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Open Thesis Final V2.Pdf
The Pennsylvania State University The Graduate School Department of the Geosciences TAXONOMIC AND ECOLOGIC IMPLICATIONS OF MAMMOTH MOLAR MORPHOLOGY AS MEASURED VIA COMPUTED TOMOGRAPHY A Thesis in Geosciences by Gregory J Smith 2015 Gregory J Smith Submitted in Partial Fulfillment of the Requirements for the Degree of Master of Science August 2015 ii The thesis of Gregory J Smith was reviewed and approved* by the following: Russell W. Graham EMS Museum Director and Professor of the Geosciences Thesis Advisor Mark Patzkowsky Professor of the Geosciences Eric Post Director of the Polar Center and Professor of Biology Timothy Ryan Associate Professor of Anthropology and Information Sciences and Technology Michael Arthur Professor of the Geosciences Interim Associate Head for Graduate Programs and Research *Signatures are on file in the Graduate School iii ABSTRACT Two Late Pleistocene species of Mammuthus, M. columbi and M. primigenius, prove difficult to identify on the basis of their third molar (M3) morphology alone due to the effects of dental wear. A newly-erupted, relatively unworn M3 exhibits drastically different characters than that tooth would after a lifetime of wear. On a highly-worn molar, the lophs that comprise the occlusal surface are more broadly spaced and the enamel ridges thicken in comparison to these respective characters on an unworn molar. Since Mammuthus taxonomy depends on the lamellar frequency (# of lophs/decimeter of occlusal surface) and enamel thickness of the third molar, given the effects of wear it becomes apparent that these taxonomic characters are variable throughout the tooth’s life. Therefore, employing static taxonomic identifications that are based on dynamic attributes is a fundamentally flawed practice. -
Recollections of William Finaughty, Elephant Hunter 1864-1875
Recollections of WILLIAM FINAUGHTY ELEPHANT HUNTER, 1864-1875 Digitized by the Internet Archive in 2012 with funding from LYRASIS Members and Sloan Foundation http://archive.org/details/recollectionsofw1916will THE RECOLLECTIONS OF WILLIAM FINAUGHTY ELEPHANT HUNTER 1864-1875 PRESS OP J. B. LIPPINCOTT COMPANY EAST WASHINGTON SQUARE PHILADELPHIA TWO HUNDRED AND FIFTY COPIES OF THIS BOOK HAVE BEEN PRINTED PREFACE In 1908 I had in my employ in Northwestern Rhodesia, William Finaughty, a son of William Finaughty who was one of the first white men to hunt elephants in Matabeleland. Mr. Selous says in one of his books that Finaughty stopped hunting because elephants were becoming too scarce, before he—Selous—first went to Africa. In 1913 I hap- pened to be in Bulawayo and, having a day to spare, I visited Finaughty on a farm nearby. He gave me the numbers of the Rhodesian Journal contain- ing his hunting experiences. One or two of the numbers are missing and as the journal has gone out of existence, I cannot replace them. His son had told me of his wonderful memory, and this I found confirmed by all who knew him. I found him a very slight old man with very white hair, much weakened by many attacks of fever. The value, if any, in the following pages, lies in the fact that they give a picture of a time now past which will never return. G. L. H. Poplar House St. Davids' Pennsylvania May, 1916 — INTRODUCTION [Just one word of introduction and explana- tion. The Editor of this paper was some months ago asked if he would undertake the task of inter- viewing and writing up for publication the remi- niscences of the famous South African big-game hunter, " Bill " Finaughty. -
Adventures in the Great Deserts;
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Disproportionate Dwarfism in a Wild Asian Elephant
Short Communication Gajah 38 (2013) 30-32 Disproportionate Dwarfism in a Wild Asian Elephant Rohan Wijesinha1, Nadika Hapuarachchi2, Brad Abbott, Jennifer Pastorini3,4 and Prithiviraj Fernando3* 1Federation of Environmental Organizations, Thimbirigasyaya, Sri Lanka 2Wildlife Conservation Society - Galle, Biodiversity Research & Education Center, Galle, Sri Lanka 3Centre for Conservation and Research, Rajagiriya, Sri Lanka 4Anthropologisches Institut, Universität Zürich, Zürich, Switzerland *Corresponding author: [email protected] Introduction and behaviourally the dwarf elephant appeared normal. Absence of previous records and body Dwarfism is unusual in wild animals. Individuals lumps from gunshot injuries indicative of crop with disproportionate dwarfism are especially raiding outside the park, suggest that he was unlikely to survive in the wild as shorter limbs not a resident but wandered into the park during would impose severe fitness costs in predators musth ranging. or prey. As social mega-herbivores without predators, Asian elephants are one of the very Forms of dwarfism few species in whom a dwarf phenotype may not be lethal. Here we report the first record of a free Two main phenotypic forms of dwarfism ranging adult wild animal - an Asian elephant occur, proportionate and disproportionate. In (Elephas maximus), with disproportionate proportionate dwarfism ‘normal’ allometric ratios dwarfism (Fig. 1). of the body and limbs are preserved, whereas in disproportionate dwarfism the limbs are Observation of dwarf elephant comparatively short. Both types of dwarfism are observed in humans. In domestic animals dwarf On July 7, 2013, at approximately 6:30 am, two phenotypes have been selectively bred, creating a elephants were observed ‘sparring’ on the main number of distinct breeds in dogs, cattle, horses, road of the Udawalawe National Park in southern rabbits and other livestock. -
{TEXTBOOK} Elephant
ELEPHANT PDF, EPUB, EBOOK Raymond Carver | 128 pages | 05 Jul 2011 | Vintage Publishing | 9780099530350 | English | London, United Kingdom Elephant - Wikipedia The seeds are typically dispersed in large amounts over great distances. This ecological niche cannot be filled by the next largest herbivore, the tapir. At Murchison Falls National Park in Uganda, the overabundance of elephants has threatened several species of small birds that depend on woodlands. Their weight can compact the soil, which causes the rain to run off , leading to erosion. Elephants typically coexist peacefully with other herbivores, which will usually stay out of their way. Some aggressive interactions between elephants and rhinoceros have been recorded. At Aberdare National Park , Kenya, a rhino attacked an elephant calf and was killed by the other elephants in the group. This is due to lower predation pressures that would otherwise kill off many of the individuals with significant parasite loads. Female elephants spend their entire lives in tight-knit matrilineal family groups, some of which are made up of more than ten members, including three mothers and their dependent offspring, and are led by the matriarch which is often the eldest female. The social circle of the female elephant does not necessarily end with the small family unit. In the case of elephants in Amboseli National Park , Kenya, a female's life involves interaction with other families, clans, and subpopulations. Families may associate and bond with each other, forming what are known as bond groups which typically made of two family groups. During the dry season, elephant families may cluster together and form another level of social organisation known as the clan. -
As Guest, Some Pages Are Restricted
O Y GH 7 B C P RI T, 1 9 3 , Y CHA R LES SCR IBNER ’S SONS Printed in the United States of A m eric a ll o a t o th book A rights reserved . N p r f is m ay be reproduced in any f orm without ’ the perm ission of Cha rles S c ribner s S ons PR EFA C E This kind of book—and it is a rare kind—is the best bo o k television we have yet discovered . It is scarcely a f at all . Never did print have such di ficulty in remain in g black or paper so nearly dissolve into a clear an d I u ndian air . It would be abs rd to discuss the writing u to as writing , as it wo ld be talk ofthe brushwork of a smile seen in the street . In short , these letters are not works of art , but n i n works of ature . Nature has certa n adva tages n n A n d n natural ess amo g others . for authe ticity there n hin i who is ot g to beat it , as those artists adm t marry their models . So here we have India as one person saw it ; the right sort of India and the right sort of person . No u politics or ologies intrude ; there are palaces , j ngles , k n mon eys , elepha ts . That is the India I was brought A lo t w up upon . ofpeople have been tinkering ith it i ever since , and I am to learn that it is st ll there beyond I fa cing T H E E L EP H A NTS A R E G ON E BA C K T O ' T H EI R J UNG L E Photograph by Barba ra Flaherty va r: Ingen an an d the tropic oce , I am glad to have a piece of it n here in my ha d . -
Dwarf Elephants on Mediterranean Islands: a Natural Experiment in Parallel Evolution
Dwarf elephants on Mediterranean islands: A natural experiment in parallel evolution Volume 1 of 2 by Victoria Louise Herridge Department of Genetics, Evolution and Environment University College London A thesis submitted for the fulfillment of the Degree of Doctor of Philosophy University College London, 2010 1 I, Victoria Louise Herridge, confirm that the work presented in this thesis is my own. Where information has been derived from other sources, I confirm that this has been indicated in the thesis. Signed: Date: 2 Abstract Mediterranean dwarf elephants represent some of the most striking examples of phyletic body- size change observed in mammals and are emblematic of the ‘island rule’, where small mammals become larger and large mammals dwarf on islands. The repeated dwarfing of mainland elephant taxa (Palaeoloxodon antiquus and Mammuthus meridionalis) on Mediterranean islands provide a ‘natural experiment’ in parallel evolution, and a unique opportunity to investigate the causes, correlates and mechanisms of island evolution and body-size change. This thesis provides the first pan-Mediterranean study that incorporates taxonomic and allometric approaches to the evolution of dwarf elephants, establishing a framework for the investigation of parallel evolution and key morphological correlates of insular dwarfism. I show that insular dwarfism has evolved independently in Mediterranean elephants at least six times, resulting in at least seven dwarf species. These species group into three, broad size-classes: ‘small- sized’ (P. falconeri, P. cypriotes and M. creticus), ‘medium-sized’ (P. mnaidriensis and P. tiliensis) and ‘large-sized’ (Palaeoloxodon sp. nov. and ‘P. antiquus’ from Crete). Size-shape similarities between independent lineages from the east and central Mediterranean indicate that homoplasy is likely among similar-sized taxa, with implications for the existence of meta-taxa. -
Tusked Elephants Challenge the Current View of Elephant Evolution
SHORT REPORT Palaeogenomes of Eurasian straight- tusked elephants challenge the current view of elephant evolution Matthias Meyer1*, Eleftheria Palkopoulou2, Sina Baleka3, Mathias Stiller1, Kirsty E H Penkman4, Kurt W Alt5,6, Yasuko Ishida7, Dietrich Mania8, Swapan Mallick2, Tom Meijer9, Harald Meller8, Sarah Nagel1, Birgit Nickel1, Sven Ostritz10, Nadin Rohland2, Karol Schauer8, Tim Schu¨ ler10, Alfred L Roca7, David Reich2,11,12, Beth Shapiro13, Michael Hofreiter3* 1Max Planck Institute for Evolutionary Anthropolgy, Leipzig, Germany; 2Department of Genetics, Harvard Medical School, Boston, United States; 3Evolutionary Adaptive Genomics, Institute for Biochemistry and Biology, Department for Mathematics and Natural Sciences, University of Potsdam, Potsdam, Germany; 4Department of Chemistry, University of York, York, United Kingdom; 5Center of Natural and Cultural History of Man, Danube Private University, Krems-Stein, Austria; 6Department of Biomedical Engineering and Integrative Prehistory and Archaeological Science, Basel University, Basel, Switzerland; 7Department of Animal Sciences, University of Illinois at Urbana-Champaign, Urbana, United States; 8State Office for Heritage Management and Archaeology Saxony-Anhalt with State Museum of Prehistory, Halle, Germany; 9Naturalis Biodiversity Center, Leiden, Netherlands; 10Thu¨ ringisches Landesamt fu¨ r Denkmalpflege und Archa¨ ologie, Weimar, Germany; 11Broad Institute of Harvard and MIT, Cambridge, United States; 12Howard Hughes Medical Institute, Harvard Medical School, Boston, United States; 13Department of Ecology and Evolutionary Biology, University of California, Santa *For correspondence: mmeyer@ Cruz, United States eva.mpg.de (MM); michael. [email protected] (MH) Competing interests: The Abstract The straight-tusked elephants Palaeoloxodon spp. were widespread across Eurasia authors declare that no during the Pleistocene. Phylogenetic reconstructions using morphological traits have grouped them competing interests exist. -
Mammoths, Woolly Rhino, Giant Elk >
QUATERNARY GEOLOGY 75 b6 Extinct Stone Age animals < mammoths, woolly rhino, giant elk > This is what we have come to: ash and shivered glass. Memorialized dead-centres without focus. Atrocious glamour / grime plastered corrosive, corroded; road salts, metals. —Geoffrey Hill, Speech! Speech! 1 Common folk have long been intrigued by the remains frozen in the tundra of animals that are no more. For millennia before the invention of elephant gun, mammoth tusks were a source of entrepreneurial ivory. Naturalists have attempted to explain the Nordic mythology of frost giants in terms of the bones of extinct mammoths and beached whales that the cold of northern lands has preserved. Unicorn horns (purported to detect and eliminate poison in food, and so valued by kings and tyrants) were brought to Medieval Europe by Vikings who kept secret for 300 years the source— the ‘mythical beast’ is the Narwhal (Monodon monoceros).6 These rarest of whales live in the frigid Arctic seas north of 70E latitude and favor Baffin Bay between Canada and Greenland.7 The spectacular spiraling tooth (length to 10 feet and weight to 22 pounds) grows on the left-hand side of the upper jaw of males (some grow a double tusk and sometimes females grow a single thin horn).8 Giant deer Megaloceros giganteus, “Irish elk” (sic: moose with broad palmate antlers; elk have thin, many branched, pointy antlers), now extinct, left their horns, which deer (family Cervidae) males characteristically grow anew each spring, strewn in Ireland and elsewhere across Eurasia.9 Neolithic people did not write (or did so on perishables) but some did leave cave paintings and rock etchings of animals.2 The antiquity of these is evident from the animals shown, some far outside their present geographic zones and some extinct and not known to have lived in historical times. -
History of the Knysna Elephants: the Southernmost Free Roaming Elephants in the World
History of the Knysna Elephants: The Southernmost Free Roaming Elephants in the World By Ryno Joubert Email: [email protected] (2018) 1 CONTENTS History (Pre-colonial – 2017): 3 Acknowledgements: 20 Bibliography: 21 Appendices: A: Photos of the Major Pretorius hunt in 1920 24 B: Knysna bull elephant photographed by F.W. Newdigate in 1940 25 C: Elephants photographed by Alan Roberts in 1962 26 D: Photos taken from the farm, Die Kop, by Johan Nel in 1968 27 E: One of the Knysna elephants photographed in 1968 by Graham Kepping 29 F: Elephant photographed by Dave Reynell (6 December 1968) in Harkerville 30 G: Aftand crossing the N2 near the Garden of Eden in May 1969 32 H: Photos of Aftand 34 I: Male calf of approximately 6 months old killed by a falling tree in the Harkerville forest in 1968 38 J: Elephant Dossiers from the 1969-1970 survey conducted by Nick Carter 40 K: Profile of Hairy Ears, animal number seven on Carter’s Elephant Dossiers 43 L: Map indicating the general boundary of the Knysna elephants in 1970 44 M: Two elephants digging at the turf near the cliff edge in Harkerville on 1 May 1969 45 N: Elephants roaming in the forest near the Garden of Eden photographed by E.M. Williams in 1977 46 O: The skeleton of an old bull found by fern-harvesters in the winter of 1983 48 P: The skeleton of the bull found in the winter of 1983 standing in the Forest Legends Museum 49 Q: Photo of a mature cow seen by a group of hikers in 1987 50 R: The skull and tusks of an old cow found dead in 1989 51 S: The ‘Matriarch’ photographed by Johan Huisamen in 1994 52 T: The imported Kruger elephants 53 U: Knysna elephant cow photographed by Hylton Herd in Nov/Dec 2008 59 V: Knysna elephant photographed by one of the Landmark Foundation’s camera traps in 2014 61 W: Maps indicating where the Knysna elephants could be found in 1983 and 1996 62 2 Pre-Colonial History Based on an estimated 3 000 elephants that may have roamed the Cape Floristic Region in pre-colonial times, it is assumed that about 1 000 elephants occupied the Outeniqua-Tsitsikamma (southern Cape) area. -
Late Pleistocene Dwarf Elephants from the Aegean Islands of Kassos and Dilos, Greece
Ann. Zool. Fennici 51: 27–42 ISSN 0003-455X (print), ISSN 1797-2450 (online) Helsinki 7 April 2014 © Finnish Zoological and Botanical Publishing Board 2014 Late Pleistocene dwarf elephants from the Aegean islands of Kassos and Dilos, Greece Sevket Sen1,*, Eric Barrier2 & Xavier Crété3 1) Laboratoire de Paléontologie du Muséum, CR2P-CNRS-UPMC, 8 rue Buffon, F-75005 Paris, France (*corresponding author’s e-mail: [email protected]) 2) iSTeP, Université Paris 6, CNRS, Case 129, 4 Place Jussieu, F-75252 Paris, France 3) 280 Chemin de la Villa des Chênes, F-30250 Aubais, France Received 17June 2013, final version received 3 Jan. 2014, accepted 20 Jan. 2014 Sen, S., Barrier, E. & Crété, X. 2014: Late Pleistocene dwarf elephants from the Aegean islands of Kassos and Dilos, Greece. — Ann. Zool. Fennici 51: 27–42. Dwarf elephants are quite well known from the western Mediterranean islands, but they are still poorly documented in most eastern Mediterranean islands. This paper reports on the discovery of a third lower molar of a dwarf elephant from the island of Kassos, which is situated in the southern Aegean, between Crete and Karpathos. This molar is determined as Palaeoloxodon aff. creutzburgi, a dwarfed species known from the Late Pleistocene caves and deposits in Crete and derived from the mainland straight-tusked elephant P. antiquus. An upper molar (M3) from the island of Dilos, previously referred to Elephas antiquus or E. mnaidriensis, is also redescribed. It is almost identical to the upper molars of a paleoloxodontine elephant from Naxos. Its small size and lack of sufficient comparative material lead us to determine it as Palae- oloxodon sp. -
First and Second Eras (To 1700 Ad) the First and Second Eras of Firearms Were Discussed in “Keep Your Powder Dry!: Unusual Firearms, Part 1.”
First and Second Eras (to 1700 ad) The first and second eras of firearms were discussed in “Keep Your Powder Dry!: Unusual Firearms, Part 1.” Third Era (1700–1860 ad) The third era of firearms saw the creation of many weapons with unbelievable destructive capacity, completely altering the way wars were fought. the weight of a pepperbox. Musket versions have Traditional fantasy weapons like the sword and been made, but the tendency of chain fire to rend spear were all but entirely obsoleted by this the hand holding the barrel grip has made the new technology. As such, use caution when larger version quite unpopular. introducing these weapons to your campaign Grapeshot Revolver. This cap & ball revolver world in large quantities. has a 9-shot cylinder that revolves around a Barrel Chain Rifle.This rifle has a barrel that single-shot smoothbore shotgun barrel. A lever is fed by chambers attached to a belt wrapped on the hammer enables the shooter to switch around the outside the gun, like a machine gun between firing bullets or the shotgun shell. belt. Where it differs from the later belt-fed The grapeshot revolver is also available as a machine guns is that the chain rifle must be carbine, which greatly increases its range. manually loaded and the chamber locked in place Gun Cane. Created as a “surprise weapon” before the bullet is fired. for self-defense, this walking cane conceals a Elephant Gun. The elephant gun is simply smoothbore single-shot carbine within its length. a large caliber rifle that was designed to bring The handle is weighted so that it can also be used down large game.