:206 Scoloplos cf. acmeceps Taiwan Hsieh, pers. eomm. Scoloplos ef. armiger (Muller, 1776) China Wu et al. 1980:122 Scoloplos (Leodamas) ?dubia Tebble, 1955 S. Vietnam Gallardo 1968:94 Scoloplos (L.) gracilis Pillai, 1961 S. Vietnam Gallardo 1968:94 Scoloplos (L.) rubra (Webster, 1879) China Yang & Sun 1988:208 Scoloplos (L.) rubra orientalis Gallardo, 1968 S. Vietnam Gallardo 1968:95 Scoloplos cf. rubra Taiwan Hsieh, pers. eomm. Scoloplos (Scoloplos) marsupialis Southern, 1921 China Yang & Sun 1988:209 Scoloplos (Scoloplos) spiniferus Gallardo, 1968 S. Vietnam Gallardo 1968:95 Scoloplos tumidus Maekie, 1991 Hong Kong Maekie 1991:38 S. Vietnam Gallardo 1968:95 as S. marsupialis Scoloplos cf. tumidus Taiwan Hsieh, pers. eomm.

Paraonidae Aricidea cf. fragilis caeca Taiwan Hsieh, pers. eomm. Paradoneis lyra (Southern, 1914) S. Vietnam Gallardo 1968:96

Spionidae Aonides sp. China Wu & Sun 1978:187 Aonides ef. oxycephala Taiwan Hsieh, pers. eomm. Dispio ef. uncinata Taiwan Hsieh, pers. eomm. Laonice cirrata (M. Sars, 1851) S. Vietnam Gallardo 1968:97 China Wu & Sun 1978:192 Hong Kong Shin 1982:167 Malacoceros (Rh.) areninc.asiaticus (Ch1ebovitseh, 1959 ) China Wu & Sun 1978:189 Malacoceros indicus (Fauve1, 1928) China Wu et al. 1980:122 Taiwan Hsieh, pers. eomm. Microspio sp. China Wu & Sun 1978:191 Minuspio cirrifera (Wiren, 1883) Hong Kong Shin 1982:167 Nerine faughni Gallardo, 1968 S. Vietnam Gallardo 1968:97 Paraprionospio pinnata (Ehlers, 1901) Hong Kong Shin 1982: 167 China Yang & Sun 1988:218 Paraprionospio sp. Taiwan Hsieh, pers. eomm. Polydorella novaegeorgiae Gibbs, 1971 Hong Kong Shin 1982: 168 Polydora cf. pilikia Ward, 1981 China Yang & Sun 1988:213 Polydora cf. socialis (Sehmarda, 1861) Hong Kong Shin 1982: 167 Polydora ef. tentaculata Blake & Kudenov,1978 Hong Kong Shin 1982: 168 Taiwan Hsieh, pers. eomm. Polydora cavitensis Pillai, 1965 Philippines Pillai 1965:152

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Polydora glycymerica Radashevsky, 1993 Manchenko & Radashevsky 1994:767 Polydora ligni Webster, 1879 Taiwan Hsieh, pers. comm. Polydora vulgaris Mohammad, 1972 Manchenko & Radashevsky 1994:767 Prionmpio africana Augener, 1918 S. Vietnam Gallardo 1968:98 Prionospio depauperata Taiwan Hsieh, pers. comm. Prionospio ehlersi Fauvel, 1928 S. Vietnam Gallardo 1968:98 China Wu et al. 1980: 122 Hong Kong Shin 1982: 168 Prionospio japonica Taiwan Hsieh, pers. comm. Prionospio komaeti Hylleberg & Nateew., 1991 Singapore Tan & Chou 1993:288 Prionospio malmgreni Claparede, 1870 China Wu & Sun 1978:189 Hong Kong Shin 1982: 168 Prionospio malayensis Caullery, 1914 S. Vietnam Gallardo 1968:99 Singapore Tan & Chou 1993:288 Prionospio paradisea Taiwan Hsieh, pers. comm. Prionospio pinnata Eh1ers, 1901 S. Vietnam Gallardo 1968:99 Prionospio cf. pulchra Taiwan Hsieh, pers. comm. Prionospio saccifera Mackie & H. 1990 Hong Kong Mackie & Hart1ey 1990:366 Prionospio sishaensis Wu & Chen, 1964 China Wu & Wu 1987:79 Pseudopolydora antennata Claparede, 1870 China Wu & Sun 1978:183 Pseudopolydora diopatra Hsieh, 1992 Taiwan Hsieh 1992:630 Pseudopolydora cf. kempi Taiwan Hsieh, pers. comm. Pygospio elegans C1aparede, 1863 China Wu & Sun 1978:186 Scolelepis ? indica Fauvel, 1928 S. Vietnam Gallardo 1968:100 Scolelepis sp. Taiwan Hsieh, pers. comm. Spiophanes bombyx (C1aparede, 1870) China Wu & Sun 1978:185 Spiophanes malayensis Caullery, 1915 China Wu et al. 1980:122 Spiophanes sp. Taiwan Hsieh, pers. comm.

Heterospionidae Heterospio sinica Wu & Chen, 1966 China Yang & Sun 1988:221

Magelonidae Magelona cf. cincta Taiwan Hsieh, pers. comm. Magelona crenulifrons Gallardo, 1968 S. Vietnam Gallardo 1968: 100 Hong Kong Shin 1982: 168 Magelona japonica Okuda, 1937 S. Vietnam Gallardo 1968:101 China Wu et a1 1980:122 Magelone koreana Okuda, 1937 S. Vietnam Gallardo 1968:101 Magelona lenticulata Gallardo, 1968 S. Vietnam Gallardo 1968: 101 Magelona cf. pacifica Taiwan Hsieh, pers. comm. Magelona sp. China Wu & Sun 1978:195

Cossuridae Cossura coasta Kitamori, 1960 Hong Kong Shin 1982: 169 Cossurella cf. diamorpha Taiwan Hsieh, pers. comm.

223 Paxton & Chou: Polychaetous annelids from the South China Sea

NAME AUTHOR COUNTRY REFERENCE Trochochaetidae Trochochaeta diverapoda (Hoagland, 1920) Hong Kong Mackie 1990:338 Trochochaeta cf. orissae (Fauvel, 1932) Hong Kong Shin 1982:168

Poecilochaetidae gallardoi Pilato & Cantone, 1976 S. Vietnam Pilato & Cantone 1976:43 S. Vietnam Gallardo 1968: 103 as P. sp. A Poecilochaetus hystricosus Mackie, 1990 Hong Kong Mackie 1990: 351 Poecilochaetus paratropicus Gallardo, 1968 S. Vietnam Gallardo 1968: 102 Poecilochaetus serpens Alien, 1904 China Wu et aI 1980:122 Singapore Tan & Chou 1993:288 Poecilochaetus spinulosus Mackie, 1990 Hong Kong Mackie 1990:342 Poecilochaetus tricirratus Mackie, 1990 Hong Kong Mackie 1990:346 Poecilochaetus tropicus Okuda, 1937 China Wu & Sun 1978:180 Poecilochaetus vietnamita Gallardo, 1968 S. Vietnam Gallardo 1968: 103 Poecilochaetus sp. Taiwan Hsieh, pers. comm.

Chaetopteridae Chaetopterus variopedatus (Renier, 1804) Singapore Tan & Chou 1993:288 China Yang & Sun 1988:224 Sasekumaria se/angora Rullier, 1976 Malaysia Ng & Sasekumar 1993:467 Spiochaetopterus costarum (Claparecte, 1868) Hong Kong Shin 1982: 168 costarum

Cirratulidae Caulleriella glabra Gallardo, 1968 S. Vietnam Gallardo 1968: 104 Caulleriella cf. typhlops (Willey, 1905) China Sun et al 1978: 163 Chaetozone flagellifera Gallardo, 1968 S. Vietnam Gallardo 1968: 105 Chaetozone maotienae Gallardo, 1968 S. Vietnam Gallardo 1968:105 Cirratulus annamensis Gallardo, 1968 S. Vietnam Gallardo 1968: 105 Cirratulus cirratulus Muller, 1776 Singapore Tan & Chou 1993:288 Cirratulus filiformis Keferstein, 1862 China Yang & Sun 1988:234 Cirriformia chrysodermoides Pill ai, 1965 Philippines Pillai 1965:156 Cirriformia cf. filiformis Taiwan Hsieh, pers. comm. Cirriformia jiligera (Delle Chiaje, 1828) China Wu 1987:79 Cirriformia punctata (Grube, 1859) China Wu et al. 1980:123 Cirriformia semicincta (Ehlers, 1905) S. Vietnam Gallardo 1968: 106 Cirriformia tentaculata (Montagu, 1808) China Yang & Sun 1988:235 Dodecaceria jewkesi Berkeley & Berk., 1954 China Yang & Sun 1988 :236 Tharyx filibranchia Day, 1961 Hong Kong Shin 1982: 169 Tharyx marioni (Saint-Joseph, 1894) S. Vietnam Gallardo 1968:1 06 Tharyx sp. Taiwan Hsieh, pers. comm.

Flabelligeridae Brada jerruginea Gallardo, 1968 S. Vietnam Gallardo 1968:107 Diplocirrus erythroporus Gallardo, 1968 S. Vietnam Gallardo 1968: 108 Flabelliderma sp. Taiwan Hsieh, pers. comm.

224 THE RAFFLES BULLETIN OF ZOOLOGY 2000 Supplement No. 8

NAME AUTHOR COUNTRY REFERENCE

Pherusa parmata (Grube, 1877) Hong Kong Mak 1982:600 China Yang & Sun 1988:241 Pherusa curvisetis (Caullery, 1944) Philippines Gallardo 1968:108 S. Vietnam Gallardo 1968: 108 Pherusa eruca indica (Fauvel, 1928) S. Vietnam Gallardo 1968:109 Pherusa monroi (Day, 1957) Singapore Tan & Chou 1993:289 Piromis congoensis (Grube, 1877) S. Vietnam Gallardo 1968:109

Scalibregmidae Scalibregma inflatum Rathke, 1843 S. Vietnam Gallardo 1968:110 China Yang & Sun 1988:271

Opheliidae Ammotrypane aulogaster Rathke, 1843 Singapore Tan & Chou 1993:289 Ammotrypane ?dubia Caullery, 1944 S. Vietnam Gallardo 1968: 111 Ammotrypane ?ehlersi Horst, 1919 S. Vietnam Gallardo 1968:111 Ammotrypane ?fauveli Caullery, 1944 S. Vietnam Gallardo 1968:111 Ammotrypane filobranchiata Gallardo, 1968 S. Vietnam Gallardo 1968:112 Ammotrypane grandis Pillai, 1961 S. Vietnam Gallardo 1968:112 Armandia ?intermedia Fauvel, 1902 S. Vietnam Gallardo 1968: 113 Armandia intermedia China Yang & Sun 1988:246 Armandia lanceolata Willey, 1905 S. Vietnam Gallardo 1968: 113 Armandia leptocirris (Grube, 1878) Philippines Grube 1878:194 S. Vietnam Gallardo 1968: 113 China Wu & Wu 1987:79 Armandia ef. leptocirris Taiwan Hsieh, pers. eomm. Armandia longicaudata (Caullery, 1944) S. Vietnam Gallardo 1968:114 Ophelina acuminata Oersted, 1843 China Yang & Sun 1988:248 Ophelina ef. acuminata Taiwan Hsieh, pers. eomm. Polyophthalmus pictus (Dujardin, 1839) Philippines Pillai 1965:157 China Sun et al. 1978: 163

Sternaspidae Sternaspis scutata (Renier, 1807) S. Vietnam Gallardo 1968:114 China Wu et al. 1980:123 Hong Kong Shin 1982:169 Taiwan Hsieh, pers. eomm.

Capitellidae Capitella capitata (Fabrieius, 1780) Singapore Tan & Chou 1993:289 Capitella sp. Taiwan Hsieh, pers. eomm. Capitellides sp. Taiwan Hsieh, pers. eomm. Capitellethus branchiferus Gallardo, 1968 S. Vietnam Gallardo 1968:115 Dasybranchus caducus (Grube, 1846) S. Vietnam Gallardo 1968:116 China Wu & Wu 1987:79 Heteromastus filiformis (Claparede, 1864) China Yang & Sun 1988:254 Heteromastus similis Southern, 1921 S. Vietnam Gallardo 1968: 117 Mastobranchus loii Gallardo, 1968 S. Vietnam Gallardo 1968:118 Mediomastus sp. Taiwan Hsieh, pers. eomm.

225

Paxton & Chou: Polychaetous annelids from the South China Sea

NAME AUTHOR COUNTRY REFERENCE

Branchiomma cf. boholense (Grube, 1878) Taiwan Hsieh, pers. comm. Branchiomma cingulata (Grube, 1870) Philippines Pillai 1965:163 Hong Kong Mak 1982:600 China Yang & Sun 1988:299 Branchiomma cf. conspersa Ehlers, 1887) Taiwan Hsieh, pers. comm. Chone ?infundibuliformis Krayer, 1856 S. Vietnam Gallardo 1968:130 Chone trilobata Gallardo, 1968 S. Vietnam Gallardo 1968:130 Hypsicomus phaeotaenia (Schmarda, 1861) China Wu et al. 1980:124 Singapore Tan & Chou 1993:290 lasmineira caducibranchiata Willey, 1905 S. Vietnam Gallardo 1968:131 Laonome albicingillum Hsieh, 1995 Taiwan Hsieh 1995:131 Megalomma intermedium (Beddard, 1889) Philippines Pillai 1965: 164 Megalomma vesiculosa (Montagu, 1815) Hong Kong Mak 1982:600 China Yang & Sun 1988:301 Megalomma sp. Taiwan Hsieh, pers. comm. Myxicola infundibulum (Renier, 1804) Hong Kong Mak 1982:600 Notaulax cL pyrroghaster Taiwan Hsieh, pers. comm. Potamilla leptochaeta Southern, 1921 Singapore Tan & Chou 1993:290 Potamilla ocalea Pillai, 1965 Philippines Pillai 1965: 164 Potamilla torelli Ma1mgren, 1866 Hong Kong Shin 1982: 170 Pseudobranchiomma sp. Taiwan Hsieh, pers. comm. Pseudopotamilla reniformis (Muller, 1771) Hong Kong Mak 1982:600 Singapore Tan & Chou 1993;600 Pseudopotamilla sp. Taiwan Hsieh, pers. comm. Sabella penicillus Linneus, 1767 China Wu et al. 1980:124 Sabella sp. Hong Kong Mak 1982:600 Sabellastarte indica (Savigny, 1818) S. Vietnam Gallardo 1968:131 Singapore Tan & Chou 1993:290 China Yang & Sun 1988:305 Taiwan Hsieh, pers. comm. Sabellastarte cf. samoensis (Grube, 1870) Taiwan Hsieh, pers. comm. Sabellastarte sanctijosephi (Gravier, 1906) China Wu et a1 1980:124

Serpulidae Apomatus enosimae Marenzeller, 1884 China Yang & Sun 1988:308 Circeis spirillum (Linnaeus, 1758) China Yang & Sun 1988:318 Dexiospira foraminosus Busch, 1904 China Yang & Sun 1988:319 Dexiospira sp. Taiwan Cheng & Chen 1994: Hydroides albiceps Grube, 1869 Hong Kong Mak 1982:601 China Yang & Sun 1988:311 Hydroides centrospina Wu & Chen, 1981 China Wu & Chen 1981:354 Hydroides dirampha March, 1863 Hong Kong Mak 1982:602 Hydroides elegans (Haswell, 1883) Hong Kong Mak 1982:602 China Yang & Sun 1988:311 Hydroides cf. elegans Taiwan Hsieh, pers. comm. Hydroides exaltata (Marenzeller, 1884) China Wu et al. 1980: 124 Hong Kong Mak 1982:603 Hydroides fusca Imajima, 1976 China Wu et al. 1980:124

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NAME AUTHOR COUNTRY REFERENCE

Hydroides cf. jusicola March, 1863 Hong Kong Mak 1982:603 Hydroides grubei Pillai, 1965 Philippines Pillai 1965:166 Hydroides inornata Pillai, 1960 Hong Kong Mak 1982:605 Hydroides longistylaris Chen & Wu, 1980 China Chen & Wu 1980:249 Hydroides nanhaiensis Wu & Chen, 1981 China Wu & Chen 1981:355 Hydroides norvegica Gunnereus, 1768 Philippines Pillai 1965: 168 Hydroides rhombobulus Chen & Wu,1980 China Chen & Wu 1980:247 Hong Kong Mak 1982:605 Hydroides trilobulus Wu & Chen, China Wu et al. 1980:125 Metavermilia acanthophora (Augener, 1914) China Wu et al. 1980:125 Hong Kong Mak 1982:606 Metavermilia annobonensis Zibrowius, 1973 China Wu et al. 1980:125 Metavermilia cf. inflata 1majima, 1977 Hong Kong Mak 1982:607 Metavermilia cf. taenia Zibrowius, 1973 China Wu et al. 1980:125 Neodexiospira cf. joraminosa Taiwan Hsieh, pers. comm. Neodexiospira cf. preacuta (Bush, 1904) Taiwan Hsieh, pers. comm. Neodexiospira cf. knightjonesi Taiwan Hsieh, pers. comm. Neodexiospira cf. steueri (Sterzinger, 1909) Taiwan Hsieh, pers. comm. Neopomatus uschakovi Pillai, 1965 Philippines Pillai 1965:170 lingayanensis Pileolaria cf. militaris C1aparecte, 1870 Taiwan Hsieh, pers. comm. Pileolaria cf. pettibonei Taiwan Hsieh, pers. comm. Pileospira koehleri (Caullery & Mesni1, 1897) China Wu et al. 1980: 125 Pomatoleios kraussii (Baird, 1865) Hong Kong Mak 1982:607 Thailand Becker 1993 Pomatoleios kraussii manilensis Pillai, 1965 Philippines Pillai 1965:169 Pomatostegus stellatus (Abi1dgaard, 1789) Hong Kong Mak 1982:608 Protohydroides elegans (Haswell, 1883) Hong Kong Shin 1982: 170 Protula tubularia tubularia (Montagu, 1803) China Wu et al. 1980:125 Protula sp. Taiwan Hsieh, pers. comm. Salmacina cf. dysteri (Hyx1ey, 1955) Taiwan Hsieh, pers. comm. (Paraserpula) sinica Wu & Chen in Wu et al.,1979 China Wu et al. 1979:92 Linnaeus, 1767 Hong Kong Mak 1982:608 China Yang & Sun 1988:312 Serpula watsoni Willey, 1905 Hong Kong Mak 1982:609 Simplicaria pseudomilitaris (Thiriot-Quievreux, 1965) Taiwan Hsieh, pers. comm. Spirobranchus giganteus (Pallas, 1766) China Wu et al. 1980: 125 Spirobranch. giganteus corniculatus ten Hove Taiwan Dai & Yang 1995: Spirobranchus jousseaumei (Gravier, 1908) China Yang & Sun 1988:314 Spirobranchus latiscapus (Marenzeller, 1884) China Wu et al. 1980:125 Spirobranchus maldivensis Pixell, 1913 China Yang & Sun 1988:315 Spirobranchus semperi March, 1863 Hong Kong Mak 1982:609 China Yang & Sun 1988:315 Spirobranchus tricornis (March, 1863) Hong Kong Mak 1982:609 China Yang & Sun 1988:317 Spirorbis (Dexiospira) treadwelli Pillai, 1965 Philippines Pillai 1965: 172 Vermiliopsis glandigerous Gravier, 1908 China Wu et al. 1980:125

229 Paxton & Chou: Polychaetous annelids from the South China Sea

NAME AUTHOR COUNTRY REFERENCE

Vermiliopsis infundibulum (Linneus, 1788) China Wu et al. 1980:125 Vermiliopsis infundibul. glandigera Imajima, 1967 Hong Kong Mak 1982:610 Vermiliopsis pygidialis (Willey, 1905) China Wu et al. 1980:125

Nerillidae Leptonerilla diplocirrata Westheide & Purschke 1996 China Westheide & Purschke 1996:587

LITERATURE CITED

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