Coleoptera, Attelabidae)

Алтайский зоологический журнал/вып. 2/2008

Волынкин А.В., Перунов Ю.Е. К фауне совок (Lepidoptera: Noctuidae s. l.) заповедника “Тигирекский” (Северо-Западный Алтай, Россия) // Эверсманния. 2007. Вып. 10. С. 42–49. Fibiger M., Lafontaine D. A review of the higher classification of the Noctuoidea (Lepidoptera) with special reference to the Holarctic fauna // Esperiana. 2004. Bd. 11. S. 7–82.

Адреса для контактов: Волынкин Антон Валерьевич (ГПЗ “Тигирекский”, г. Барнаул, Россия) E-mail: [email protected]

УДК 595.768.1

A.A. Legalov To the knowledge of the leaf-rolling weevils of subfamily Attelabinae (Coleoptera, Attelabidae)

А.А. Легалов. К познанию жуков-трубковертов подсемейства Attelabinae (Coleoptera, Attelabidae)

Key words: Coleoptera, Attelabidae, Attelabinae, sp. n. (Бразилия), Omolabus (Pseudomolabus) bicolor Euopsini, Euscelini, Hybolabini, new subgenus, new spe- Legalov, sp. n. (Бразилия) и Omolabus (Neomolabus) cies, new synonym, fauna, New Guinea, Neotropic Area. spinicollis Legalov, sp. n. (Французская Гвиана и Гайа- Ключевые слова: Coleoptera, Attelabidae, Atte- на). Подтриба Alleuselina Legalov, 2003, syn. n. све- labinae, Euopsini, Euscelini, Hybolabini, новый подрод, дена в синонимы к подтрибе Euscelina Voss, 1925. новые виды, новый синоним, фауна, Неотропическая Alleuscelus violaceipennis Voss, 1937 впервые указан область. для фауны Бразилии. Приводятся материалы для ред- ких видов. Abstract. In the paper new subgenus Piloseuscelus Legalov, subgen. n. (type species: Euscelus landanguinus Introduction Legalov, sp. n.) of the genus Euscelus Schoenherr, 1833 The leaf-rolling weevils are widespread and 7 new species: Euopsidius (Pascoeuops) lorentzien- in the world fauna. The family includes two sis Legalov, sp. n. (Indonesia: New Guinea), Euscelus subfamilies (Attelabinae and Apoderinae). (Meteuscelus) pseudbinotatus Legalov, sp. n. (Brazil), Euscelus (Piloseuscelus) landanguinus Legalov, sp. Representatives of the subfamily Attelabinae n. (Ecuador), Omolabus (s. str.) bifoveatoides Legalov, roll tubes with double cut [Legalov, 2004a]. sp. n. (French Guiana), Omolabus (Pseudomolabus) Growing interest in studies of systematic lagoasensis Legalov, sp. n. (Brazil), Omolabus of this subfamily [Hamilton, 1994, 2001, 2005, (Pseudomolabus) bicolor Legalov, sp. n. (Brazil) and 2007; Legalov, 2003, 2004b, 2007; Riedel, Omolabus (Neomolabus) spinicollis Legalov, sp. n. (French Guiana, Guyana) are described. New synonym: 2002] in the world fauna can be seen in recent Alleuselina Legalov, 2003, syn. n. for Euscelina Voss, years. This work continues studying of this 1925 is presented. Alleuscelus violaceipennis Voss, 1937 subfamily of the world fauna by the author. is for the first time revealed in fauna of Brazil. Materials for rare species are resulted. Material and methods Резюме. В статье описаны новый подрод Piloseuscelus Legalov, subgen. n. (типовой вид: Types and specimens are kept in the following Euscelus (Pascoeuops) landanguinus Legalov, sp. n.) collections and museums: ISNB – Institut Royal рода Euscelus Schoenherr, 1833 и 7 новых видов: des Sciences Naturelles de Belgique (Belgium: Euopsidius lorentziensis Legalov, sp. n. (Индонезия: Brussels); MCSN – Museo Civico di Storia Новая Гвинея), Euscelus (Meteuscelus) pseudbinotatus Naturale “Giacomo Doria” (Italy: Genova); Legalov, sp. n. (Бразилия), Euscelus (Piloseuscelus) landanguinus Legalov, sp. n. (Эквадор), Omolabus (s. MZLU – Lund University (Sweden: Lund); str.) bifoveatoides Legalov, sp. n. (Французская Гвиа- SMTD – Staatliches Museum für Tierkunde на), Omolabus (Pseudomolabus) lagoasensis Legalov, (Germany: Dresden); ZIN – Zoological Institute

69 A.A. Legalov / ЭНТОМОЛОГИЯ of Russian Academy of Sciences (Russia: St. weakly convex. Striae clear. Points in them Petersburg); ZMAN – Zoologisch Museum, thick and large. Instituut voor Taxonomische Zoologie, Prothorax smooth. Precoxal part wide. Universiteit van Amsterdam (Netherlands: Postcoxal part narrow. Mesothorax and Amsterdam); ZMN – Zoological Museum, mesepisternum matte. Metathorax and Institute of Animal Systematics and Ecology metepisternum sparsely and small punctate. (Russia: Novosibirsk); ZMUC – Zoological Abdomen convex, weakly flattened on the Museum, University of Copenhagen (Denmark: middle, weakly rugosity-punctate. 1st – 4th Copenhagen). ventrites wide, almost identical width. 5th ventrite narrow, considerably narrower than Results 4th ventrite. Pygidium weakly convex, densely Family Attelabidae Billberg, 1820 punctate. Subfamily Attelabinae Billberg, 1820 Legs long. Forelegs strongly elongated. Supertribe Attelabitae Billberg, 1820 Profemora thick with small protuberances on Tribe Euopsini Voss, 1925 inside. Meso- and metafemora weakly widened. Subtribe Parasynaptopsisina Legalov, Mesofemora with weak tooth ventrally in first 2007 third. Protibiae long, curved, small crenate on Genus Euopsidius Legalov, 2003 internal edge, with long mucro. Meso- and Subgenus Pascoeuops Legalov, 2007 metatibiae shorter, biconcave. Tarsi long. 1st segment long triangular. 2nd segment flattened- Euopsidius (Pascoeuops) lorentziensis triangular. 3rd segment bilobed. Clausal segment Legalov, sp. n. (Tab: II: a, Fig. 1f) elongated. Claws long. Material: Holotype – male (ZMAN), Length of body: 4.3 mm. “Z. Nieuw Guinea, Lorentz, 1909-10, Bivak Diagnosis: This new species is close Eiland”, “143”, “gen. ? sp. ? Attelabid varium, to E. divisus (Pascoe, 1874) but differs by the Det. K.M. Heller, 1913”. weak teeth on mesofemora, weaker clavate Description: Male: Body yellow, naked. profemora, large dark stain on elytra, light Elytra brown except the basic third. Head and clava of antennae, form of basal sclerite of the antennae yellowy-brown. endophallus. Head elongated. Rostrum short, widened Distribution: Indonesia (New Guinea). to apex, smooth. Antennae attached before the Etymology: The name is formed from the rostrum basis. Eyes large, convex. Forehead very location river “Lorentz” – “lorentziensis”. narrow. Temples weakly extends to pronotum, long. Vertex with transversal striae, smooth. Tribe Euscelini Voss, 1925 Antennae long, reaching humeri. Scapus Subtribe Clinolabina Legalov, 2003 and 1st segment oval. Scapus hardly longer than Genus Iseuscelus Voss, 1925 1st segment. 2nd–4th segments long trapezoid, almost equal length. 5th–7th segments shorter, Iseuscelus flohri Voss, 1925 (Tab.: II: fig. i) trapezoid. 7th segment shorter than 6th segment. Iseuscelus flohri Voss, 1925: 71 Clava elongated, compact. 1st segment Material: Female (MCSN), “Durango, trapezoid. 2nd segment rectangular, equal length Mexico”. to 1st segment. 3rd segment elongated, pointed, Distribution: Mexico. equal length to the previous segments. Pronotum almost campaniform, 1.1 times Subtribe Euscelina Voss, 1925 wider than length. Grooves weak. Sides weakly Attelabini subtribe Euscelina Voss, 1925: 28 rounded. Disk convex, almost smooth. Greatest type genus: Euscelus Schoenherr, 1833 width in first third and at the basis. Scutellum Alleuselina Legalov, 2003: 411, syn. n. trapezoid. type genus: Alleuscelus Voss, 1937 Elytra almost rectangular, 1.14 times longer Remarks: The subtribe has been described than width, weakly narrowed to apex. Greatest based on characters: metallic elytra and rows of width in humeri. Humeri convex. Intervals erect setae on abdomen of males. By the author 70 Алтайский зоологический журнал/вып. 2/2008 is found these characters in other genera (rows of lenhal, 1839) setae at male of Emphyleuscelus (Eleuscelinus) Attelabus binotatus Gyllenhal, 1839: 316 ruber Legalov, 2007 and metallic elytra at Material: Female (ZMN), Costa Rica Ala., Parelleuscelus boliviensis Legalov, 2007). 20 km S of Upala, 21.II.1991, F.D. Parker; female Because of this, the author is placed subtribe (ISNB), El Salvador, Apaneca (Ahuachapan), Alleuselina Legalov, 2003, syn.n. in synonyms 14–15.VII.1959, J. Bechyne; female (SMTD), to subtribe Euscelina Voss, 1925. “Mexico”, “Collection Wagner”; male (SMTD), “Mexico”. Genus Alleuscelus Voss, 1937 Distribution: Belize, Costa Rica, Guatemala, Honduras, El Salvador, Mexico, Alleuscelus violaceipennis Voss, 1937 Nicaragua, Panama. Alleuscelus violaceipennis Voss, 1937: 159 Euscelus (Meteuscelus) pseudbinotatus Material: Male (SMTD), “Jatahy, Brasilia”, Legalov, sp.n. (Tab. II: b Figs. 1a–1b) “1906, 23”; male (SMTD), “Amazonas, Jatahi”, Material: Holotype – male (MCSN), “Gerh. W. Müller, Vermächt. 1909”; male “Amazzono, Coll. Jekel”. (ZMUC), [Brazil] “Lagoa Santa, Reichardt”; Description: Male: female (MCSN), [Brazil] “Jatahy, Donckler”, Body red-brown. Coxa, basis of meso- and “typus, female”; female (MCSN), [Brazil] metafemora, scapus and funicle of antennae, “Jatahy, Goyaz”, “typus, female”. convex stains on elytra yellowy-brown. Distribution: Peru, Brazil. Head elongated, almost conic. Rostrum Remarks: This species is for the first time short, widened to apex, small punctate. Antennae revealed in fauna of Brazil. attached in first third of the rostrum. Eyes weakly convex. Forehead flat, smooth, narrower than Genus Euscelus Schoenherr, 1833 eye. Temples long, weakly transversal-wrinkled. Subgenus Meteuscelus Voss, 1925 Vertex convex, small punctate. Antennae long, reaching pronotum first line. Euscelus (Meteuscelus) binotatus (Gyl- Scapus and funicle segments almost trapezoid.

Fig. 1. Aedeagus: a – Euscelus pseudbinotatus Legalov, sp. n., b – E. pseudbinotatus Legalov, sp. n. (laterally), c – Omolabus bifoveatoides Legalov, sp. n., d – O. spinicollis Legalov, sp. n., e – O. lagoasensis Legalov, sp. n., f - Euopsidius lorentziensis Legalov, sp. n. Рис. 1. Эдеагус: a – Euscelus pseudbinotatus Legalov, sp. n., b – E. pseudbinotatus Legalov, sp. n. (вид сбоку), c – Omolabus bifoveatoides Legalov, sp. n., d – O. spinicollis Legalov, sp. n., e – O. lagoasensis Legalov, sp. n., f - Euopsidius lorentziensis Legalov, sp. n. 71 A.A. Legalov / ЭНТОМОЛОГИЯ

1st segment shorter than scapus. 2nd segment Euscelus (Meteuscelus) elliptiguttatus longer than 1st segment. 5th segment shorter Voss, 1925 than 4th segment. 6th segment much shorter Euscelus elliptiguttatus Voss, 1925: 35 than 5th segment. 7th segment hardly wider Material: Female (SMTD), “Costa Rica, than 6th segment. Clava elongated, shorter than A.H. Fassl '13N 30”, “Onosi, 1500 m”. funicle. 1st segment elongated. 2nd segment Distribution: Colombia, Costa Rica, much shorter than 1st segment. 3rd segment Peru. pointed, longer than 1st segment. Pronotum wide, 1.13 times wider than Euscelus (Meteuscelus) fenestratus length. Greatest width in first third. Disk convex, (Sharp, 1889) smooth, with two transversal striae. Pronotal Attelabus fenestratus Sharp, 1889: 14 groove sharp. Sides almost direct. Scutellum Material: Female (SMTD), “Collection triangular. Schild-Burgdorf, Costa Rica, Turrialaba”. Elytra almost rectangular, extend to apex, Distribution: Costa Rica, Mexico, 1.3 times wider than length. Greatest width Panama. behind the middle. Humeri weakly smoothed. Intervals wide, convex, smooth. 2nd interval Subgenus Meteusceliodes Legalov, 2007 in first third with convex round stain. Points in striae large, sometimes merging. Euscelus (Meteusceliodes) mundanoides Precoxal and postcoxal parts of the prothorax Legalov, 2007 equal length. Meso- and metathorax with Euscelus mundanoides Legalov, 2007: 256 episternum sparsely punctate. Metepisternum Material: Female (MCSN), “Colombia, wide. Abdomen convex, small punctate- Coll. Jekel”, “Coscineuscelus hispoides in lit. wrinkled. 1st – 3rd ventrites wide. 1st ventrite Chevr.” without blades. 4th and 5th ventrites narrower. Distribution: Brazil, Columbia. Pygidium convex, densely punctate. Legs long. Forelegs strongly elongated. Subgenus Piloseuscelus Legalov, Profemora clavate, with small and long teeth. subgen.n. Meso- and metafemora shorter, weakly widened, Type species: Euscelus landanguinus without teeth. Protibiae long, narrow, curved, Legalov, sp.n. with long mucro at apex. Meso- and metatibiae Description: Body red-brown, with short, weakly biconcave, wider. Tarsi long. semierect long setae. Head elongated. Rostrum Protarsi stronger elongated. short. Antennae attached before the rostrum Length of body: 5.0 mm. basis. Eyes very weakly convex. Forehead Diagnosis: This new species is very close weakly pressed, dense rugosity-punctate. to E. binotatus (Gyllenhal, 1839) but differs by Temples long. Vertex convex, rugosity-punctate. the apex of elytra stronger widened, aedeagus Antennae long, reaching pronotum first line. apex, and form of the basal sclerite. Pronotum wide. Greatest width in first third and Distribution: Brazil. on the basis. Disk convex, densely transversal- Etymology: The name is formed by wrinkled. Elytra almost rectangular, wide. addition of the prefix “pseudo-” to “binotatus”. Greatest width on the middle. Intervals wide, convex, densely rugosity-punctate. 8th interval Euscelus (Meteuscelus) cruralis (Sharp, carinate in first half. Thorax rugosity-punctate. 1889) Abdomen convex, small punctate-wrinkled. Attelabus cruralis Sharp, 1889: 14 Legs long. Forelegs elongated. Profemora Material: Male (SMTD), female (SMTD), clavate, with long tooth of females. Meso- and “Jalapa, Mexico”. metafemora shorter, weakly widened. Meso- Distribution: Guatemala, Mexico, femora with tooth of females. Tibiae wide, with Panama. mucro and uncus at apex of females. Tarsi long. Length of body: 4.2 mm. Diagnosis: This new subgenus is similar to 72 Алтайский зоологический журнал/вып. 2/2008 subgenus Eusceloides Legalov, 2003 but differs tooth. Protibiae elongated, wide, weakly by the body with semierect setae, wide elytra, curved, with mucro and uncus at apex. Meso- mesofemora with tooth, humeri without tooth, and metatibiae short, weakly biconcave. Tarsi 8th interval of the elytra carinate in first half. long. Etymology: The name is formed addition Length of body: 4.2 mm. “hairy” – “pilosus” to “euscelus”. Distribution: Ecuador. Etymology: The name is formed from the Euscelus (Piloseuscelus) landanguinus location “Landangui” – “landanguinus”. Legalov, sp.n. (Tab. II: c) Material: Holotype – female (SMTD), Genus Neoeuscelus Voss, 1925 [Ecuador] “Landangui, Dr. Ohaus”, “1, 1907”. Subgenus Neoeuscelus s. str. Description: Female: Body red-brown, with semierect long setae. Neoeuscelus (s. str.) longimanus (Olivier, Antennae, legs and elytra more light. 1789) Head elongated, almost conic. Rostrum Attelabus longimanus Olivier, 1789: 278 short, widened to apex, dense punctate. Antennae Material: Female (ISNB), [French Guiana] located before the rostrum basis. Eyes very “Cayenne, F. Pillaut”; male (ISNB), “Bahia”, weakly convex. Forehead densely rugosity- “Coll. Castelnau, Coll. Roelofs”; male (SMTD), punctate, weakly pressed. Temples long, weakly “S Bras., Staudg.”; transversal-wrinkled. Vertex convex, rugosity Distribution: Brazil, French Guiana. punctate. Antennae long, reaching pronotum first Subgenus Neoeusceloides Legalov, 2007 line. Scapus and 1st segment of the funicle almost tear-shaped, wider than 2nd segment. Neoeuscelus (Neoeusceloides) atratus 1st segment shorter than scapus. 2nd – 7th (Voss, 1925) segments almost trapezoid, short. 2nd segment Euscelus atratus Voss, 1925: 40 shorter than 1st segment. 6th and 7th segments Material: Male (SMTD), [French Guiana] transversal. Clava elongated, little shorter than “Cayenne, Märxel”; male (SMTD), [Brazil] funicle. 1st segment elongated. 2nd segment “Bahia, Dohrn”. little shorter than 1st segment. 3rd segment Distribution: Brazil, French Guiana. weakly pointed, equal to 2nd segment. Pronotum wide, 1.41 times wider than Genus Pheleuscelus Jekel, 1860 length. Greatest width in first third and on Subgenus Pheleuscelus s. str. the basis. Disk convex, densely transversal- wrinkled. Pronotal groove weak. Sides almost Pheleuscelus (s. str.) innotatus (Voss, direct. Scutellum triangular. 1925) Elytra almost rectangular, wide, 1.14 times Euscelus innotatus Voss, 1925: 37 wider than length. Greatest width on the middle. Material: 2 males (SMTD), [Brazil] Humeri weakly smoothed. Intervals wide, “Corumba, Matt Grosso”. convex, densely rugosity-punctate. 8th interval Distribution: Argentina, Brazil. carinate in first half. Points in striae large. Precoxal and postcoxal parts of the Subgenus Pheleusceloides Legalov, 2007 prothorax equal length. Thorax rugosity- punctate. Pheleuscelus (Pheleusceloides) subimp- Abdomen convex, small punctate- ressus (Voss, 1925) wrinkled. 1st – 3rd ventrites wide. 1st ventrite Euscelus subimpressus Voss, 1925: 37 without blades. 4th ventrite narrow. 5th ventrite Material: Male (ZIN), [Brazil] “Fontebou, narrower. Pygidium convex, densely punctate. fl. Amazon”, “17328”. Legs long. Forelegs elongated. Profemora Distribution: Brazil. clavate, with long tooth. Meso- and metafemora shorter, weakly widened. Mesofemora with 73 A.A. Legalov / ЭНТОМОЛОГИЯ

Genus Chryseuscelus Voss, 1925 Material: Male (MCSN), “Guatemala, L. Conradt”. Chryseuscelus sexmaculatus (Chevrolat, Distribution: Bolivia, Columbien, Costa 1876) Rica, El Salvador, Guatemala, Honduras, Attelabus sexmaculatus Chevrolat, 1876: Mexico, Panama, Peru. 228 Material: Female (ZMUC), “Porto Rico”, Omolabus (s. str.) bifoveatoides Legalov, “Mus. Westerm”. sp.n. (Tab. II: d, Fig. 1: c) Distribution: Cuba, P. Rico, Virgin Is. Material: Holotype – male (ISNB), “Guyane Francaise, Cayenne, Coll. Roelofs”, Genus Emphyleuscelus Voss, 1925 “Attelabus placidus mihi, h. Cayennae, D. Subgenus Eleuscelinus Legalov, 2007 Lacordaire”. Description: Male: Body dark, red-brown, Emphyleuscelus (Eleuscelinus) ruber lustrous, naked. Rostrum, antennae, elytra, Legalov, 2007 abdomen and legs more light. Emphyleuscelus ruber Legalov, 2007: 262 Head elongated. Rostrum weakly Material: Male (ZMUC), [Brazil] “Mus. elongated, widened to apex, small punctate. Hauschild, 12-9-1914”, “Melanco “ryphce” Antennae attached in first third of the rostrum. aff., Lagoa Santa”. Eyes large, convex. Forehead narrow, flat, with Distribution: Brazil. 2 longitudinal striae. Temples weakly elongated, smooth. Vertex with elongated 2 cambers on the Subgenus Eleuscelus Voss, 1925 basis. Prementum with 2 teeth. Antennae long, reaching pronotum middle. Emphyleuscelus (Eleuscelus) vicinus Scapus and 1st segment oval. Scapus longer (Voss, 1925) than 1st segment. 2nd – 4th segments elongated, Euscelus vicinus Voss, 1925: 39 narrower than 1st segment. 5th segment almost Material: Female (ISNB), [French Guiana] trapezoid. 6th and 7th segments short trapezoid. “Cayenne”, “Coll. Dejean, Coll. Roelofs”, Clava elongated, compact, little shorter than “Attelabus relucens Lacordaire, h. Cayennae, funicle. 1st segment longer than 2nd segment. D. Lacordaire”. 3rd segment pointed, equal to 2nd segment. Distribution: Brazil, French Guiana. Pronotum almost campaniform, 1.39 times wider than length. Grooves weak. Sides almost Tribe Hybolabini Voss, 1925 direct. Disk convex, almost smooth. Scutellum Subtribe Omolabina Legalov, 2003 almost rectangular, large, wide, smooth. Genus Omolabus Jekel, 1860 Elytra square. Greatest width in humeri. Subgenus Omolabus s. str. Humeri prominent convex, with small tooth. Intervals almost flat, wide, very weakly Omolabus (s. str.) bifoveatus (Jekel, transversal-wrinkled. Striae clear. Points in 1860) them rare and not deep. Attelabus bifoveatus Jekel, 1860: 203 Prothorax wrinkled, with teeth on first Material: Male (ZMUC), “Brasil, line, wrinkled. Precoxal part of the prothorax Reinhardt”, “Mus. Westerm.”; male (ZMUC), strongly elongated. Mesothorax largely “Brasils”, “Mus. Westerm.”; female (ZMUC), punctate. Episternum and metathorax small [Brazil] “Lagoa Santa, Reinhardt”; 2 males punctate. Abdomen convex, small rugosity- (SMTD), [Brazil] “Amazonas, Jatahi”; male punctate. 1st – 3rd ventrites wide. 4th ventrites (SMTD), “Brasilein”. narrow. 5th ventrite narrower. Pygidium weakly Distribution: Brazil. convex, densely punctate. Legs long. Forelegs elongated. Profemora Omolabus (s. str.) ligulatus (Sharp, widened. Meso- and metafemora weaker 1889) widened. Protibiae long, weakly curved, crenate Attelabus ligulatus Sharp, 1889: 10 on internal edge, with mucro at apex. Meso- and 74 Алтайский зоологический журнал/вып. 2/2008 metatibiae shorter, weakly biconcave, weakly widened to apex, very small punctate. Antennae widened to apex. Tarsi long. Length of body: located in first third of the rostrum. Eyes large, 4.6 mm. not protruding from contour of head. Forehead Diagnosis: This new species is close to narrow, weakly pressed, with 2 longitudinal striae. O. bifoveatus (Jekel, 1860) but differs by the Temples weakly elongated, weakly transversal- larger sizes, narrower body, and strong teeth on wrinkled. Vertex convex, weakly transversal- prothorax, one-colour elytra, and armament of wrinkled. Prementum without long teeth. the endophallus. Antennae short, reaching pronotum first line. Distribution: French Guiana. Scapus and 1st segment oval. Scapus longer than Etymology: The name is formed by 1st segment. 2nd segment trapezoid, narrower and addition of the ending “-oides” to “bifoveatus”. shorter than 1st segment. 3rd and 4th segments elongated trapezoid. 3rd segment much longer Subgenus Pseudomolabus Legalov, 2004 than 2nd segment. 4th segment hardly shorter than 3rd segment. 5th – 7th segments short Omolabus (Pseudomolabus) angulipennis trapezoid. Clava elongated, compact, shorter than (Sharp, 1889) funicle. 1st segment longer than 2nd segment. Attelabus angulipennis Sharp, 1889: 8 3rd segment pointed, longer than 2nd segment Material: Female (MZLU), Costa Rica, and shorter than 1st segment. Heredia, Porrosati, 2000 m, 2.III.1997, C. Pronotum almost campaniform, 1.65 times Hansason. wider than length. Grooves weak. Sides almost Distribution: Costa Rica, El Salvador, direct. Disk convex, small smooth. Scutellum Guatemala, Mexico, Panama. almost rectangular. Elytra almost square, 1.09 times wider Omolabus (Pseudomolabus) corumbaensis than length. Greatest width in humeri and Voss, 1929 behind the middle. Humeri convex, with small Omolabus corumbaensis Voss, 1929: 213 tooth. Intervals almost flat, wide, very weakly Material: Female (ZIN), [Brazil] transversal-wrinkled. Striae clear. Points in them “Fontebou, fl. Amazon”, “17328”. not deep. Distribution: Brazil. Prothorax largely punctate. Precoxal part of the prothorax strongly elongated, wrinkled. Omolabus (Pseudomolabus) rugicollis Mesothorax densely punctate. Episternum (Jekel, 1860) and metathorax densely and largely punctate. Attelabus rugicollis Jekel, 1860: 192 Metepisternum partially smooth. Abdomen Material: Female (ISNB), [Brazil] “Minas convex, on middle weakly flattened, small Geraes”, “Coll. Castelnau, Coll. Roelofs”. rugosity-punctate. 1st – 3rd ventrites wide. 4th Distribution: Brazil, French Guiana. ventrites narrow. 5th ventrite narrower. Pygidium weakly convex, densely punctate. Omolabus (Pseudomolabus) subrugosus Legs long. Forelegs elongated. Profemora Voss, 1925 widened. Meso- and metafemora weaker Omolabus subrugosus Voss, 1925: 281 widened. Protibiae long, weakly curved, small Material: Male (SMTD), [Brazil] crenate on internal edge, with mucro at apex. “Amazonas”. Meso- and metatibiae shorter, weakly biconcave, Distribution: Brazil. weakly widened to apex. Tarsi long. Length of body: 4.4 mm. Omolabus (Pseudomolabus) lagoasensis Diagnosis: This new species is Legalov, sp.n. (Tab. II: g, Fig. 1e) O. angulipennis (Sharp, 1889) but differs by Material: Holotype – male (ZMUC), the small punctate pronotum, very weak convex [Brazil] “Sete Lagoas, Reinhardt”. intervals of the elytra and armament of the Description: Male: Body red-brown, endophallus. lustrous, and naked. Distribution: Brazil. Head elongated. Rostrum weakly elongated, Etymology: The name is formed from the 75 A.A. Legalov / ЭНТОМОЛОГИЯ location “Lagoas” – “lagoasensis”. humeri with teeth, wider elytra, pronotum more strongly narrowed to apex and light antennae. Omolabus (Pseudomolabus) bicolor Distribution: Brazil. Legalov, sp.n. (Tab. II: h) Etymology: The name is formed from the Material: Holotype – female (ZMUC), word for “bicoloured” – “bicolor”. [Brazil] “Lagoa Santa, Reinhardt”. Subgenus Phyletobius Voss, 1925 Description: Male: Body dark red-brown, lustrous, naked. Antennae, head, pronotum, Omolabus (Phyletobius) equestris (Voss, scutellum and tarsi red. 1925) Head elongated. Rostrum weakly elongated, Phyletobius equestris Voss, 1925: 265 widened to apex, small punctate. Antennae Material: Male (ISNB), San Salvador, La attached before the rostrum basis. Eyes not Libertad, 15.VI.1960, J. Bechyne. protruding from contour of head. Forehead Distribution: Costa Rica, El Salvador. narrow, flat, smooth. Temples weakly elongated, weakly transversal-wrinkled. Vertex convex, Subgenus Sharpilabus Legalov, 2007 smooth. Prementum without long teeth. Antennae short, reaching pronotum middle. Omolabus (Sharpilabus) quadratus Scapus and 1st segment oval, equal length. (Sharp, 1889) 2nd and 4th segments elongated trapezoid. 2nd Attelabus quadratus Sharp, 1889: 11 segment narrower and shorter than 1st segment. Material: Male (MCSN), “Panama, 3rd segment longer than 2nd segment. 4th segment Coll. Jekel”; male (SMTD), female (SMTD), equal to 3rd segment. 5th – 7th segments short “Collection Schild-Burgdorf, Costa Rica, trapezoid. Clava elongated, compact, shorter than Turrialaba”, “16402”, “Attelabus elatus n. sp., funicle. 1st segment weakly elongated, longer Det. E. Voss”. than 2nd segment. 3rd segment pointed, hardly Distribution: Costa Rica, Nicaragua, longer than 1st segment. Panama. Pronotum almost campaniform, 1.33 times wider than length. Grooves weak. Sides almost Subgenus Sternolabus Jekel, 1860 direct. Disk convex, largely punctate, with two weak deepenings. Scutellum almost rectangular, Omolabus (Sternolabus) longirostris small punctate. (Jekel, 1860) Elytra almost square, 1.14 times wider Attelabus longirostris Jekel, 1860: 207 than length. Greatest width in humeri and Material: Male (ISNB), “Colombia”, behind the middle. Humeri convex, with small “Coll. Roelofs”. tooth. Intervals almost flat, wide, very weakly Distribution: Columbia. transversal-wrinkled. Striae clear. Points in them not deep. Subgenus Mesitolabus Voss, 1936 Thorax sparsely punctate. Precoxal part of the prothorax weakly elongated. Abdomen Omolabus (Mesitolabus) aeneicollis Voss, convex, rugosity-punctate. 1st – 3rd ventrites 1925 wide. 4th ventrites narrow. 5th ventrite narrower. Omolabus aeneicollis Voss, 1925: 280 Pygidium weakly convex, densely punctate. Material: Male (ISNB), “Colombie”, Legs long. Forelegs weakly elongated. “Coll. Roelofs”. Profemora widened. Meso- and metafemora Distribution: Columbia. weaker widened. Protibiae shorter than profemora, weakly curved, crenate on internal Subgenus Thyreolabus Jekel, 1860 edge, with mucro and uncus at apex. Meso- and metatibiae shorter. Tarsi long. Length of body: Omolabus (Thyreolabus) corniculatus 3.1 mm. (Gyllenhal, 1839) Diagnosis: This new species is similar Attelabus corniculatus Gyllenhal, 1839: to O. mulicus Legalov, 2007 but differs by the 306 76 Алтайский зоологический журнал/вып. 2/2008

Material: Male (SMTD), “Dr. Ohaus, than 1st segment. 3rd segment elongated Brasilien, 13348”; male (SMTD), [Brazil] trapezoid, 2 times longer than 2nd segment. “Santos, J. Melz”; female (SMTD), “Brasilen, 4th segment hardly longer than 3rd segment. Heyne”. 5th – 7th segments short trapezoid. 7th segment Distribution: Brazil. wider. Clava elongated, compact, shorter than Subgenus Neomolabus Legalov, 2004 funicle. 1st segment longer than 2nd segment. Omolabus (Neomolabus) deceptor (Jekel, 3rd segment pointed, hardly longer than 1st 1860) segment. Attelabus deceptor Jekel, 1860: 207 Pronotum almost campaniform, 1.22–1.25 Material: Male (ISNB), [Brazil] “Minas times wider than length. Grooves weak. Sides Geraes”, “Coll. Roelofs”; male (ISNB), “Brasil”, almost direct. Disk convex, largely rugosity- “Coll. Dejean, Coll. Roelofs”, “Attelabus punctate. Scutellum almost rectangular, large, luridus mihi, h. in Brasilia, D. Lacordaire”; male wide, small punctate. (ZMUC), [Brazil] “Lagoa Santa, Reichardt”. Elytra almost square, almost equal length Distribution: Brazil. and width. Greatest width in humeri and behind the middle. Humeri convex, with small tooth. Omolabus (Neomolabus) piceus (Germar, Intervals flat, wide, transversal-wrinkled. Striae 1824) clear. Points in them small. Rhynchites piceus Germar, 1824: 187 Prothorax wrinkled, with teeth on first line. Material: Male (ISNB), “Brazil, Bresil, Precoxal part of the prothorax strongly elongated. ex. coll.Latreille, Coll. Dejean, Coll. Roelofs”, Meso- and metathorax with episternum largely “rugicollis Dej.”, “Attelabus rugicollis mihi, h. and densely punctate. Abdomen convex, weakly in Brasilia, D. Lacordaire”. flattened on middle, rugosity-punctate. 1st – 2nd Distribution: Brazil. ventrites wide. 3rd and 4th ventrites narrower. 5th ventrite very narrow. Pygidium weakly Omolabus (Neomolabus) spinicollis convex, densely punctate. Legalov, sp.n. (Tab. II: e–f, Fig. 1d) Legs long. Forelegs elongated. Profemora Material: Holotype – male (ISNB), widened. Meso- and metafemora weaker [French Guiana] “Cayenne”, “Coll. Dejean, widened. Protibiae long, curved, crenate on Coll. Roelofs”, “spinicollis Dej.”, “Attelabus internal edge, with mucro at apex. Meso- and spinicollis mihi, h. Cayenne”. Paratype – male metatibiae shorter, weakly biconcave, weakly (ZMUC), [Guyana] “Essequibo Smidt, Mus. widened to apex. Tarsi long. Length of body: S.j.T.L.”. 3.8–4.6 mm. Description: Male: Body red-brown, Diagnosis: This new species is close to lustrous, naked. Antennae, tibiae and tarsi more O. gibbiphorus Voss, 1925 but differs by the light. longer body, stronger convex humeri, narrower Head elongated. Rostrum elongated, pronotum, prothorax with long teeth, vertex widened to apex, small punctate. Antennae with teeth and an armament of the endophallus. located in first third of the rostrum. Eyes Distribution: French Guiana, Guyana. large, weakly convex. Forehead narrow, flat, with 2 longitudinal striae. Temples elongated, Acknowledgements transversal wrinkled. Vertex with middle striae, I wish to thank B. Brugge (Amsredam), with teeth directed sideways. Prementum with R. Danielsson (Lund), O. Jaeger (Dresden), 2 long teeth. K.-D. Klass (Dresden), P. Limbourg Antennae long, reaching pronotum first (Bruxelles), B.A. Korotyaev (Saint Petersburg), line. Scapus and 1st segment oval. Scapus O. Martin (Copenhagen), R. Poggi (Genova), little longer than 1st segment. 2nd segment A. Solodovnicov (Copenhagen) and J. Willers trapezoid, short, much shorter and narrower (Berlin), for help with the work. References Chevrolat L.A.A. Description de Curculionites provenant des captures de M. Le docteur

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Gundlach a l’ile de Porto-Rico // Bulletin de la Société entomologique de France. 1876. Ser. 5. T. 6. P. 227–229. Germar E.F. Insectorum species novae aut minus cognitae, descriptionibus illustratae. 1824. Vol. 1. Coleoptera. Halae: J.C. Hendelii et filii. xxiv + 624 pp. Gyllenhal L. In: Schoenherr C.J. Genera et species curculionidum, cum synonymia hujus familiae, species novae aut hactenus minus cognitae, descriptionibus a Dom. Leonardo Gyllenhal, C.H. Boheman, et entomologiis aliis illustratae. 1839. T. 5. Pars 1. Paris. 456 pp. Hamilton R.W. Revision of the new world genus Pilolabus Jekel (Coleoptera: Attelabidae) // Transactions of the American Entomological Society. 1994. Vol. 120. No. 4. P. 369–411. Hamilton R.W. New species of Euscelus Schoenherr (Coleoptera, Attelabidae) from Central America // The Coleopterists Bulletin. 2001. Vol. 55. No. 4. P. 453–470. Hamilton R.W. Omolabus Jekel in north and central America (Coleoptera: Attelabidae) // Zootaxa. 2005. No. 986. P. 1–60. Hamilton R.W. Euscelus species of the West Indies (Coleoptera: Attelabidae) // Zootaxa. 2007. No. 1495. P. 1–34. Jekel H. Insecta Saundersiana: or characters of undescribed insects in the collection of William Wilson Saunders, Esq. Coleoptera. Curculionoides 2. 1860. London: John van Voorst. P. 155–244. Legalov A.A. Taxonomy, classification and phylogeny of the leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) of the world fauna. 2003. Novosibirsk. CD-R. № 0320301200. 733+350 p. (641 Mb.). [in Russian] Legalov A.A. A new classification of ecological groups of the leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) // Eurasian Entomological Journal. 2004a. Vol. 3. № 1. P. 43–45. [in Russian] Legalov A.A. New data of the leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) of the world fauna with description of 35 new taxons // Baltic Journal of Coleopterology. 2004b. Vol. 4. № 1. P. 63–88. Legalov A.A. Leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) of the world fauna. 2007. Novosibirsk: Agro-Siberia. 523 pp. Olivier A.G. Encyclopédie méthodique. Historie Naturelle. 4. Insectes. 1798. Paris. ccclxxii + 331 pp. Riedel A. Taxonomy, phylogeny, and zoogeography of the weevil genus Euops (Insecta: Coleoptera: Curculionoidea) in the Papuan Region. Ph.Diss. 2002. Muenchen 216 pp. Sharp D. Insecta. Coleoptera. Rhynchophora. Curculionidae. Attelabinae, Pterocolinae, Allocoryninae, Apioninae, Thecesterinae, Otiorhynchinae. 1889. Pp. 1–40. In: Sharp D., Champion G.C. Biologia Central-Americana. 1889–1911. Vol. 4. Part 3. vi = 354 pp. + 15 pl. Voss E. Die Unterfamilien Attelabinae und Apoderinae (Col. Curc.) (18. Beitrag zur Kenntnis der Curculioniden) // Stettiner Entomologische Zeitung. 1925. Jg. 85. Heft 1–2. S. 1–78, 191– 304. Voss E. Einige bisher unbeschriebene Rhynchitinen der paläarktischen Region (Col. Curc.) (27. Beitrag zur Kenntnis der Curculioniden) // Entomologische Blätter. 1929. Jg. 26. S. 24–29. Voss E. Beschreibung neu bekannt gewordener Attelabinen aus der neotropischen Region (Col.) (60. Beitrag zur Kenntnis der Curculioniden) // Revista de Entomologia. 1937. Vol. 7. No. 2–3. P. 154–164.

Адреса для контактов: Legalov Andrei Institute of Animal Systematics and Ecology, Zoological Museum 630091, Novosibirsk, Frunze street, 11; e-mail: [email protected]

82 Цветная таблица 2

80 Алтайский зоологический журнал/вып. 2/2008

Обозначения к цветной таблице 2. a. Euopsidius lorentziensis Legalov, sp. n. b. Euscelus pseudbinotatus Legalov, sp. n. c. E. landanguinus Legalov, sp. n. d. Omolabus bifoveatoides Legalov, sp. n. e. O. spinicollis Legalov, sp. n. (major form) f. O. spinicollis Legalov, sp. n. (minor form) g. O. lagoasensis Legalov, sp. n. h. O. bicolor Legalov, sp. n. i. Iseuscelus flohri Voss, 1925.