Research Article Biology and Medicine, 3 (2) Special Issue: 320-323, 2011

eISSN: 09748369, www.biolmedonline.com

A review of parasitoids (: ) of Uttar Pradesh and Uttarakhand, India

*Akhtar MS1, Rafi U2, Usmani MK2, Dey D1 1Division of Entomology, Indian Agricultural Research Institute, New Delhi 110012 (India). 2Department of Zoology, Aligarh Muslim University, Aligarh 202002 (India).

*Corresponding Author: [email protected] Abstract A review of aphidiine parasitoids (Hymenoptera: Braconidae) associated with various species occurring in Uttar Pradesh and Uttrakhand, India was compiled. The fauna of Uttar Pradesh and Uttrakhand is represented by 40 species under 13 genera. This information is further reinforced with host names. Keywords: Aphid; ; distribution; Uttar Pradesh; Uttarakhand.

Introduction eglanteriae Haliday, A. matricariae Haliday, A. Aphid pest problems are known worldwide. polycostulari Das and Chakrabarti, A. qadrii Most of the aphid parasitoids belong to the (Shuja-Uddin), A. rosae Haliday, A. smithi subfamily Aphidiinae which can be Sharma and Subba Rao, A. staryi Das and distinguished structurally by their medium body Chakrabarti, A. urticae Haliday, A. size usually about 1.5-3.5 mm long, females uzbekistanicus Luzhetzki] having the highest with short ovipositor, metasomal tergum number of species followed by Binodoxys [8 weakly sclerotized (Achterberg, 1997). species viz., B. centaureae (Haliday), B. Approximately 400 species placed under 60 indicus (Subba Rao and Sharma), B. genera have been described (Dolphine and joshimathensis (Das and Chakrabarti), B. Quicke, 2001), of them 123 species under 23 mackaueri (Das and Chakrabarti), B. oregama genera, which is around 31% of the world (Agarwala, Saha and Mahapatra), B. rubicola fauna found in India (Dey and Akhtar, 2007). (Shuja-Uddin), B. sinensis (Mackauer), B. This subfamily was represented by four tribes tomentosae (Das and Chakrabarti)]. Analysis viz., Ephederini, Praini, Aphidiini and Trioxini. made on the host indicated that the While analyzing the existing information, we Aphis is parasitized by 7 species [Binodoxys concluded that it was absolutely necessary to sinensis (Mackauer), Binodoxys indicus bring together, summarize, check and compile (Subba Rao and Sharma), Lipolexis oregmae all the existing information in a compact (Gahan), Ephedrus plagiator (Nees), presentation. Therefore, an attempt was made Binodoxys rubicola (Shuja-Uddin), Aphidius to compile an inventory of Aphidiinae colemani Viereck, Cristicaudus indicus Das parasitoids associated with aphids from Uttar and Chakrabarti] followed by genus Pradesh and Uttrakhand. Macrosiphum [Binodoxys joshimathensis (Das and Chakrabarti), Ephedrus lacertosus Materials and Methods (Haliday), Aphidius rosae Haliday, Praon Host data for Uttar Pradesh and Uttrakhand volucre (Haliday), Aphidius polycostulari Das records were compiled for each species. The and Chakrabarti, Aphidius uzbekistanicus information were collected from many primary Luzhetzki] and genus Myzus with 5 species and secondary sources of information material, [Aphidius staryi Das and Chakrabarti, both electronic and non-electronic viz., Review (M'Intosh), Lipolexis of Applied Entomology, CAB Pest CD, oregmae (Gahan), Aphidius eglanteriae Zoological Records, Thompson Catalogue, Haliday, Binodoxys tomentosae (Das and Hymenopterorum Catalogus and various other Chakrabarti)]. related books, journals, bulletins, newsletters, etc. were accessed. Acknowledgement We would like to thank Dr. V.V. Ramamurthy, Results and Discussion Principal Investigator, Network Project on The fauna of Uttar Pradesh and Uttrakhand is Biosystematics (NPIB 21-17), Division represented by 40 species under 13 genera of Entomology, Indian Agricultural Research (Table 1). Analysis of data reveals that Institute, New Delhi for providing necessary Aphidius [12 species viz., A. absinthii Marshall, facility and Indian Council of Agricultural A. cingulatus Ruthe, A. colemani Viereck, A.

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Research (ICAR), New Delhi for providing Dey D, Akhtar MS, 2007. Diversity of natural financial assistance to NPIB 21-17. enemies of aphids belonging to Aphidiinae (Hymenoptera: Braconidae) in India. Journal of Asia-Pacific Entomology, 10: 281-296. References Dolphin K, Quicke DLJ, 2001. Estimating the global Achterberg CV, 1997. Subfamily Aphidiinae. In: species richness of an incompletely described Wharton, R. A., Marsh, P. M., and Sharkey, M. J. taxon: an example using parasitoid (Eds.), Manual of the New World Genera of the (Hymenoptera: Braconidae). Biological Journal of Family Braconidae (Hymenoptera). International the Linnean Society, 73: 279-286. Society of Hymenopterists, Washington D.C., 1: 119-132.

Table 1. A list of Aphid Parasitoids (Hymenoptera: Aphidiinae) of Uttar Pradesh and Uttarakhand. S. Parasitoids Hosts No. 1. Aphidius absinthii Macrosiphoniella yomogifoliae on Artemisia sp.; M. Marshall kikungshana on Artemisia sp.; M. pseudoartemisae on Artemisia vulgaris; M. sanborni on sp.; Macrosiphoniella sp. on Chrysanthemum sp. 2. Aphidius cingulatus Pterocomma sp. on Populus citiata, Salix tetrasperma Ruthe 3. Aphidius colemani Aphis sp. on Rumex sp.; Hyalopterus pruni on Prunus persicae Viereck 4. Aphidius Myzus sorbi on Sorbaria tomentosa; Macrosiphoniella eglanteriae Haliday pseudoartemisiae on Artemisia vulgaris; Capitophorus formosartemisiae on Artemisia sp. 5. Aphidius helichrysi on Prunus persicae matricariae Haliday 6. Aphidius Macrosiphum (Sitobion) sp. on Rosa sp. polycostulari Das and Chakrabarti 7. Aphidius qadrii Brachycaudus helichrysi on Anaphalis sp.; Brachycaudus sp. on (Shuja-Uddin) Gnaphalium sp.; Capitophorus sp. on Anaphalis cinnamomea; Unidentified aphid on Artocarpus heterophyllus 8. Aphidius rosae Macrosiphum (Sitobion) rosaeformis on Rosa sp.; Macrosiphum Haliday (Sitobion) rosae on Rosa sp. 9. Aphidius smithi Acyrthosiphon pisum on Pisum sativum, Medicago sativa, Vicia Sharma and Subba faba Rao 10. Aphidius staryi Das Myzus cornutus on Prunus cornuta and Chakrabarti 11. Aphidius urticae Acyrthosiphon sp. on Euphorbia pilosa Haliday 12. Aphidius Acyrthosiphon sp. on Bothriochloa sp.; Macrosiphum miscanthi uzbekistanicus on Hordeum vulgaris; Schzaphis sp. on Grass, Macrosiphum Luzhetzki (Sitobion) miscanthi; Macrosiphum (Sitobion) sp. on Avena sativa, Grass, Penisetum falccidum 13. Archaphidus Greenidea ficicola on Psidium guajava; G. formosana heeri on greenideae Stary Syzygium guajava and Schlinger

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14. Betuloxys Capitophorus formosartemisiae on Artemisia vulgaris, intermedius (Shuja- Artemisia sp Uddin) 15. Binodoxys Senecio sp. centaureae (Haliday) 16. Binodoxys indicus Aphis craccivora on Solanum nigrum; A. gossypii on Bidens (Subba Rao and pilosa, Capsicum sp., Cestrum nocturnom, Chrysanthemum sp., Sharma) Coccinia cardifolia, Cucumis melo, Cuscuta reflexa, Durant sp., Gossypium herbaceum, G. hirsutum, Gynura angutosa, Hibiscus rosasinensis, Psidium guajava, Solanum melongena, Tinospora sp.; A. nerii on Calotropis procera; Aphis sp. on Solanum sp.; Brachycaudus helichrysi on Pyrus sp. 17. Binodoxys Macrosiphum (Sitobion) miscanthi on Bothriochloa sp. joshimathensis (Das and Chakrabarti) 18. Binodoxys Melanaphis meghalayensis meghalayensis on Bamboo mackaueri (Das and Chakrabarti) 19. Binodoxys oregama Ceratovacuna sp.? perglandulosa on Grass (Agarwala, Saha and Mahapatra) 20. Binodoxys rubicola Aphis ruborum longisetosus on Rubus ellipticus (Shuja-Uddin) 21. Binodoxys sinensis Aphis citricola and A. gossypii on Cestrum nocturnum (Mackauer) 22. Binodoxys Myzus sorbi on Sorbaria tomentosa tomentosae (Das and Chakrabarti) 23. Cristicaudus Capitophorus formosartemisiae on Artimisia vestita garhwalensis, Das and Chakrabarti 24. Cristicaudus indicus Aphis sp. on Glochoin Das and Chakrabarti 25. Diaeretiella rapae Brachycaudus helichrysi on Prunus amygdalus; Brevicoryne (M'Intosh) brassicae on Raphanus sativus; Hayhurstia atriplicis on Chenopodium album; Lipaphis erysimi on Brassica campestris var. cauliflower, Mustard; Metopolophium (Metopolophium) dirhodum; Myzus persicae 26. Ephedrus lacertosus Macrosiphum (Sitobion) miscanthi; Takecallis himalayensis on (Haliday) Bamboo 27. Ephedrus laevicollis Cavariella aegopodii on Salix hastate (Thomson) 28. Ephedrus nacheri Hayhurstia atriplicis on Chenopodium album; Melanaphis Quilis arundinariae on Pyrus pashia 29. Ephedrus plagiator Aphis fabae on Tagetes erecta (Nees) 30. Ephedrus Prociphilus sp. on Lonicera quinquelocularis; Unidentified srinagarensis Stary aphids on Cotoneaster sp. and Bhagat

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31. Kashmiria aphidis Prociphilus sp. on Lonicera quinquelocularis Stary and Bhagat 32. Lipolexis oregmae Aphis craccivora on Benincasa hispida, Cassia fistula; A. (Gahan) gossypii on Rumex sp.; A. nasturtii on Luffa cylindrical, Tectona grandis; Myzus persicae on Capsicum frutescens, Solanum melongena 33. Lysiphlebia mirzai Hyalopterus pruni on Phragmites kerke; Longiungus sacchari Shuja-Uddin on Sugarcane; Melanaphis sacchari on Pennisetum glaucum, Saccharum officinarum, Sorghum bicolor, Zea mayz, Rhopalosiphum maidis on Pennisetum glaucum, Triticum aestivum, Zea mays; Schizaphis graminum on Penninsetum glaucum, Zea mays 34. Lysiphlebus Melanaphis sp. on Grass; Rhopalosiphum maidis on Pennisetum delhiensis (Subba glaucum, Triticum aestivum, Zea mays; Schizaphis graminum Rao and Sharma) on Pennisetum glaucum, Zea mays 35. Pauesia lachniella Lachnus tropicalis on Quercus dealbata Das and Chakrabarti 36. Pauesia orientalis Cinara maculipes on Pinus excelsa Das and Chakrabarti 37. Praon himalayensis Callaphis juglandis on Juglans regia Das and Chakrabarti 38. Praon orientale Uroleucon sp. on Sanchus aevensis and Sanchus sp. Stary and Schlinger 39. Praon pubescens Nasonovia sp. on Sitobilanthes sp. Stary 40. Praon volucre Macrosiphum (Sitobion) sp. on Rosa sp. (Haliday)

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