Revision of the Genus Trachyarus Thomson

©Zoologische Staatssammlung München/Verlag Friedrich Pfeil; download www.pfeil-verlag.de

SPIXIANA 30 1 65–83 München, 1. Mai 2007 ISSN 0341–8391

Revision of the genus Trachyarus Thomson

(Insecta, Hymenoptera, Ichneumonidae, Alomyini)

Vladimir E. Gokhman

Gokhman, V. E. (2007): Revision of the genus Trachyarus Thomson (Insecta, Hymenoptera, Ichneumonidae, Alomyini). – Spixiana 30/1: 65-83 A revision of the Palaearctic genus Trachyarus is presented. This genus includes 15 described and valid species (including nine species described in this paper): Trachyarus bacillatus, spec. nov., T. decipiens, spec. nov., T. edilleri, spec. nov., T. hemichneumonoides, spec. nov., T. khrulevae, spec. nov., T. parvipennis, spec. nov., T. punctigaster, spec. nov., T. solyanikovi, spec. nov. and T. subtilipiceus, spec. nov.). Trachyarus specularis Roman is considered a junior synonym of T. anceps (Berthou- mieu). Vladimir E. Gokhman, Botanical Garden, Moscow State University, Moscow 119992, Russia; e-mail: [email protected]

Introduction of Sciences, St. Petersburg, Russia; ZMMU – Zoological Museum, Moscow State University, Moscow, Russia; Trachyarus is a medium-sized Palaearctic genus of ZSM – Zoologische Staatssammlung, Munich, Ger- the tribe Alomyini (= Phaeogenini sensu Perkins many. Unless otherwise stated, all type material of 1959, Wahl & Mason 1995). The genus and its type newly described species is deposited in the ZSM collection. species were described at the end of the nineteenth century (Thomson 1891). However, it is poorly known although several other species now belong- Genus Trachyarus Thomson ing to Trachyarus were discovered since that time (Berthoumieu 1906, Roman 1918, Diller 1988, 1989). Trachyarus Thomson, 1891: 1612. The examination of the extensive material from collections of the Zoologische Staatssammlung (Munich, Type species. Trachyarus corvinus Thomson, 1891: Germany), Zoological Institute, Russian Academy 1612, by monotypy. of Sciences (St. Petersburg, Russia), and the Zoo- logical Museum of the Moscow University (Moscow, Description Russia) resulted in the discovery of several new Female. Head conspicuously inflated. Clypeus species. Moreover, a revision of Trachyarus is neces- strongly convex, well separated from the face. sary because of the considerable intraspeciﬁ c varia- Lower edge of the clypeus thin, usually more or less tion in this genus, especially regarding wing struc- rounded, with an apical margin. Clypeus about twice ture. as wide as high. Face about twice as wide as high, median part rising above the lateral ones. Antennal Abbreviations used: MNHN – Muséum National D’Histoire Naturelle, Paris, France; NHM – Natural scrobe weak. Frons obviously convex. Head trans- History Museum, London, UK; NHRM – Naturhistor- verse, sometimes narrowed behind eyes in dorsal iska Riksmuseet, Stockholm, Sweden; UZI – Museum view. Temples usually enlarged in proﬁ le, with their of Zoology and Entomology, Lund University, Lund, maximum length somewhat equal to transverse Sweden; ZIN – Zoological Institute, Russian Academy ocular diameter. Malar space more or less equal to

65 ©Zoologische Staatssammlung München/Verlag Friedrich Pfeil; download www.pfeil-verlag.de width of mandibular base. Scape oviform, its apical Biology. All known hosts belong to Psychidae truncation inclined. Maximum length of the scape (Lepidoptera). slightly exceeds its width. Flagellum fi liform, with Differential diagnosis. Members of a few genera 21-27 segments. Mandibles sometimes narrowed, of the tribe Alomyini, namely those of Hemichneumon their upper tooth longer or shorter than the lower Wesmael, Epitomus Förster and Tycherus Förster tooth. (fuscicornis species group), are closest to Trachyarus. Mesoscutum convex. Scutellum distinctly con- Species of Trachyarus (as well as those of Hemichneu- vex, its lateral carinae usually very short. Sternauli mon and Epitomus), however, differ from Tycherus stretch almost to the middle of mesopleura. Propo- by having areolet of the fore wing open, its occlud- deum formed by weakly inclined short basal part ing vein spectral; from Epitomus – by having propo- and strongly inclined long apical one (except T. bra- deum sloping from anterior margin of area supero- chypterator and T. khrulevae). Area basalis of propo- media; from Hemichneumon – by having large thy- deum short and indistinct. Area superomedia heart- ridia, rugose mesopleura and short but distinct area shaped, narrowed basally, separated from area superomedia of the propodeum; the former area is posteromedia by weak and indistinct carina. Area often separated from area posteromedia by more or petiolaris bordered by weak diverging carinae, wide, less developed carina (see Selfa & Diller 1994). distinctly concave. Side carinae of area posteromedia hardly visible and very indistinct, sometimes absent. Wings fully developed or reduced. Areolet open, its 1. Trachyarus anceps (Berthoumieu) occluding vein spectral. Second recurrent vein with Figs 1, 2 2 bullae. Nervellus usually vertical. Coxal armature absent. Hemichneumon anceps Berthoumieu, 1906: 60. Holotype: Metasoma oval to longish-oval. Petiolus more W, France, “Val d’Isère” (MNHN, studied). or less compressed dorsoventrally. Thyridia often Trachyarus anceps: Diller, 1989: 288. wide and slightly inclined. Protruding part of the Trachyarus specularis Roman, 1918: 8. Lectotype: W, Swe- ovipositor sheath very short. Sterna of metasomal den, “Lapp. suecica merid.” (NHRM, designated here, segments II-IV with a median fold. Hypopygium selected by E. Diller). Syn. nov. short, far from overlapping the ovipositor slit. Material. 12WW, 6MM (MNHN, NHRM, ZIN, ZSM) in- Body weakly to moderately sculptured. Head cluding holotype (W, France, “Val d’Isère”, “type”, “Hem- shining, side parts of face more or less densely ichneumon anceps n. sp. (Berth. vid.)”, “Museum Paris”, punctate, punctures of the median part of the face “Holotypus Hemichneumon anceps Berth. W Diller 89”, fi ner, clypeus fi nely punctate. Frons with fi ne scat- “Trachyarus anceps (Berth.) W det. E. Diller 89”, “montage tered punctures. Temples very sparsely and fi nely sur Polypore le 23.III.1989 M. Lachaise”) (MNHN). – punctate. Malar space with a chagrinate groove. Paratypes: 12WW, 6MM, same data. Mesoscutum and scutellum usually shining, the Additional material. Two females and a male reared in former with comparatively dense and fi ne punctures, the Moscow region from Siederia spp. (including S. lis- the latter more sparsely punctate. Mesopleura usu- terella; ZIN) probably also belong to this species. ally rugose, speculum more or less smooth. Propo- deum rugose (areae superoexternae less sculptured). Differential diagnosis. The female of T. anceps is Base of metasoma usually chagrinate, sculpture closest to that of T. brevipennis, but in T. anceps the disappears towards apex of metasoma. fl agellum is more slender (fi rst fl agellar segment Body usually brown. Head brown, clypeus, 3.6-4.0 × as long as wide) and the metasomal sculp- median part of face and mandibles usually lighter. ture is less developed (especially on tergum III). Antennae brown to reddish brown, palps light Among Trachyarus males, T. anceps is the only species brown. Thorax brown. Stigma and veins reddish. having a combination of the following characters: Legs predominantly brown. Metasoma usually upper mandibular tooth longer than the lower tooth; brown with hind margins of terga II-III yellowish fl agellum without tyloids; metasomal tergum III and tergum VII reddish brown. 1.6-1.7 × as wide as long, its apical half smooth with Body length 3.5-6.0 mm, fore wing length 0.3- obscure punctures. 4.3 mm. Note. Another species of the genus, Trachyarus Male. Similar to female. Tyloids, if developed, specularis Roman, appeared to be a synonym of oval, usually present before the middle of fl agel- T. anceps. Contrary to Diller (1988: 371), this species lum. is not a synonym of T. fuscipes (lectotypes com- pared). Distribution. Palaearctis.

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2 3 4 5 6 1

8 10 9 11 Figs 1-11. Trachyarus spp., WW. 1, 3. Base of ﬂ agellum. 1. T. anceps (Berthoumieu). 3. T. brevipennis Roman. 2. T. anceps, mandible. 4-6, 8-11. Metasomal terga II and III. 4. T. corvinus Thomson. 5. T. fuscipes (Thomson). 6. T. prominulus Diller. 8. T. edilleri, spec. nov. 9. T. hemichneumonoides, spec. nov. 10. T. parvipennis, spec. nov. 11. T. solyanikovi, spec. nov. 7. T. bacillatus, spec. nov., mandible. – Scale equals 0.25 mm for Figs 1-3, 7 and 0.5 mm for Figs 4-6, 8-11.

Description fi nely and sparsely punctate. Upper halves of the pronotal lobes shining, with fi ne scattered punctures, Female. Lower edge of the clypeus very slightly the lower ones longitudinally rugose. Mesopleura rounded, apical margin hardly visible. Head slight- almost smooth to rugose-punctate, sometimes only ly narrowed behind eyes in dorsal view. Temples in their apical half, speculum smooth with a few more or less enlarged ventrad. Flagellum slightly very fi ne punctures. Metasomal tergum I coarsely enlarged in apical half, with 23-26 segments, fi rst punctate, postpetiolus irregularly longitudinally segment 3.6-4.0 × as long as wide (Fig. 1), 12th or 13th striate, tergum II and sometimes basal fourth of segment square in lateral view. Mandibles somewhat tergum III obviously chagrinate, with obscure punc- narrowed, upper mandibular tooth distinctly long- tures, the others sparsely punctate, sculpture almost er than the lower tooth (Fig. 2). Hypostomal carina disappears towards apex of metasoma. meets genal carina before the mandibular base at Body brown. Head brown, clypeus and some- half its width. times cheeks and most part of face lighter, mandibles Distinct notauli present in the basal third of (except teeth) yellowish brown. Antennae brown, mesoscutum. Area superomedia of propodeum slightly reddish beneath, palps light brown. Tegulae about 1.3 × as wide as long. Area petiolaris about 3 × whitish. Legs brown, apices of front and middle as long as area superomedia. Wings fully developed. femora and stripes on inner sides of the front and Nervellus broken in its lower third. Middle tibiae middle tibiae reddish, the latters sometimes en- sometimes with a transverse shallow subapical tirely reddish, all tarsi sometimes reddish. Metasoma impression on outer sides. Hind femora 3.5-3.6 × as brown with thyridia and hind margins of terga II-III long as wide. (IV-V) yellowish, tergum VII sometimes reddish Metasoma longish-oval. Petiolus slightly com- brown. pressed dorsoventrally. Distance between spiracles Body length 3.5-5.0 mm, fore wing length 2.9- of the fi rst metasomal segment twice as long as the 3.6 mm. distance between the spiracle and hind corner of the Male similar to female. Flagellum with 23-24 segment. Thyridia medially removed from the base segments, fi rst segment 2.8-2.9 × as long as wide, of the second tergum by about half their width, ap- several segments before apex of fl agellum square in proximately twice as wide as the interthyridial space. lateral view, tyloids absent. Sculpture of metasoma Metasomal tergum II 1.0-1.1 × as wide as long, ter- more robust. Body brown, clypeus sometimes whit- gum III 1.6-1.7 × as wide as long. Protruding part of ish, mandibles sometimes with whitish markings at the ovipositor sheath more or less visible from above. the middle. Body weakly sculptured. Frons shining, very Body length 4.5 mm, fore wing length 3.3 mm.

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Biology. Parasitoid of Dahlica charlottae (Meier), Metasomal tergum II 1.2-1.3 × as wide as long, ter- D. klimeschi (Sieder), D. triquetrella (Hübner) and gum III 1.8-1.9 × as wide as long. Siederia listerella (L.). Body moderately sculptured. Head weakly shining, fi nely chagrinate. Thorax chagrinate. Metaso- Distribution. Ukraine (Transcarpathia), Finland, mal terga I-III chagrinate, sculpture disappears to- Sweden, Austria, Germany (Bavaria), France (Isère). wards apex of metasoma. A boreal-montane species. Body yellowish brown. Head brown, mandibles (except teeth), clypeus and median part of face yellowish brown. Scape and pedicel brown, fl agellum 2. Trachyarus brachypterator Diller reddish ventrally, 1st to 4th fl agellar segments reddish, 5th to 7th with white markings, other segments brown, Trachyarus brachypterator Diller, 1988: 372. palps light brown. Thorax yellowish brown. Legs Holotype: W, Austria, “Niederösterreich, Wimpassing, yellowish, middle and hind femora somewhat an Schwarzföhre” (ZSM, not examined). darker. Metasomal terga I-II, basal third and hind Material: Paratype: W, Austria, “Paratypus”, “Österreich”, margin of tergum III yellowish brown, other terga “Parasit e Psychidae, leg. H. Meier”, “NÖ, Wimpassing, brown, tergum VII reddish brown. an Schwarzföhre, Mai 1974, Sied. pineti Z.”, “Paratypus Body length 3.5 mm, fore wing length 0.3 mm. W Trachyarus brachypterator W Diller, 87” (ZSM). Male unknown.

Differential diagnosis. Among Trachyarus species, Biology. Parasitoid of Siederia listerella. the wings are also strongly reduced in T. parvipennis, spec. nov., from which T. brachypterator well differs Distribution. Austria, Hungary (Diller 1988). by the structure of mandibles, number of fl agellar segments, proportions of metasomal terga and general colour of the body and legs. 3. Trachyarus brevipennis Roman Fig. 3 Description Trachyarus brevipennis Roman, 1918: 7. Lectotype: W, Female. Lower edge of the clypeus slightly round- Sweden, “Lapp. suecica meridionalis” (NHRM, ed, with a visible apical margin. Head moderately designated here, selected by E. Diller). transverse, not narrowed behind eyes in dorsal view. Temples prominently enlarged ventrad. Flagellum Material. 12WW, 7MM (NHRM, ZIN, ZSM), including lectotype (W, Sweden, “Lp.m.”, “Bhm”, “28/7”, “60 81”, slightly enlarged in apical half, with 21 segments, “Riksmuseum Stockholm”, “Lectotypus Trachyarus bre- th th fi rst segment 1.7-1.8 × as long as wide, 4 or 5 seg- vipennis Roman W Diller 1981”, “Naturhistoriska Riks- ment square in lateral view. Upper mandibular tooth museet Stockholm Loan no 574/94”) (NHRM) and pa- distinctly longer and wider than the lower tooth. ralectotype (W, Sweden, “Lp.m.”, “Bhm”, “28/7”, “61 81”, Hypostomal carina meets the genal carina before “Riksmuseum Stockholm”, “Paralectotypus Trachyarus mandibular base at about half its width. brevipennis Roman W Diller 1981”, “Naturhistoriska Distinct notauli present in the basal third of Riksmuseet Stockholm loan no 575/94”) (NHRM). mesoscutum. Scutellum comparatively small, its Differential diagnosis. The female of T. brevipennis lateral carinae developed in its basal half. Propodeum is closest to that of T. anceps, but in T. brevipennis the formed by comparatively long and horizontal, con- fl agellum is more slender (fi rst fl agellar segment vex basal part and strongly inclined, slightly concave 2.2-2.3 × as long as wide) and the metasomal tergum long apical part, the latter about 1.2 × as long as the III is distinctly shagrinate in its basal half. The male former. Areolation of the propodeum very indistinct of T. brevipennis is closest to that of Trachyarus sp. C, or absent. Area superomedia very inconspicuous, but in T. brevipennis fl agellum and metasomal tergum rounded, about 1.5 × as long as wide. Wings ves- II are stouter (17th or 18th fl agellar segment square tigial, hardly reaching base of propodeum. Hind in lateral view; tergum II as long as wide). femora 3.1-3.2 × as long as wide. Metasoma oval. Petiolus slightly compressed Note. T. brevipennis was previously considered to be dorsoventrally. Distance between spiracles of the the commonest European species of that genus, but fi rst metasomal segment about 1.2 × as long as dis- the majority of specimens deposited in ZSM actu- tance between the spiracle and hind corner of the ally belongs to T. edilleri, spec. nov., T. decipiens, spec. segment. Thyridia medially removed from base of nov., and T. solyanikovi, spec. nov. (see below). second tergum by about one-third their width, approximately 1.5 × as wide as the interthyridial space.

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Description Male (described provisionally) similar to female. Flagellum with 23-24 segments, fi rst segment 2.4 × Female. Lower edge of the clypeus very slightly as long as wide, 17th or 18th segment square in lat- rounded, straight medially, with a visible apical eral view. Short-oval tyloids present on 10th-11th margin. Head slightly narrowed behind eyes in fl agellar segment. Clypeus (except its lower margin) dorsal view. Temples moderately enlarged ventrad, and mandibles (except teeth) sometimes whitish. with their maximum length somewhat less than Body length 3.5-4.5 mm, fore wing length 2.8- transverse ocular diameter. Flagellum with 23-24 3.6 mm. segments, fi rst segment 2.2-2.3 × as long as wide (Fig. 3), 9th or 10th segment square in lateral view. Biology. Parasitoid of Anaproutia comitella (Bruand), Upper mandibular tooth distinctly longer but not A. raiblensis (Mann), Brevantennia siederi (Sauter) as wider than the lower tooth. Hypostomal carina meets well as of Dahlica and Siederia spp., including the genal carina before mandibular base at a distance D. charlottae and S. listerella. of two-thirds its width. Distinct notauli present in the basal third of Distribution. Sweden, Russia (Centre and Far East), mesoscutum. Area superomedia 1.4-1.5 × as wide as Austria, Italy (Alps), Switzerland, Germany (Bavaria). long (up to equal length and width in the paralec- totype). Area petiolaris about 2.5 × as long as area superomedia. Wings from fully to partially devel- 4. Trachyarus corvinus Thomson oped, in the latter case they may hardly reach apex Fig. 4 of propodeum. Hind femora 3.3-3.4 × as long as wide. Metasoma oval. Distance between spiracles of Trachyarus corvinus Thomson, 1891: 1612. Lectotype: W, the fi rst metasomal segment approximately 1.5 × as Sweden, “Pålsjö” (UZI, studied). long as distance between the spiracle and hind Material. 8WW, 3MM (NHM, UZI, ZIN, ZSM), including corner of the segment. Thyridia medially removed lectotype (W, Sweden, “Pål”, “corvinus m”, “1978 163”, from base of the second tergum by about one-third “Lectotypus Trachyarus corvinus Thm. Tow’58”, “1981 their width, approximately 1.5 × as wide as interthy- 765”, “1994 201”) (UZI). ridial space. Metasomal tergum II as wide as long, tergum III 1.4-1.5 × as wide as long. Differential diagnosis. T. corvinus is most closely Body moderately sculptured. Head shining, side related to T. anceps, T. brevipennis and Trachyarus parts of face more or less densely punctate and sp. C from which it differs by having a stouter fl agellum (7th or 8th fl agellar segment (in the female) and obscurely chagrinate, median part of face and th th clypeus more fi nely punctate and chagrinate. Frons 11 or 12 segment (in the male) square in lateral very fi nely punctate and sometimes obscurely cha- view). grinate. Mesoscutum and scutellum fi nely and Note. A similar form (13WW, 7MM, ZIN) also occurs sparsely punctate; pronotal lobes punctate and in the Russian Far East: female fl agellum with 23-25 chagrinate, longitudinally rugose posteriorly. Mes- segments, facial orbits and usually central part of opleura longitudinally rugose, speculum smooth. the face in the male with whitish markings. Metasomal tergum I chagrinate, petiolus rugose, tergum II and basal half of tergum III (sometimes Description terga II-III and basal half of tergum IV) obviously chagrinate with obscure scattered punctures, sculp- Female. Lower edge of the clypeus very slightly ture becomes more smooth towards apex of meta- rounded, straight medially, with a visible apical soma. margin. Head distinctly narrowed behind eyes in Body brown. Head brown, clypeus, median part dorsal view. Temples slightly enlarged at the middle, of face and sometimes cheeks somewhat lighter, with their maximum length equal to transverse mandibles (except base and teeth) yellowish brown. ocular diameter. Flagellum with 22-24 segments, th th Antennae brownish, scape and pedicel usually yel- fi rst segment 2.4-2.5 × as long as wide, 7 or 8 seg- lowish brown, palps light brown. Tegulae whitish. ment square in lateral view. Upper mandibular tooth Legs reddish, last segments of middle and hind distinctly longer and hardly wider than the lower tarsi sometimes infuscate. Metasoma brown with tooth. Hypostomal carina meets the genal carina the very apex of the fi rst segment medially, thy- before the mandibular base at half its width. ridia and hind margins of terga II-III yellowish, Distinct notauli present in the basal third of tergum VII reddish brown. mesoscutum. Area superomedia heart-shaped to Body length 3.5-4.5 mm, fore wing length 0.9- reniform, about twice as wide as long. Area petiola- 3.0 mm. ris about 5 × as long as area superomedia. Wings

69 ©Zoologische Staatssammlung München/Verlag Friedrich Pfeil; download www.pfeil-verlag.de fully developed. Nervellus broken slightly below 5. Trachyarus fuscipes (Thomson) the middle. Middle tibiae sometimes with a trans- Fig. 5 verse shallow subapical impression on outer sides. Hind femora 3.5-3.6 × as long as wide. Hemichneumon fuscipes Thomson, 1891: 1612. Holotype: Metasoma oval. Petiolus distinctly compressed W, Sweden, “Öland” (UZI, studied). dorsoventrally. Distance between spiracles of the Trachyarus fuscipes, Diller 1988: 371. fi rst metasomal segment twice as long as distance Material. 67WW, 126MM (UZI, ZIN, ZSM), including between the spiracle and hind corner of the segment. holotype (W, Sweden, “Ö”, “fuscipes m”, “Hemichneumon Thyridia medially removed from the base of the fuscipes Ths. 1891 Holotype W det. M. G. Fitton 1978”, second tergum by about one-third their width, ap- “1981 757”, “1994 198”) (UZI). proximately 1.5 × as wide as the interthyridial space. Differential diagnosis. T. fuscipes is most closely Metasomal tergum II 1.2-1.3 × as wide as long, ter- related to T. prominulus, from which it differs by gum III 1.8-1.9 × as wide as long (Fig. 4). having more slender metasoma, whitish tegulae and Body moderately sculptured. Frons very ob- no tyloids on the male fl agellum. scurely chagrinate and sparsely punctate. Mesoscu- tum and scutellum weakly shining, the former Note. T. fuscipes appears to be the commonest re- chagrinate, comparatively roughly punctate, the corded species of its genus in Western Europe. latter much more fi nely punctate; pronotal lobes strongly and coarsely rugose-punctate, longitudi- Description nally rugose in their lower half. Mesopleura rugose, speculum smooth. Postpetiolus of the fi rst metaso- Female. Lower edge of the clypeus very slightly mal segment irregularly longitudinally striate, petio- rounded, apical margin hardly visible. Head hardly lus rugose-punctate. Metasomal terga II-III and narrowed behind eyes in dorsal view. Temples basal two-thirds of tergum IV obviously chagrinate moderately enlarged at the middle, with their with obscure scattered punctures, sculpture becomes maximum length slightly less than transverse ocular more smooth towards apex of metasoma. diameter. Flagellum distinctly enlarged in apical Body brown. Head brown, clypeus and median half, with 25-27 segments, fi rst segment 2.6-2.7 × as th part of face a little lighter, mandibles (except base long as wide, 10 or 11th segment approximately and teeth) yellowish brown. Antennae brown, base square in lateral view. Upper mandibular tooth of fl agellum sometimes slightly reddish ventrally, distinctly longer and hardly wider than the lower palps light brown. Tegulae reddish brown. Legs tooth. Hypostomal carina meets the genal carina brown, apices of front femora and all tibiae and before the mandibular base at two-thirds its width. tarsi reddish; legs sometimes entirely reddish. Meta- Distinct notauli present in the basal fourth of soma brown with the very apex of metasomal tergum mesoscutum. Area superomedia about 1.3 × as wide I medially, thyridia and hind margins of terga II-III as long. Area posteromedia bordered by hardly vis- (sometimes also of terga IV-V) yellowish, VII tergum ible although distinct carinae, approximately 3 × as reddish brown. long as area superomedia. Wings fully developed. Body length 4.0-4.5 mm, fore wing length 3.1- Nervellus broken in its lower third. Hind femora 3.4 mm. 3.4-3.5 × as long as wide. Male similar to female. Flagelum with 22 seg- Metasoma longish-oval. Petiolus hardly com- ments, fi rst segment 2.8 × as long as wide, 11th or 12th pressed dorsoventrally. Distance between spiracles segment square in lateral view, longish-oval tyloids of the fi rst metasomal segment about twice as long present on (7)8th-10th segment. Metasoma more as distance between the spiracle and hind corner of slender (metasomal tergum II 1.0-1.1 × as long as the segment. Thyridia wide, medially removed from wide, tergum III 1.5-1.6 × as wide as long), its sculp- the base of the second tergum by about two-thirds ture more robust. Body brown, clypeus often whit- their width, 1.5 × as wide as the interthyridial space. ish, apex of fore femora and stripes on inner sides Metasomal tergum II 1.2-1.5 × (rarely to 1.1 ×) as long of all tibiae reddish. as wide, tergum III 1.1-1.3 × as wide as long (Fig. 5). Body length 4.0-4.5 mm, fore wing length 3.1- Body weakly sculptured. Mesoscutum and scu- 3.3 mm. tellum fi nely and sparsely punctate; pronotal lobes shining, rugose ventrally. Mesopleura longitudi- Biology. Parasitoid of Luffi a ferchaultella (Stephens), nally rugose (sometimes only in their apical half), Narycia duplicella (Goeze) and Siederia sp. speculum smooth. Petiolus of the fi rst metasomal segment chagrinate, postpetiolus weakly longitudi- Distribution. Sweden, Poland, United Kingdom, nally striate. Metasomal tergum II and basal third the Netherlands. of tergum III obviously chagrinate with obscure

70 ©Zoologische Staatssammlung München/Verlag Friedrich Pfeil; download www.pfeil-verlag.de scattered punctures, sculpture becomes more smooth Differential diagnosis. T. prominulus is most close- towards apex of metasoma. ly related to T. fuscipes, from which it differs by Body brown. Head brown, clypeus and median having more stout metasoma, reddish brown tegu- part of face somewhat lighter, mandibles (except lae and presence of tyloids on the male fl agellum. base and teeth) yellowish brown. Antennae brown, fl agellum reddish ventrally, palps light brown. Description Tegulae whitish. Legs reddish brown, apices of fore Female. Clypeus weakly fl attened in its apical third. and middle femora and stripes on inner sides of fore Lower edge of the clypeus very slightly rounded, and middle tibiae reddish, all tarsi reddish, their last almost straight medially, apical margin hardly vis- segments infuscate. Metasoma brown with the very ible. Head slightly narrowed behind eyes in dorsal apex of tergum I medially, thyridia and hind margins view. Temples moderately enlarged at the middle, of tergum II and usually also of tergum III yellowish, with their maximum length slightly less than trans- tergum VII reddish brown. verse ocular diameter. Flagellum with 24 segments Body length 4.0-5.0 mm, fore wing length 2.9- (23-25 according to the original description), fi rst 3.5 mm. segment 2.5-2.6 × as long as wide, 11th or 12th segment Male similar to female. Flagellum with 23-26 square in lateral view. Upper mandibular tooth segments, fi rst segment 2.6-2.7 × as long as wide, obviously longer and wider than the lower tooth. one or two segments before apex square in lateral Hypostomal carina meets the genal carina before the view. No visible tyloids were found. Metasomal mandibular base at a distance hardly less than width tergum II 1.3-1.4 × as long as wide, tergum III 1.1- of the latter. 1.2 × as wide as long. Clypeus (usually), central part Distinct notauli present in the basal third of of mandibles and markings on the lower side of mesoscutum. Area superomedia about 1.3 × as wide scape often whitish. as long. Area petiolaris about 3 × as long as area Body length 3.5-5.0 mm, fore wing length 3.1- superomedia. Wings fully developed. Nervellus 3.9 mm. broken hardly below the middle. Hind femora 3.3- 3.4 × as long as wide. Biology. Parasitoid of Anaproutia raiblensis, Bacotia Metasoma longish-oval. Distance between spir- claustrella (Bruand), Brevantennia styriaca Meier, acles of the fi rst metasomal segment about twice as Siederia listerella as well as of Dahlica, Eosolenobia and long as distance between the spiracle and hind Siederia spp. (including D. charlottae, D. goppenstein- corner of the segment. Thyridia medially removed ensis (Sauter), D. nickerlii (Heinemann), D. triquetrel- from the base of second metasomal tergum by about la, E. manni Zeller and S. alpicolella (Rebel)) and two-thirds their width, twice as wide as the inter- Tale poria tubulosa (Retzius). thyridial space. Metasomal tergum II 1.1-1.2 × as Distribution. Sweden, Finland, Georgia, Italy, long as wide, tergum III 1.4-1.5 × as wide as long Austria, Switzerland, Germany (Lower Saxony, (Fig. 6). Bavaria). A boreal-montane species. Body moderately sculptured. Frons shining, very fi nely and sparsely punctate. Pronotal lobes punctate, rugose ventrally. Mesopleura longitudinally rugose 6. Trachyarus prominulus Diller to rugose-punctate, speculum smooth with more or Fig. 6 less distinct punctures. Postpetiolus of the fi rst metasomal segment rugose, petiolus rugose-punc- Trachyarus prominulus Diller, 1989: 282. Holotype: W, tate. Metasomal tergum II and basal half of tergum Austria, “Steiermark, Österreich”, “Frauenalpe” III obviously chagrinate, terga II-IV(V) punctate with (ZSM, not studied). large but obscure punctures, sculpture disappears Material. Paratypes: 2WW, Austria, “Paratypus”, “Steier- towards apex of metasoma. mark, Österreich, Parasit e Psychidae, leg. H. Meier”, Body brown. Head brown, clypeus and median “Frauenalpe, b. Muren, Mai 1966, 2000 m S. meierella part of face a little lighter, mandibles (except base Sied.”, “Paratypus Trachyarus prominulus W Diller, 1989” and teeth) yellowish brown. Antennae brown, fl ag- (ZSM); 2MM, Austria, “Paratypus”, “Steiermark, Öster- ellum slightly reddish ventrally, palps light brown. reich, Parasit e Psychidae, leg. H. Meier”, “Frauenalpe Tegulae reddish brown. Legs brown, apices of fore e S. meierella Sied.”, “Paratypus Trachyarus prominulus M femora and all tibiae and tarsi reddish (except last Diller, 1989” (ZSM). segments), fore tibiae somewhat darker on outer Additional material. 10MM from Hampshire (United sides. Metasoma brown with the very apex of tergum Kingdom, NHM) probably also belong to this species. I medially, thyridia and hind margins of terga II-III segments yellowish, apex of tergum VII reddish brown.

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Body length 5.0 mm, fore wing length 3.3- Distinct notauli present in the basal third of 3.5 mm. mesoscutum. Area superomedia heart-shaped, some- Male. Similar to female. Apical third of the times not separated from area posteromedia, about clypeus almost not fl attened. Flagellum with 22-24 as long as wide. Area petiolaris about twice as long segments, fi rst segment 2.7-2.8 × as long as wide, as area superomedia. Wings fully developed or re- longish-oval tyloids present on (8)9th-10(11)th seg- duced, in the latter case stretching to the middle of ment. Side carinae of area posteromedia more or less the second metasomal segment. Nervellus broken visible. Hind femora 3.7-3.8 × as long as wide. Meta- slightly below the middle. Hind femora 3.7-3.8 × as somal tergum II 1.0-1.1 × as long as wide, tergum III long as wide. 1.1-1.3 × as wide as long. Sculpture more developed Metasoma longish-oval. Petiolus not compressed (i.e., more dense and coarse) than in female. Body dorsoventrally. Distance between spiracles of the brown to dark brown. Clypeus often whitish or with fi rst metasomal segment twice as long as distance white markings. Fore tibiae entirely reddish. Last between the spiracle and hind corner of the segment. metasomal tergum brown. Thyridia medially removed from the base of meta- 2 Body length 4.0-5.5 mm, fore wing length 2.9- somal tergum II by about /3 their width, slightly 4.2 mm. wider than the interthyridial space. Metasomal tergum II 1.2-1.3 × as long as wide, tergum III 1.1- Biology. Parasitoid of Siederia meierella (Sieder). 1.2 × as wide as long. Body moderately sculptured. Frons shagrinated Distribution. Austria. with obscure scattered punctures. Mesoscutum and scutellum shagrinate, whole scutellum and medio-

apical part of mesoscutum also sparsely punctate, 7. Trachyarus bacillatus Gokhman, spec. nov. pronotal lobes rugose. Mesopleura rugose, speculum Fig. 7 smooth with a few punctures. Postpetiolus of the fi rst metasomal segment irregularly longitudinally Types. Holotype: W, Russia, Kavalerovo, Primorsky district, from Eosolenobia sp., 25.V.1983 (Solyanikov). – striate, petiolus shagrinate. Metasomal terga II-V Paratypes: 1W, Russia, Primorsky district, Kentsuche distinctly shagrinate, sculpture becomes more River, from Eosolenobia sp., without date (Solyanikov); smooth towards apex of metasoma. 2MM, same data; 1M, Russia, Kavalerovo, Primorsky Body dark brown. Head dark brown, clypeus district, from Eosolenobia sp., 13.V.1983 (Solyanikov); 1M, and mandibles a little lighter. Antennae reddish same locality, from Eosolenobia sp. (?), 10.V.1983 (Soly- brown, palpi light brown. Tegulae reddish. Legs anikov). The material is deposited in the ZIN collec- reddish brown, middle and hind coxae brownish. tion. Metasoma brown with the very apex of tergum I Differential diagnosis. Among Trachyarus species medially, thyridia and hind margins of metasomal having lower mandibular tooth longer than the up- terga II-III (sometimes also of terga IV-V) and apex per tooth, T. bacillatus, spec. nov. is most closely of tergum VII reddish brown. related to T. edilleri, spec. nov. and T. hemichneumono- Body length 5.5 mm, fore wing length 2.0- ides, spec. nov., from which it differs by having more 3.6 mm. fl attened clypeus (especially in its basal part) and Male similar to female. Flagellum with 24-25 larger body size. segments, fi rst segment 2.8-2.9 × as long as wide, 9th-11(12)th fl agellar segment with longish-oval ty- th th Description loids, 17 or 18 segment square in lateral view. Hypostomal carina meets the genal carina just at the Female. Clypeus somewhat fl attened in its apical corner of the mandibular base. Metasomal tergum half. Lower edge of the clypeus hardly rounded, II 1.3-1.4 × as long as wide, tergum III 1.1-1.2 × as straight medially, with a narrow apical margin. Head wide as long. Body sculpture more robust than in slightly narrowed behind eyes in dorsal view. Tem- female. Clypeus, apex of scape (ventrally) and often ples slightly enlarged at the middle, with their central parts of mandibles whitish. maximum length a little less than transverse ocular Body length 5.0-5.5 mm, fore wing length 3.7- diameter. Flagellum with 25 segments, fi rst segment 4.9 mm. 2.5-2.6 × as long as wide, 10th or 11th segment square in lateral view. Lower mandibular tooth distinctly Biology. Parasitoid of Eosolenobia sp. longer than the upper tooth (Fig. 7). Hypostomal carina meets the genal carina before the mandibular Distribution. Russia (Primorsky district). base at its width.

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8. Trachyarus decipiens Gokhman, spec. nov. (rarely with 23) segments (10th or 11th segment square in lateral view), metasomal tergum II not longer than Types. Holotype: W, Austria, “Steiermark, Österreich, its width and tergum III 1.3-1.4 × as wide as long. Parasit e Psychidae, leg. H. Meier”, “Seckauer Alpen, The male of T. decipiens, spec. nov. is closest to that Sechs Zinken, 2000 m, 1959, Sol. triquetrella Hbn.”. – of T. solyanikovi, spec. nov., from which it differs by Paratypes: Austria: 1W, 1M, same data as holotype; 1W, having longer fl agellum with 24-25 segments, wid- same data, “Trachyarus brevipennis Rom. W det. E. Diller er mandibles and more slender metasomal tergum 87”; 2MM, “Österreich, Seckauer Zinken, 2000 m, Juni III (1.3-1.4 × as wide as long). 1957, leg. Meier”, “ex Sol. triquetrella”; M, “Steiermark, Österreich, Parasit e Psychidae, leg. H. Meier”, “Leoben, Description Häuslberg, 1960, e Brev. styriaca Meier”; 2MM, “Steier- mark, Österreich, Parasit e Psychidae, leg. H. Meier”, Female. Lower edge of the clypeus very slightly “Permegg, 1960, Sol. triquetrella Hbn. parth.”; 1W, “Steier- rounded, straight medially, with a visible apical mark, Österreich, Parasit e Psychidae, leg. H. Meier”, margin. Head slightly narrowed behind eyes in “Utschgraben b. Bruck, 1963, B. styriaca Meier”; 1W, dorsal view. Temples moderately enlarged at the “Steiermark, Österreich, Parasit e Psychidae, leg. H. middle, with their maximum length distinctly less ” “ ” Meier , Frohnleiten, Buche, 1960, S. nickerlii Hein. ; 1W, than transverse ocular diameter. Flagellum hardly “Parasit e Psychidae, leg. H. Meier”, “Knittelfeld, Steier- enlarged in apical half, with 24-26 (rarely 23) seg- mark, Österreich”, “Eppenstein, 1964, Sol. charlottae th Meier”; 2MM, “Österreich, Steiermark, Reiting, 1500 m, ments, fi rst segment 2.5-2.6 × as long as wide, 10 or th 1956, leg. Sieder, ex Sol. klimeschi”; 1W, “Österreich, 11 segment square in lateral view. Lower man- Parasit e Psychidae, leg. H. Meier”, “Kärnten, Koralpe, dibular tooth distinctly longer and wider than the 1800-2000 m, 1967, S. triquetrella Hbn.”; 1M, “Österreich, upper tooth. Hypostomal carina meets the genal Kärnten, Urbitz Kogel, 200 m, 13.VI.1951, leg. Sieder”, carina before the mandibular base at two-thirds its “ex 2 jährig Sol. thurneri”; 1W, “Österreich, Parasit e width. Psychidae, leg. H. Meier”, “Linz, St. Magdalena, 1962, Distinct notauli present in the basal third of Sol. triquetrella Hbn.”; 2MM, “Österreich, Parasit e Psychi- mesoscutum. Area superomedia as wide as long, dae, leg. H. Meier”, “Linz, Kaltenberg, 1962, Sol. trique- weakly separated from area posteromedia. Area trella Hbn.”, 1W, same data, “Trachyarus brevipennis Rom. petiolaris about 2.5 × as long as area superomedia. W det. E. Diller 87”; 1W, “Waidhofen/Y., Austria, leg. Wings fully developed. Nervellus broken in its Lichtenberger”, “ex Solenobia triquetr. parth.”, “Trachya- rus corvinus Thom. W det. E. Diller 84”; 1M, “ex Solenobia lower third. Hind femora 3.4-3.5 × as long as wide. triquetr. parth., 1.4.1978, ex 14.5.78”, “Waidhofen/Y., Metasoma longish-oval. Distance between spir- Austria, leg. Lichtenberger”. Germany: 1W, “Bayern- acles of the fi rst metasomal segment about 2.5 × as München, Aubinger Lohe, 1.VIII.1987, leg. Diller 550 m”; long as distance between the spiracle and hind 1M, “Bayern, Deggendorf, A.V.1963, leg. Schätz”; 1M, corner of the segment. Thyridia medially removed “Bayern, Nürnberg, Altdorf, 3.V.73, leg. Schätz”; 1M, from the base of metasomal tergum II by about two- “Bav. mer. Ammergauer Berge, Frieder Gebiet, 1700- thirds their width, approximately twice as wide as 2000 m, e-e.12.VI.1949, F. Daniel leg.”. Switzerland: 1W, the interthyridial space. Metasomal tergum II 1.0- “Parasit e Psychidae, leg. H. Meier”, “Schweiz, Wallis, 1.1 × as wide as long, tergum III 1.3-1.4 × as wide as Simplon, 1963, S. goppensteinensis Saut.”. Italy: 2MM, long. “Parasit e Psychidae, leg. H. Meier”, “Matajur, 1961, Sol. Body moderately sculptured. Head shining, side juliella Rbl.”; 1W, same data, “Trachyarus brevipennis Rom. W det. E. Diller 87”; 1W, “Parasit e Psychidae, leg. H. parts of face coarsely punctate, median part of the Meier”, “Italien, Mt. Grappa, 1960, 1000-1700 m, e Sied. face fi nely and sparsely punctate, clypeus with a few meierella Sied.”; 2WW, same data, “Trachyarus brevipennis fi ne punctures. Frons fi nely and sparsely punctate. Rom. W det. E. Diller 87”; 1M, “Italia (Bolzano), Sarntal, Mesoscutum and scutellum fi nely and sparsely 1250 m, 24-VI.1976, C. J. Zwakhals”; 2MM, “Italien, Kur- punctate; pronotal lobes more densely and coarsely tatsch, Oberfennberg, 1300 m, 1.6.1976, Diller leg.”. Slo- punctate, rugose ventrally. Mesopleura longitudi- venia: 1W, “Istrien, Snežnik, 1971, 1200-1300 m, S. tri- nally rugose, speculum smooth with a few punctures. quetrella Hbn.”. Romania: 2WW, 3MM, “Parasit e Psychi- Metasomal tergum I rugose-punctate, petiolus dae, leg. H. Meier”, “Rumanien, Süd-Karpaten, Bombes- weakly striate. Terga II-III (sometimes also the very ti-Jiu 300 m”, “Südabhang Pietroasa, Sol. triquetrella base of tergum IV) obviously chagrinate, sculpture Hbn.”; 1W, same data, “Trachyarus brevipennis Rom. W becomes more smooth towards apex of metasoma. det. E. Diller 87”. Body brown. Head dark brown, mandibles Differential diagnosis. Among Trachyarus species (except teeth) and often clypeus reddish brown. having the lower mandibular tooth longer than the Antennae brownish, fl agellum reddish ventrally, upper tooth, T. decipiens, spec. nov. is the only species palps light brown. Tegulae reddish to whitish. Legs having a combination of the following characters in reddish, hind coxae, all femora, trochanters I and the female: long and slender fl agellum with 24-26 last tarsal segments infuscate. Metasoma brown with

73 ©Zoologische Staatssammlung München/Verlag Friedrich Pfeil; download www.pfeil-verlag.de the very apex of tergum I medially, thyridia and 1960, Sol. triquetrella Hbn. parth.”; 1M, “Steiermark, Ös- hind margins of terga II-III and often of tergum IV terreich, Parasit e Psychidae, leg. H. Meier”, “Proleb, yellowish, tergum VII reddish brown. 1956, Sied. pineti Z.”; 1M, “Steiermark, Österreich, Parasit Body length 4.0-5.0 mm, fore wing length 3.4- e Psychidae, leg. H. Meier”, “Hinterlobening, Stefan 3.5 mm. Serpentin, e S. nickerli Hein.”; 1M, “Steiermark, Öster- reich, Parasit e Psychidae, leg. H. Meier”, “Seebergkogl, Male similar to female. Flagellum with 24-25 Hochalpe, 1900 m, e Sol. triquetrella Hbn.”; 1M, “Steier- segments, fi rst fl agellar segment 2.5 × as long as mark, Österreich, Parasit e Psychidae, leg. H. Meier”, wide, all fl agellar segments longer than wide in “Seckauer Alpen, Sechs Zinken, 2000 m, 1954, Sol. trique- th th lateral view, 8 -11 segment with longish-oval ty- trella Hbn.”; 1M, “Steiermark, Österreich, Parasit e Psy- loids. Body sculpture more robust. Clypeus and chidae, leg. H. Meier”, “Ameringkogel, 1900 m, 1967, manbibles (except teeth) white. Sol. triquetrella Hbn.”; 1M, “Steiermark, Österreich, Para- Body length 4.5-5.5 mm, fore wing length 3.7- sit e Psychidae, leg. H. Meier”, “Judenburg, 1954, Sol. 4.1 mm. klimeschi Sied.”; 1M, “Österreich, Parasit e Psychidae, leg. H. Meier”, “Hohe Wand, 1965, Sied. pineti Z.”; 1M, “Öster- Biology. Parasitoid of Brevantennia styriaca, Dahlica reich, Parasit e Psychidae, leg. H. Meier”, “Kärnt. Pölling, charlottae, D. goppensteinensis, D. klimeschi, D. nicker- 1959, Brev. reliqua Sied.”; 1M, “Österreich, Villach, Warm- bad, Mai 1957, leg. Meier”, “Brev. santicensis”; 1M, “Öster- lii, D. triquetrella, Postsolenobia juliella (Rebel) and reich, Knittelfeld, 1957, leg. Meier”, “ex Brev. styriaca”. Siederia meierella. Italy: 1W, 2MM, “Parasit e Psychidae, leg. H. Meier”, Distribution. Austria, Germany (Bavaria), Switzer- “Matajur, 1961, Sol. juliella”; 1W, “Italien-Nord, Matajur land, Italy (Alps), Romania (Carpathians), Slovenia 150 m, 14.V.1961, leg. Sieder”, “ex Bank. juliella Rbl.”, (Istria). A montane species. “Trachyarus corvinus Thoms. det. E. Diller 1977 W”; 1W, 2MM, “Italien, Mt. Generoso, 1700 m, Juni 1957, ex Sol. siederi, leg. Meier”. Romania: 1M, “Rumänien, Süd-Kar- paten, Bombesti-Jiu 300 m”, “Südabhang Pietroasa, Sol. 9. Trachyarus edilleri Gokhman, spec. nov. triquetrella Hbn.”, “Parasit e Psychidae, leg. H. Meier”; Fig. 8 1W, same data, “Trachyarus brevipennis Rom. W det. E. Diller 87”. One W paratype is deposited in the ZIN col- Types. Holotype: W, Austria, “Steiermark, Österreich, lection. Parasit e Psychidae, leg. H. Meier”, “Leoben, Häuslberg, Differential diagnosis. The female of T. edilleri, 1960, e Brev. styriaca Meier”. – Paratypes: Austria: 1W, “Steiermark, Österreich, Parasit e Psychidae, leg. H. spec. nov. is closest to that of T. hemichneumonoides, Meier”, “Österreich, Knittelfeld, 1957, leg. Meier”; “ex spec. nov., but in T. edilleri fl agellum and metaso- th th Brev. styriaca”; 1W, 2MM, “Österreich, Steiermark, Leoben, mal tergum II are stouter (9 or 10 fl agellar segment 1956, leg. Sieder ex Brev. styriaca”, 7MM, same data, “Kai- square in lateral view; tergum II 1.1-1.2 × as long as sersberg”; 2WW, same data, “Kaisersberg”, “Trachyarus wide). The male is similar to that of T. bacillatus, spec. brevipennis Rom. W det. E. Diller 87”; 1M, “Österreich, nov., but smaller, with evenly convex clypeus. Leoben, Kaisersberg, Mai 1957, leg. Meier”, “Sol. pineti”; 1M, “Österreich, Steiermark, Seiz, Liesingtal, 1956, leg. Description Sieder ex Brev. styriaca”; 1M, “Steiermark, Österreich, Female. Lower edge of the clypeus very slightly Parasit e Psychidae, leg. H. Meier”, “Leoben, Seiz, 1956, Brev. styriaca Meier”; 1M, “Steiermark, Österreich, Parasit rounded, straight medially, with a visible apical e Psychidae, leg. H. Meier”, “Leoben, St. Stefan, 1954, margin. Head slightly narrowed behind eyes in Sied. pineti Z.”; 1M, “Steiermark, Österreich, Parasit e dorsal view. Temples moderately enlarged at the Psychidae, leg. H. Meier”, “Leoben, Trabach, 1954, Brev. middle, with their maximum length distinctly less styriaca Meier”; 2MM, “Steiermark, Österreich, Parasit e than transverse ocular diameter. Flagellum with Psychidae, leg. H. Meier”, “Preggraben, 1958, B. styriaca 23-26 segments, fi rst segment 2.4-2.5 × as long as Meier”; 1M, “Österreich, Steiermark, Oberzeiring, 1956, wide, 9th or 10th segment square in lateral view. leg. Sieder”; 1M, “Österreich, Steiermark, Kofl ach, leg. Lower mandibular tooth distinctly longer and Meier, ex Brev. styriaca”; 1W and 15MM, “Steiermark, wider than the upper tooth. Hypostomal carina meets Österreich, Parasit e Psychidae, leg. H. Meier”, “Leoben, the genal carina before the mandibular base at a Häuslberg, 1960, e Brev. styriaca Meier”, 1W, 1M, “Steier- distance of its width. mark, Österreich, Parasit e Psychidae, leg. H. Meier”, “ Häuslberg – Leoben, 1963, B. styriaca Meier”; 2WW, Distinct notauli present in the basal third of “Steiermark, Österreich, Parasit e Psychidae, leg. H. mesoscutum. Area superomedia about as wide as Meier”; 1M, “Österreich, Steiermark, St. Michael bei long, weakly separated from area posteromedia. Leoben, 1956, leg. Sieder ex Sol. klimeschi”; 2WW. 1M, Area petiolaris about 2.5 × as long as area supero- “Steiermark, Österreich, Parasit e Psychidae, leg. H. media. Wings fully developed. Nervellus broken in Meier”, “Gulsen, 1956, Sol. pineti Z.”; 1M, “Steiermark, its lower third. Hind femora 3.5-3.6 × as long as Österreich, Parasit e Psychidae, leg. H. Meier”, “Pernegg, wide.

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Metasoma longish-oval. Distance between spir- 10. Trachyarus hemichneumonoides Gokhman, acles of the fi rst metasomal segment about twice as spec. nov. long as distance between the spiracle and hind Fig. 9 corner of the segment. Thyridia medially removed from the base of metasomal tergum II by about two- Holotype: W, Austria, “Koralmschutzhaus m 1940, Ko- thirds their width, approximately 1.5 × as wide as ralpe, K., 8.VI.1957, K. Rath”. the interthyridial space. Metasomal tergum II 1.1- Differential diagnosis. T. hemichneumonoides, spec. 1.2 × as long as wide, tergum III 1.1-1.2 × as wide as nov. is most closely related to T. edilleri, spec. nov., long (Fig. 8). but in T. hemichneumonoides, spec. nov. fl agellum and Body moderately sculptured. Head shining, side metasomal tergum II are more slender (11th or 12th parts of face densely and coarsely punctate, median fl agellar segment square in lateral view, tergum II part of the face and clypeus more fi nely punctate. 1.3-1.4 × as long as wide). Frons fi nely punctate. Mesoscutum and scutellum fi nely and sparsely punctate; pronotal lobes shining Description and fi nely punctate in their upper part, longitudinally rugose ventrally. Mesopleura longitudinally Female. Clypeus distinctly convex. Lower edge of rugose, speculum smooth, sometimes with obscure the clypeus slightly rounded, with a visible apical rugosity. Metasomal segment I chagrinate, petiolus margin. Head almost not narrowed behind eyes in longitudinally rugose. Metasomal tergum II and dorsal view. Temples very slightly enlarged ventrad, basal half of tergum III (sometimes also the very with their maximum length distinctly less than base of tergum IV) obviously chagrinate, sculpture transverse ocular diameter. Flagellum fi liform, with becomes more smooth towards apex of metasoma. 25 segments, fi rst segment 2.3-2.4 × as long as wide, Body brown. Head brown, clypeus somewhat 11th or 12th segment square in lateral view. Lower lighter, mandibles (except base and teeth) yellowish mandibular tooth distinctly longer and slightly brown. Antennae brownish, fl agellum slightly red- wider than the upper tooth. Hypostomal carina meets dish ventrally, palps light brown. Tegulae reddish the genal carina before the mandibular base at ap- to whitish. Legs reddish, middle and hind coxae, all proximately one-third its width. femora and last tarsal segments slightly infuscate. Distinct notauli present in the basal fourth of Metasoma brown with the very apex of tergum I mesoscutum. Area superomedia slightly longer than medially, thyridia and hind margins of terga II-III wide, weakly separated from area posteromedia. and often of tergum IV yellowish, tergum VII reddish Area petiolaris about twice as long as area supero- brown. media. Wings fully developed. Nervellus broken at Body length 3.5-5.0 mm, fore wing length 2.3- the middle. Hind femora about 3.5 × as long as 3.4 mm. wide. Male similar to female. Flagellum with 23-24 Metasoma longish-oval. Distance between spir- segments, fi rst segment 2.8 × as long as wide, 8th- acles of the fi rst metasomal segment twice as long 11(12)th segment with longish-oval tyloids. Body as distance between the spiracle and hind corner of sculpture more robust than in female. Clypeus and the segment. Thyridia medially removed from the often mandibles (except teeth) white, scape with base of metasomal tergum II by about two-thirds whitish marking. Legs infuscate (except for reddish their width, approximately twice as wide as the inner surfaces of fore femora, the very apex of mid- interthyridial space. Metasomal tergum II 1.3-1.4 × dle femora, fore and middle tibia). as long as wide, tergum III 1.0-1.1 × as wide as long Body length 3.5-4.5 mm, fore wing length 2.8- (Fig. 9). 3.1 mm. Body moderately sculptured. Head shining, side parts of face moderately punctate, median part of Biology. Parasitoid of Brevantennia reliqua Sieder, the face fi nely and densely punctate, clypeus with B. siederi, B. styriaca, Dahlica klimeschi, D. nickerlii, a few punctures. Frons fi nely and sparsely punctate. D. triquetrella, Postsolenobia juliella and Siederia lis- Mesoscutum and scutellum fi nely and sparsely terella. punctate; pronotal lobes more densely and coarsely punctate, rugose in their lower half. Mesopleura Distribution. Austria, Italy, Romania. A montane longitudinally rugose, speculum rugose-punctate. species. Metasomal tergum I shagrinate, postpetiolus later- ally striate. Tergum II and basal two-thirds of tergum III obviously chagrinate, sculpture becomes more smooth towards apex of metasoma.

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Body brown. Head dark brown, mandibles pressed dorsoventrally. Distance between spiracles (except teeth) and clypeus reddish brown. Antennae of the fi rst metasomal segment twice as long as reddish brown, their scape, pedicel and a few basal distance between the spiracle and hind corner of the segments of fl agellum infuscate, palps light brown. segment. Thyridia medially removed from the base Tegulae whitish. Legs reddish brown, hind coxae of metasomal tergum II by about one-third their and all last tarsal segments infuscate. Metasoma width, approximately 1.5 × as wide as the interthy- brown with the very apex of metasomal tergum I ridial space. Metasomal tergum II 1.2-1.3 × as long medially and hind margins of terga II-IV reddish, as wide, tergum III 1.2-1.3 × as wide as long. tergum VII reddish brown. Body weakly sculptured. Frons smooth and Body length 4.5 mm, fore wing length 3.1 mm. strongly shining. Mesoscutum and scutellum strong- Male unknown. ly shining, very obscurely and fi nely punctate; pronotal lobes shining, longitudinally rugose in their Biology. Parasitoid of an undetermined Psychidae. lower half. Mesopleura weakly rugose, speculum smooth. Postpetiolus of the fi rst metasomal segment Distribution. Austria. longitudinally striate, petiolus rugose-punctate. Metasomal tergum II and basal half of tergum III obviously chagrinate, sculpture becomes more 11. Trachyarus khrulevae Gokhman, spec. nov. smooth towards apex of metasoma. Body brown. Head brown, clypeus, mandibles Types. Holotype: W, Russia, Wrangel Island, 31.VII.1983 (O. Khruleva). – Paratypes: 4WW, same locality and col- (except teeth) lighter, up to yellowish brown. Anten- lector, 1-23.VIII.1983; W, Russia, Buryatiya, Vitim River, nae brown, scape and pedicel sometimes reddish Baissa, 7.VII.1983 (V. Zherikhin, D. Shcherbakov). The ventrally, palps light brown. Mesoscutum and mes- material is deposited in the ZMMU collection. osternum sometimes reddish brown. Tegulae light reddish. Legs reddish. Metasoma brown with the Differential diagnosis. Among Trachyarus species very apex of tergum I medially, thyridia and hind having the lower mandibular tooth longer than the margins of terga II-III (sometimes also of terga IV-V) upper tooth, T. khrulevae, spec. nov. is the only spe- yellowish, tergum VII reddish brown. cies having a combination of the following characters: Body length 4.5-5.5 mm, fore wing length 0.8- metasomal tergum II longer than its width and stout 1.3 mm. th th fl agellum (5 or 6 fl agellar segment square in lat- Male unknown. eral view). Biology. Unknown. Description Distribution. Russia (Wrangel Island and Buryati- Female. Lower edge of the clypeus moderately ya). rounded, with a visible apical margin. Head not narrowed behind eyes in dorsal view. Temples distinctly enlarged ventrad, with their maximum 12. Trachyarus parvipennis Gokhman, spec. nov. length slightly exceeding transverse ocular diameter. Fig. 10 Flagellum with 22-24 segments, fi rst segment 1.6- 1.7 × as long as wide, 5th or 6th segment square in Holotype: W, Russia, Moscow region, Bronnitsy station, lateral view. Lower mandibular tooth distinctly from Solenobia sp., collected 1.V.1982, emerged 6.V.1982 longer and wider than the upper tooth. Hypos- (Solyanikov). Holotype deposited in the ZIN collec- tomal carina meets the genal carina before the tion. mandibular base at about its width. Differential diagnosis. Among the described Tra- Distinct notauli present in the basal fourth of chyarus species, wings are also strongly reduced only mesoscutum. Horizontal part of propodeum com- in T. brachypterator, from which T. parvipennis, spec. paratively long. Area superomedia about 1.2-1.3 × nov. well differs by the structure of mandibles, as long as wide. Area petiolaris about 1.5 × as long number of fl agellar segments, proportions of meta- as area superomedia. Wings partly reduced, not somal terga and general colour of the body and extending beyond apex of the fi rst metasomal seg- legs. ment. Wing venation variable, 2 to 3 closed cells present in the fore wing, pterostigma subapical. Description Nervellus reclivous, continually curved. Hind femora 3.1-3.2 × as long as wide. Female. Lower edge of the clypeus rounded, with Metasoma longish-oval. Petiolus hardly com- a visible apical margin. Head not narrowed behind

76 ©Zoologische Staatssammlung München/Verlag Friedrich Pfeil; download www.pfeil-verlag.de eyes in dorsal view. Temples distinctly enlarged 13. Trachyarus punctigaster Gokhman, spec. nov. ventrad, with their maximum length equal to transverse ocular diameter. Flagellum slightly enlarged Types. Holotype: W, Italy, “Parasit e Psychidae, leg. H. in apical half, with 23 segments, fi rst segment 2.2- Meier”, “Matajur, 1961, S. juliella Rbl.”. – Paratypes: 1W, 2.3 × as long as wide, 5th or 6th segment square in same data as holotype, “Trachyarus brevipennis Rom. lateral view. Lower mandibular tooth distinctly W det E. Diller 1987”; 1W, “Parasit e Psychidae, leg. H. longer than the upper tooth. Hypostomal carina Meier”, “Tessin, Mt Generoso, 1963, B. siederi Sauter”. meets the genal carina before the mandibular base Differential diagnosis. T. punctigaster, spec. nov. is at two-thirds its width. most closely related to T. subtilipiceus, spec. nov., but Mesoscutum moderately transverse, distinct in T. punctigaster, spec. nov. the fl agellum is stouter notauli present in its basal third. Area superomedia (11th or 12th flagellar segment square in lateral about as wide as long. Area posteromedia approxi- view). mately twice as long as area superomedia. Wings strongly reduced, not extending beyond apex of Description propodeum. Fore wing bears small triangular stigma on the top and the only closed cell evidently formed Female. Clypeus convex. Lower edge of the clypeus by median and cubital veins. Nervellus reclivous, very slightly rounded, straight medially, apical continually curved. Hind femora 3.3-3.4 × as long as margin hardly visible. Head somewhat infl ated, wide. almost not narrowed behind eyes in dorsal view. Metasoma oval. Petiolus distinctly compressed Temples moderately enlarged at the middle, with dorsoventrally. Postpetiolus wide, distance between their maximum length approximately equal to spiracles of the fi rst metasomal segment about 3 × transverse ocular diameter. Flagellum fi liform, with as long as distance between the spiracle and hind 22-23 segments, fi rst segment 2.7-2.8 × as long as th th corner of the segment. Thyridia not inclined, very wide, 11 or 12 segment square in lateral view. wide, removed from the base of the metasomal Lower mandibular tooth distinctly longer and tergum II by about one-third their width, approxi- wider than the upper tooth. Hypostomal carina meets mately 3 × as wide as the interthyridial space. Meta- the genal carina before the mandibular base at half somal tergum II 1.4-1.5 × as wide as long, tergum III its width. 2.5-2.6 × as wide as long (Fig. 10). Distinct notauli present in the basal fourth of Body shining, moderately sculptured. Side parts mesoscutum. Area superomedia about as wide as of face densely and obscurely punctate, frons hard- long, weakly separated from area posteromedia. ly chagrinate. Mesoscutum and scutellum weakly Area petiolaris approximately twice as long as area shining, pronotal lobes chagrinate above, longitudi- superomedia. Wings fully developed. Nervellus nally rugose ventrally. Mesopleura longitudinally broken in its lower third. Hind femora 3.0-3.1 × as rugose, speculum smooth. Petiolus of the fi rst meta- long as wide. somal segment chagrinate, postpetiolus irregularly Metasoma longish-oval. Distance between spir- longitudinally striate. Metasomal terga II-III and acles of the fi rst metasomal segment about twice as apical half of tergum IV roughly chagrinate, sculp- long as distance between the spiracle and hind ture becomes more smooth towards apex of meta- corner of the segment. Thyridia medially removed soma. from the base of metasomal tergum II by about half Body brown. Head brown, mandibles (except their width, approximately equal to the interthy- teeth) and clypeus yellowish brown. Antennae ridial space. Metasomal tergum II 1.1-1.2 × as wide brown (scape reddish ventrally), palps whitish. as long, tergum III 1.8-1.9 × as wide as long. Thorax brown with yellow spots on pronotal lobes Body moderately sculptured. Head shining, side (above their lower corner), mesopleura (ventrad to parts of face coarsely punctate, median part of the sternauli) and propodeum near areae posteroexter- face fi nely and sparsely punctate, clypeus with a few nae. Tegulae white. Legs yellowish brown. Metaso- fi ne punctures. Frons almost smooth. Mesoscutum ma brown with thyridia and hind margins of terga and scutellum fi nely and sparsely punctate; prono- II-IV yellowish brown, tergum VII reddish brown. tal lobes rugose in their lower two-thirds. Mesop- Body length 3.5 mm, fore wing length 0.7 mm. leura longitudinally rugose, speculum smooth. Male unknown. Metasomal tergum I chagrinate, postpetiolus sometimes striate. Terga II-III more or less coarsely Biology. Parasitoid of Dahlica or Siederia sp. punctate apically and obviously chagrinate in their basal halves (tergum II sometimes fully chagrinate), Distribution. Russia (Moscow region). sculpture becomes more smooth towards apex of metasoma.

Body brown. Head dark brown, mandibles Differential diagnosis. Among Trachyarus species (except teeth) and clypeus reddish brown. Antennae having the lower mandibular tooth longer than the brownish, fl agellum reddish beneath, palps light upper tooth, the female of T. solyanikovi, spec. nov. brown. Tegulae reddish. Legs brownish, stripes on is closest to those of T. subtilipiceus, spec. nov. and inner sides of fore tibiae reddish. Metasoma brown T. punctigaster, spec. nov. from which it differs by with hind margins of tergum II and often of tergum having a stouter fl agellum (7th or 8th fl agellar segment III and thyridia yellowish, tergum VII reddish square in lateral view), narrowed mandibles as well brown. as by occurrence of brachypterous specimens. The Body length 3.0-4.0 mm, fore wing length 2.3- male of T. solyanikovi, spec. nov. is closest to that of 2.6 mm. T. decipiens, spec. nov., from which it differs by hav- Male unknown. ing narrowed mandibles, shorter fl agellum with 21-23 segments and stouter metasomal tergum III Biology. Parasitoid of Brevantennia siederi and Post- (1.5-1.6 × as wide as long). solenobia juliella. Description Distribution. Italy. Female. Lower edge of the clypeus very slightly rounded, straight medially, apical margin hardly 14. Trachyarus solyanikovi Gokhman, spec. nov. visible. Head weakly narrowed behind eyes in dor- Fig. 11 sal view. Temples moderately enlarged ventrad, with their maximum length distinctly less than Types. Holotype: W, Russia, Moscow region, Bronnitsy transverse ocular diameter. Flagellum slightly en- station, from Solenobia sp., 26.IV.1982 (Solyanikov). – larged in apical half, with 21-23 segments, fi rst seg- Paratypes: Russia: 2WW, same data; 1M, same locality ment 2.2-2.3 × as long as wide, 7th or 8th segment and collector, from Solenobia cembrella, IV.1973; 1M, same square in lateral view. Mandibles narrowed, lower locality, host and collector, 1-19.V.1974, 1973-1977; 2MM, mandibular tooth distinctly longer and wider than same locality, host and collector, 22.IV.1974; 3MM, same the upper tooth. Hypostomal carina meets the genal locality and collector, from Solenobia sp., 24.IV.1982; 2WW, 3MM, same locality, host and collector, 25.IV.1982; carina before the mandibular base at one-third its 1M, same locality, host and collector, 29.IV.1982; 1M, width. same locality, host and collector, 30.IV.1982; 4WW, same Distinct notauli present in the basal fourth to locality, host and collector, coll. 1.V, emerged 6.V.1982; third of mesoscutum. Area superomedia about 1.5 × 1W, same locality, host and collector, 1982; 1W, same as wide as long, weakly separated from area pos- data, “21”, “Tycherus sp.”; 2WW, 2MM, Moscow region, teromedia. Area petiolaris about 3 × as long as area Lyubertsy station, 5.V.1982 (Solyanikov); M, Moscow superomedia. Wings of variable length, from fully region, VNIIPriroda, 14.VI.1982 (Butovsky); Ukraine: developed to reduced, in the latter case stretching W, Ivano-Frankovsk region, Mt. Pozhizhevskaya, 1500 m, to the apex of propodeum or even to the apex of the 26.7.1989 (Kasparyan); Hungary: W, “Parasit e Psychi- fi rst metasomal segment. Pterostigma absent only dae, leg. H. Meier”, “Ungarn, Bükk-Geb., 1974, An Bäu- in the shortest fore wings. Nervellus broken in its men, Sol. nickerli Hein.”; Austria: W, “Knittelfeld, Steier- mark, Österreich”, “Parasit e Psychidae, leg. H. Meier”, lower third. Hind femora 3.2-3.3 × as long as wide. “Eppenstein, 1964, Sol. charlottae Meier”, “Trachyarus Metasoma oval. Distance between spiracles of brevipennis Rom. W det. E. Diller 87”; M, “Steiermark, St. the fi rst metasomal segment about 2 to 3 × as long [illegible]”, “Leoben, 15.V.1955, leg. [illegible]”, “Soleno- as distance between the spiracle and hind corner of bia pineti Z.”; M, “Österreich, Parasit e Psychidae, leg. H. the segment. Thyridia medially removed from the Meier”, “Linz, Puchenau, 1962, Sol. triquetrella Hbn.”; base of metasomal tergum II by about half their 2MM, “Bernstein (Burgenland)”, “17.5.55, leg. H. Meier”; width, approximately 1.5 × as wide as the inter- 4MM, same locality and collector, 18.5.55; 3MM, same lo- thyridial space. Metasomal tergum II 1.1-1.2 × as cality and collector, 20.5.55; M, same data, “Staatssamm- wide as long, tergum III 1.7-1.8 × as wide as long lung München”, “Trachyarus corvinus Thoms. (Ichneu- (Fig. 11). monidae, Ichneumoninae)”, “det. Hedwig”; Germany: Body moderately sculptured. Head shining, side M, “5.VI.28 el, Bacc. sepium, Barrien, Brem., Jäckh.”, “[illegible] nicht B. sepium, [illegible] Sol. triquetrella”, “Di- parts of face sparsely punctate, median part of the caelotus pusillus Holmgr. W”[!], “det. Haberm.”. W, “Bay- face and clypeus almost smooth. Frons smooth. ern-Süd, Irlbasch, VI.1962, leg. Diller”; The Nether- Mesoscutum and scutellum smooth; pronotal lobes lands: W, “Ede E.L., V.1967, C. J. Zwakhals”; W, “Putten weakly chagrinate. Mesopleura weakly rugose, E.L., V.1967, C. J. Zwakhals”. Material from Russia and speculum smooth. Metasomal tergum I weakly the Ukraine is deposited in the ZMMU (1 paratype M) chagrinate (more strongly at petiolus). Terga II-III and ZIN collections. (sometimes also base of tergum IV) obviously cha-

78 ©Zoologische Staatssammlung München/Verlag Friedrich Pfeil; download www.pfeil-verlag.de grinate, sculpture becomes more smooth towards mandibular tooth distinctly longer and wider than apex of metasoma. the upper tooth. Hypostomal carina meets the genal Body brown. Head dark brown, clypeus and carina approximately at the mandibular base. mandibles (except teeth) yellowish brown. Antennae Distinct notauli present in the basal third of brownish, fl agellum reddish ventrally, palps light mesoscutum. Sternauli weak but stretch almost to brown. Thorax from dark to yellowish brown, tegu- the middle of mesopleura. Area superomedia about lae whitish. Legs reddish, hind coxae sometimes as wide as long, weakly separated from area pos- slightly infuscate, last segments of all tarsi infuscate. teromedia. Area petiolaris about 2.5 × as long as area Metasoma brown with the very apex of tergum I superomedia. Wings fully developed. Nervellus medially, thyridia and hind margins of terga II-III broken in its lower third. Hind femora about 4 × as and often of tergum IV yellowish; terga II-IV some- long as wide. times lighter. Metasoma longish-oval. Distance between spir- Body length 3.5-4.5 mm, fore wing length 0.7- acles of the fi rst metasomal segment about twice as 3.1 mm. long as distance between the spiracle and hind Male similar to female. Flagelum with 21-23 corner of the segment. Thyridia medially removed segments, fi rst segment 2.3-2.4 × as long as wide, 17th from the base of metasomal tergum II by about their or 18th segment square in lateral view, longish-oval width, slightly narrower than the interthyridial tyloids present on (7)8th-10th segment. Metasoma space. Metasomal tergum II as long as wide, tergum more slender (tergum II 1.0-1.1 × as long as wide, III 1.4-1.5 × as wide as long. tergum III 1.5-1.6 × as wide as long), its sculpture Body moderately sculptured. Head shining, side more robust. Body brown, clypeus often whitish. parts of face sparsely punctate, median part of the All coxae, trochanters I, femora (fore femora only face and clypeus almost smooth. Frons smooth. on their outer sides), tarsi and hind tibiae infuscate. Mesoscutum and scutellum fi nely and sparsely Apex of metasoma brown. punctate; pronotal lobes rugose except for their up- Body length 3.5-4.5 mm, fore wing length 3.0- per part. Mesopleura longitudinally rugose, specu- 3.7 mm. lum smooth. Metasomal terga I-II strongly chagrinate except for the apical margin of tergum II. Tergum Biology. Parasitoid of Dahlica and Siederia spp., III chagrinate in its basal half, sculpture becomes including D. charlottae, D. nickerlii, D. triquetrella and more smooth towards apex of metasoma. S. listerella. Body brown. Head dark brown, clypeus somewhat lighter, mandibles with reddish markings. Distribution. Russia (Moscow region), Ukraine, Antennae brownish, palps light brown. Tegulae Hungary, Austria, Germany (Lower Saxony, Ba- whitish. Legs brownish, apices of fore femora and varia), the Netherlands. stripes on inner sides of fore tibiae reddish. Meta- soma brown with hind margin of tergum II yellowish, tergum VII reddish brown. 15. Trachyarus subtilipiceus Gokhman, spec. nov. Body length 3.5 mm, fore wing length 2.7 mm. Male unknown. Holotype: W, Italy, “Italia, Mte Baldo, Prada, 1300 m, el M 5.1967, leg. Burmann”. Biology. Parasitoid of Brevantennia siederi. Differential diagnosis. T. subtilipiceus, spec. nov. is most closely related to T. punctigaster, spec. nov., but Distribution. Italy. in T. subtilipiceus, spec. nov. fl agellum is more slender (17th or 18th fl agellar segment square in lateral view). Trachyarus sp. A. Material. Finland: 1M, “Kohde R/K, Leivo A., Puulaju Description L/P Koivu”, 1M, same data, “no brevipennis M det. E. Female. Lower edge of the clypeus very slightly Diller 2002”; 1M, “Suomi, U. Askola, 1971, T. Suoma- rounded, almost straight medially, apical margin lainen”, “Trachyarus corvinus Ths. M, Karvonen det.”, hardly visible. Head slightly narrowed behind eyes “Ranin”; 1M, “Suomi, U: Tuusula, 6665: 389, *16.5.1975, E.O. Peltonen leg”, “Solenobia Dup. fumosella n Hein. ? in dorsal view. Temples moderately enlarged at the loinen”, “Ranin”; Austria: 1M, “Österreich, Innsbruck, middle, with their maximum length somewhat equal Kranebilten, 500 m”, “A.IV.1966, leg. Burmann”, “e.p. to transverse ocular diameter. Flagellum thin, with 19.IV.1966, host: Talaeporia oder Solenobia”. All material 23 segments, fi rst segment about 3 × as long as wide, is deposited in the ZIN collection. 17th or 18th segment square in lateral view. Lower

Differential diagnosis. Trachyarus sp. A is most brownish, apices of all femora (and inner surfaces closely related to Trachyarus sp. B, but Trachyarus sp. of fore femora), tibiae and tarsi reddish, hind tibiae A has a longer fl agellum with 24-25 segments and and last segments of all tarsi infuscate. Metasoma the area superomedia of propodeum separated from brown with the very apex of tergum I medially, the area petiolaris by a strong and distinct carina. thyridia and hind margin of tergum II reddish. Body length 4.0-4.5 mm, fore wing length 3.5- Note. A closely related female from Finland (W, 3.7 mm. “Suomi, U. Askola, 1971, T. Suomalainen”, “Trachy- arus corvinus Ths., Karvonen det.”, “Ranin”) with Biology. Parasitoid of Dahlica lazuri (Clerck) and very broad postpetiolus probably belongs to this undetermined Psychidae. species. Distribution. Austria, Finland. A boreal-montane Description species. Female unknown. Male. Lower edge of the clypeus very slightly Trachyarus sp. B. rounded, almost straight medially. Head slightly narrowed behind eyes in dorsal view. Temples al- Material. 2MM, Russia, Primorsky district, “GTS” (Mon- most parallel, with their maximum length distinctly tane Forest Station), ex Solenobia, 28.IV.1983 (Solyanik- less than transverse ocular diameter. Flagellum with ov); 1M, same date, locality and collector, ex Eosolenobia 24-25 segments, fi rst segment 2.9-3.0 × as long as suifunella. All material is deposited in the ZIN collection. wide, a few subapical segments square in lateral Differential diagnosis. Trachyarus sp. B is most view, tyloids absent. Upper mandibular tooth dis- closely related to Trachyarus sp. A, but Trachyarus tinctly longer and hardly wider than the lower tooth. sp. B has a shorter fl agellum with 21-23 segments Hypostomal carina meets the genal carina before the and the area superomedia of propodeum separated mandibular base at about one-third its width. from the area petiolaris by a weak and indistinct Distinct notauli converge in the basal third of carina. mesoscutum. Area superomedia heart-shaped, bordered by strong carinae, about as wide as long. Area Description petiolaris approximately twice as long as area superomedia. Wings fully developed. Nervellus broken Female unknown. slightly below the middle. Hind femora 3.2-3.3 × as Male. Lower edge of the clypeus slightly round- long as wide. ed. Head almost not narrowed behind eyes in dorsal Metasoma longish-oval. Petiolus distinctly com- view. Temples parallel, their maximum length a pressed dorsoventrally. Distance between spiracles little less than transverse ocular diameter. Flagellum of the fi rst metasomal segment twice as long as with 21-23 segments, fi rst segment 3.2-3.3 × as long distance between the spiracle and hind corner of the as wide, a few subapical segments almost square in segment. Thyridia medially removed from the base lateral view, tyloids absent. Upper mandibular tooth of metasomal tergum II by about one-third their distinctly longer and wider than the lower tooth. width, approximately 3 × as wide as the interthy- Hypostomal carina meets the genal carina before the ridial space. Metasomal tergum II 1.2-1.3 × as wide mandibular base at about half its width. as long, tergum III 1.7-1.8 × as wide as long. Distinct notauli converge in the basal third of Body moderately sculptured. Frons obviously mesoscutum. Area superomedia heart-shaped, about chagrinate. Mesoscutum chagrinate and compara- as wide as long. Area petiolaris approximately 2.5 × tively roughly punctate, scutellum weakly shining as long as area superomedia. Wings fully developed. and more fi nely punctate; pronotal lobes longitudi- Nervellus broken in its lower third. Hind femora nally rugose in their lower half. Mesopleura rugose, 3.6-3.7 × as long as wide. speculum smooth. Postpetiolus of the fi rst metaso- Metasoma longish-oval. Distance between spir- mal segment rugose-punctate and chagrinate, peti- acles of the fi rst metasomal segment twice as long olus chagrinate. Metasomal terga II-III and basal half as distance between the spiracle and hind corner of of tergum IV obviously chagrinate with obscure the segment. Thyridia medially removed from the scattered punctures, sculpture becomes more smooth base of metasomal tergum II by about half their towards apex of metasoma. width, approximately twice as wide as the interthy- Body brown. Head brown, clypeus (except its ridial space. Metasomal tergum II 1.1-1.2 × as wide lower margin), ventral markings of scape and some- as long, tergum III 1.5-1.6 × as wide as long. times mandibles (except teeth) whitish. Antennae Body moderately sculptured. Frons distinctly brown, palps whitish. Tegulae brownish. Legs chagrinate. Mesoscutum chagrinate (except for lat-

80 ©Zoologische Staatssammlung München/Verlag Friedrich Pfeil; download www.pfeil-verlag.de eral lobes) and comparatively densely punctate, Distinct notauli present in the basal third of scutellum punctate and more weakly chagrinate. mesoscutum. Area superomedia heart-shaped, bor- Pronotal lobes chagrinate, longitudinally rugose in dered by strong carinae, about as wide as long. Area their lower half. Mesopleura rugose-punctate and petiolaris approximately 2.5 × as long as area su- chagrinate, speculum smooth. Postpetiolus of the peromedia. Wings fully developed. Nervellus broken fi rst metasomal segment rugose-punctate and cha- slightly below the middle. Hind femora 3.9-4.0 × as grinate, petiolus chagrinate. Metasomal terga II-IV long as wide. obviously chagrinate with obscure scattered punc- Metasoma longish-oval. Distance between spir- tures, sculpture becomes more smooth towards apex acles of the fi rst metasomal segment about twice as of metasoma. long as distance between the spiracle and hind Body brown. Head brown, clypeus sometimes corner of the segment. Thyridia medially removed light brown to whitish (except its lower margin), from the base of metasomal tergum II by about half mandibles whitish (except teeth). Antennae brown, their width, approximately 2.5 × as wide as the in- palps brownish to whitish. Tegulae brownish. Legs terthyridial space. Metasomal tergum II 1.2-1.3 × as brownish, apices of all femora (and inner surfaces long as wide, tergum III 1.5-1.6 × as wide as long. of fore femora), tibiae and tarsi (except hind tarsi) Body moderately sculptured. Frons obviously reddish, last segments of all tarsi infuscate. Meta- chagrinate. Mesoscutum and scutellum fi nely punc- soma brown with the very apex of tergum I medi- tate and shining; pronotal lobes rugose centrally. ally, thyridia, all margins of terga II-III and hind Mesopleura rugose-punctate, speculum smooth. margin of tergum IV yellowish brown. Metasomal tergum I rugose-punctate and chagrinate. Body length 4.0-4.5 mm, fore wing length 3.6- Terga II-V obviously chagrinate with dense but 4.1 mm. obscure punctures, sculpture becomes more smooth towards apex of metasoma. Biology. Parasitoid of Eosolenobia suifunella (Chris- Body dark brown. Head brown, mandibles and toph) and Dahlica or Siederia sp. clypeus somewhat lighter, clypeus sometimes whitish. Antennae and palps brown. Tegulae brownish. Distribution. Russia (Primorsky district). Legs brownish, apices of all femora (and inner sur- face of fore femora), fore and middle tibiae reddish. Metasoma brown with the very apex of tergum I Trachyarus sp. C. medially and hind margins of terga II-III (IV) reddish. Material. 1M, Germany, “Bayern, Penzberg, IV.1976, e Psychidae, leg. Hinterholzer”; 1M, Yugoslavia, “Parasit Body length 3.5-4.0 mm, fore wing length 2.7- e Psychidae, leg. H. Meier”, “Jugoslavien, Steiner Alpen, 2.9 mm. 1000-2000 m, Sied. meierella Sied.”. All material is deposited in the ZIN collection. Biology. Parasitoid of Siederia meierella and an undetermined Psychidae. Differential diagnosis. Trachyarus sp. C is most closely related to T. brevipennis Roman, but in Tra- Distribution. Germany (Bavaria), Yugoslavia (Alps). chyarus sp. C the fl agellum is more slender (just a A boreal-montane species. few subapical segments square in lateral view) and metasomal tergum II longer than its width. Table for the identifi cation of the species Description of the genus Trachyarus

Female unknown. 1. Females (unknown for a few species) ...... 2. Male. Lower edge of the clypeus very slightly – Males (unknown for many species) ...... 16. rounded, almost straight medially. Head slightly narrowed behind eyes in dorsal view. Temples al- 2. Upper mandibular tooth longer than the lower most parallel, with their maximum length about 1.5 × tooth (Fig. 2) ...... 3. less than transverse ocular diameter. Flagellum with – Upper mandibular tooth shorter than the lower 22-23 segments, ﬁ rst segment 2.8 × as long as wide, tooth (Fig. 7) ...... 8. a few subapical segments square in lateral view, longish-oval tyloids present on 8th-11th ﬂ agellar seg- 3. Wings very short, not stretched beyond base of ment. Upper mandibular tooth distinctly longer and propodeum ...... brachypterator Diller hardly wider than the lower tooth. Hypostomal – Wings fully developed, if short, obviously carina meets the genal carina before the mandibular stretched beyond base of propodeum ...... 4. base at about one-third its width.

4. Metasomal tergum II 1.1-1.5 × as long as wide 13. Flagellum with 24-26 (rarely with 23) segments. (Figs 5, 6) ...... 5. 10th or 11th fl agellar segment square in lateral – Metasomal tergum II not longer than wide view. Mandibles not narrowed. Wings fully (Fig. 4) ...... 6. developed. Metasomal tergum III 1.3-1.4 × as wide as long ...... decipiens, spec. nov. 5. Metasomal tergum III 1.1-1.3 × as wide as long – Flagellum with 21-23 segments. If approxi- (Fig. 5). Tegulae whitish ...... fuscipes (Thomson) mately 11th fl agellar segment square in lateral – Metasomal tergum III 1.4-1.5 × as wide as long view, then metasomal tergum III 1.8-1.9 × as wide (Fig. 6). Tegulae reddish brown ...... as long ...... 14...... prominulus Diller 14. 7th or 8th fl agellar segment square in lateral view. 6. 7th or 8th fl agellar segment square in lateral view. Mandibles narrowed. Wings sometimes partly Tegulae usually reddish brown ...... reduced ...... solyanikovi, spec. nov...... corvinus Thomson – 7th or 8th fl agellar segment longer than wide. – 7th or 8th fl agellar segment longer than its width. Mandibles not narrowed. Wings fully devel- Tegulae usually whitish ...... 7. oped ...... 15. 7. First fl agellar segment 3.6-4.0 × as long as wide 15. 17th or 18th fl agellar segment square in lateral (Fig. 1). Metasomal tergum III smooth or shagri- view ...... subtilipiceus, spec. nov. nate at the very base. Wings fully developed .... – 11th or 12th fl agellar segment square in lateral ...... anceps (Berthoumieu) view ...... punctigaster, spec. nov. – First fl agellar segment 2.2-2.3 × as long as wide (Fig. 3). Metasomal tergum III distinctly shagri- 16. Upper mandibular tooth longer than the lower nate in its basal half. Often brachypterous ...... tooth ...... 17...... brevipennis Roman – Upper mandibular tooth shorter than the lower tooth. Flagellum with tyloids ...... 24. 8. Metasomal tergum III 2.5-2.6 × as wide as long (Fig. 10). Wings strongly reduced ...... 17. Width/length ratio of metasomal tergum III = ...... parvipennis, spec. nov. 1.5-1.7 ...... 18. – Metasomal tergum III at least 1.8 × as wide as – Width/length ratio of metasomal tergum III = long (Figs 8, 9). Wings fully developed or re- 1.1-1.3 ...... 23. duced ...... 9. 18. Flagellum with tyloids on (7)8th-10th segment .... 9. Metasomal tergum II longer than wide (Figs ...... 19. 8-9) ...... 10. – Flagellum without tyloids ...... 21. – Metasomal tergum II not longer than wide 19. Metasomal tergum II 1.0-1.1 × as wide as long. (Fig. 11) ...... 13. 11th or 12th fl agellar segment square in lateral 10. 5th or 6th fl agellar segment square in lateral view. view ...... corvinus Thomson Brachypterous ...... khrulevae, spec. nov. – Flagellum and metasoma more slender ...... 20. – 5th or 6th fl agellar segment distinctly longer than 20. Metasomal tergum II as long as wide. 17th or 18th wide ...... 11. fl agellar segment square in lateral view ...... 11. Clypeus obviously fl attened in apical half. Brach- ...... brevipennis Roman ypterous. Body length 5.0 mm and more ...... – Metasomal tergum II 1.2-1.3 × as long as wide...... bacillatus, spec. nov. A few subapical fl agellar segments square in – Clypeus convex. Fully winged. Body length lateral view ...... sp. C 5.0 mm or less ...... 12. 21. Apical half of metasomal tergum III smooth, 12. Flagellum more stout, 9th or 10th fl agellar seg- with obscure punctures ...... ments square in lateral view. Metasomal tergum ...... anceps (Berthoumieu) II 1.1-1.2 × as long as wide (Fig. 8) ...... – Apical half of metasomal tergum III obviously ...... edilleri, spec. nov. chagrinate ...... 22. – Flagellum more slender, 11th or 12th fl agellar seg- 22. Area superomedia separated from area postero- ments square in lateral view. Metasomal tergum media by strong carina. Flagellum with 24-25 II 1.3-1.4 × as long as wide (Fig. 9) ...... segments ...... sp. A ...... hemichneumonoides, spec. nov.

– Area superomedia separated from area postero- Drs. Janine Casewitz-Weulersse and Claire Villemant media by weak and indistinct carina. Flagellum (MNHN), Dr. Dmitry Kasparyan (ZIN), Dr. Michael with 21-23 segments ...... sp. B Fitton (NHM) and Dr. Alexandr Antropov (ZMMU) for providing access to/loaning material from various col- 23. Flagellum with tyloids on (8)9th-10(11)th segment. lections and museums. I am also very grateful to E. Length/width ratio of metasomal tergum II = Diller and S. Schmidt for their generous and helpful 1.0-1.1 ...... prominulus Diller advice. – Flagellum without tyloids. Length/width ratio of metasomal tergum II = 1.3-1.4 ...... fuscipes (Thomson) References 24. Width/length ratio of metasomal tergum III = Berthoumieu, V. (1906). Supplement aux Ichneumo- 1.1-1.2 ...... 25. niens d’Europe et d’Algerie. – Echange 22: 59-60 Diller, E. (1988). Vorläuﬁ ge Resultate einer Bearbeitung – Width/length ratio of metasomal tergum III = der Gattungen Hemichneumon Wesmael, 1857, und 1.3-1.6 ...... 26. Trachyarus Thomson, 1891 (Hymenoptera, Ichneu- 25. Larger (body length 5.0 mm and more) ...... monidae, Phaeogenini). – Entomofauna 9: 369-380 ...... bacillatus, spec. nov. – – (1989). Eine neue Art der Gattung Trachyarus Thom- son, 1891, und Bemerkungen zu Hemichneumon – Smaller (body length 4.5 mm or less) ...... Wesmael, 1857 (Hymenoptera, Ichneumonidae, ...... edilleri, spec. nov. Phaeogenini). – Entomofauna 10: 281-289 Perkins, J. F. (1959). Hymenoptera, Ichneumonoidea, 26. Flagellum with 21-23 segments. Mandibles Ichneumonidae, Ichneumoninae I. – Handbooks somewhat narrowed. Width/length ratio of for the Identiﬁ cation of British Insects 7 (2) (ai): metasomal tergum III = 1.5-1.6 ...... 1-116 ...... solyanikovi, spec. nov. Roman, A. (1918). Beiträge zu den schwedischen Ich- – Flagellum with 24-25 segments. Mandibles not neumones pentagoni. – Arkiv för Zoologi 12: 1-32 narrowed. Width/length ratio of metasomal Selfa, J. & E. Diller (1994). Illustrated key to the Western tergum III = 1.3-1.4 ...... decipiens, spec. nov. Palearctic genera of Phaeogenini (Hymenoptera, Ichneumonidae, Ichneumoninae). – Entomofauna 15: 237-252 Thomson, C. G. (1891). – Opuscula entomologica 67: Acknowledgements 1603-1656 Wahl, D. B. & W. R. M. Mason (1995). The family-group The author is very grateful to Mr. Erich Diller and Dr. names of the Ichneumoninae. – Journal of Hy- Stefan Schmidt (ZSM), Dr. Roy Danielsson (UZI), Drs. menoptera Research 4: 285-293 Karl-Johan Hedquist and Fredrik Ronquist (NHRM),

Buchbesprechungen

8. Richarz, K.: Fledermäuse beobachten, erkennen und 9. Rabitsch, W. (Hrsg.): Hug the bug – For love of true schützen. – Franckh-Kosmos Verlags-GmbH & Co. KG, bugs. – Festschrift zum 70. Geburtstag von Ernst Stuttgart, 2004. 126 S., 173 Abb., ISBN 3-440-09691-2 Heiss. Denisia Bd. 19, Linz, 2006. 1184 S. ISBN Klaus Richarz, seit langer Zeit aktiv im Fledermausschutz 3-85474-161-8 tätig und für sein Engagement bereits mit mehreren Der vorliegende Monumentalband ist eine Liebeser- Preisen ausgezeichnet, ist Autor des vorliegenden Ta- klärung an die sonst so oft geschmähten Wanzen und schenbuches über europäische Fledermäuse. Diesen zugleich eine Hommage für den bekannten Wanzenlieb- angeblich unheimlichen oder unglückbringenden “Ge- haber und Wissenschaftler Dr. Ernst Heiss. Seine Begeis- schöpfen der Nacht”, die die Erde seit rund 50 Millionen terung hat vielfach angesteckt, seine Hilfsbereitschaft ist Jahren bevölkern, begegnen viele Menschen noch immer in diesem Werk vielfach bekundet und seine wissen- mit Mißtrauen und zahlreichen mythologisch behafteten schaftliche Leistung ist zweifelsohne beeidruckend. Fehlinformationen. Richarz liefert nun sorgfältig recher- In dem Werk haben fast einhundert Autoren aus 25 chierte Beiträge zu ihrer Lebensweise, portraitiert alle Ländern einem Beitrag zur Entomologie der Wanzen mit uns lebenden Arten und erklärt, wie man sie am verfaßt. Im ersten Teil der Arbeiten sind Artikel zusam- besten beobachten und schützen kann. So wird mit Si- mengestellt, die dem Leben und Werk von Ernst Heiss cherheit mit vielen Vorurteilen gegenüber diesen faszi- gewidmet sind, sowie eine Geschichte und Bibliographie nierenden und mittlerweile stark gefährdeten Tieren der österreichischen Wanzenkunde und ein unterhaltsa- gründlich aufgeräumt. mer Artikel über “ ... ernstes und kurioses - ein hetero- Das erste Kapitel “Echos aus einer versunkenen Welt” pterologisches Panoptikum”. Mehrere Arbeiten beschäf- beleuchtet kurz die Evolution der Chiroptera und die tigen sich mit den Aradidae, der Wanzenfamilie, der Entwicklung ihrer ungewöhnlich vielfältigen, erfolgrei- E. Heiss in besonderer Weise verbunden ist. Diese abge- chen Ernährungsstrategien. Die folgenden vier Abschnit- fl achten Wanzen, zu deutsch “Rindenwanzen”, leben te begleiten die Fledermäuse durch die Jahreszeiten bevorzugt (aber nicht immer) unter der Rinde von Bäu- Frühling, Sommer, Herbst und Winter und liefern gleich- men und saugen an Pilzmycelien. Hier, wie in dem ge- zeitig wichtige Details zu ihren bevorzugten Lebensräu- samten Werk, faszinieren die vielen farbigen Abbildun- men, zur Echoortung durch Ultraschall und zum Erken- gen, mit denen insbesondere die Tiere im Habitus vor- nen der Arten im Gelände durch Beobachtung und gestellt werden, sowie die schönen Tafeln mit REM Fledermaus-Detektoren. Sie beschreiben die Aufzucht Abbildungen. Beeindruckend ist zum Beispiel die mit der Jungen, informieren über die verschiedenen Tages- herrlichen REM-Tafeln illustrierte Beschreibung einer und Wochenstuben-Quartiere, ihre Erhaltung, Gestaltung atypischen micropteren Aradide, die zu Ehren von Ernst und Betreuung und streifen auch kurz die Methoden, Heiss Breviscutheissia ernsti genannt und in Südafrika an Fledermäuse zu beringen und zu erforschen. Ein weite- sukkulenten Kletterpfl anzen gefunden wurde. Der größ- res Kapitel stellt die 32 in Europa und im Mittelmeerraum te Teil des Bandes enthält taxonomische Arbeiten über heimischen Arten mit anschaulichen Bildern und einem verschiedene Wanzentaxa: Tingidae, Miridae, Coreidae, kurzen Text vor. Sogar der auf Zypern und im äußersten Pentatomidae, Pyrrhocoridae und so fort. Viele Arten Süden der Türkei vorkommende Nilfl ughund wird be- wurden neu beschrieben, mehrere Arbeiten sind mehr rücksichtigt. vergleichend morphologisch (z. B. Genitaluntersuchun- Empfehlungen zum Schutz der stark bedrohten gen) orientiert. Es wundert nicht, daß die meisten neuen Fledermäuse ziehen sich wie ein roter Faden durch das Namen dem Jubilar gewidmet sind, aber wie viele das gesamte Werk. Ein “Service für Fledermausschützer”, sind, das erstaunt dann doch! Es werden in dem Band Literatur-Tips, eine Liste nützlicher Adressen und ein eine Unterfamilie, 18 Gattungen und Untergattungen 116 informatives Register runden dieses sehr empfehlens- Arten und Unterarten beschrieben. Im weiteren werden werte, dabei preiswerte Buch ab, das durch seine schöne noch eine Reihe von Arbeiten zur Biogeographie und zur Aufbereitung und ansprechende Illustration nicht nur Biologie der Wanzen vorgestellt. ältere, sondern besonders auch junge Fledermausfreun- In dem Werk sind praktisch alle derzeit aktiven de begeistern wird, die für den künftigen Schutz unserer Taxonomen der vielgestaltigen Heteropteren vertreten. Natur und Umwelt Verantwortung übernehmen wol- Das vorliegende Werk ist mit weit mehr als tausend len. J. Diller Seiten und unzähligen Farbabbildungen ein Prachtband und ein Meilenstein der aktuellen Wanzentaxonomie und das alles zu einem überraschend moderaten Preis. Ein Werk, zu dem man Autoren und Herausgeber nur herzlich gratulieren kann, besonders aber jenem, dem das Werk gewidmet ist: Ernst Heiss. Ad multos annos! K. Schönitzer und T. Kothe