Hawaiian Islands Animals: Updated November 29, 1999 Listed and Candidate Species , As Designated Under the U.S

HAWAIIAN ISLANDS ANIMALS: UPDATED NOVEMBER 29, 1999 LISTED AND CANDIDATE SPECIES , AS DESIGNATED UNDER THE U.S. ENDANGERED SPECIES ACT

Species status by island: E=endangered; T=threatened; P=formally proposed as E or T; C=candidate for listing; R=recommended for DISTRIBUTION listing (L) as E or T, or for candidacy (C); SOC=species of concern; *=possibly extinct; #=extremely rare; (CH)=critical Habitat STATUS N.W. Islands‡, designated. (listing Hawai'i Maui L~na'i Moloka'i O'ahu Kaua'i Kaho'olawe, ‡=N.W. Hawaiian Islands: Fr. Frigate,Kure, Laysan, Midway, Necker, Nihoa, Pearl & Hermes priority) Ni'ihau, or Oceanic

VERTEBRATES (36 Endangered + 4 Threatened + 2 PE + 2 C = 44 taxa + 5 SOC)

LISTED MAMMALS (4 Endangered taxa)

Lasiurus cinereus semotus Bat, Hawaiian hoary; 'Æpe'ape'a E T T T T

Megaptera novaeangliae Whale, humpback; Kohol~ E O Monachus schauinslandi (CH) Hawaiian monk seal; '¦lio-holo-i-ka-uaua E T T T T T T L, M, Ni, N, F Physeter macrocephalus Whale, sperm; Palaoa (uncommon) E O

LISTED BIRDS (30 Endangered + 1 Threatened = 31 taxa) Acrocephalus familiaris kingi Millerbird, Nihoa E N

Anas laysanensis Duck, Laysan E L Anas wyvilliana Duck, Hawaiian; Koloa maoli E T T* T T Buteo solitarius Hawk, Hawaiian; 'Io E T

Corvus hawaiiensis Crow, Hawaiian; 'Alal~ E T# Fulica alai Coot, Hawaiian; 'Alae ke'oke'o E T T T T T T Gallinula chloropus sandvicensis Moorhen, Common; Hawiian gallinule; 'Alae 'ula E T* T* T* T T

Hemignathus lucidus affinus Nuku pu'u, Maui E T* Hemignathus lucidus hanapepe Nuku pu'u, Kaua'i E T* Hemignathus munroi 'Akia pÇl~'au E T

Hemignathus procerus 'Akia loa, Kaua'i E T* Himantopus mexicanus knudseni Stilt, Black-necked; Hawaiian stilt; Ae'o E T T T T T T Ni Loxioides bailleui (CH) Palila E T

Loxops coccineus coccineus '}kepa, Hawai'i; 'Akakane E T Loxops coccineus ochraceus '}kepa, Maui; 'Akepeu'ie E T* Melamprosops phaeosoma Po'ouli E T#

Moho braccatus 'Æ'Ç, Kaua'i; 'Æ'Ç '~'~ E T* Myadestes lanaiensis rutha Thrush, Moloka'i; Moloka'i Oloma'o E T* Myadestes myadestinus Thrush, Large Kaua'i; K~ma'o E T*

Myadestes palmeri Solitare, Small Kaua'i; Puaiohi E T# Nesochen sandvicensis Goose, Hawaiian; N‘n‘ E T T T Oreomystis mana Creeper, Hawai'i; Hawai'i 'Alauahio E T

Palmeria dolei Honeycreeper, Crested; '}kohekohe E T Paroreomyza flammea Creeper, Moloka'i; K~k~wahie E T* Paroreomyza maculata Creeper, O'ahu; O'ahu 'Alauahio E T#

Pseudonestor xanthophrys Parrotbill, Maui E T Psittirostra psittacea 'Æ'ã E T* T* T* T* T* T*

Page 1 HAWAIIAN ISLANDS ANIMALS: UPDATED NOVEMBER 29, 1999 LISTED AND CANDIDATE SPECIES , AS DESIGNATED UNDER THE U.S. ENDANGERED SPECIES ACT

Pterodroma phaeopygia sandwichensis Petrel, Dark-rumped; Hawaiian petrel; 'Ua'u E T T T T Puffinus auricularis Shearwater, Newell's T T T T Telespiza cantans Finch, Laysan E L

Telespiza ultima Finch, Nihoa E N

PROPOSED AND CANDIDATE BIRDS (2 PE + 2 C = 4 taxa)

Chasiempis sandwichensis ibidis 'Elepaio, O'ahu PE(3) T Phoebastria albatrus Albatross, short-tailed PE(3) M, L Oceanodroma castro Petrel, band-rumped storm C(3) T T T Oreomystis bairdi Creeper, Kaua'i; `Akikiki C(5) T

SPECIES OF CONCERN: BIRDS (4 taxa) Asio flammeus sandwichensis Owl, Hawaiian short-eared; Pueo SOC T T T T T T Loxops caeruleirostris '}kepa, Kaua'i; `Akeakea SOC T Moho bishopi 'Æ'Ç, Bishop's SOC ?; T* T* Numenius tahitiensis Curlew, Bristle-thighed SOC T T M, L

LISTED REPTILES (2 Endangered + 3 Threatened = 5 taxa) Caretta caretta Turtle, Loggerhead sea; (incidental in Hawai'i) T T T T T T T O+all islands Chelonia mydas Turtle, Green sea; Honu T T T T T T T O+all islands Dermochelys coriaceae Turtle, Leatherback sea; (incidental in Hawai'i) E T T T T T T O+all islands Eretmochelys imbricata Turtle, Hawksbill; 'Ea E T T T T T T O+all islands Lepidochelys olivacea Turtle, Olive ridley sea; (incidental in Hawai'i) T T T T T T T O+all islands

SPECIES OF CONCERN: FISHES (1 taxon) Lentipes concolor O'opu alamo'o SOC T T T T T

INVERTEBRATES (41 Endangered + 4 Proposed + 27 C + 875 SOC = 947 taxa; 37 RL) LISTED SNAILS (41 Endangered taxa) Achatinella abbreviata Snail, O'ahu tree; Pãpã kani oe E T* Achatinella apexfulva Snail, O'ahu tree; Pãpã kani oe E T# Achatinella bellula Snail, O'ahu tree; Pãpã kani oe E T# Achatinella buddii Snail, O'ahu tree; Pãpã kani oe E T* Achatinella bulimoides Snail, O'ahu tree; Pãpã kani oe E T# Achatinella byronii Snail, O'ahu tree; Pupu kani oe E T#

Page 2 HAWAIIAN ISLANDS ANIMALS: UPDATED NOVEMBER 29, 1999 LISTED AND CANDIDATE SPECIES , AS DESIGNATED UNDER THE U.S. ENDANGERED SPECIES ACT

Achatinella caesia Snail, O'ahu tree; Pupu kani oe E T* Achatinella casta Snail, O'ahu tree; Pupu kani oe E T* Achatinella cestus Snail, O'ahu tree; Pupu kani oe E T# Achatinella concavospira Snail, O'ahu tree; Pupu kani oe E T# Achatinella curta Snail, O'ahu tree; Pupu kani oe E T# Achatinella decipiens Snail, O'ahu tree; Pupu kani oe E T# Achatinella decora Snail, O'ahu tree; Pupu kani oe E T* Achatinella dimorpha Snail, O'ahu tree; Pupu kani oe E T# Achatinella elegans Snail, O'ahu tree; Pupu kani oe E T* Achatinella fulgens Snail, O'ahu tree; Pupu kani oe E T# Achatinella fuscobasis Snail, O'ahu tree; Pupu kani oe E T# Achatinella juddii Snail, O'ahu tree; Pupu kani oe E T* Achatinella juncea Snail, O'ahu tree; Pupu kani oe E T* Achatinella lehuiensis Snail, O'ahu tree; Pupu kani oe E T* Achatinella leucorrhaphe Snail, O'ahu tree; Pupu kani oe E T# Achatinella lila Snail, O'ahu tree; Pupu kani oe E T# Achatinella livida Snail, O'ahu tree; Pupu kani oe E T# Achatinella lorata Snail, O'ahu tree; Pupu kani oe E T# Achatinella mustelina Snail, O'ahu tree; Pupu kani oe E T Achatinella papyracea Snail, O'ahu tree; Pupu kani oe E T* Achatinella phaeozona Snail, O'ahu tree; Pupu kani oe E T# Achatinella pulcherrima Snail, O'ahu tree; Pupu kani oe E T* Achatinella pupukanioe Snail, O'ahu tree; Pupu kani oe E T# Achatinella rosea Snail, O'ahu tree; Pupu kani oe E T* Achatinella sowerbyana Snail, O'ahu tree; Pupu kani oe E T Achatinella spaldingi Snail, O'ahu tree; Pupu kani oe E T* Achatinella stewartii Snail, O'ahu tree; Pupu kani oe E T* Achatinella swiftii Snail, O'ahu tree; Pupu kani oe E T# Achatinella taeniolata Snail, O'ahu tree; Pupu kani oe E T# Achatinella thaanumi Snail, O'ahu tree; Pupu kani oe E T* Achatinella turgida Snail, O'ahu tree; Pupu kani oe E T# Achatinella valida Snail, O'ahu tree; Pupu kani oe E T* Achatinella viridans Snail, O'ahu tree; Pãpã kani oe E T# Achatinella vittata Snail, O'ahu tree; Pãpã kani oe E T* Achatinella vulpina Snail, O'ahu tree; Pãpã kani oe E T#

Page 3 HAWAIIAN ISLANDS ANIMALS: UPDATED NOVEMBER 29, 1999 LISTED AND CANDIDATE SPECIES , AS DESIGNATED UNDER THE U.S. ENDANGERED SPECIES ACT

PROPOSED SNAILS (1 taxon; 1 PT) Erinna newcombi Snail, Newcomb's; Pupu wai lani PT(7) T

CANDIDATE SNAILS (3 taxa) Newcombia cumingi Snail, Newcomb's tree C(5) T Partulina semicarinata Snail, Lana’i tree; Pãpã kani oe C(2) T Partulina variabilis Snail, Lana’i tree; Pãpã kani oe C(2) T

SPECIES OF CONCERN: SNAILS (595 taxa) Amastra - 149 Hawaiian species Snail (No common name) SOC T T T T T T Armsia petasus Snail (No common name) SOC T Carelia - 21 Hawaiian species Snail (No common name) SOC T Ni Catinella - 6 Hawaiian species Snail (No common name) SOC T T T T T Clithon - 2 Hawaiian species Snail (No common name); pipiwai SOC T T T T T T Columella - 3 Hawaiian species Snail (No common name) SOC T T Cookeconcha - 17 Hawaiian species Snail (No common name) SOC T T T T T T Endodonta - 11 Hawaiian species Snail (No common name) SOC T T T T T Euconulus - 6 Hawaiian species Snail (No common name) SOC T T T T Georissa - 2 Hawaiian species Snail (No common name) SOC T Godwinia - 3 Hawaiian species Snail (No common name) SOC T Hiona - 13 hawaiian species Snail (No common name) SOC T T T T T T Kaala subrutila Snail (No common name) SOC T Laminella sanguinea Snail (No common name) SOC T Laminella - 14 Hawaiian species Snail (No common name) SOC T T T T Laxisuccinea - 2 Hawaiian species Snail (No common name) SOC T Leptachatina - 129 Hawaiian species Snail (No common name) SOC T T T T T T K Lymnaea aulacospira Snail, Hawaiian freshwater (No common name) SOC T T T T Lymnaea producta Snail, Hawaiian freshwater (No common name) SOC T T Ni Lymnaea rubella Snail, Hawaiian freshwater (No common name) SOC T T T T T Ni Lyropupa - 22 Hawaiian species Snail (No common name) SOC T T T T T T K Neritilia hawaiiensis Snail, Anchialine pool (No common name) SOC T T T T T Neritina vespertina Snail, Hawaiian stream; hihiwai SOC T T T T T T Neritina granosa Snail, Hawaiian stream, hapawai SOC T T T T T T Nesophila - 4 Kauai species Snail (No common name) SOC T

Page 4 HAWAIIAN ISLANDS ANIMALS: UPDATED NOVEMBER 29, 1999 LISTED AND CANDIDATE SPECIES , AS DESIGNATED UNDER THE U.S. ENDANGERED SPECIES ACT

Nesopupa - 20 Hawaiian species Snail (No common name) SOC T T T T T T K Nesovitrea - 3 Hawaiian species Snail (No common name) SOC T T T T Newcombia canaliculata Snail, Newcomb's tree SOC T Newcombia lirata Snail, Newcomb's tree SOC T Newcombia perkinsi Snail, Newcomb's tree SOC T Newcombia pfeifferi Snail, Newcomb's tree SOC T Newcombia sulcata Snail, Newcomb's tree SOC T Neritilia hawaiiensis Snail, anchialine pond SOC T T T T T Orobophana - 6 Hawaiian species Snail, No common name SOC T T Panamastra - 3 Hawaiian species Snail (No common name) SOC T Partulina anceyana Snail, Maui tree; Pãpã kani oe SOC T Partulina carnicolor Snail, Maui tree; Pãpã kani oe SOC T Partulina confusa Snail, Hawai'i tree; Pãpã Kani oe SOC T Partulina crassa Snail, L~na'i tree; Pãpã Kani oe SOC T* Partulina crocea Snail, Maui tree; Pãpã kani oe SOC T Partulina dolei Snail, Maui tree; Pupu kani oe SOC T Partulina dubia Snail, O'ahu partulina tree; Pupu kani oe SOC T Partulina dwightii Snail, Moloka'i tree; Pupu kani oe SOC T Partulina fusoidea Snail, Maui tree; Pupu kani oe SOC T Partulina germana Snail, Maui tree; Pupu kani oe SOC T Partulina grisea Snail, Maui tree; Pupu kani oe SOC T Partulina horneri Snail, Hawai'i tree; Pupu kani oe SOC T Partulina induta Snail, Maui tree; Pupu kani oe SOC T Partulina kaaeana Snail, Maui tree; Pupu kani oe SOC T Partulina lemmoni Snail, Maui tree; Pupu kani oe SOC T Partulina marmorata Snail, Maui tree; Pupu kani oe SOC T Partulina mighelsiana Snail, Moloka'i tree; Pupu kani oe SOC T Partulina mucida Snail, Moloka'i tree; Pupu kani oe SOC T Partulina mutabilis Snail, Maui tree; Pupu kani oe SOC T Partulina nattii Snail, Maui tree; Pupu kani oe SOC T Partulina nivea Snail, Maui tree; Pupu kani oe SOC T Partulina perdix Snail, Maui tree; Pupu kani oe SOC T Partulina physa Snail, Hawai'i tree; Pupu kani oe SOC T Partulina plumbea Snail, Maui tree; Pupu kani oe SOC T Partulina porcellana Snail, Maui tree; Pupu kani oe SOC T

Page 5 HAWAIIAN ISLANDS ANIMALS: UPDATED NOVEMBER 29, 1999 LISTED AND CANDIDATE SPECIES , AS DESIGNATED UNDER THE U.S. ENDANGERED SPECIES ACT

Partulina proxima Snail, Moloka'i tree; Pupu kani oe SOC T Partulina radiata Snail, Maui tree; Pupu kani oe SOC T Partulina redfieldi Snail, Moloka'i tree; Pupu kani oe SOC T Partulina rufa Snail, Moloka'i tree; Pupu kani oe SOC T Partulina splendida Snail, Maui tree; Pupu kani oe SOC T Partulina subpolita Snail, Moloka'i tree; Pupu kani oe SOC T Partulina talpina (gouldi) Snail, Maui tree; Pupu kani oe SOC T Partulina tappaniana Snail, Maui tree; Pupu kani oe SOC T Partulina terebra Snail, Maui tree; Pupu kani oe SOC T Partulina tessellata Snail, Moloka'i tree; Pupu kani oe SOC T Partulina thaanumiana Snail, Maui tree; Pupu kani oe SOC T Partulina theodorei Snail, Moloka'i tree; Pupu kani oe SOC T Partulina ustulata Snail, Maui tree; Pãpã kani oe SOC T Partulina virgulata Snail, Moloka'i tree; Pãpã kani oe SOC T Partulina winniei Snail, Maui tree; Pãpã kani oe SOC T Pauahia - 3 hawaiian species Snail (No common nam3) SOC T Perdicella carinella Snail (No common name) SOC T Perdicella fulgurans Snail (No common name) SOC T Perdicella helena Snail (No common name) SOC T Perdicella kuhnsi Snail (No common name) SOC T Perdicella mauiensis (maniensis) Snail (No common name) SOC T Perdicella ornata Snail (No common name) SOC T Perdicella thwingi Snail (No common name) SOC T Perdicella zebra Snail (No common name) SOC T Perdicella zebrina Snail (No common name) SOC T Philonesia - 40 Hawaiian species Snail (No common name) SOC T T T T T T K Pleuropoma - 8 Hawaiian species Snail (No common name) SOC T T T T T T Nii, K Striatura - 3 Hawaiian species Snail (No common name) SOC T T T T T T Succinea - 34 Hawaiian species Snail (No common name) SOC T T T T T T Tropidoptera - 5 Hawaiina species Snail (No common name) SOC T T Vitrina tenella Snail (No common name) SOC T T T

LISTED ARTHROPODS (0 taxa)

PROPOSED ARTHROPODS (3 taxa; 3 PE)

Page 6 HAWAIIAN ISLANDS ANIMALS: UPDATED NOVEMBER 29, 1999 LISTED AND CANDIDATE SPECIES , AS DESIGNATED UNDER THE U.S. ENDANGERED SPECIES ACT

Adelocosa anops Wolf spider, Kaua'i cave PE T# Manduca blackburni Moth, Blackburn's sphinx PE T T# T* T* T* T* K Spelaeorchestia koloana Amphipod, Kaua'i cave PE T#

CANDIDIATE ARTHROPODS (30 taxa; 18 RL) DRAGONFLIES AND DAMSELFLIES (Order Odonata; 6 taxa; 6 RL) Megalagrion leptodemus Damselfly, Leptodemas megalagrion C(2) T Megalagrion nesiotes Damselfly, Nesiotes megalagrion C(2) T T Megalagrion nigrohamatum nigrolineatum Damselfly, blackline megalagrion C(3) T T Megalagrion oceanicum Damselfly, oceanic megalagrion C(2) T Megalagrion pacificum Damselfly, Pacific megalagrion C(2) T T T T T T Megalagrion xanthomelas Damselfly, orangeblack megalagrion C(8) T T T T T T

FLIES (Order Diptera; 16 taxa; 12 RL) Drosophila aglaia Pomace fly, Oahu C(2) T Drosophila attigua Pomace fly, Kauai C(2) T Drosophila differens Pomace fly, Molokai C(2) T Drosophila digressa Pomace fly, Hawaii C(2) T Drosophila hemipeza Pomace fly, Oahu C(2) T Drosophila heteroneura Pomace fly, Hawaii C(2) T Drosophila montgomeryi Pomace fly, Oahu C(2) T Drosophila mulli Pomace fly, Hawaii C(2) T Drosophila musaphila Pomace fly, Kauai C(2) T Drosophila neoclavisetae Pomace fly, Maui C(2) T Drosophila obatai Pomace fly, Oahu C(2) T Drosophila ochrobasis Pomace fly, Hawaii C(2) T Drosophila substenoptera Pomace fly, Oahu C(2) T Drosophila tarphytrichia Pomace fly, Oahu C(2) T Drosophila toxochaeta Pomace fly, Molokai C(2) T Phaeogramma sp. Gall fly, Po'olanui C(5) T

BUTTERFLIES AND MOTHS (Order Lepidoptera; 1 taxon) Tinostoma smaragditis Moth, Fabulous green sphinx C(1) T

TRUE BUGS (Order Hemiptera; 1 taxon)

Page 7 HAWAIIAN ISLANDS ANIMALS: UPDATED NOVEMBER 29, 1999 LISTED AND CANDIDATE SPECIES , AS DESIGNATED UNDER THE U.S. ENDANGERED SPECIES ACT

Nysius wekiuicola Seed bug, Wekiu C(2) T

SPIDERS (Order Aranea; 0 taxa)

AMPHIPODS (Order Amphipoda; 0 taxa)

SHRIMP (Order Decapoda; 6 taxa) Antecaridina lauensis Shrimp, anchialine pool C(2) T T JPN,MOZ,SA Calliasmata pholidota Shrimp, anchialine pool C(2) T T FUN,SA,SIN,TUV Metabetaeus lohena Shrimp, ancialine pool C(2) T T T Palaemonella burnsi Shrimp, anchialine pool C(2) T T Procaris hawaiana Shrimp, anchialine pool C(2) T T Vetericaris chaceorum Shrimp,,anchialine pool C(2) T

SPECIES OF CONCERN: ARTHROPODS (275 taxa; 2 RL) ROCKHOPPERS AND BRISTLETAILS (Order Archeognatha; 2 taxa) Neomachilis (=Machiloides) heteropus Bristletail, Hawaiian long-palp SOC T T T T T Neomachilis (=Machiloides) perkinsi Bristletail, Perkins' club-palp SOC T

DRAGONFLIES AND DAMSELFLIES (Order Odonata; 6 taxa) Megalagrion adytum Damselfly, Adytum megalagrion SOC T Megalagrion amaurodytum peles Damselfly, Pele megalagrion SOC T* Megalagrion amaurodytum waianaeanum Damselfly, Waianae megalagrion SOC T* Megalagrion molokaiense Damselfly, Moloka'i megalagrion SOC T Megalagrion nigrohamatum Damselfly, nigrohamatum megalagrion SOC T T T T Megalagrion oahuense Damselfly, O'ahu megalagrion SOC T

GRASSHOPPERS AND ALLIES (Order Orthoptera; 7 taxa) Banza nihoa Conehead katydid, Nihoa banza SOC N Caconemobius howarthi Cave cricket, Howarth's SOC T Caconemobius schauinslandi Bush cricket, Schauinsland's SOC T Caconemobius varius Cave cricket, Kaumana SOC T Leptogryllus deceptor Bush cricket, O'ahu deceptor SOC T T Thaumatogryllus cavicola Cave cricket, volcanoes SOC T Thaumatogryllus variegatus Bush cricket, Kaua'i thinfooted SOC T

Page 8 HAWAIIAN ISLANDS ANIMALS: UPDATED NOVEMBER 29, 1999 LISTED AND CANDIDATE SPECIES , AS DESIGNATED UNDER THE U.S. ENDANGERED SPECIES ACT

ZORAPTERANS (Order Zoraptera; 1 taxon) Zorotypus swezeyi Zorapteran, Swezey's SOC T T

TRUE BUGS (Order Hemiptera; 31 taxa) Cavaticovelia aaa Water treader bug, aaa SOC T Coleotichus blackburniae Koa bug SOC T T T T T T Empicoris pulcher Thread bug, Pulchrus SOC T* Engytatus sp 1 Plant bug, Abutilon SOC T Engytatus sp 2 Plant bug, `Anunu SOC T Engytatus sp 3 Plant bug, Wilkesia SOC T Ithamar annectans Rhopalid bug, annectans SOC T T T Ithamar hawaiiensis Rhopalid bug, Hawaiian SOC T T T T T Kalania sp. Leaf bug, O'ahu kalanian SOC T Kalania hawaiiensis Leaf bug, L~na'i kalanian SOC T T Metrarga obscura Seed bug, Mauna Loa metrargan SOC T T Neseis alternata Seed bug, Kaua'i band-legged SOC T Neseis haleakalae Seed bug, Mt. Haleakala SOC T Nesidiolestes ana Thread bug, ana wingless SOC T Nesidiolestes insularis Thread bug, Mt. Tantalus wingless SOC T T T Nesidiolestes roberti Thread bug, Robert's wingless SOC T Nesidiolestes selium Thread bug, Selium wingless SOC T Nesocryptias villosa Seed bug, villosan flightless SOC T T T T Nesomiris sp. (ohe) Plant bug, ohe SOC T Nysius frigatensis Seed bug, French Frigate Shoal SOC F Nysius fullawayi Seed bug, Fullaway's SOC M, L Nysius neckerensis Seed bug, Necker goosefoot SOC Ne Nysius nihoae Seed bug, Nihoa nysius SOC N Nysius suffusus Seed bug, Necker bunchgrass SOC N Oceanides bryani Seed bug, Bryan's oceanides SOC T Oceanides perkinsi Seed bug, Perkins' oceanides SOC T Oceanides rugosiceps Seed bug, Rough-headed oceanides SOC T Oechalia grisea Stink bug, Gray oechalia SOC T T Oechalia patruelis Stink bug, Patruelis oechalia SOC T Saicella smithi Reduviid, Smith's saicellan SOC T

Page 9 HAWAIIAN ISLANDS ANIMALS: UPDATED NOVEMBER 29, 1999 LISTED AND CANDIDATE SPECIES , AS DESIGNATED UNDER THE U.S. ENDANGERED SPECIES ACT

CICADAS AND ALLIES (Order Homoptera; 22 taxa) Clavicoccus erinaceus Mealybug, O'ahu abutilon clavicoccus SOC T* Clavicoccus tribulus Mealybug, O'ahu ke'oke'o clavicoccus SOC T Nesorestias filicicola Planthopper, Mt Tantalus short-wing SOC T Nesosydne acuta Planthopper, Iao Valley nesosydne SOC T Nesosydne bridwelli Planthopper, Bridwell's nesosydne SOC T Nesosydne cyrtandrae Planthopper, Nahiku nesosydne SOC T Nesosydne cyrtandricola Planthopper, Glenwood nesosydne SOC T Nesosydne kuschei Planthopper, Kusche's nesosydne SOC T Nesosydne leahi Planthopper, Diamond Head nesosydne SOC T T T Nesosydne longipes Planthopper, long-footed nesosydne SOC T Nesosydne sulcata Planthopper, Keanae nesosydne SOC T Oliarus consimilis Planthopper, Kaua'i parti-colored oliarus SOC T Oliarus discrepans Planthopper, wild cotton oliarus SOC T Oliarus lihue Planthopper, Lihue oliarus SOC T Oliarus lorettae Planthopper, Loretta Lynn’s oliarus SOC T Oliarus myoporicola Planthopper, Barber's Pt. oliarus SOC T Oliarus priola Planthopper, priolan oliarus SOC T Oliarus sp 1 Planthopper, Doc Bellou cave oliarus SOC T Oliarus sp 2 Planthopper, Pahoa cave oliarus SOC T Paurotriozana adaptata Psyllid, O'ahu holio gall SOC T Phyllococcus oahuensis Mealybug, Opuhe gall SOC T T Pseudococcus antricolens Mealybug, O'ahu iliahi gall SOC T

LACEWINGS AND ALLIES (Order Neuroptera; 6 taxa) Distolean (=Eidolean) perjurus Antlion, Moloka'i SOC T T T T T Micromus (=Nesothauma) haleakalae Brown lacewing, Haleakala SOC T Micromus (=Pseudospectra) cookeorum Brown lacewing, Cooke's SOC T Micromus (=Pseudospectra) lobipennis Brown lacewing, Lobe-wing SOC T Micromus (=Pseudospectra) swezeyi Brown lacewing, Swezey's SOC T Micromus (=Pseudospectra) usingeri Brown lacewing, Usinger's SOC T

BEETLES (Order Coleoptera; 108 taxa) Anchotefflus gracilis Ground beetle, Gracile anchotefflus SOC T

Page 10 HAWAIIAN ISLANDS ANIMALS: UPDATED NOVEMBER 29, 1999 LISTED AND CANDIDATE SPECIES , AS DESIGNATED UNDER THE U.S. ENDANGERED SPECIES ACT

Apterocyclus honoluluensis Stag beetle, Kaua'i flightless SOC T Atelothrus transiens Ground beetle, Transient atelothrus SOC T* Blackburnia insignis Ground beetle, O'ahu blackburnia SOC T* Deinocossonus nesiotes Weevil, O'ahu nesiotes SOC T T T Deropristus deroderus Ground beetle, Haleakala deropristus SOC T Disenochus micantipennis Ground beetle, Kaua'i micantipennis SOC T* Eopenthes ambiguus Click beetle, Ambiguous eopenthes SOC T Eopenthes arduus Click beetle, Arduous eopenthes SOC T T Eopenthes auratus Click beetle, Golden eopenthes SOC T Eopenthes celatus Click beetle, Hidden eopenthes SOC T Eopenthes cognatus Click beetle, Cognatus eopenthes SOC T Eopenthes debilis Click beetle, Weak eopenthes SOC T Eopenthes deceptor Click beetle, Deceptive eopenthes SOC T Eopenthes funebris Click beetle, Death eopenthes SOC T Eopenthes germanus Click beetle, Germanus eopenthes SOC T Eopenthes kauaiensis Click beetle, Kaua'i eopenthes SOC T Eopenthes muticus Click beetle, Muticus eopenthes SOC T Eopenthes pallipes Click beetle, Pallipes eopenthes SOC T Eopenthes plebeius Click beetle, Common eopenthes SOC T T Eopenthes politus Click beetle, Politus eopenthes SOC T Eopenthes tarsalis Click beetle, Tarsalis eopenthes SOC T Eopenthes tinctus Click beetle, Tinged eopenthes SOC T Eopenthes unicolor Click beetle, Unicolored eopenthes SOC T Heteramphus filicum Weevil, O'ahu heteramphus fern SOC T Holcobius pikoensis Anobiid beetle, Piko SOC T Itodacnus novicornis Click beetle, Necker itodacnus SOC Ne Itodacnus paradoxus Click beetle, strange itodacnus SOC Ne Metromenus bardus Ground beetle, Heavy metromenus SOC T Metromenus cuneipennis Ground beetle, Wedge-winged metromenus SOC T* Metromenus oceanicus Ground beetle, Oceanic metromenus SOC T* Nesotocus giffardi Weevil, Giffard's nesotocus SOC T T T T Nesotocus kauaiensis Weevil, Kaua'i nesotocus SOC T* Nesotocus munroi Weevil, Munro's nesotocus SOC T T T Oodemas breviscapum Weevil, Nihoa oodemas SOC N Oodemas erro Weevil, Wandering oodemas SOC N

Page 11 HAWAIIAN ISLANDS ANIMALS: UPDATED NOVEMBER 29, 1999 LISTED AND CANDIDATE SPECIES , AS DESIGNATED UNDER THE U.S. ENDANGERED SPECIES ACT

Oodemas laysanensis Weevil, Laysan oodemas SOC N, L, M, Ne Oodemas neckeri Weevil, Necker oodemas SOC Ne Pentarthrum blackburni Weevil, Blackburn's pentarthrum SOC T L Pentarthrum obscurum Weevil, obscure pentarthrum SOC T T M Plagithmysus alani Long-horned beetle, Maui alani SOC T Plagithmysus annectans Long-horned beetle, Kaua'i annectant SOC T Plagithmysus bidensae Long-horned beetle, Bidens SOC T Plagithmysus bridwelli Long-horned beetle, Bridwell's SOC T Plagithmysus claviger Long-horned beetle, Hawai'i clubbed SOC T Plagithmysus decorus Long-horned beetle, Hawai'i decorus SOC T Plagithmysus dubautianus Long-horned beetle, Maui dubautia SOC T Plagithmysus elegans Long-horned beetle, Hawai'i elegant SOC T Plagithmysus forbesii Long-horned beetle, Forbes' Maui SOC T Plagithmysus forbesianus Long-horned beetle, Forbes' Kaua'i SOC T Plagithmysus fractus Long-horned beetle, Moloka'i fractured SOC T Plagithmysus greenwelli Long-horned beetle, Greenwell's SOC T Plagithmysus haasi Long-horned beetle, Haas' iliahi SOC T Plagithmysus ignotus Long-horned beetle, Kaua'i kalia SOC T Plagithmysus koae Long-horned beetle, Maui koa SOC T Plagithmysus kohalae Long-horned beetle, Kohala SOC T Plagithmysus kraussi Long-horned beetle, Krauss' SOC T Plagithmysus kuhnsi Long-horned beetle, Kuhns' O'ahu SOC T Plagithmysus lanaiensis Long-horned beetle, L~na'i 'ohi'a SOC T Plagithmysus laticollis Long-horned beetle, Maui wide-necked SOC T Plagithmysus longicollis Long-horned beetle, Long-necked SOC T T Plagithmysus mezoneuri Long-horned beetle, Hawai'i uhiuhi SOC T Plagithmysus muiri Long-horned beetle, Muir's ala'a SOC T Plagithmysus nihoae Long-horned beetle, Nihoa SOC N Plagithmysus paludis Long-horned beetle, Kaua'i swamp SOC T Plagithmysus permundus Long-horned beetle, Kaua'i 'ahakea SOC T Plagithmysus pipturicola Long-horned beetle, Maui mamaki SOC T Plagithmysus platydesmae Long-horned beetle, Pilo kea SOC T T Plagithmysus podagricus Long-horned beetle, Hawai'i podagricus SOC T Plagithmysus polystictus Long-horned beetle, Kaua'i holio SOC T Plagithmysus pulvillatus Long-horned beetle, 'Ohi'a SOC T T T

Page 12 HAWAIIAN ISLANDS ANIMALS: UPDATED NOVEMBER 29, 1999 LISTED AND CANDIDATE SPECIES , AS DESIGNATED UNDER THE U.S. ENDANGERED SPECIES ACT

Plagithmysus rubi Long-horned beetle, Maui 'akala SOC T Plagithmysus simplicollis Long-horned beetle, Simple-necked SOC T Plagithmysus simillimus Long-horned beetle, Maui similar SOC T T Plagithmysus speculifer Long-horned beetle, Maui speculifer SOC T Plagithmysus sulphurescens Long-horned beetle, Hawai'i opuhe SOC T Plagithmysus superstes Long-horned beetle, O'ahu super SOC T Plagiithmysus swezeyanus Long-horned beetle, Ahinahina SOC T T Plagithmysus swezeyi Long-horned beetle, Swezey's SOC T T Plagithmysus sylvai Long-horned beetle, Maui forest SOC T Plagithmysus vicinus Long-horned beetle, Hawai'i alani SOC T Proterhinus 72 spp. Proterhinid beetles, Hawaiian SOC T T T T T T Pseudobroscus lentus Ground beetle, Haleakala pseudobroscus SOC T* Rhyncogonus biformis Weevil, Necker rhyncogonus SOC Ne Rhyncogonus blackburni Weevil, Blackburn's rhyncogonus SOC T Rhyncogonus bryani Weevil, Laysan rhyncogonus SOC L* Rhyncogonus extraneus Weevil, Kunia rhyncogonus SOC T* Rhyncogonus exsul Weevil, Nihoa rhyncogonus SOC N Rhyncogonus freycinetiae Weevil, 'Ie'ie rhyncogonus SOC T T T Rhyncogonus giffardi Weevil, Giffard's rhyncogonus SOC T# Rhyncogonus koebelei Weevil, Koebele's rhyncogonus SOC T Rhyncogonus lahainae Weevil, Lahaina rhyncogonus SOC T Rhyncogonus lanaiensis Weevil, L~na'i rhyncogonus SOC T Rhyncogonus molokaiensis Weevil, Moloka'i rhyncogonus SOC T Rhyncogonus mutatus Weevil, Mutated rhyncogonus SOC T Rhyncogonus obsoletus Weevil, Obsolete rhyncogonus SOC T Rhyncogonus oleae Weevil, Olopua rhyncogonus SOC T Rhyncogonus segnis segnis Weevil, Slow rhyncogonus SOC T Rhyncogonus segnis fordi Weevil, Ford's rhyncogonus SOC T Rhyncogonus sharpi Weevil, Sharp's rhyncogonus SOC T Rhyncogonus simplex Weevil, Simple rhyncogonus SOC T T Rhyncogonus squamiger Weevil, Scaley rhyncogonus SOC T Rhyncogonus stygius Weevil, Black rhyncogonus SOC T Rhyncogonus sylvicola Weevil, Kaua'i forest rhyncogonus SOC T Rhyncogonus tuberculatus Weevil, Tubercled rhyncogonus SOC T Rhyncogonus vittatus Weevil, Striped rhyncogonus SOC T*

Page 13 HAWAIIAN ISLANDS ANIMALS: UPDATED NOVEMBER 29, 1999 LISTED AND CANDIDATE SPECIES , AS DESIGNATED UNDER THE U.S. ENDANGERED SPECIES ACT

Rhyncogonus welchii Weevil, Welch's rhyncogonus SOC T# Stenotrupis pritchardiae Weevil, Nihoa stenotrupis SOC N

FLIES (Order Diptera; 5 taxa) Bryania bipunctata Asteriid fly, Nihoa two-spotted SOC T T N, L Chersodromia hawaiiensis Dance fly, Hawaiian chersodromian SOC T Drosophila lanaiensis Pomace fly, L~na'i SOC T* T Drosophila reynoldsiae Pomace fly, Oahu SOC T Drosophila touchardiae Pomace fly, Oahu SOC T

BUTTERFLIES AND MOTHS (Order Lepidoptera; 32 taxa) Agrotis cremata Moth, Cremata agrotis noctuid SOC T* T* Agrotis melanoneura Moth, Black-veined agrotis noctuid SOC T* T* Agrotis microreas Moth, Microreas agrotis noctuid SOC T* Agrotis potophila Moth, Potophila agrotis noctuid SOC T* Anomis vulpicolor Moth, Red anomis noctuid SOC T* T* T* Carposina (=Heterocrossa) viridis Moth, green carposinid SOC T T Glyphodes (=Margaronia) cyanomichla Moth, blue glyphodes SOC T T T T Helicoverpa confusa Moth, confused helicoverpan noctuid SOC T* T* T* T* T* Hyposmocoma neckerensis Moth, Necker hyposmocoman leaf miner SOC Ne* Kaua'iina (=Fletcherana) ioxantha Moth, ioxanthan looper SOC T Omiodes (=Hedylepta) anastrepta Moth, Moloka'i sedge omiodes SOC T* T* T* Omiodes (=Hedylepta) anastreptoides Moth, Kohala Mts. omiodes SOC T Omiodes (=Hedylepta) asaphombra Moth, 'ohe omiodes SOC T* T* T* T* Omiodes (=Hedylepta) euryprora Moth, Ola'a banana omiodes SOC T* Omiodes (=Hedylepta) fullawayi Moth, Fullaway's banana omiodes SOC T* Omiodes (=Hedylepta) giffardi Moth, Giffard's 'ohe omiodes SOC T Omiodes (=Hedylepta) iridias Moth, Kilauea pa'inia omiodes SOC T* Omiodes (=Hedylepta) meyricki Moth, Meyrick's banana omiodes SOC T* Omiodes (=Hedylepta) monogona Moth, Hawaiian bean leafroller SOC T* T* T* T* T* T* Omiodes (=Hedylepta) musicola Moth, Maui banana omiodes SOC T* T* Omiodes (=Hedylepta) pritchardii Moth, Hawaiian o'ulu omiodes SOC T Philodoria naenaeiella Moth, Kaua'i na'ena'e philodoria SOC T* Philodoria sp. Moth, O'ahu hesperomannia philodoria SOC T* Philodoria ureraella Moth, O'ahu opuhe leaf mining SOC T*

Page 14 HAWAIIAN ISLANDS ANIMALS: UPDATED NOVEMBER 29, 1999 LISTED AND CANDIDATE SPECIES , AS DESIGNATED UNDER THE U.S. ENDANGERED SPECIES ACT

Plutella capparidis Moth, O'ahu capper SOC T* Spheterista oheoheana Moth, 'ohe'ohe leafroller SOC T Spheterista pterotropiana Moth, greenbanded 'ohe'ohe leafroller SOC T Spheterista reynoldsiana Moth, Wailupe leafroller SOC T Stemorrhages (=Margaronia) exaula Moth, green stemorrhages SOC T T T T T Thyrocopa apatela Moth, Haleakala flightless thyrocopa SOC T Thyrocopa sapindiella Moth, O'ahu aulu thyrocopa SOC T* Udea (=Oeobia) dryadopa Moth, 'Ohenaupaka udean SOC T T T L

ANTS, BEES AND WASPS (Order Hymenoptera; 54 taxa) Deinomimesa hawaiiensis Wasp, Hawaiian deinomimesan sphecid SOC T Deinomimesa punae Wasp, Puna deinomimesan sphecid SOC T Ectemnius bidecoratus Wasp, bidecoratus sphecid SOC T Ectemnius curtipes Wasp, short-footed ectemnius sphecid SOC T T Ectemnius fulvicrus Wasp, brown cross ectemnius sphecid SOC T Ectemnius haleakalae Wasp, Haleakala ectemnius sphecid SOC T Ectemnius rubrocaudatus Wasp, redtail ectemnius sphecid SOC T* Ectemnius yoshimotoi Wasp, Yoshimoto's ectemnius sphecid SOC T Eupelmus nihoaensis Wasp, Nihoa eupelmus SOC N Nesomimesa kauaiensis Wasp, Kaua'i nesomimesan sphecid SOC T Nesomimesa perkinsi Wasp, Perkins' nesomimesan sphecid SOC T Nesomimesa sciopteryx Wasp, shade-winged sphecid SOC T Hylaeus (=Nesoprosopis) andrenoides Bee, andrenoid yellow-faced SOC T* Hylaeus (=Nesoprosopis) anomala Bee, anomalous yellow-faced SOC T Hylaeus (=Nesoprosopis) anthracina Bee, anthracinan yellow-faced SOC T# T# T# T# Ni# Hylaeus (=Nesoprosopis) assimulans Bee, assimulans yellow-faced SOC T* T* T* Hylaeus (=Nesoprosopis) caeruleipennis Bee, bluewing yellow-faced SOC T* T* Hylaeus (=Nesoprosopis) chlorosticta Bee, chlorostictan yellow-faced SOC T* Hylaeus (=Nesoprosopis) comes Bee, comes yellow-faced SOC T* T* Hylaeus (=Nesoprosopis) coniceps Bee, conehead yellow-faced SOC T* Hylaeus (=Nesoprosopis) crabronoides Bee, crabronoid yellow-faced SOC T* Hylaeus (=Nesoprosopis) difficilis Bee, difficult yellow-faced SOC T* T* T* T* T* Hylaeus (=Nesoprosopis) dimidiata Bee, dimidiatan yellow-faced SOC T* Hylaeus (=Nesoprosopis) facilis Bee, easy yellow-faced SOC T* T* T* T* T* Hylaeus (=Nesoprosopis) filicum Bee, fern yellow-faced SOC T* T* T* T*

Page 15 HAWAIIAN ISLANDS ANIMALS: UPDATED NOVEMBER 29, 1999 LISTED AND CANDIDATE SPECIES , AS DESIGNATED UNDER THE U.S. ENDANGERED SPECIES ACT

Hylaeus (=Nesoprosopis) flavifrons Bee, very yellow-faced SOC T Hylaeus (=Nesoprosopis) flavipes Bee, yellow-footed yellow-faced SOC T* Hylaeus (=Nesoprosopis) fuscipennis Bee, darkwig yellow-faced SOC T Hylaeus (=Nesoprosopis) haleakalae Bee, Haleakala yellow-faced SOC T* Hylaeus (=Nesoprosopis) hirsutula Bee, hirsute yellow-faced SOC T* Hylaeus (=Nesoprosopis) hostilis Bee, hostile yellow-faced SOC T* Hylaeus (=Nesoprosopis) hula Bee, hulan yellow-faced SOC T* Hylaeus (=Nesoprosopis) insignis Bee, insignis yellow-faced SOC T* Hylaeus (=Nesoprosopis) kauaiensis Bee, Kaua'i yellow-faced SOC T* Hylaeus (=Nesoprosopis) koae Bee, koa yellow-faced SOC T* Hylaeus (=Nesoprosopis) kona Bee, Kona yellow-faced SOC T* Hylaeus (=Nesoprosopis) laeta Bee, laetan yellow-faced SOC T* T* T* T* T* Hylaeus (=Nesoprosopis) longiceps Bee, longhead yellow-faced SOC T* T* T* T* K* Hylaeus (=Nesoprosopis) obscurata Bee, obscuratan yellow-faced SOC T* T* T* Hylaeus (=Nesoprosopis) ombrias Bee, ombrias yellow-faced SOC T* Hylaeus (=Nesoprosopis) perkinsiana Bee, Perkins' yellow-faced SOC N Hylaeus (=Nesoprosopis) pubescens Bee, furry yellow-faced SOC T Hylaeus (=Nesoprosopis) satellus Bee, satellus yellow-faced SOC T* T* T* Hylaeus (=Nesoprosopis) simplex Bee, simple yellow-faced SOC T* Hylaeus (=Nesoprosopis) specularis Bee, specular yellow-faced SOC T* Hylaeus (=Nesoprosopis) sphecodoides Bee, sphecodoid yellow-faced SOC T* Hylaeus (=Nesoprosopis) unica Bee, unique yellow-faced SOC T* Hylaeus (=Nesoprosopis) vicina Bee, vicinan yellow-faced SOC T* T* T* T* T* Hylaeus (=Nesoprosopis) volatilis Bee, volatile yellow-faced SOC T* T* T* T* T* Odynerus niihauensis Wasp, Niihau odynerus vespid SOC Ni Odynerus nigripennis Wasp, Black-winged odynerus vespid SOC T T T T T Odynerus radula Wasp, Radulan odynerus vespid SOC T* Odynerus soror Wasp, soror odynerus vespid SOC T Ni Sclerodermus nihoaensis Wasp, Nihoa sclerodermus SOC N

SHRIMP (Order Decapoda; 2 taxa) Halocaridina palehemo Shrimp, no common name SOC T Halocaridina rubra Shrimp, no common name SOC T T T T

HYDROIDS (Order Cnidaria; 1 taxon)

Page 16 HAWAIIAN ISLANDS ANIMALS: UPDATED NOVEMBER 29, 1999 LISTED AND CANDIDATE SPECIES , AS DESIGNATED UNDER THE U.S. ENDANGERED SPECIES ACT

Ostromouvia horii No common name SOC T

Page 17