Ipl;xr,q" dlsnonrldsuof, euuruvl JeaI WW tllnrpuapouv eanf,unsrp oAr (8002 '7a 7a q8wg aos) gIVf Jo oru{u?rSordqns >lz1r\er"S Jo aeerery eqr Jo l,red se >lu-,r\urvs ur >lJo \pIOg Surqzrrepun elnl6 'eleqa.asla pue Epuns w euop ag ol srwuor i{f,mu'(ssatd ul'aodog) sarceds wql puulsJepun o] epetu uoeq suq lrvls e el.rtl,&\ 'poquf,srpun aq or readde pIOg rqt q paralunof,uo stuzld 1o t{tto[ew eiit ]zrp tleJ aqt uodn 'lanerzg pu? pu?Ircq1 'zrsdqel{ w1nsrrru3cl ui >lrorKpleg uo paszq '1sea1 ]ou pve 'vavqtaq q pue 'amtzJetll ul sorueu Jo uonzcrlddeslur Jo uonrpzJl 3uo1 z'sad&uds uox"t-pexrur uodn poszq JJu seuruu f,veut wW peJ aql 'saddl ptrrrolsrq ]soru Jo 'uonTpuol puz 'arnteu alznbepuur eq] dq papunodruo) ete sruelqord eqJ, 'pleg aq] q sruqd o] seru"u eseq] Jo uonzf,rtddz aqr prc8at ul drpcggrp olqzreprsuof, slsrxo ererp ]nq 's1uzl W te setueu peqs[qnd y1 aw ereq+ dpuarrn3 '(uzuu.i uralsaaqlnos sE rpJou r"J "rr1ql su ?rm{3opul pu"'pu"Lruql'sawdd[n{cl eW SpqS zpuns aql ol polrulsar sailqdrdalurar{ snua8 st lunqilappil ;o Wtus "

tu flJp a ap p aV' evatety : spt o ra Ay

'sesEq ?rrllu"I awpJof, dpuolsrsuoc aqt dq puz 'slooqs parrSuola 3o sdp arp rlo Suuarnog fqoptulot 'Vprw aa{o 'V qtoq uro4 srelJlp lng'srseq}ue elerJqlv PolJosxe lou srerFus eq] Jo tefrerew aq] seJsgs osIY sffaa?l\wu tuftJpaappav 'srseqtu? eryw tb Stnpuetxe tou slueurclll uew?ts etp puz 'acuecseroguds prouo8ro8 Fpourlq arp vr au{{o 'V :o.oq sroJJlp aprltlet tl/tupaappuv '(erererung) aa$o 'V o] J?Trurrs lsoru are daql snocnq8 f,11vrxvqe vwwvl JseI eql dg 'paruasnlp pu" pequf,sep arv 'suuanTauv pue alztulot 'V (11vzorEwS rnory a/flJpaappuv l{ev o/LL lf,?Irsqv

Sual urs 8uo/N puu ardog'J retocl a

}IvtrVUVS WOUf, WNAQMIOVNV IIIAON OA"I. :I OIINUOfl f,O (WUTVUV) WUUU,TSNOru NO SilO0ff 286 STUDIES ONMONSTERBAE (ARACEAE) OFBORNEO I PETERC l

adaia)ly m^tte olive green (in striking contrast to all other glaucous, Anadendrii Botneensibus < Sarawak species in which leaf laminae adzxtzlly glossy medium ^te adultis basaliter cuneads, , to dark green, while abanalTy palet and slighdy shiny), and with the gorgonoidiis bimodulis petiolar sheath extending to the base of the apical pulvinus, were (Sumatera) accedit, petioJ: encouritered, both usually, but not exclusively, in association with exserta distinguitur. limestone. Trpus: lvlalay'sia, Sa Anadendrum Schott, Bonplandia 5: 45 (1857); Mayo, Bogner & 54.8"r- 110o 08' 08.2", 45 Boyce, Genera of Araceae 11,3,Map.11, Pl. 11 (1997). \2ngAR-2450 (SAR, hoio

climbing hemiepipfutes. I-.eaues opposite on climbing shoots, Snall C /i n bi ng h eniepip fu'le rc' usually congested-distichous on flowering shoots; petiole pulvinate florvering shoots; climb^: apically and basaliy, sheathed to the apical pulvinus in most species, green. Itaues on flot'ern: sheath persistent or marcesc ent; lamina obliquely ovato-oblong, entire; petiole 5 - 33 cm long. ou primary Latenl veins pinnate, running into matginal vein, higher ca 1. - 2 cm long; pefio,'-;" -', order venation reticulate. Inflorescences 1' - 7 rn each flo:a| sympodium; conspicuous, long-persr::= peduncle relatively long. Spathe oblongo-ovate, boat-shaped, inflating- of sheath membranou:-: ; gaping at female anthesis, thence opening and remaining erect or the extreme margl ::--=:. reflexing dudng male anthesis, gteenish vrhite to yellowish, rz;ely to oblong lanceolate. and much ''i: internally purple-stained, or bright green, rostrate apt'cdly 8-34x3-i-lcrr.. -.., overtopping the spadix, caducous after anthesis, rately marcescent. compared with the lel=: cylindric. Flowers bisexual, perigoniate; perigone Spadix stipitate, aderrallv dull. mame o--.-- , membranaceous, cup-like, tepals truncate, equalling or shottet than '-,11 '':r-erlile plants. base c-:: gynoecium; stamens 4,free, filaments relatively short, broad, spathulate, :::'r^-,-:lre shoots,. aper ;f sometimes extending during male anthesis to taise the anthers above - - :-.:- - .::*: onman- .=:=: thecae lineat-elliptic, dehiscing -: jer the gynoecium; connective slender, :t-.t-.i .:::i ie ar-e s. - sLit. Pollen lrr pert:ur^te, subspheroidal, small (mean - -, r,-='- by longrtudinal =br:'-- . -;1ia:::::13:-,- -: 22 pm.), exine psilate or subretiptlate, ptJae spinulose tipped and - f=l -:-' -l-l=l:lilj;l united into group s 2 4 ot more. Glnoecium wtrh t: solitary, or of - rhe :::i-:-r ": --; -: l- - olrary obconic or obpyramidal, subquadrangrtlar, 1-iocular, ovule 1, fttr'c "i:--. :i I 1-=1 :l-.-r'i-: -:: anatropous, funicle short, placenta basal, stylar region as broad as - toi;-:.a: j. : l-::- -: ov^ry, stigma transversely oblong. Fruits a beny, distincdy truncate subte lc.: -- -: : :: ripening from gteen through medium yellow to apicalTy, subglobose, each hi- - t--: --:1---: : -t t,t- orange red. Seed rounded, subglobose, testa smooth, glossy, - scarlet of x 2 cm: r,i.1.':..: .:- -r . - . embryo Iarge, endospetm absent. oldest irt -:- :: : . : iu,. = -: Anad,end,rum calci'cola Boyce & Wong, sp. noa, 'Lb allt' and the u'hoie .,. :::: - r: : : -:

THE SARA\rAK MUSEUM JOURNAI TTNIIOOf NInSSNW >IVINVWS AHI Tv',\UlC

{roR"rurquof, E dq pe)uldsp ueq} uoryo sef,uorseJoguds aloqn eqt pue {p qV 'aou '{s '8uotrtr 'srsaqluu q8norqr passzd s"q tsa.&\ou eq] eroJeq tmrJ 3lruo n( ur rsaplo

egr qryn'parnp ord dgequanbe s'rag1s3o1 ol dn s at u ar a og[u t irttt Z I "t 7 7 7 'dsso13'qloours u1sa1'asoq< or ldqdord alnlorruor draqlzel e dq papurlqns of,uef,selogur r{r?e or noilad ruilpaur q8nolp 'sldqdzlzc puu sldqdord 8uo1qo-ateloeruvl g ot dn dq pepuelqns JwrunJ] dprrrysp 'f,tnq e" oFpoiu qrua 'acuacsatoguds prouo8toS rcFpouqq e w reqlaSol 1 s" pzoJq sz uor8ar lz1-us ' vt sanarang[u1'EvfinvIaq] u?qt re>lJup dlpe>1wru uogo puz'runlncqar '1 ap-r.o'repro11'wp8ue esooi snonrrdsuoc e SrmrrroJ srfial reprc-Z iogg vt ]e qu-pruJ eql 'etortt WW t//ft?aou[2 to 1 ruory Sursus A.II ry AT.I qroq luaqr qtn\ pasredsrolur direln8ar ; pue paddn esopwds er;rc puz sE ]ueurLrrotd sz er (laD sura,r dwunrdrelw if,11exeqv ATCI ;pq ueaur) Iprus'pproraqdsqn pesr"J dlsnonridsuoc 'd;pxzpz passardrur 'salzel r3ll€ws uo J3ra.e; Surcsrqap'rpdrfia-reaull aE r.If,nur 1nq 'aprs rad o] dn suiel prrlul drzrurpd f8uol urur 11 C\T.D 7 - e^oqs sJerpu? aql ssruJ 01 s g Ef, oJf,nur wpqnt snoncrdsuol z qtpN elnfz xade '(slooqs eprarr.nfur 'elepqlzds'pzorq'lroqs r1a dpo ranq eqr) arzpror dpluandrel o] atueunr as?q's1uqd elnre.tn(uo u"qt ralJoqs ro Surgznba 'r dlrqnrnred 'dar3-snornzp dgepzqv '.vaat? elrlo a]]pur '1np dlzxepe auo7uad lalsruo8uad'Trns zurruzl iwauq ]sourle sJs?f, oureltxe uI 'ql8ua1 ar{} q}la paredruoc ']uef saoJzur f,1smt'srsaqtu lo.orruu drarr esrrrrq ar{} qtrlyr ueuo ssprarrnf 'w el - e x ne - g qf,ntu puz dlerrdu Jl?lsor wTrnzI qrrzn sruqd Suuenog 'anb4qo dpq8qs 'alzloaruzl 8uo1qo or f,1a:r-l'qsralolJad ot alrq$ J]zlooJuzl dlarorrzu f,iatt ouuual fluacsecrzur ur3teu eureJlxa erp Jo lleJa Suruwurar puz 3t 'draqleal ]sLIA\auJos ra ddn'snour8ulqr"f, - snouu qtu ar{s o rrzd rqweru J -Sunu gur 'padeqs-lzoq 'aru. lrud ]sernol 'epIA w) Z'l or dn s8ur.r, 'tualsrsred-8uo1 'snonlrdsuol iurnrpodwds prog r{lua rn. 'snw,qnd aqr Jo essq oq] ol 8tm{cuer qpaqs "toJo4ad fBvol {uf, Z - L v) raqSrq 'urarr pur8ruru oru saum,Jn{ Jan{u 'f,11exde pue dluseq areur,qnd '3uo1 tur tE - g apgad iarnua'3uo1qo-ole'ro dlanbr :]ustslp ruf, g E v) ol pererl?f,s stooqs Sur.ranrog vo saflorl 'uaal8 - 'serrads ]soru ur snur,Lpd 1 eizd auzru ''zlp ruru t e) ot slooqs 8.{qqp fslootls 8ur.ra.r.og aruur,rlnd apga{ islooqs 3r 'seJleru uo 'uTp ruf, I et 'oletef srualS g ot aqfqQ{a1ruaq 3a7qu72 'slooqs 8,r1qql, uo elrsoc 'II a]elcl'(olo,{\tVO 7gtZ-EV 7il^ 'Q-ee) n 'rd u1g 7u0711 arrflg ')'cI '6002 kW '1sz ru gl ',,2'g0.g0 :,,g'lg ry e o0l L ry rsuSog'ode1q :0"Sgt) St ,VZ ol0 'ur8uy zng 'neg €rm4rn;4'lenereg'ersdqu;4 :snd,i1- 'rnrmSunsrp srJesxa qlIA uoBulfossu w '-i1a rr=r -uou srurursls ]e suelsrsred 3uo1 uulSerl qopsd 'lrpalle (zrarzumg) ala.n 'snultlnd pcrde ar4r slu$o ru%paallpav snrsodsrp s{npounq snprouo8ro8 ;ragyp eq] qlla pue '(drm4s +q& srrluoJseroguds u1 sarnld sBuof,serogur ta 'srteauntr ralrpszq sDlnpz (unrpeur dsso18 dlzxzpu a. sIuIruEI 'sicne18 sntqns sntoJ uon?urquof snqrsuaauJog rupaapoav Jerpo [" o] lseJ]uor SurTu

LqZ cNa NIS CNO/A. ONV Sl Og I UAffId I OEQIOS IO f PFI:I STUDIES ON MONSTERE,AE (ARACEAE) OF BORNEO I - module of the primary arjs; pedwncle extending during development, up to 17 cmlong, ca 3 mm wide duting anthesis, lztetally somewhat .o-pre.sed, p"1. green, occasionally slighdy glaucous. Spadix stipiiate; stipe to 1.5 cm long, 3 mm wide, very slighdy longitudinally aigled;fertileportion of spadix cylindrical, obliquely insetted on stipe, Z Z.S x ca 0.7 cm, pale yellow at anthesis, becoming darker yellow, - at and thence gfeen as fruit begin to develop . spathe inflating-gaping female anthlsis, 8 X 1 cm, pale green, then at onset of male anthesis X swiftly opening, lanceolate, and conspicuously beaked, 10 3 cm, pale yellowish white, the interiof \r/axy, exterior pale glaucous gfeen' iprth. late in anthesis reflexing and blackening before citcumscissile ,i spathe/spadix inserrion, and fall,ing;Flowers 4 x 6 mm;perigone mwch shorter than gynoecium, translucent very pale green; gynoecium pale cfeamy yellov/, stigma ca 1 mm, lineat, transverse' gynoecium latet yellow, later still gfeen, with stigm^thc fem Ifis blackening; -.dio* anthers stamens with filaments fiot extending during male anthesis, dehiscing while still concealed between adiacent flowers' Infrurtesnnce up to 10 together, spreading to efect; spadix dark green;fruits flpening datk green through yellow to scadet. Distribution: To date endemic to Satawak, but there widespread and with collections from very close to the Brunei and Kalim^nt^frborder such that it is almos t certarnTy distributed thtoughout N' Borneo as fat north as Brunei. Hzbitat: climbing hemiepiphyte in pethumid dense fotest, often on, but not restricted to, limestone, 35 - 450 m alt' Notes: A highly distinctive species bv r.irtue of the large (fot the numefous genus) leu.res with the laminae abaxialll'glaucous, and the inflorescences produced in a strongly bimodulat gotgonoid similat in overall to synfloresce nce. Anadercdrum ca/cico/ais ^ppeaf^nce A,ffineSchott(Sumatera)butteadilydistinguishedbythepersistent p.tiol^, sheath, and the anthers not exseftttlg at anthesis. Juvenile plrrr,. are remarkable by the long, naffoq occasionally almost l-ineaf, lezf lamina.

THE, SARA\(/AK MUSEUM JO U RNAL TSN}INOf NIrISSOW >IV/(VuVS SHI THN]LIOI

g g i,,g' gV, : ',,9' l,y oZI I 9 g o I 0 1u q€qas rs?aqeU veurvl lldz;E (UVS) gg{-EV laryJ 7a uoQ[ ry losD[ ryp paplaf 'afig 7'cI 'V002 l*WZt' ,,g'll 'rvevrl lsourle dgzuotszrro '-u JE oeTI i,,L'I1 ,ZV ol0'vpvr1vg dwv3 o] pzor 99 ur{ 'uorssecuo3 epra.tnf 'srsaqlu? tz Sunras poolA. 8uz{ag 1ee1 z8ue51 'r1dz;4 :uorsrlr6l rldey '(UVS) lg5 t-VV '1o 'J luatrsrsrad aqr dq peqsm8uns 7a atfog cI'9002 ra6[ I '.,9'gg ,62 olLl',,,1'g0.gg ol0 '1a>1peg n161 ol eruerzedde gz;ea.o uI JuLrL le>llres :(UVS) 09t t-EV 7o 7a affog ')'cI '9002 roCI 9 ,,I'69 ,gg ol I I prouo8ro8 w1npourq {3 i,,g'1e ,fg ol0 'loszrag'ue1z4:uorsrarcl Ie>llrzs'(UVS) l{/t-EV 70 snoJaurnu eql puz 'sno:n213 7a ufog')d'9002 q]r"][ gl ',,8'gt ,09 o0II i,,Z'gZ.Lg o00 'rnqzlag aqr ro3) a3re1 aqr antrr.\ Sunrrng '.{olrg Sundwr;1 'so18uory 'uepeg :(W$ gGZ t-EV r/rod Jo ((t/t/X ryo aztals T afflg')'cI'9002 [pfgZ'qvhlw8rmg 'nq1'rm4rr4 'rlz ru t 'ra6[ 'uzpeg :(UVS) ggg-Ev %od ryo tlnltx antllxs ry atftg 27t't002 g 'uo uago 1saro; asuep Prum 'oqtuog p8ung'rmtrrr4'uuuag :(UVS) 6r/-HVltroclryo qnlDI uoil/ls 2g, ufrg 27I'y002 gl '3uwa;4 un8ng 'urnpzpe; Sunung 'rm4rl4 ^oN su oaurog '51 rnoq8norqr p; 'uzuag :(UVS) g/t-EV lpsDl VCI puala[ ry uftg 7'cI 't002 tZ ^oN JsPto q veluvTJIw> rerlrl ',,2'lZ ,Lt, o0ll:,,2'90.60 ol0 'ue8undury Sunung 'uzuag :uorsrlrq Puz pue pza.rdsaplin ararll tnq T ueryrvurvs'(UVS) 991Z-EV lpsry 7p puoJa[ ry anfig 77I '9002 qeJ L',1'lLgI o0ll:,,2'll .6I oI0 'tuerg unfraa rry'urzrg Sundruq'1 'uurnqrg '3u1qcn;,1 :(WS) g I/ t-EV lpsn rya puaJaf ry ufrg 2'd '9002 Srmredu srm{ iuaet? lrsp "y r{f,rel{ L 'WqnI p8ung 'o3org Sundruz;4 'nzg :(gyg) Z6g t-VVFaq auarapru{u1'sJeaog tuerelpr u1g7uo41ry lasDlrya puoJaf tufi7 7'cI'9002 'qed I ',,t'lL ,gI o1tL:,,2'll sreqlu? 'stsaqluz apu.r Suur ,61 ol0 lwer"g un{ra1 rry'ruel.g Sunduley'uurnqrg'3rmtrrn;4 :(WS) iSurualcqq suluureJ cPeru8r .rc1 9G, fVV 7o 7a atfog ')'d '9002 gZ ' ,,2' L0 .90 o0l I i,,L'gV ,gZ ol0 Jalzl uJruJJoudB'asralsu"rl '"t\n[ Sunrmg 'neg :(gyg) 0g{ t-VV 'Ja 7a affog ')'cI '9002'ldag 9t elvd runuaou[F iuaat? epd i ',,g'Bl .80 o0l I i,,g'ZZ ,Zt ol0'srpy re8uns 'lros 'Suo8ag Stmdruz;'1 qtnut auoZua { iuxn ) x y u a,n 'nzg :(gyg) gZ{ I-EV 'Jo 7a atfog ')d '9002 E V 0l ',,8'89 .80 o0l I a1rssrf sulnf,nf 3roJa q Srmrar 1,,6'00 ,Zt, oL1 'Suo8ag Sundurq4'neg i(gyg) S tOt-UVlaryIry? ,//0q[ 'uaar8 snocnulS apd rouarr (P3>lzaq T lpslxrya puoJaf btfrg 2'a'g00zri]re]{ ? ',,6'tg,gg o0l l:,,g'ZZ.g l oI0 '.ur g x 0l ,{1snon '8uog w8rmg'sn83uua Sundwe;'1 '3uo1or;1 :(UVS) tgd-Evloslx Te slserpuB aFLu Jo lssuo 1z u: 'd< pual af 4, .raaax X.'rfr g 27I' g00Z uef ZI',,L' 81, g0 o0 I I :,,1' IE,IZ oL1 re 8urdz8-8qq"Hul aqudS 'p8nqya Sunung '3z3og Sundrue;'1 'nug :(gyg) ,tg-EV t/oaraq!)'I,f 'zno11ad Je1wp Stmuoreq'srs ry afflg')'cI'i00Z 'ro6[ 8'uesnquref 'nzg:(gyg) 1ZFVVlos[ryo uoQ[ 'adns uo polJrsur dlenbqqo o ry ut6g 7'cI'V002 eUn[ Z wSung n1n 'Suo8ag Sundwe) 'nzg .(Tluurpnrr8uol dpq8qs i:ar "lpv iunuzqrall luauruzdaq dnsarod >lE.,lNErsS sl UVS (UVS) t I t-VV losy xlpnds'snorne18 dpq81s ryo paoJa[ ry nfog ')'rI '1002 11rdy 7 'rqvraoteg Sundruq'1 'uzsnquel t?ql\etuos f,yenw1'sts aqtw 'ozg :uorsrlr6l Surqrn;4 '1e.,rnerug ueos suaurrceds req]O ']ueurdolalep Surmp Surpu

682 cNg NISCNOIN CINVIII Og'lugrild IOANUO8IO T\ STUDIES ON MONSTEREAE (ARACEAE) OF BORNEO I PETER C. I

13 Dec. 2004, P.C. Boyq Jeland ak Kisai & M. Gibernau AR-872 (SAR); particulatly on juvenile p1a Belaga,BeIaga road, SungarUnan, 02" 55' 40.0"; 1.13" 44' 1.5.4", 12 but half this number on s- Oct. 2005, P.C. Bo1a, Jeland ak Kisai & Jipom ak Tisai AR-|469 (SAF.). rais ed abaxtd)y ; interprima Binnrlu Division: , Bukit Sarang, Grand Perfect field station, * regularly interspersed xir path to Gua Lubang Batu Rusa L-45, 1,2 Jdy 2006, Skornickoua / 14 mid-db at ca 60";2-order sab. AR-20l5 (SAR). Mid Division: Niah Suai, Niah National Park, Inforescences ca 3 together Madu Tlat7,03o 48' 57.9";1,1,3" 46'18.3",13July 2006,P.C. Bolce et pnman shoots; fitst nvo a/. AR-/ 882 (SAR); Marudi, Long Lamz, Mulu N.P., Trail to Gunung termnal inflorescence suL . >: rl.[]-1 Muiu Summit, 04" 02' L8.7";1,1,4o 49' 44.2",29 Sept. 2007 , P.C. Bolce :,t, cmiinflorescei rur-enile et a/. AR-|957 (SAP*); Marudi, Long Lama, Mulu N.P., Nightwalk l:: fruit before the Tnr7,04" 02' 42.2"; 114" 48' 55.5", 3 Oct. 2007, P.C. Boye et a/. AR- s.,-ii.orescence alv'ar-s Cs; 1999 (SAP.). Division: Nanga Medamit, Mulu N.P., Sungai ::rmar\- axts; pedunc/t ex--e Empangau, tributary from Sungai Mendalam,04o 13' 41.6";1,1,4" 52' -rlg. ca 2,5 mm lride :i 50.5", 30 Sept. 2007, P.C. Bojtce et a/. AR-224, (SAR). ::edium on Anad,end.rum mulaensis Boyce & Wong, sp. noa. Anandrum 'nd. muluensis differt ab speciebus ceteris laminis foliorum adaxrzY glaucis, basalis cordatis, caulis florenti foliis non distichus congestis et staminis non-exserta. Anandrwn cordatum accedit sed ab eo foliis subtus glaucis atque in synflorescentiis flotenti ad apicem surculorum non-specialis differens. Typus: , , , Marudi, Long Lama, Mulu N.P., -Trail to Deer Cave,}4" 02' 23.8";114o 48' 54.6",60 m asl, 27 Sept.2007, P.C. Boye, et al. AR-|929 (SAR, holo). Plate III.

Climbing heniepipfute to two metres but usually much less. Stems weakly compressed, ca 5 mm flowering shoots, climbing shoots to c - 3 mm dia. glaucous medium green; I-naues scattered to ca 3 - 5 cm distant; petiok 2 - 9 cm long, canaltculate, pulvinate aprcally and basally, apical puluinus ca 1..5 cm long, strongiy D-shaped in cross section, the angles sharply defined; petiolar sheath extending to the base of the apical pulvinus, conspicuous, pale green, turning yellow, then soon-marcescent, wings up to 3 mmwide.; /aninabtoadly oblongo lanceolate to broadly cordate, 3 - 12 x 3.5 - 6 cm, slighdy oblique, base sub-truncate (and then very slighdy cordate at petiole D-=-- - -: - :. :::,'- . '.- .i. insertion) to cordate, apex acute with a conspicuous tubular mucro ca Fl:-:::.:: :: - ::' 1 mm long; adaxially dull, matte olive green; abaxtally glaucous-grey, ---:: - 1.^---^^ - L-ii]!:--1-f- -,-:-

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Notes: Anadendrum muluensis is unusual in the genus in floweting on undifferentiated shoots (as compared with most species where flowering occurs on shoots with fans comprised of congested- distichous leaves). The leaf lamtnae cordate basaliy and giaucous abaxjally, and stameris non-exserted during anthesis readily differentiates A. mu/uensis from all other Sarawak species. In leaf form, and in carqnng inflorescences on shoots with scattered leaves, A. mu/uensis approaches A. cordatum Schott (Sumatera), and A. m o n ta n u m var. c o rd atu m Ridl. @eninsular Malaysia), from both it differs by the leaves glaucous abanally and the anthers not exserting during anthesis.

ACKNOWILEDGEMENTS The collaboration and support of the Sarawak Foresfty Department, the Forest Research Centre (I(uching), The Semenggoh Botanical Research Cente, and the Sarawak Biodiversity Centre are much appreciated. This study was carded out under Sarawak Forestry Department Research Permit Nos NPSfl907.4.2Q)-101 & NCCD.907.4Q{t-41,, and Park Permit Nos 58/2007 & 37 /2009.

REFERENCES

Boyce, P.C. In Anadendrum (Araceae: Monsteroideae: Anadendreae) in press Thailand. Thai Forest Bulktin (Botary)

Haigh, A., Bogner,J., Boyce, P.C., Gral-um, M.H., Kostelac, C., Croat, T.B., Hay, A., Hetterscheid, $flL.A., Keating, R., Lay, L., Mayo, SJ., Mora, M., Plare II: ^ln&rcdlrrn uXt --:-- Reynolds, L., Sellaro, M., & \fong, S.Y :--. ;<-^ :3:\'; : ij:C: - : ;:.:iu:-.):C S-::: 2008 A new website fot Araceae taxonomy on www.cate-^r^ce2e. =: - -i'-jj iii!::r-Er t-:or€-.cctces i,ar otg. Aroideana 31: 148 - 154. =- -::<: :: -t',e .l:-iresi-. b'€it- E. T-i,,1";5ci:ci.ar ler:";-e a: !--- L.---^--:i -'-= ;:'i:------i:- \::= =i-i;:;j-::-.- =:: s:ari:x .e.i ::'r e i::le.i-.- lAe fre rndir-i*:; i:u-*s- H- 1-:r ,eares. A-G: --J,--'+;ii- H: -{

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Plate III: Anadendrum muluensds Boyce & Wong. A. Plant in habitat. B. Flowering shoot. Note that there is no congested {an of leaves. C. Synflorescence, ventral view: D. Inflotescence at male anthesis. E. Leaf lamina, abaial view to show glaucous surface. A-E: AR-/ 929. Images @Peter C. Boyce.

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