JOURNAL OF VIROLOGY

VOLUME 32 0 NUMBER 2 0 NOVEMBER 1979 EDITORIAL BOARD

Robert R. Wagner, Editor-in-Chief (1982) University of Virginia School of Medicine, Charlottesville Dwight L. Anderson, Editor (1983) Harold S. Ginsberg, Editor (1984) School of Dentistry, Columbia University University of Minnesota, New York, N. Y. Minneapolis David T. Denhardt, Editor (1982) Edward M. Scoliick, Editor (1982) McGill University National Cancer Institute Montreal, Que., Canada Bethesda, Md. John P. Bader (1979) Alice S. Huang (1981) Fred Rapp (1981) David Baltimore (1981) Paul J. Kaesberg (1979) Dan S. Ray (1980) Amiya K. Banerjee (1979) Joseph Kates (1979) Bernard E. Reilly (1980) David H. L. Bishop (1979) D. C. Kelly (1979) William S. Robinson (1980) David Botstein (1979) George Khoury (1981) Bernard Roizman (1979) Michael A. Bratt (1980) Jonathan A. King (1981) Roland R. Rueckert (1979) Dennis T. Brown (1981) David W. Kingsbury (1979) Norman P. Salzman (1981) Ahmad I. Bukhari (1981) Lloyd M. Kozloff (1979) Joseph Sambrook (1979) Purnell Choppin (1980) Robert M. Krug (1980) Priscilla A. Schaffer (1981) John M. Coffin (1980) Robert A. Lazzarini 1981) Sondra Schlesinger (1980) Richard W. Compans (1979) Richard A. Lerner (1981) Phillip A. Sharp (1979) Geoffrey M. Cooper (1981) Myron Levine (1979) Aaron J. Shatkin (1979) Nicholas R. Cozzarelli (1980) Tomas Lindahl (1981) Lee D. Simon (1981) Clive Dickson (1981) David M. Livingston (1980) Kai Simons (1981) Walter Doerfler (1980) Ronald B. Luftig (1981) Alan E. Smith (1980) Harrison Echols (1981) Brian McAuslan (1979) Patricia G. Spear (1981) Elvera Ehrenfeld (1980) Robert Martin (1981) F. William Studier (1981) Suzanne U. Emerson (1980) Christopher K. Mathews (1979) Donald F. Summers (1979) Lynn Enquist (1981) Robert C. Miller (1980) William C. Summers (1980) Erwin J. Fleissner (1980) Gisela Mosig (1979) John M. Taylor (1981) S. Jane Flint (1981) Bernard Moss (1980) Natalie M. Teich (1979) Robert M. Friedman (1979) Joseph S. Pagano (1979) Howard M. Temin (1979) Hidesaburo Hanafusa (1980) J. Thomas Parsons (1980) David C. Ward (1980) William S. Hayward (1981) Ulf G. Petterson (1980) Robin Weiss (1979) John J. Holland (1981) Lennart Philipson (1981) John S. Wiberg (1979) Nancy Hopkins (1980) Peter G. W. Plagemann (1980) Ernest Winocour (1979) Calderon Howe (1979) Craig R. Pringle (1980) Julius Youngner (1979)

Robert A. Day, Managing Editor Gisella Pollock, Assistant Managing Editor Susan Birch, Production Editor 1913 I St., NW, Washington, DC 20006 EX OFFICIO Wiflis A. Wood, President (1979-1980) Albert Balows, President-Elect (1979-1980) J. Mehsen Joseph, Secretary Brinton M. Miller, Treasurer

The Journal of Virology (ISSN 0022-538X), a publication of Second-class postage paid at Washington, DC 20006, and at the American Society for Microbiology, 1913 I St., N.W., Wash- additional mailing offices. ington, DC 20006, is devoted to the dissemination offundamental Made in the U.S.A. knowledge concerning viruses of bacteria, plants, and animals. Copyright © 1979, American Society for Microbiology. Investigators are invited to submit reports of original research in All Rights Reserved. all areas of basic virology, including biochemistry, biophysics, genetics, immunology, morphology, and physiology. Instructions to Authors are published in the January issue each year. The The code at the top of the first page of an article in this journal Journal is issued monthly, four volumes per year. The nonmem- indicates the copyright owner's consent that copies of the article ber subscription price is $90 per year; single copies are $8. The may be made for personal use, or for personal use of specific member subscription price is $21 per year. Correspondence clients. This consent is given on the condition, however, that the the Clear- relating to subscriptions, reprints, defective copies, availability of copier pay the stated per-copy fee through Copyright of submitted manu- ance Center, Inc., P.O. Box 765, Schenectady, New York 12301, back issues, lost or late proofs, disposition that Sections 107 and 108 of and editorial matters should be directed to the for copying beyond permitted by scripts, general consent does not extend to other ASM Publications 1913 I St., N.W., Washington, DC the U.S. Copyright Law. This Office, such as for distribution, for 20006 (area 202 833-9680). kinds of copying, copying general advertising or promotional purposes, for creating new collective El * : idO V 'I t lL '.r %k ( P1]4Ut works, or for resale. Author Index Anderson, S. J., 507 Hampar, Berge, 676 Nayak, Debi P., 697 Hanafusa, H., 546 Nehorayan, Angela, 648 Baugh, C. M., 497 Hattman, Stanley, 468 Neubauer, Russell H., 676 Becht, Hermann, 593 Heilman, Conrad, J., Jr., 676 Nilsson, Maj-Greth, 386 Bendig, Mary M., 530 Highfield, Peter E., 567 Berg, Paul, 523 Hill, Barry J., 593 Ojima, Setsuko, 379 Bolen, Joseph B., 679 Hoffmann, Peter J., 449 Bowen, J. M., 507 Holland, John J., 420 Padgett, Billie L., 476 Brady, J. N., 640 Hopkins, Ralph F., III, 676 Parsons, J. Thomas, 567 Bruenn, Jeremy A., 692 Hunter, Eric, 412 Pauley, Robert J., 557 Brusca, John, 623 Hutchinson, Mary A., 517 Payne, Lendon G., 614 Burck, Kathy Bauman, 606 Pietras, Dennis F., 692 Iverson, Linda, 404 Preston, C. M., 357 Carlin, Nils, 583 Cheng, Yung-chi, 449 Janda, J. Michael, 697 Rabin, Harvey, 676 Chinnadurai, G., 623 Johnson, Paul H., 629 Rose, John K., 404 Chinnadurai, Shanbagam, 623 Cohen, J. Craig, 483 Kane, William P., 692 Sawada, Yukiharu, 379 Consigli, R. A., 640 Karess, R. E., 546 Scraba, Douglas G., 606 Consigli, Richard A., 679 Kasamatsu, Harumi, 648 Seifried, Adele S., 688 Crosby, L. K., 497 Kells, David T. C., 593 Semler, Bert L., 420 Kelly, Samuel V., 629 Shimojo, Hiroto, 379 Davis, J., 507 Kendall, J. D., 640 Shiroki, Kazuko, 379 Dierks, Peter M., 567 Kozloff, L. M., 497 Socher, Susan H., 557 Dobos, Peter, 593 Kristenson, Krister, 614 Sprague, Judy, 688 Kuhnert, Lee K., 536 Stohlman, Stephen A., 672 Eckhart, Walter, 517 Stohrer, Robert, 412 Edson, Clark M., 458 Lai, Michael M. C., 672 Stoltzfus, C. Martin, 536 Eron, Lawrence J., 688 Lancaster, Wayne D., 684 Svenson, Stefan B., 583 Lindberg, Alf A., 583 Flint, S. J., 394 Locker, Hilla, 429 Teninges, Danielle, 593 Folk, William R., 530 Lonngren, Jorgen, 583 Thorley-Lawson, David A., 458 Frenkel, Niza, 429 Friis, Robert R., 370 Magnusson, Goran, 386, 523 Varmus, Harold E., 483 Frisque, Richard J., 476 Majors, John E., 483 Fujinaga, Kei, 379 Martin, Jonathan D., 476 Walker, Duard L., 476 Medina, Daniel, 557 Wang, L.-H., 546 Goorha, Rakesh, 442 Miller, Michael J., 629 Wells, Robert D., 517 Granoff, Allan, 442 Miller, Robert C., Jr., 606 Wild, Elizabeth, 629 Grossman, Lawrence I., 629 Milstien, Julie B., 688 Willis, Dawn B., 442 Gurney, Elizabeth G., 661 Moelling, Karin, 370 Gurney, Theodore, Jr., 661 Moscovici, C., 546 Yang, V. W., 394 Hallett, Ross, 593 Naso, R. B., 507 Zweig, Martin, 676