Bulletin Zoologisch Museum

I ■ 0JNIVERSITEIT VAN AMSTERDAM

Vol. 10 No. 10 1985

Two new species of Dicranomyia Stephens, 1829, with notes on related species

(Diptera, Limoniidae)

Willy Geiger

Abstract

(D.) lorettae Dicranomyia sp.n. and Dicranomyia (D.) mattheyi sp.n. are described. Comments are

the of the and the chorea Lackschewitz & given on relationship new species group sensu Pagast,

1941. The of and mitis synonymy Dicranomyia hygropetrica Vaillant, 1952, Dicranomyia (Meigen,

1830) is established.

INTRODUCTION The species mentioned by Lackschewitz (1928)

the are following: livornica Lackschewitz, 1928,

his work the Lacksche- In basic on Limoniidae, mitis (Meigen, 1830), conchifera Strobl, 1901,

witz (1928) regrouped the species of the genus luteipennis Goetghebuer, 1920, chorea (Meigen,

Dicranomyia into categories named after a cha- 1818), incisurata Lackschewitz, 1928, and zernyi

racteristic representative. These groups, defin- Lackschewitz, 1928. The latter species was sub-

of and referred the ed by the general aspect the species the sequently to hyalinata group by

the male structure of genitalia, are no longer Lackschewitz & Pagast (1941). In their publica-

strobli used in the present-day taxonomy of the genus. tion they add to this list Pagast,

have been They replaced by genera or subgenera, 1941, madarensis (wollaston, 1858), micronychia

which since. have been created or reinstated Loew/Pagast, 1941, and signata Lackschewitz,

still 1941. Also Lackschewitz' s “chorea group", however, Dicranomyia (D.) michaeli Theowald,

remarkable All the be considered of constitutes a unit. species 1977, can part this group, as

included in it by Lackschewitz (1928) and subse- well as two new species, Dicranomyia (D.) loret-

quently added by Lackschewitz & Pagast (1941) tae sp.n. and Dicranomyia (D.) mattheyi sp.n. have been referred to the nominate subgenus, Eight species of this group inhabit Central and form a natural group. and Western Europe: chorea, mitis, conchife-

ra, luteipennis, lorettae, mattheyi, strobli and

in incisurata. Some species occur only southern l is ) This paper part of the author's doctor-

work Europe: madarensis (Madeira), signata (Corsica, . 54

Yugoslavia, Italy), michaeli (Azores, Madeira, corresponding segments; flagellum yellow tend-

and livornica and more Canary Is.) (Italy). D. microny- ing to greyish-brown, scape pedicel

chia is only known from two unlabelled speci- definitely yellow (only the right antenna is

cited & of the left the and mens by Lackschewitz Pagast, 1941, as complete, one only scape

“micronychia Loew". the pedicel remain); rostrum and palpi greyish-

In Central Europe two species, strobli and yellow. All the thoracic tergites yellow, with

their incisurata, are easily identified, male a grey pollinosity, silky; praescutum with a

characteristic. The other genitalia being very not very clear central stripe, slightly darker

similar: species are very a comparative study than the rest of the thorax; what can be seen

is in view of the necessary, especially descrip- of the pleurae, pierced by the pin, yellow with

tion of the two new Central European species. grey pollinosity, though less than the ter-

iridis- gites; coxae yellow; wings transparent,

Dicranomyia (D.) chorea (Meigen, 1818) cent, stigma practically invisible, no spots on

Figs. 1, 3. the front margin of the wing, ending in C

level at the about half be- Rs-base, way

Limonia chorea 1818: tween the base of the and no hairs on Meigen, 134. wing Sc^, Limonia lutea Meigen, 1804: 55 (nomen oblitum). Sc, distal cell closed, m-cu at its base; Limnobia quadra Meigen, 1838: 29.

Dicranomyia flavicollis Becker, 1908: 82. length of wing 6.4 mm.

Dicranomyia vidua Santos, 1923: 19. f. lutescens and the Dicranomyia longipes Santos, 1923: 38.

small forms ; Santos, 1923: 18; Lacksche- very light-coloured of D. mitis are

witz, in Frey, 1937: 7. much alike: without the help of the characteris- Dicranomyia chorea f. lutescens Lackschewitz,

1928: 211. tics of the without legs and, more important,

Dicranomyia chorea f. grisescens Lackschewitz, an examination of the structure of the male 1928: 211.

genitalia, it is impossible to classify a speci- Non :

Limnobia Wiedemann 1818: 134. modesta in Meigen, men with certainty. Two details, however, en-

able us to decide that the specimen of lutea in

Notes on synonymy.- the Meigen collection is probably identical

Mendl of with (1979) gave a chronological account chorea. First the Sc of mitis has hairs

lutea Meigen, the status of which for a long (Edwards, 1938; Mendl, 1979), while that of

time had not been clear. It has been is smooth and like lutea It regarded chorea hairless .

either as a species in itself, as a variety of must, however, be noted, that this criterion

of chorea. Mendl settles does be for mitis or as a synonym not seem to constant mitis, or at

for this last hypothesis on the basis of the least it is not always clearly visible. Further-

the of original drawings by Meigen, reproduced by more antennae mitis are entirely brown,

(1976). Lackschewitz (1928) had those of also of Morge already while chorea, and lutea, are

the come to same conclusion, establishing the greyish-yellow, at least in the first segments.

of with synonymy Limonia lutea Meigen Dicranomy- Probably therefore lutea Meigen is- identical

f. Lackschewitz. with ia chorea lutescens chorea Meigen and not with mitis Meigen,

Examination of the of only specimen Limonia as already suggested by Lackschewitz (1928) and

lutea in the Meigen collection in the Muséum Mendl (1979).

National d'Histoire Naturelle, Paris, leads us to confirm Lackschewitz' s and Mendl's state- Descriptive remarks.- ments. The male specimen is lacking the abdomen A species of variable coloration and size, and legs and the of the wing. There a apex right presenting a seasonal dimorphism: spring

labels: are two one dark grey, the (7-8 mm) with bearing form, large , grey, spotted wings number 440 40, and reddish with another one the (f. and an autumnal smaller grisescens) , form, indication "lutea c?". (5-6 nun) with no clear on the , yellow, spots

Description of L. lutea: Head yellow, with wings (f. lutescens).

antennal a grey pollinosity; segments spherical Genitalia ti: 9th tergite with slight indenta-

verticils to than the with oval basal oblong, slightly longer tion; cylindrical basistyle, an 55

lobe; od two times larger than the basistyle, Description. -

with slender spherical to oblong, a relatively General appearance as D. luteipennis Goetghe-

with two in rostrum very short, parallel spines buer; body brown, with a light brownish-grey

the middle. little brown pollinosity; wings transparent, a

with well visible and infuscated, a stigma

Distribution.- transverse veins with dark spots.

North Africa From to Scandinavia, eastwards Head dark brown, with a grey pollinosity;

far the North as as Urals; Canary Islands; rostrum and palpi brownish; antennae 14-segment-

America. and ed; scape brown, pedicel flagellum yellow-

brown; antennal segments spherical to oval, the

(D.) 1920 last verticils lon- Dicranomyia luteipennis Goetghebuer, seven being spindle-shaped;

Figs. 2, 4. ger than the corresponding segments. Thorax

dark brown; praescutum dark brown, with a grey

line Dicranomyia luteipennis Goetghebuer, 1920: 108, pollinosity and without any or stripe;

figs. 1, 4 (wing, genitalia d). pronotum, scutum, scutellum and postscutellum

trochanter and brown; pleurae brownish; coxae,

Descriptive remarks.- legs yellowish; femora without dark ring; wings

A reddish-yellow species; size 7 mm; wings transparent, with a yellow-brown infuscation, with well-marked stigma and with three brown veins yellowish-brown, ending in front at

spots, transverse veins clearly infuscated with Rs-base, half-way along Se, discal cell brown. with its closed, m-cu at base, stigma rectangu-

Genitalia <5: 9th tergite with slight indenta- brown and well brown lar, visible, a spot over tion; cylindrical basistyle with oval basal and Rs-base and another at the bifurcation

od almost 1.5 times lobe; spherical, as long as of Rs in R "4+5' this last spot is 2 +3 the basistyle, with a very short rostrum, bear- blended with the stigma, transverse veins of

ing two straight spines at its base, spines discal cell and m-cu have a vertical brown almost parallel and 1.5 times as long as the dark vertical strokes spot, forming two on api- rostrum. cal part of wing; halteres yellow. Abdomen

brownish.

Distribution. - Genitalia i: 9th tergite with a light exci-

Some localities in Central and Western sion on dorsal posterior margin; 9th sternite

Europe and in Italy. The distribution is most small the of is a oblong plate at base the

still known. probably very incompletely penis; basistyle yellowish-brown with an oval

ventral lobe; outer style yellowish, oval to

Dicranomyia (D.) mattheyi sp.n. spherical with a short rostrum, bearing at its

Figs. 5-9. base two dark brown straight spines, slightly

divergent, measuring 1.5 times the length of

Material.- the rostrum; ratio basistyle : outer style,

Id: Fuorn Holotype. Il (CH-GR), 1790 m, 6:7; inner style quite strong, bent at 90°; Swiss coordinates 812/171 7-8.VII1.1980, light

Musée trap (LT). W. Geiger & C. Dufour leg. in penis and parameres as in figs. 7-9.

d'Histoire naturelle de la Ville de Neuchâtel, 6.5 7.8 Body length mm; wing length mm; preserved in 70% alcohol.

Id: halteres 1.8 Paratypes. as holotype in Natur-Museum mm.

Luzern; Id: as holotype, 17-25.IX.1980 in Natur- 9 unknown. historisches Museum, Chur, coll. Swiss National

in Park; Id: as holotype, 9-11.VIII.1980 Zoôlo-

gisch Museum Amsterdam; Id: Fully (CH-VS),

550 m, LT, 13-19.X. 1980, W. Geiger & C. Dufour leg. in Musée d'Histoire naturelle de la Ville

Neuchâtel; Id: Altdorf (CH-UR), 465 de m, LT, Distribution .-

25.X. 1980, L. Rezbanyai leg. in coll. Dr. J. Switzerland, inner Central part of valleys Stary, Olomouc. All material is preserved in

707o alcohol. of the Alps. Figs. 1-16. 1: Dicranomyia chorea: genitalia ￿, dorsal view; 2: D. luteipennis: genitalia ￿,

dorsal view; 3: D. chorea: right od, detail; 4: D. luteipennis: right od, detail; 5:

D. mattheyi: right od, detail; 6: D. mattheyi: genitalia ￿, dorsal view; 7: D.

mattheyi: genitalia ￿, lateral view; 8: D. mattheyi: genitalia ￿, ventral view; 9:

D. left mitis: D. mattheyi: paramere; 10: Dicranomyia genitalia ￿, dorsal view; 11:

mitis: tarsal claws; 12: D. lorettae: genitalia ￿, dorsal view; 13: D. lorettae:

genitalia ￿, ventral view; 14: D. lorettae: genitalia ￿, lateral view; 15: D.

left 16: D. detail of the lorettae: paramere; lorettae: right of, (enlargement all figures: 100x). 57

Remarks.- Constantine, Rhumel, avril 1950 on another. The

This species is easily distinguished from two labels are white. This slide shows two

luteipennis by the dark brown head and praescu- wings, two 6 abdomens, a head with part of the

ratio of This second slide tum and the length basistyle/length thorax and three legs. is

outer style (od), being almost 1:1 in mattheyi probably the holotype (Vaillant, 1952: 250,

and 2:3 in luteipennis. "Type récolté, à l'état larvaire, dans les

gorges du Rummel, à Constantine").

Derivatio nominis.- The differential characteristics between mi—-

I am pleased to dedicate this species to my tis and hygropetrica proposed by Vaillant are

professor and master Dr. Willy Matthey, Neuchâ- the following: mitis has the praescutum with

the tel. three brown stripes, varying in width;

small proximal teeth of the tarsal claws very

Dicranomyia (D.) mitis (Meigen, 1830) and the wing spots pale but clear; hygropetrica

median the Figs. 10-11 has praescutum with one stripe,

proximal teeth of the tarsal claws well-develop-

Limnobia affinis Schummel, 1829: 127 (nomen ob- ed .

1 it um). Besides these characteristics a few details Limnobia mitis Meigen, 1830: 278

Limnobia excisa Walker, 1848: 47 in the original description of hygropetrica are 1920: 378, Fig. 1 ; Kuntze, times different from mitis: od 1.2-1.5 as long (genitalia d).

of the Diaranomyia mitis var. lutea Lackschewitz, 1928: as wide ( mitis 2.0 times); coloration

213. body ochraceous ( mitis greyish-yellow to brown- Diaranomyia mitis var. infuscata Lackschewitz,

1928: 213. ish-yellow) .

mitis imbecilla Lackschewitz Dicranomyia var. & The coloration of mitis is very variable, as Pagast, 1941: 30, Tafel VII, Fig. 56 (genitalia 6). is the ornamentation of the praescutum, which

mitis var. Lackschewitz & Dicranomyia affinis; all. can have one to three stripes or none at Pagast, 1941: 30.

The be Dicranomyia hygropetrica Vaillant, 1952: 250, wing spots may clearly marked, just

fig. 5 (wing), fig. 11 (tarsal claws), fig. 13— visible absent for the The or except stigma. 14 (genitalia rf) (syn. nov.).

only differential criteria that can be accepted Non :

Limnobia didyma Meigen, 1818: 135. are the structure of the d genitalia and the

claws of the tarsus. The material available,

dissected and does allow Notes on synonymy.- on slides, not any

Vaillant (1952) described Dicranomyia hygro- account to be taken of coloration. Precise

with larvae from measurements show that the od the three male petrica, a species hygropetric of

North This also is to 1.95 times Africa (Algeria). species oc- genitalia 1.75 as long as

in Maritimes the at which the curs France, Alpes (Vaillant, wide, depending on point

The and the draw- measurements are taken (the in 1956) . original description genitalia being

ings accompanying it, do not permit to separate resin, the od has undergone changes in its

this species from mitis. Thanks to the kindness original form). These measurements correspond

Vaillant those taken No difference is to of professor F. (Grenoble) however, we to on mitis

have been able to examine two microscopic be observed between the other parts of the

of both Nor does slides with some types of D. hygropetrica. 1. genitalia species. a compari-

of the claws of the of A slide with the indication Dicranomyia hygrope- son tarsus hygropetrica

on with several of mitis reveal re- trica Vaillant one label and Dicranomyia, specimens any

Eel. 4.4.51 another. The markable difference: the of the Kerrata, Elevage, on largest proxi-

mal teeth is less than half two labels are white. The slide shows two always as long as

the tooth and the others wings, the end of a leg and a A abdomen. 2. A principal (fig. 11)

smaller. These observations slide with the indication Dicranomyia hygrope- are progressively

Vaillant label make it that Vaillant, trica on one and Dicranomyia, clear, hygropetrica 58

1952 = mitis (Meigen, 1830). Head yellowish-brown, rostrum pale yellow-

ish, palpi yellowish-brown, head with two

remarks.- Descriptive diffuse brown spots on the posterior part be-

This is variable in size species strongly hind the eyes; antennae 14-segmented, scape,

and coloration. There of the and are some specimens pedical flagellum brownish; pedicel spheri-

size of chorea f. lutescens, very light, and cal, other segments oblong to spindle-shaped;

other much larger of the size of conchifera, verticils shorter than the respective segments.

and brown. The be Thorax wing spots may very clearly yellow-brown; pronotum yellow, praescu-

marked absent It is the with central brown or altogether. only tum yellow-brown a stripe;

species of this group which sometimes has a scutum brownish, scutellum yellowish, postscu-

hairy Sc. tellum brown; pleurae yellowish; coxae pale

Genitalia with 6: 9th tergite slight indenta- yellow, legs yellow with a dark brown ring at

with tion; basistyle cylindrical, an oval basal tip of femora; wings transparent with yellowish-

od 1.5 the in front of lobe; times as long as basistyle brown veins, ending Rs-base,

with short rostrum, bearing two long distal half-way along Sc; stigma oval, yellowish-

times the of the dark brown and spines, spines 2.5 length brown; two spots on on

and bent. small base of rostrum, parallel slightly Rs-fork, a light-brown spot at

fork ,/R. ; discal cell closed, m-cu at R„ ' 2+3' 4+5',.

base of discal cell, transverse veins and tip Distribution.-

of R "1+2' M M a liçht brown Palaearctic region. 2+3' 3' 4

halo; halteres pale yellow. Abdomen brownish.

Genitalia d: 9th tergite with a V-shaped

dorsal 9th Dicranomyia (D.) lorettae sp. n. excision on posterior margin; ster-

12-16 basi- Figs. nite is a small plate at base of penis;

with style yellowish, an oblong ventral lobe;

outer style (od) yellowish, larger at base that

Material.- at tip; rostrum short, slightly bent with at

Holotype. le!: Vezia (CH-T1), 410 m, Swiss the base two yellowish-brown spines, slightly coordinates 716.400/98.500, light trap (LT), 8-

14.X.1979, W. Geiger & C. Dufour leg. in Musée divergent, measuring 1.5 times the length of

d'Histoire naturelle de la ViLle de Neuchâtel the rostrum; ratio basistyle: outer style is (MNHVN), preserved in 70% alcohol.

Ici: 6-12 .VIII 1979 inner bent and Paratypes. as holotype, . 6:7; style slight, regularly

in coll. Dr. J. Stary, Oloraouc; 1

(CH-TI) 340 m, LT, 21-30. IX. 1979, L. and parameres as in 10-13. , Rezbanyai figs.

leg in MHNVN; 1

330 m, 6-18. VIII. 1979, W. Geiger & C. Dufour 1.3 mm.

leg. in MHNVN; 16: ibid. 27 .VII 1-2. IX. 1979 and 9 unknown. 16: ibid. 20-26 .VIII 1979 both in Id . MHNVN;

460 Vernayaz (CH-VS), m, 18. VI. 1979, W. Geiger

in 16: 550 leg. MHNVN; Fully (CH-VS), m, LT,

1-7. IX. 1980 in Zoologisch Museum Amsterdam; 36 Remarks.-

ibid, 8-14. IX.1980, 16 ibid, 22-28. IX 1980 and This species is easily distinguished from D. 3 6 ibid, 6-9.X.1980 all in MHNVN; 16 Verschiez

540 W. & mitis the excision of the 9th (CH-VD), m, LT, 9-15.VI.1980, Geiger by V-shaped ter-

C. Dufour leg. in MHNVN; 16: Altdorf (CH-UR), gite and the spines of the rostrum being 1.5 465 m, LT, 15.IX.1979, L. Rezbanyia leg. in

times MHNVN; Ici: Kleinbosingen (CH-FR), 490 m, as long as the rostrum.

W. in MHNVN. All 2.IX.1980, Geiger leg. pre-

served in 70% alcohol.

Distribution. -

Switzerland, particularly Ticino and Valais. Description. -

General ba- appearance as D. mitis (Meigen) ;

sic coloration of body yellowish; wings trans- Derivatio nominis.-

well visible and three parent, with a stigma It is with gratitude that I dedicate this brown the costal spots on margin. species to my wife Lorette. 59

Figs. 17-19. 17: Dicranomyia conchifera: genitalia ￿, dorsal view; 18: D. strobli: genitalia ￿,

incisurata: dorsal view; 19: D. genitalia ￿, dorsal view (enlargement of all the

figures: 100x).

Dicranomyia (D.) conchifera (Strobl, 1901) 2. Posterior margin of the 9th tergite with

thorax at most a slight indentation; Fig. 17 yellowish-grey to yellowish-brown, or

light yellow 3

Posterior margin of the 9th tergite with Limnobia conchifera Strobl, 1901: 189. and thorax dark a deep narrow excision;

brown with a grey pollinosity with silve-

sheen strobli ry (fig. 18) Pagast Descriptive remarks.-

3. Large light yellow species; transverse Large species (10.5 mm); light yellow; wings veins clearly infuscated; od 3 times as

the with four small brown spots, transverse long as the basistyle (fig. 17) conchifera Strobl veins infuscated. Smaller species; wings with spots hardly

9th the small Genitalia d: very voluminous; tergite marked or absent; forms yellow,

the od 1.5 larger ones brownish-yellow; with slight indentation; cylindrical basistyle times as long as the basistyle (figs. 10, with oval ventral lobe; od cylindrical, three 11) mitis (Meigen)

short 4. with times as long as the basistyle, rostrum Posterior margin of the 9th tergite

a deep V-shaped excision 5 with two long slightly curved spines. Posterior of the 9th with -. margin tergite

at most a slight indentation 6

Distribution.- 5. Wings without spots, stigma very pale;

thorax dark brown with silvery-grey polli- Central and Western Europe, Italy, Balkans. nosity; spines of the rostrum straight

and very divergent (figs. 19) incisurata Lackschewitz

Anterior margin of the wing with stigma

KEY FOR THE IDENTIFICATION OF THE CENTRAL AND and three fairly well marked spots; tho-

rax yellowish-brown; od slightly conical; WESTERN EUROPEAN SPECIES OF THE DICRANOMYIA 1.5 spines times as long as the rostrum,

CHOREA GROUP implanted at its base, slightly divergent

lorettae (figs. 12-16) sp. n.

6. Spines implanted in the centre of the 1. Spines of rostrum very long (2.5. times rostrum, very short 7 the as long as rostrum), slightly bent; base the -. Spines implanted at the of ros- twice od cylindrical to reniform, as long the 8 trum, 1.5 times as long as rostrum.... as wide 2

7. anterior -. Spines shorter (1.5 times as long as the Spring form; greyish-brown; mar-

with and rostrum or less), straight; od spherical gin of the wing stigma three

times wide to ovoid, 1.5 as long as or well-marked spots (figs. 1, 3) less 4 chorea t. grisescens Lackschewitz 60

Autumnal form; yellow; wings -with stigma LACKSCHEWITZ, P., 1928. Die palaearktischen Lim-

and marked absent spots hardly or (figs. nobiinen des Wiener naturhistorischen Mu-

chorea lutescens 1, 3) f. Lackschewitz seum.- Annln Naturh. Mus. Wien, 42: 195-244.

1937. Limoniidae, in FREY, R., Die the brown , 8. Od as long as basistyle; thorax Dipterenfauna der Kanarischen inseln und 5-9) (figs. mattheyi sp.n. ihre Problème.- Comm. Biol. Soc. Sci. Fenni- Od 1.5 times as long as the basistyle; ca, 6: 6-8. thorax yellowish-brown to yellowish-red & F. Pagast, 1941. Limoniidae. Fliegen (figs. 2, 4) luteipennis Goetghebuer palaearkt. Reg., 139: 1-64.

MEIGEN, J.W., 1804. Klassifikazion und Beschrei-

bung der europaischen zweif liigeligen Insec-

ten, 1_: i-xxviii, 1-152, pi. 1-8, i-vi, 153- ACKNOWLEDGEMENTS 314, pi. 9-15 (Braunschweig).

,1818. Systematische Beschreibung der be-

kannten europâischen zweiflugeligen Insek-

should thank I like to Dr. W. Matthey (Neuchâ- xxxvi + 333 1-11 (Aachen). ten, 1_: pp., pis.

1830. 6: iv + 401 55-66 , Idem, pp., pis. tel) h.c. , Dr. H. Mendl Dr. Theowald (Kempten) , (Hamm).

van Leeuwen (Amsterdam), Dr. J. Stary (Olomouc) 1838. Idem, + 67- , _7* pp., pis. 74 (Hamm). for all their valuable advices, and Dr. L. Ma- MENDL, H., 1979. Die Limoniiden-AbbildungenMei-

tile and F. Vaillant for (Paris) Dr. (Grenoble) gens.- Beitr. Ent., 319-341.

MORGE, G. 1976. Dipteren-FarbtafeIn nach den available the material of , kindly making to me bisher nicht veroffent1ichten Original-Hand- lutea D. Meigen and D. hygropetrica Vaillant, zeignungen Meigens: ":

Abbildungen der europâischen zweif luge 1igen respectively. My thanks are also due to Madame Insekten nach der Natur".- Beitr. Ent., 26:

A. L'Eplattenier (Neuchâtel) for translating 543, pi. 161-305.

SANTOS ABREU, E., 1923. Monografia de los Limo- the article into English, and to all those who nidos de las Islas Canarias.- Mem. Real Ac.

have the material of the helped me to collect Cienc. Artes Barcelona, J_8: 35-164.

SCHUMMEL, T.E., 1829. Beschreibung der, in two new species, especially Dr. C. Dufour (Neu- Schlesien einheimischen, Arten einiger Dipte-

châtel) and Dr. L. Reser (Luzern). ren-Gattungen. 1 Limnobia Meigen.- Beitr.

Ent. Breslau, 97-201, pis. 1-5.

STROBL, G., 1901. Tief's dipterologischer Nach-

lass aus Kârnten und Oesterr.- Schlesien.-

Jb. naturh. Landesmus. Karnten, 48: 171-246.

REFERENCES VAILLANT, F., 1952. Quelques Limoniiden à

larves hygropétriquesRevue fr. Ent., 19:

244-251.

1908. 1956. BECKER, T., der Kanarischen In- Recherches sur la faune madicole Dipteren ,

seln.- Mitt. Mus. 4: zool. Berlin, 1-180. (hygropétriques s.l.) de France, de Corse et

British EDWARDS, F.W., 1938. short-palped crane- d'afrique du Nord.- Mém. Mus. natn. Hist,

. Taxonomy of adults.- Trans. Soc. Br. nat., Paris, [N.S.] 1_1 : 1-258.

Ent., 5: 1-168. WALKER, F., 1848. List of the specimens of 1920. KUNTZE, A., Limoniidae.- Zool. Jb. Syst., dipterous in the collection of the

43: 371-432. British Museum, 1: 1-229 (London).

W. Geiger,

Institut de Zoologie,

Université de Neuchâtel, Received : 3.1.1984 Neuchâtel ,

Switzerland. Distributed : 29.111.1985