Western North American Naturalist

Volume 72 Number 1 Article 17

4-5-2012

Obituary: Wilmer Webster Tanner, 1909–2011

Stanley L. Welsh Brigham Young University, Provo, UT, [email protected]

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Recommended Citation Welsh, Stanley L. (2012) "Obituary: Wilmer Webster Tanner, 1909–2011," Western North American Naturalist: Vol. 72 : No. 1 , Article 17. Available at: https://scholarsarchive.byu.edu/wnan/vol72/iss1/17

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OBITUARY: WILMER WEBSTER TANNER, 1909–2011

Wilmer W. Tanner in British Columbia, summer 1998

On 28 October 2011, two months short of his parents were John Myron and Lois Ann Tan- 102nd birthday, Dr. Wilmer W. Tanner passed ner, descendants of early Utah pioneers, who away, leaving a lifetime of accomplishments. resided on a ranch in the open and meadow- His most enduring achievement was his mar- bottomed valley at Indianola in Sanpete County. riage to Helen Brown on 4 January 1935, which They had 5 children: Vasco, Ray, Laura, Wilmer, resulted in a family consisting of Lynn, David, and Jean. and Mary Ann. Helen died in 1995, following Wilmer attended public schools in Sanpete 60 years of marriage. At age 90, Wilmer, always County, Utah. Following graduation from high an optimist, married again, to Otella Tyndal school, he served as a missionary between Devey, who died in 1999. Wilmer was a caring 1929 and 1932 for the Church of Jesus Christ person, who showed consideration and kind- of Latter-day Saints in the Netherlands, where ness throughout his life to each of his wives he learned Dutch by immersion, there being and to his 3 children, 10 grandchildren, and 24 no language training prior to the mission. great-grandchildren. Wilmer considered his Upon his return to Utah, he attended Brigham posterity to be his greatest legacy. He was Young University (BYU), where he earned a proud of them! Wilmer’s career success came B.A. degree in 1936, and an M.S. degree in about in large part because of the sacrifices 1937. His earliest publications dealing with and hard work of Helen and the children. date from 1939 and give indication of a Wilmer was born 17 December 1909 in life-long interest in herpetology. Economic Fairview, Utah, the fourth of five children. His conditions of the time were difficult. Helen

118 2012] OBITUARIES 119 worked at a laundry in Provo for $7.00 per to a bonafide teaching and research institu- week to aid in keeping the family economi- tion. These new faculty members worked along cally viable. Wilmer, meantime, had completed with the amazing trio to build research collec- requirements necessary to become a biology tions and to teach literally tens of thousands of teacher and was eventually employed by the students who entered to learn and went forth Provo School District. Among his students to serve as productive individuals. The faculty was one Joseph Richard (Dick) Murdock, who also devoted time to numerous graduate stu- would ultimately serve as an assistant director dents, who became important teachers and of the Monte L. Bean Life Science Museum. researchers at various institutions across the How Wilmer survived live-wire students such country and beyond. Vasco developed an as Dick Murdock and his close friend Tom Pur- collection to be envied (curated later by vance is open to consideration. “They didn’t Wood) and an ichthyological collection; Hay- fool me!” Wilmer said of the boys’ shenanigans. ward built representative collections of birds “I knew what they were up to!” And he smiled! and mammals; Beck (and later Allred) added Work began on a Ph.D., but exigencies of greatly to the collections of ticks and fleas; and the time, mainly World War II, intervened. In Wilmer assembled a sizeable collection of 1946, he began studies with Edward Harrison amphibians and . Not one of these Taylor at the University of Kansas. And in dedicated professionals ever had the collec- 1948, Wilmer completed his dissertation on tion of zoological specimens formally listed as comparative anatomy of slamanders of Mexico part of their job descriptions! They built the and Central Mexico. collections in addition to their assignments as Vasco M. Tanner, the eldest child of John classroom instructors, committee members, and Lois Tanner, was some 17 years older than and student advisors. Their devotion and last- Wilmer and served as a role model for him. ing legacies to the University, the community Vasco earned a Ph.D. from Stanford Univer- at large, their students, and their families speak sity and joined the faculty at BYU in the early volumes (including hundreds of published 1920s, where he founded the zoological natu- papers in their respective disciplines). One ral history collections and the Department of wonders how they achieved balance in their Zoology and Entomology. Vasco was the one lives, but they did! person always available, Saturdays included, Wilmer’s name appeared sometime during in his campus office on the west side of the the 1970s in a sensational news article with a second floor of the Brimhall Building. He kept title that read, “Brigham Young University Pro- track of his former students, followed their fessor Missing in Mexico.” Wilmer was on a col- careers, and found employment for many of lecting trip to Barranca del Cobre, a deep gorge them as opportunities surfaced through his in northwestern Mexico, and the party had not connections. returned at the specified time. A search was Wilmer officially joined the BYU zoology undertaken and there was much to-do, but faculty in 1950, but he was teaching summer according to Wilmer, the group was never in school at the university in 1949 when one of danger and all were completely unaware that his students and life-long associates, Stanley they had been reported missing. Everything L. Welsh, completed a genetics class taught by came out well, except for short-lasting orders him. The faculty he joined was championed by from the Dean of Biology and Agriculture that an awesome trio: Wilmer’s larger-than-life all personnel had to check with him prior to brother Vasco, D Elden Beck, and C. Lynn leaving campus—an edict ignored at first and Hayward. Among them, they taught a bevy of later discarded as unnecessary. courses that covered the entire breadth of By the 1970s, it became apparent that the zoology, an assignment whose difficulty in - invertebrate/vertebrate collections, then housed creased vastly with the postwar arrival of with the Department of Zoology and Entomol- thousands of young men, who desired an edu- ogy, and the herbarium, then under direction cation and a chance to get on with their lives. of the Department of Botany, required a cen- Wilmer was soon joined by numerous newly tralized location apart from their departments. minted Ph.D.s—including Steven L. Wood Rationale for a change in administration of the and Dorald M. Allred—as the University tran- collections involved their perpetuity, with his- sitioned from a teacher-preparation institution tory of departmental collections being degraded 120 WESTERN NORTH AMERICAN NATURALIST [Volume 72 and sometimes destroyed by nonresearch re - Ephraim Hatch. Plans were drawn up and lated activities. Around 1970, Dr. Kent H. presented to Monte and Birdie; and construc- McKnight, professor of botany, was given the tion began in the mid-1970s. Money for con- task of exploring development of a collections struction, some $3.5 million, was donated by related facility separate from the departments. Monte L. Bean and his wife, who also estab- Upon his removal to the eastern , lished an endowment fund for the museum. Dr. Wilmer Tanner was chosen as his succes- The Monte L. Bean Life Science Museum was sor. He selected Dr. Stephen L. Wood, then occupied in fall 1977 and officially opened in curator of and related groups, and Dr. spring 1978. It now houses research collec- Stanley L. Welsh, then curator of the herbar- tions of arthropods, mollusks, fish, birds, ium, as committee members. They worked mammals, reptiles/amphibians, vascular plants, together to outline goals and procedures for and nonvascular cryptogams, as well as a mas- establishment of a separate entity to initially sive inventory of trophy from through- oversee and ultimately house the collections. out the world. The individual specimens in The committee was already functioning the museum now number in the millions. under established guidelines when a letter Monte, in a meeting with the committee in arrived at the BYU Development Office where the physical plant offices, looked at Wilmer Helen, Wilmer’s wife, was working as a secre- and stated, “I know that you have sufficient tary. That letter was from a person in Seattle room for the trophy animals, but do you have who desired to divest himself (actually at his sufficient room for the natural history collec- wife’s insistence) of his collection of trophy tions?” He then stated that the trophy mounts animals. The letter was shunted aside! No one were ephemeral and could and should be dis- would wish to receive such a collection of carded ultimately, but the natural history col- “stuffed animals,” they decided. Helen re - lections were the most important. trieved the letter, which was otherwise headed Without Helen’s interception of Monte’s for the discard file, and with permission asked letter, there is a very real possibility there to show it to her husband. Wilmer saw at once would be no life science museum at BYU, or if an opportunity! “Could this be the very open- so, a much smaller one relegated to a transient ing we have been hoping for that will allow existence in older buildings. segregation of the and plant collections Wilmer served as the first director of the in a separate facility?” Again with permission, museum, with Dorald Allred and Dick Mur- Wilmer contacted the letter’s author, Monte L. dock as assistant directors. Forced by law to Bean, whose wife, Birdie, had been the prime retire at age 65, Wilmer continued to visit the mover in getting him to release his collection museum almost daily to follow its growth and of animals from a life-long passion for hunting. development. He served for decades on a Soon Wilmer, Helen, Monte, and Birdie, were fundraising committee for the museum, and together in person; and shortly thereafter the traveled to California and other places to meet collection of trophy mounts arrived and was with potential or actual donors. He thought displayed temporarily in a portion of the main always of the museum! Wilmer continued to reading room of the [Heber J.] Library, actively write for publication until the time of its collections having been moved to a new his death. Three days prior to his passing, he facility. The east end of the reading room contacted the present museum director, Dr. housed the herbarium, which had been moved Larry L. St. Clair, about meeting to talk about from the third floor of the Brimhall Building future publications, and some of his work will some time earlier. Now, there were 2 parts to indeed be published posthumously. His body the puzzle in place. What was required was a wore out, but his mind never did! building large enough and sufficiently modern He will be greatly missed by all who knew to house the collections of arthropods, rep- him. tiles/amphibians, birds, and mammals. Wilmer Wilmer was a member of the American and Monte, and members of the committee, Society of Ichthyologists and Herpetologists, worked together to define what would be the American Association for the Advancement required. Monte listened and agreed. Also of Science, and the Herpetologist’s League working with the committee were Physical (Editor of Herpetologica, 1960–1967; Secre- Plant Vice President Fred Schwendiman and tary-Treasurer, ex officio, 1960; Vice President, 2012] OBITUARIES 121

1968–1969; President, 1970–1971; Executive ______. 1953. A study of and phylogeny of Lam- Council, 1972–1977). propeltis pyromelana Cope. Great Basin Naturalist 13:47–66. The following list of publications speaks ______. 1954. Additional note on the with volumes for the scientific legacy of Wilmer W. a description of a new subspecies. Herpetologica Tanner. 10:54–56. ______. 1954. Herpetological notes concerning some rep- tiles of Utah and Arizona. Herpetologica 10:92–96. Stanley L. Welsh ______. 1955. A new Sceloporus magister from eastern Utah. Monte L. Bean Life Science Museum Great Basin Naturalist 15:32–34. Brigham Young University TANNER, W.W., AND R.C. LOOMIS. 1957. A taxonomic and Provo, UT 84602 distributional study of the western subspecies of the milk , Lampropeltis doliata. Transactions of E-mail: [email protected] the Kansas Academy of Science 60(1):12–42. TANNER, W.W. 1957. A new Xantusia from southeastern Utah. Herpetologica 13:5–11. ______. 1957. Notes on a collection of amphibians and rep- ILMER ANNER IBLIOGRAPHY W W. T B tiles from southern Mexico, with a description of a (arranged in chronological order) new Hyla. Great Basin Naturalist 17:52–56. ______. 1957. A new skink of the multivirgatus group from TANNER W. W. 1939. Reptiles of Utah County. Utah Acad. Chihuahua. Great Basin Naturalist 17:111–117. of Sci. 16:105. ______. 1957. A taxonomic and ecological study of the west- ______. 1939. The status of the Utah Gopher Snake. Utah ern skink (Eumeces skiltonianus). Great Basin Natural- Acad. of Sci. 16:107. ist 17:59–94. TANNER, V.M., AND W. W. T ANNER. 1939. Notes on Charina ______. 1958. Herpetology of Glen Canyon of the Upper bottae in Utah: reproduction. Great Basin Naturalist Colorado River basin. Herpetologica 14:193–195. 1:27–30. ______. 1958. Herpetological range extensions. Herpeto- TANNER, W.W. 1940. Notes on the herpetological speci- logica 14:195–196. mens added to the Brigham Young University Verte- ______. 1958. Two new skinks from Durango, Mexico. Great brate Collection during 1939. Great Basin Naturalist Basin Naturalist 18:57–62. 1:138–146. HAYWARD, C.L., D ELDON BECK, AND W. W. T ANNER. 1958. ______. 1941. A study of the variation in the less common Zoology of the Upper Colorado River basin. I. The snakes of Utah. Great Basin Naturalist 2:16–28. biotic communities. Brigham Young University Science ______. 1941. The reptiles and amphibians of Idaho No. I. Bulletin, Biological Series 1(3):1–74. Great Basin Naturalist 2:87–97. HANSEN, A.M., AND W. W. T ANNER. 1958. A comparative ______. 1943. Two new of Hypsiglena from western osteological study of certain species belonging to the North America. Great Basin Naturalist 4:49–54. genus Bolitoglossa (Amphibia). Great Basin Naturalist ______. 1943. Notes on the life history of Eumeces skilto- 18:85–100. nianus skiltonianus. Great Basin Naturalist 4:81–88. TANNER, W.W. 1959. A new Thamnophis from western ______. 1944. A taxonomic study of the genus Hypsiglena. Chihuahua with notes on four other species. Herpeto- Great Basin Naturalist 5:25–92. logica 15(4):165–172. SMITH, H.M., AND W. W. T ANNER. 1944. Description of a ______. 1959. The status of Gerrhonotus in Utah. Herpeto- new snake from Mexico. Copeia 1944:131–136. logica 15:178–180. TANNER, W.W. 1949. Food of the wandering garter snake, TANNER, W.W., AND W. G. R OBISON JR. 1959. A collection Thamnophis elegans vagrans (Baird & Girard), in Utah. of herptiles from Urique, Chihuahua. Great Basin Herpetologica 5:85–86. Naturalist 19:75–82. ______. 1950. Notes on the habits of Microhyla carolinensis ______. 1960. Herpetological notes for northwestern Jalisco, olivacea (Hallowell). Herpetologica 6:47–48. Mexico. Herpetologica 16:59–62. ______. 1950. A new genus of plethodontid salamander from ______. 1960. New and unusual serpents from Chihuahua, Mexico. Great Basin Naturalist 10:37–44. Mexico. Herpetologica 16:67–70. ______. 1950. Variation in the scale and color pattern of TANNER, W.W. 1960. Crotalus mitchilli pyrrhus Cope in the wandering garter snake, in Utah and southern Utah. Herpetologica 16:140. Idaho. Herpetologica 6:194–196. LEVITON, A.E., AND W. W. T ANNER. 1960. The generic allo- FITCH, H.S., AND W. W. T ANNER. 1951. Remarks concerning cation of Hypsiglena slevini Tanner (Serpentes: Colu- the systematics of the collared , Crotaphytus col- bridae). Occasional Papers of the California Academy laris, with a description of a new subspecies. Kansas of Sciences 27:1–7. Academy of Science 54(4):548–559. TANNER, W.W., AND W. G. R OBISON JR. 1960. New name TANNER, W.W. 1951. Lampropeltis doliata rediscovered in for a Chihuahua lizard. Herpetologica 16:114. Arizona. Herpetologica 7:180. TANNER, W.W. 1960. Sceloporus p. poinsetti taken in Sonora. ______. 1952. A comparative study of the throat muscula- Herpetologica 16:235. ture of the Plethodontidae of Mexico and Central ______. 1961. A new subspecies of Conopsis nasus from America. University of Kansas Science Bulletin 34(2): Chihuahua, Mexico. Herpetologica 17(1):13–18. 583–677. TANNER, W.W., AND A.H. BRAHME JR. 1961. Description of ______. 1952. Diadophis regalis regalis (B. & G.) found in a new species of salamander from Panama. Great Basin Nevada. Great Basin Naturalist 12:63–64. Naturalist 21:23–26. ______. 1953. Notes on the life history of Plethopsis wrighti ROBISON, W.G., JR., AND W. W. T ANNER. 1962. A compara- Bishop. Herpetologica 9:139–140. tive study of the species of the genus Crotaphytus 122 WESTERN NORTH AMERICAN NATURALIST [Volume 72

Holbrook (Iquanidae). Brigham Young University BULLOCK, R.E., AND W. W. T ANNER. 1966. A comparative Science Bulletin, Biological Series 2(1): 1–31. osteological study of two species of (Pitu- TANNER, W.W. 1962. A new Bolitoglossa (salamander) from ophis and Thamnophis). Brigham Young University southern Panama. Herpetologica. 18.1:18–20. Science Bulletin, Biological Series 8(3):1–29. TANNER, W.W., AND B.H. BANTA. 1962. Description of a TANNER, W.W. 1966. A new rattlesnake from western new Hypsiglena from San Martín Island, México, Mexico. Herpetologica 22(4):298–302. with a resumé of the fauna of the island. Her- TANNER, W.W., AND B.H. BANTA. 1966. A systematic review petologica 18(1):21–25. of the Great Basin reptiles in the collections of Brig- ______. 1962. The distribution of Tantilla utahensis Blan- ham Young University and the University of Utah. chard. Great Basin Naturalist 22:116–118. Great Basin Naturalist 26:87–135. JORGENSEN, C.D., A.M. ORTON, AND W. W. T ANNER. 1963. TANNER, W.W. 1967. An albino wandering garter snake. Voice of the leopard lizard Crotaphytus wislizeni Baird Proceedings of the Utah Academy of Sciences, Arts and Girard. Proceedings of Utah Academy of Sciences, and Letters 43(1):163. Arts, and Letters 40(1):115–116. ______. 1967. Contia tenuis Baird and Girard in continental TANNER, W.W. 1963. Natural radiation effects on vertebrate British Columbia, Canada. Herpetologica 23(4):323. animals inhabiting the uranium areas of southeastern JENKINS, R.L., AND W. W. T ANNER. 1968. Osteology and Utah. Pages 325–326 in Vincent Schultz and Alfred mycology of Phrynosoma p. platyrhinos Girard and W. Klement, editors, Radioecology: Proceedings of the Phrynosoma d. hernandesi Girard. Brigham Young First National Symposium on Radioecology. Colorado University Science Bulletin, Biological Series 9(4): State University, Fort Collins, CO; 10–15 Sep tember 1–34. 1961. Reinhold Publ. Corp., , NY, and Ameri- BANTA, B.H. AND W. W. T ANNER. 1968. The systematics of can Institute of Biol. Sci., Washington, DC. 746 pp. Crotaphytus wislizeni, the leopard (Sauria: JORGENSEN, C.D., AND W. W. T ANNER. 1963. The application Iguanidae), Part II: a review of the status of the Baja of the density probability function to determine the California peninsular populations and a description home ranges of Uta stansburiana stansburiana and of a new subspecies from Cedros Island. Great Basin Cnemidophorus tigris tigris. Herpetologica 19(2): Naturalist 28:183–194. 105–115. TANNER, W.W. 1969. New records and distributional notes TANNER, W.W., AND C.D. JORGESEN. 1963. Reptiles of the for reptiles of the Nevada Test Site. Great Basin Natu- Nevada Test Site. Brigham Young University Science ralist 29:31–34. Bulletin, Biological Series 3(3):1–31. MORRIS, R.L., AND W. W. T ANNER. 1969. The ecology of TANNER, W.W., AND B.H. BANTA. 1963. The systematics of the western spotted frog, Rana pretiosa pretiosa Baird Crotaphytus wislizeni the leopard lizards, Part I: A and Girard, a life history study. Great Basin Natural- redescription of Crotaphytus wislizeni wislizeni Baird ist 29:45–81. and Girard, and a description of a new subspecies from TANNER, W.W. 1969. Presidential Address, AAS Banquet, the Upper Colorado River basin. Great Basin Natural- Boston. Pages 65–68 in Directory and Proceedings, ist 23:129–148. Association of Academics of Science 1969–70. TANNER, W.W. 1963. Obituary, Charles E. Burt. Copeia PACK, L.E., JR., AND W. W. T ANNER. 1970. A taxonomic 1963:722–723. comparison of Uta stansburiana of the Great Basin TANNER, W.W., AND J.W. HEINRICHS. 1964. An extension and the Upper Colorado River basin in Utah, with a of Arizona e. philipi and Rhinocheilus l. lecontei into description of a new subspecies. Great Basin Natu- south central Utah. Southwestern Naturalist 9:45–46. ralist 30:71–90. TANNER, W.W., AND D.F. AVERY. 1964. A new Sauromalus FISHER, D.L., AND W. W. T ANNER. 1970. Osteological and obesus from the Upper Colorado Basin of Utah. mylogical comparisons of the head and thorax regions Herpetologica. 20.1:38–42. of Cnemidophorus tigris septentrionalis Burger and BANTA B.H., AND W. W. T ANNER. 1964. A brief historical Ameiva undulata parva Barbour and Noble (family résumé of herpetological studies in the Great Basin Teiidae). Brigham Young University Science Bulletin, of the western United States Part I: The reptiles. Biological Series 11(1):1–41. Great Basin Naturalist 24:37–57. AVERY, D.F., AND W. W. T ANNER. 1970. Speciation in the AVERY, D.F., AND W. W. T ANNER. 1964. The osteology and Fijian and Tongan iguana Brachylophus (Sauria, Igua - myology of the head and thorax regions of the obesus nidae) with the description of a new species. Great group of the genus Sauromalus Dumeril (Iguanidae). Basin Naturalist 30:166–172. Brigham Young University Science Bulletin, Biologi- NASH, D.F., AND W. W. T ANNER. 1970. A comparative study cal Series 5(3):1–30. of the head and thoracic osteology and myology of BECK, D.E., D.M. ALLRED, J.R. MURDOCK, C.D. JORGENSEN, skinks Eumeces gilberti van Denburgh and Eumeces C.L. HAYWARD, AND W. W. T ANNER. 1964. Nevada skiltonianus (Baird and Girard). Brigham Young Uni- Test Site desert ecology. Proceedings of Utah Academy versity Science Bulletin, Biological Series 12(2):1–32. of Sciences, Arts and Letters 41(2):202–210. TANNER, W.W. 1970. A catalogue of the fish, amphibian, TANNER, W.W. 1965. A comparative population study of and reptile types in the Brigham Young University small vertebrates in the uranium areas of the Upper Museum of Natural History. Great Basin Naturalist Colorado River basin of Utah. Brigham Young Univer- 30:219–226. sity Science Bulletin, Biological Series 7(1):1–31. AVERY, D.F., AND W. W. T ANNER. 1971. Evolution of the ______. 1966. A re-evaluation of the genus Tantilla in the iguanine lizards (Sauria, Iguanidae) as determined southwestern United States and northwestern Mexico. by osteological and myological characters. Brigham Herpetologica 22(2):134–152. Young University Science Bulletin, Biological Series ______. 1966. The night snakes of Baja California. Transac- 12(3):1–79. tions of the San Diego Society of Natural History INGRAM, W., III, AND W. W. T ANNER. 1971. A taxonomic 14(15):189–196. study of Crotaphytus collaris between the Rio Grande 2012] OBITUARIES 123

and Colorado Rivers. Brigham Young University Sci- TANNER, W.W., AND J.E. KROGH. 1975. Ecology of the ence Bulletin, Biological Series 13(2):1–29. zebra-tailed Callisaurus draconoides at the Nevada TANNER, W.W., D.L. FISHER AND T.J. W ILLIS. 1971. Test Site. Herpetologica 31:302–316. Notes on the life history of Ambystoma tigrinum FANGHELLA, C., D.F. AVERY, AND W. W. T ANNER. 1975. nebulosum Hallowell in Utah. Great Basin Naturalist Urosaurus and its phylogenetic relationship to Uta 31:213–222. as determined by osteology and myology (Reptilia: TANNER, W.W., AND J.M. HOPKIN. 1972. Ecology of Scelo- Iguanidae). Great Basin Naturalist 35:245–268. porus occidentalis longipes Baird and Uta stansburiana SMITHM N.M., AND W. W. T ANNER. 1976. Index to Herpeto- stansburiana Baird and Girard on Rainier Mesa, logica 1956–1975 (Volumes 12–31). 165 pp. Nevada Test Site Nye County, Nevada. Brigham TANNER, W.W., AND N.M. SMITH. 1976. Cumulative Index Young University Science Bulletin, Biological Series to Brigham Young University Science Bulletin, Bio- 15(4):1–39. logical Series 1955–1970 (Volumes 1–20). 24 pp. KROGH, J.E., AND W. W. T ANNER. 1972. The hyobranchium COX, D.C., AND W. W. T ANNER. 1977. Osteology and myol- and throat mycology of the adult Ambystomidae of ogy of the head and neck regions of Callisaurus, the United States and northern Mexico. Brigham Cophosaurus, Holbrookia, and Uma (Reptilia: Igua- Young University Science Bulletin, Biological Series nidae). Great Basin Naturalist 37:35–56. 16(1):1–69. TANNER, W.W., AND B.H. BANTA. 1977. The systematics TANNER, W.W., J.R. DIXON, AND H.S. HARRIS JR. 1972. A of Crotaphytus wislizeni, the leopard lizards. Part new subspecies of Crotalus lepidus from western III. The leopard lizards of the Great Basin and Mexico. Great Basin Naturalist 32:16–24. adjoining areas, with a description of a new sub- SMITH, N.M., AND W. W. T ANNER. 1972. Two new subspecies species from the Lahontan Basin. Great Basin Natu- of Crotaphytus (Sauria: Iguanidae). Great Basin Natu- ralist 37:225–240. ralist 32:25–34. TANNER, W.W. 1978. Zoogeography of reptiles and amphibi - PEARCE, R.C., AND W. W. T ANNER. 1973. Helminths of ans in the Intermountain Region. Great Basin Natu- Sceloporus lizards in the Great Basin and Upper ralist Memoirs 2:43–53. Colorado Plateau of Utah. Great Basin Naturalist OTTLEY, J.R., AND W. W. T ANNER. 1978. New range and a 33:1–18. new subspecies for the snake Eridiphas slevini. Great TANNER, W.W., AND J.E. KROGH. 1973. Ecology of Scelo- Basin Naturalist 38:406–410. porus magister at the Nevada Test Site, Nye County, TANNER, W.W. 1981. A new Hypsiglena from Tiburon Island, Nevada. Great Basin Naturalist 33:133–146. Sonora, Mexico. Great Basin Naturalist 41:139–142. ______. 1973. Ecology of Phrynosoma platyrhinos at the TANNER, W.W., AND J.R. OTTLEY. 1981. Reproduction in Nevada Test Site, Nye County, Nevada. Herpetologica Hypsiglena. Great Basin Naturalist 41:310. 29(4):327–342. TANNER, W.W., AND D.C. COX. 1981. Reproduction in the ______. 1974. Ecology of the leopard lizard, Crotaphtus snake Lampropeltis pyromelana. Great Basin Natu- wislizeni at the Nevada test site, Nye County, Nevada. ralist 41:314–316. Herpetologica 30(1):63–72. PRITCHETT, C.L., H.H., FROST, AND W. W. T ANNER. 1981. LARSEN, K.R., AND W. W. T ANNER. 1974. Numeric analysis Terrestrial vertebrates in the environs of Utah Lake. of the lizard genus Sceloporus with special reference Great Basin Naturalist Memoirs 5:128–168. to cranial osteology. Great Basin Naturalist 34:1–41. TANNER, W.W. 1982. Herpetological notes from the Nevada SMITH, N.M., AND W. W. T ANNER. 1974. A taxonomic study Test Site. Great Basin Naturalist 42:219–222. of the western collared lizards, Crotaphytus collaris TANNER, W.W., AND D.F. AVERY. 1982. Buccal floor of rep- and Crotaphytus insularis. Brigham Young University tiles, a summary. Great Basin Naturalist 42:273–349. Science Bulletin, Biological Series 19(4):1–29. TANNER, W.W. 1983. Major Chapman Grant. Herpetologica TANNER, W.W., AND J.E. KROGH. 1974. Variations in activity 39:331. as seen in four sympatric lizard species of southern ______. 1983. Lampropeltis pyromelana. Reptilia: Squa- Nevada. Herpetologica 30(3):303–308. mata: Serpentes: Colubridae. Catalogue of American TANNER, V.M., AND W. W. T ANNER. 1974. Additional Amphibians and Reptiles 342(2). records of Coleoptera collected at the Nevada Test ______. 1984. Reptiles and amphibians of Idaho, No 2. Site, Mercury, Nevada. Great Basin Naturalist 34: Great Basin Naturalist 44:111–112. 218–220. ______. 1985. Snakes of western Chihuahua. Great Basin GROGAN, W.L., JR., AND W. W. T ANNER. 1974. Range exten- Naturalist 45:615–676. sion of the long-nosed snake, Rhinocheilus l. lecontei, ______. 1987. Lizards and turtles of western Chihuahua. into east-central Utah. Great Basin Naturalist 34: Great Basin Naturalist 47:383–421. 238–240. ______. 1988. Status of Thamnophis sirtalis in Chihuahua, BURKHOLDER, G.L., AND W. W. T ANNER. 1974. Life history Mexico (Reptilia: Colubridae). Great Basin Naturalist and ecology of the Great Basin sagebrush swift, Scelo - 48:499–507. porus graciosus graciosus Baird and Girard, 1852. ______. 1988. Eumeces skiltonianus. Catalogue of American Brigham Young University Science Bulletin, Biological Amphibians and Reptiles 447:1–4. Series 19(5):1–44. ______. 1989. Amphibians of western Chihuahua. Great ______. 1974. A new gland in Sceloporus graciosus males Basin Naturalist 49:38–70. (Sauria: Iguanidae). Herpetologica 30(4):368–371. COX, D.D., AND W. W. T ANNER. 1989. Hyobranchial appara- LARSEN, K.R., AND W. W. T ANNER. 1975. Evolution of the tus of the Cryptobranchoidea (Amphibia). Great Basin sceloporine lizards (Iguanidae). Great Basin Naturalist Naturalist 49:482–490. 35:1–20. TANNER, W.W. 1989. Status of Spea stagnalis Cope (1975), TANNER, W.W. 1975. 1975 Checklist of Utah amphibians Spea intermontanus Cope (1889), and systematic and reptiles. Proceedings of the Utah Academy of Sci- review of Spea hammondii Baird (1839) (Amphibia: ences, Arts and Letters 52(1):4–8. Anura). Great Basin Naturalist 49:503–510. 124 WESTERN NORTH AMERICAN NATURALIST [Volume 72

TANNER, W. W. AND C.H. LOWE. 1989. Variations in Tham- ______. 1997. Utah reptiles occurring only in southern Utah. nophis elegans with descriptions of new subspecies. Great Basin Naturalist 57:369–370. Great Basin Naturalist 49:511–516. FROST, H.H., AND W. W. T ANNER. 1999. C[harles] Lynn COX, D.C., AND W. W. T ANNER. 1995. Snakes of Utah. Hayward. Great Basin Naturalist 59:201–203. Monte L. Bean Museum, Provo, UT. 92 pp. TANNER, W.W. 1999. Distribution of the species Phryno- TANNER, W.W. 1996. James William Bee, 1913–1996. soma platyrhinos in Utah. Great Basin Naturalist Great Basin Naturalist 56:279–280. 59:295–296. ______. 1997. Dorald M. Allred, 1923–1996. Great Basin ______. 1999. Additional comments on the nesting behavior Naturalist 57:70–73. of Batrachoseps wrighti (Bishop). Great Basin Natu- ______. 1997. Type locality restriction of Hypsiglena torquata ralist 59:387–390. Gűnther. Great Basin Naturalist 57:79–82.