Jourwul of’ M’icropalaeontolo~v,14: 59-65 0262-821 X/95 $07.00 0British Micropalaeontological Society.
An annotated check-list of British Pleistocene, Holocene and modern freshwater ostracods
H. I. GRIFFITHS’ & J. G. EVANS’ ‘Delpartmlent of Genetics, School of Biological Sciences. University of Leeds, Woodhouse Road, Leeds LS2 9JT, UK. ‘School of History & Archaeology, University of Wales College of Cardiff, PO Box 909, Cardiff CFI 3XU. UK.
ABSTRACT - A revised check-list of British Pleistocene, Holocene and Kecent freshwater ostracods is given, and notes provided on the taxonomy and occurrence of some rare and newly-reported species. Stmocvpria ,fischeri (Lilljeborg. 1883) and the hypogean species Pseudocuntiona cf. eremito (Vecijvosky. 18x2) and Pseudocandoncr hrruili (Paris, 1920) arc reported from Britain for the first time. J. Micro,palawn/ol. 14( I): 59-65. April 1995.
INTRODUCTION work accrued. Despite this, J. E. Robinson’s Pleistocene and Over recent years there has been renewed interest in Holocene studies continued to document ostracod faunas, as freshwater ostracods, particularly because of their potential occasionally have archaeological (e.g. Griffiths & Mount, in palaeolimnological and palaeohydrological studies (see 1993) and modern synecological works (e.g. Ham, 1982). reviews by Delorme, 1989; De Deckker & Forester, 1988: An updated check-list of freshwater Ostracoda was made Carbonel et a/. 1988; Holmes, in press). This increase in as part of the Institute of Terrestrial Ecology’s attempt to ecological interest has been accompanied by a spate of compile a complete listing of British freshwater animals taxonomic and systematic research that has done much to (Maitland, 1977). This was not based upon the work of a increase our understanding of evolutionary patterns and single individual, but gleaned from a variety of sources, processes in freshwater Ostracoda. Some freshwater largely without reappraisal. More recently, a new faunal ostracod lineages are evolutionarily dynamic, and diver- work on modern British freshwater ostracods has been gence within rhe Holocene has been demonstrated for at published as part of the Linnean Society’s ‘Synopses of the least onc cluster of lake-specific taxa (Martens, 1990). It is British Fauna’ series (Henderson, 1990). Unfortunately, this possible that the examination of ostracod valves in extended important benchmark was delayed in press for almost five lake corm (which often cover periods of several thousands years (Henderson, pers. comm.) and, as a result, over 30% of years) may prove to be a powerful tool to the of the specific and generic nomenclature used by Henderson evolutioi-iary biologist (Evans Rr Griffiths, 1993). However, had been superseded by the time of publication. The advances in ostracod palaeobiology do require continuity in undesirable result is a loss of continuity with current systematic nomenclature. European taxonomic usage (e.g. Wouters. 1989: Meisch et al. 1990). With the increasing interest in the group by both palaeoecologists and freshwater biologists, it seems timely to provide a check-list of the British species that includes not A BRIEF HISTORY OF BRITISH OSTRACOD only recent revisions, but also records of Recent species ST UDI E,:S omitted by Henderson (1990), plus a small number of Some of the key works in ostracod taxonomy were based on species that are thus far known only from Pleistocene and studies of the British and Irish faunas during the last Holocene contexts. century, notably through the work of G. S. Brady and colleagues (e.g. Brady & Robertson, 1869; Brady & NOTES ON SOME OF THE SPECIES Norman, 1889; Brady, 1910). As a result, comparatively Changes of generic assignation and specific nomenclature comprehcnsive check-lists of the faunas of both Britain have come about as the result of formal revisions, the details (Scourfield. 1904) and Ireland (Norman, 1905) were of which are beyond the scope of this account. Key works available within the first years of this century. Subsequently. include those of Broodbakker (1983), Carbonnel (1963, interest in native ostracod faunas waned, and scientific Colin & Danielopol (1980), Danielopol (1 978), Danielopol attention was turned to the description of species from & McKenzie (1977), Danielopol et al. (1989). Marmonier et overseas. Some workers continued to study British a/. (1989), Martens (1989, 1992) and Meisch (1984, 1985. freshwater species, notably P. F. Holmes, A. G. Lowndes, 1991). Some species do require additional comment. G. Fryer and H. M. Fox, however most had a broad interest however. in freshwater Crustacea, rathcr than a specific interest in Cyprirfopsis hamberi was originally described from a ostracods. Hence, prior to the 1980s. there were remarkably spring in Cornwall by Hendcrson (1986). Professor T. K. few published contributions, and no substantial body of Petkovski (pers. comm.) initially suggested the possible
59 Griffiths & Evans synonymy of C. hamheri with C. brincki Petkovski, 1963, air-dried material from Regent’s Park Lake, London (leg. J. described from wells in the Azores and also known from E. Robinson). The species is also known from British late Germany, Portugal and Macedonia (Petkovski; 1963). We Devensian and Holocene deposits as P. maculata Alm, 1914 examined type material of C. bamheri lodged in the (see list of synonyms in Meisch, 1985). collections of the Natural History Museum in London Eucypris elliptica. This is one of Britain’s less well-known (BMNH) and reached a similar conclusion. More recently, species; Henderson (1990) knew of no definite locations Petkovski et ul. (1993) have shown C. hrincki to be a junior for E. elliptica in the UK. In December 1990, we collected synonym of C. lusatica Shaffer, 1943, and the synonymy of two females from the shallows of Llangorse Lake in the C. humheri has been confirmed by Dr. C. Meisch of the Brecon Beacons National Park, Wales, and Professor D. Natural History Museum of Luxemburg (Meisch, pers. D. Williams has recently provided further specimens comm.). collected from the Island of Bardsey off the North Wales Eucypris anglicu was posthumously described as a British coast. endemic by Fox (1967) from sites in Buckinghamshire and Eucypris lilljeborgi. Again, Henderson (1990) knew of no Hertfordshire, but has since only been reported as a single definite records for this ‘exceedingly rare’ species, specimen from Hampshire (Ham, 1982). Syntypes are although E. cf. lilljeborgi had been reported from maintained in the BM(NH)’s Fox Collection (accession nos Holocene tufas (Preece & Robinson, 1984; Willing, 1985). 1967.4.3.1,2,3,4), the material consisting of four sealed We have since found E. lilljeborgi living in great numbers microscope slides bearing valves, two intact individuals, and in a run-off and rain-fed, grassy, seasonally-inundated intact specimens decalcified in potassium hydroxide. All are meadow adjacent to a Phrugmites-rich pond at Thornhill mounted in glycerine jelly or euparal. These specimens in Cardiff. Eucypris lilljehorgi first appears when the compare well with illustrated descriptions of E. crassa in habitat is inundated in October or November, and then Klie (1938) and Sywula (1974), although are a little larger seems to breed continuously until the site dries out in than usually cited (i.e. 2.2 mm). At our request, Professor T. early summer. The species rarely occurs in the reed beds K. Petkovski has re-examined material of E. anglica which border the inundated grasses, and males are absent. originally sent to him by Fox, and concluded that the species Carapace length is more variable than usually believed; we is indeed distinct. We therefore maintain E. anglica on the have collected gravid females ranging from 1.42-1.88 mm British list, although reinvestigation of ncw material of this long, although clearly identifiable as E. lilljeborgi on the species would be welcome. basis of soft-part anatomy. Ilyocypris hiplicata is well-known in Quaternary palaeon- Trajancypris serrata. There has been considerable confu- tology, and has been recorded in the modern faunas of sion over the taxonomic status of denticulate eucypridines. Canada (Delorme, 1970) and France (Meisch et ul., 1990). Some are assignable to Prionocypris serrata (Norman, The status of the species has long been the source of debate; 1861) of which we have modern and Holocene material Scourfield (1904) stated that 1. hiplicata was common in from the Test Valley, Hampshire. Martens (1 989) erected Britain, although it seems possible that he may have been genus Trajancypris to accommodate subclavate eucyprid- confused between I. biplicata and 1. gihha. Sywula (1974) ines with pronounced lists and selvages on the anterior lists 1. hiplicata as a subspecies of I. gihba. Palaeontologists inner margins, including Eucypris serrnta G. W. Miiller, have long accepted I. hiplicata (sensu Diebel & Pietrzeniuk, 1900 and E. clavata, the latter having often been recorded 1969: pl 7, figs 1-3), and Van Harten (1979) provides a from the Pleistocene as Sclerocypris? clavata prisca Diebel valve-based diagnosis of the species. Ilyocypris hiplicata is a & Pietrzeniuk, 1969. Because of the confusion over bland ilyocypridid, in which surface ornamentation is nomenclature, it now seems almost impossible to say with reduced. The carapace is sub-rectangular, posteriorally and confidence which denticulate eucypridine was actually anteriorally rounded, and with ventral and posterior margins meant by many earlier authors. Hence, we provisionally running almost parallel. In Britain the species has often maintain T. serruta in the British check-list, despite having been recorded at Pleistocene sites, and we have material seen neither subfossil or modern material. from Holocene deposits at West Overton in Wiltshire Paralimnocythere spp. There has been some confusion (Griffiths & Mount, 1993). As soft-part diagnosis remains over the correct specific assignation of non-Balkan species elusive (Meisch, 1988), it would seem that 1. gibha and I. of Paralimnocylhere, now resolved by Martens (1992). biplicata would benefit from examination by either Subfossil British material has usually been referred to P. molecular or genitalia-based taxonomic techniques (cf. compressa or P. cf. dieheli. Parulimnocythere compressa is Martens, 1991). Until such a time as they can either be known from Mid Pleistocene, late Devensian and Holo- synonymized or verified, it seems prudent to maintain I. cene deposits in Britain, although it has not been collected biplicata as a valid species. alive for over a century (Martens, 1992). Paralimnocythere Western European species of Potamocypris have been ciieheli was originally described from Macedonia (Pet- extensively revised by Meisch (1984, 1985) and the kovski, 1969) and a variant, Parulimnocythere cf. diebeli, nomenclature used here differs somewhat from that of was described from the German Mid Pleistocene (Diebel Henderson (1 990). Potamocypris arcuuta was first recorded & Pietrzeniuk. 1978: fig.2, pl. 52, figs 8-11), in the modern British fauna by Griifiths & Evans (1992) Paralimnocythere cf. dieheli has been reported from late from a temporary, groundwater-fed pool in Hampshire. Devensian deposits at Kildale, Yorkshire (Keen et al., Since that time we have also found the species amongst 1984). Paralimnocythere relicta is only known from the
60 British freshwater ostracods
modern fauna, having been reported from Hampshire by Candona lozeki has been reported from the British Henderson (1990). Holocene (Willing, 1985: Mount, 1991) and from the Mid Fabaeformiscandona siliquosa. This is an unusual species Devensian (Gibbard et al., 1981), whilst locations for C. which appears to be a British endemic. Some authors have tricicatricosa are all Mid Pleistocene. Fuhrmann (1991) has doubted this: Nuchterlein (1965): 246) believed F. siliquosa suggested that C. IozekiC. tricicatricosa are synonymous. If to be synonymous with F. caudata. Henderson (1990) this is the case, this would remove the stratigraphic value maintains F. siliquosa as a valid species, listing several of both taxa. sites in the New Forest of Hampshire. We have also Pseudocandona breuili was first described from a cave in collected F. si/iquosa from permanent ponds in the New Spain (Paris, 1920) and, as Candona breuili, is known Forest arid cornpared it with specimens of F. caudutu from from the German Quaternary (Diebel & Pietrzeniuk, France (i'eg. P. Marmonier). The two species appear very 1984) and the Belgian Holocene (Van Frausum & different, and we have retained them both as distinct Wouters, 1990). Definitive determination of this species is species. difficult without soft-parts, although we have collected it in Candona lactea was reviewed by Brady (1910) and considerable numbers from Holocene deposits at West retained as a valid species by Henderson (1990). Although Overton, Wiltshire and from the Test Valley, Hampshire. we have not examined type material of this taxon, we It appears that the species lived interstitially, and havc collected material that appears similar to C. lactea, Danielopol (1978) lists P. breuili as a hypogean species. In but have never encountered mature individuals. We some cases at least, it seems that 1'. hreuili has been therefore maintain C. lactea in the British list until formal erroneously identified as the juvenile moult stages of revision is made, but believe that it may be a synonym Psychrodromus olivaceus. mistakenly erected upon juvenile material of another Pseudocandona eremita is one of a cluster of hypogean species. Further investigation is required to validate or ostracods that display a high degree of local endemism, deny this, suggestion. and represent a distinct lineage within Pseudocandona Pseudocandona elongata was initially described from (Danielopol, 1082). These are difficult to identify with Lakes Windermere and Ohrid (Holmes, 1937) although no precision without soft parts, hence we have cited our type material was nominated, and none has been located taxon as P. cf. eremita. The species occurs in Holocene (Henderson, pers. comm.). Petkovski (pers. comm.) has sediments from West Overton, Wiltshire and Bossington, failed to record the species despite many years collecting Hampshire, where it seems to have existed interstitially. at Lake Ohricl. Within recent years P. elongata has been Pseudocandonu erernita has a distinctive triangulate reported from Lake Windermere by Horne et al. (1990), carapace, quite unlike any other British candonid. but subsequent collections have failed to provide further Stenocypria fischeri is only known in Britain from specimens (Horne, pers. comm.). This species also Holocene material from West Overton, Wiltshire (Griffiths requires further investigation before it can be validated. & Mount, 1993). Stenocypriu fischeri is illustrated in several European faunal works (e.g. Klie, 1938: 124, figs PLEISTOCENE AND HOLOCENE SPECIES 416-418: Sywula, 1974: 211, fig. 105, pls 17 g-h). In addition to a diverse modern British fauna, a small Nannocandona faba. The absence of this species from the number of species are known exclusively from Pleistocene fauna of Modern mainland Britain is rather puzzling: the and Holocene deposits. although some exist only as single species is quite widespread in Pleistocene and Holocene records. These include Leucocythere baltica (as deposits, furthermore Nannocandona sp. has been re- L,irnnocythere baltica), Limnocythrre falcata, Lirnnocytherr ported from Modern Ireland (Douglas & Healey, 1991). stationis, Limnocythere cf. usenensis, Ilyocypris As Nunnocandona faba often occurs in interstitial quinculminata, I. papillata, I. schwarzhachi, Candona contexts, especially in rivers (Marmonier & Danielopol, Icvanderi, Candona lozeki, Candona tricicatricosa, Fahu- 1988), it is possible that it may yet be found in modern e,formiscundona fabellu (as Candona fabella), Fabaefortnis- Britain, where interstitial habitats remain largely candona balatonica (as Candona balatonica), Pseudocan- unexplored. dona hreuili, Pseudocandona cf. erernitu, Nunnocandonu faha, Scottia browniana, S. tunzida, Eucypris dulcifrons, E. SYSTEMATIC CHECK-LIST heinrich,;, Herpetocypris ehrin,gsdorfensis and Strnocypriu In the following species list, higher-level systematic .fischeri. In addition, the Tertiary species Eucypris cf. nomenclature follows Bowman & Abele (1982), and familial gernella has been reported from Pleistocene deposits at nomenclature largely conforms to Hartmann & Puri (1974). Holderness (Catt & Penny, 1962) and a few taxa, e.g. The nomenclature of rankings below the familial level Candono hrevicornis Klie, 1925 have been reported from the conforms to current European usage (Meisch et al., 1990). Irish Republic (Preece et al., 1986). Some taxa are believed Taxonomic authorities are drawn from Kempf's index to have stratigraphic value (Robinson, 1978), notably: (1980a, b). Species are broadly provcnanced by the Candomi tricicatricosu, C. lozeki, Lirnnocythrre falcata, following superscripts: P = Pleistocene, H = Holocene, R = Leucocyrhere haltica, Ilyocypris quinculminata, I. Recent. Those taxa which have been recorded in the British schwarzbachi, and possibly I. papillata (see Robinson, 1990). fauna, but whose status is here considered questionable, are The binstratigaphic importansce of Scottia spp. has been prcfixed by a question mark. Cyprideis torosu is included, discussed in detail by Kempf (1971). although more typically a species of brackish waters.
61 Griffiths & Evans
Phylum or sub-phylum Crustacea Pennant, 1777 Pseudocandona compressa (Koch, 1 838)"'K Class Ostracoda Latreille, I806 Pseudocandona cf. eremita (Vcjdovsky, 1882)" Sub-class Podocopa G. W. Muller, 1894 Pseudocandona insculpta (G. W.Miiller, 1900)' Order Podocopida Sars, 1866 Pseudocandona lobipes (Hartwig, 1900)H" Sub-order Podocopina, Sars 1866 Pseudocandona marchica (Hartwig, 1899)'" Pseudocandona pratensis (Hartwig, 1 90 1 )'I I Superfamily Darwinuloidea Brady & Norman, 1889 Pseudocandona rostrata (Brady & Norman, 1889)H Family Darwinulidae Brady & Norman. 1889 Pseudocandona sarsi (Hartwig, 1899)R Genus Darwinula Brady & Robertson, 1885 Pseudocandona sragnalis (Sars, 1 X90)R Darwinula steuensoni (Brady & Norman, 1 870)"'R Genus Candonopsis VBvra, 1891 Superfamily Cytheroidea Raird, 1850 Candonopsis kingsleii (Brady & Robertson, 1870)PHR Family Limnocytheridae Klie, 1938 Candonopsis scourjieldi Brady, I 910R Sub-family Limnocytherinae Klie. 1938 Genus Nunnocandona Ekman, 1914 Genus Leucocythere Kaufmann. 1900 Nannocandonafaba Ekman, 19 14"' Leucocythere balrica (Diebel, 1965)' Genus Paracandona Hartwig, 1899 Genus Limnocyrhere Brady, 1867 Paracandona euplecrella (Robertson, 1 889)"' Limnocythere falcata Dicbcl, 1968" Sub-family Cyclocypridinae Kaufmann, 1900 Limnocythere inopinara (Baird, 1843)"" Genus Cyclocypris Brady & Norman, 1889 Limnocythere sancriparricii (Brady & Robertson, CycIocypris globosa (Sars. 1x63)~ I X69)PH' Cyclocypris Iaeuis (0.F. Muller, 1776)'H" Limnocythere starionis Vavra, 1891 Cyclocypris ovum (Jurine, 1820)P"R Limnocythere cf. usenensis Karmischina, 1970' Cyclocypris serena (Koch, 1838)P"R Genus Paralimnocythere Carbonnel, 1965. Genus Cypria (Zenkcr, 1854) Paralimnocythere compressa (Brady & Norman, 1889)""' Cypria ewsculpra Fischer, 1855" Paralimnocythere cf. diebeli (Petkovrki, 1969) Diebel & Cypria ophtalmica (J u ri nc, 1 820)"H Pietrzeniuk, 1978' Family Ilyocyprididae Kaufmann, 1900 Paralimnocythere relicta (Lilljeborg, I 86.3)' Genus Ilyocypris Brady & Norman, 1889 Sub-family Timiriaseviinae Mandelstam, 1960 Ilyocypris biplicata Koch, 1 838''' Genus Metacypris Brady & Robertson, 1870 IIyocypris bradyi Sars, 1890"'~ Meracypris cordara Brady & Robertson, 1 870"HK Ilyocypris decipiens Masi, 190SR Family Cytherideidae Sars, 1925 Ilyocypris getica Masi, 1906R Sub-family Cytherideinae Sars, 1925 Zlyocypris gibba (Ramdohr, 1808)PHR Genus Cytherissa (Sars, 1863) Ilyocypris inermis Kaufmann, 1900PHK Cytherissa lacusrris (Sars, 1863)"H" llyocypris lacustris Kaufmann, 1900'' Genus Cyprideis (Joncs, 1850) Ilyocypris monstrifica (Norman, 1862)1'R Cyprideis rorosa (Jones, 1850)"H" Ilyocypris papillata Robinson, 1990' Superfamily Cypridoidea Baird, 1845 Ilyocypris quinculminata Sylvester-Bradley, 1973' Family Candonidae Kaufmann, 1900 Ilyocypris schwarzbachi Kcmpf, 1967' Sub-family Candoninae Kaufmann, 1900 Family Notodromatidae Kaufmann, 1900 Genus Candona ss. Baird, 1845 Sub-family Notodromatinae Kaufmann. 1900 Candona angulata G. W. Muller, 1900'R Genus Notodromas Lilljehorg, 1853 Candona brevicornis Klie, 1925" Notodromas monacha (0.F. Muller, 1776)" Candona candida (0.F. Muller, 1776)""" Sub-family Cyproidinae Hartmann. 1963 ? Candona lactea Baird, 1850R Genus Cyprois Zenker, 18.54 Candona leuanderi Hirschmann, 1912'' Cyprois marginata (Straus, 1821 ? Candona lozeki Absolon, 19731'H Family Cyprididae Baird, 1845 Candona neglecra Sars, 1887P"R Sub-family Cypricercinae McKenzie, 1971 Candona rricicatricosa Dicbcl & Pietrxniuk, 1969' Genus Bradleystrandesia Broodbakkcr, 1983 Genus Cryprocandona Kaufmann, 1900 Bradleystrandesia fuscara (Jurine, 1820)"R Cryprocandona reducra (Ah, 1913)' Bradleystrandesia reticulata (Zaddach, 1 H44)IlR Cryprocandona uaurai Kaufmann, 1 900H" Genus Strandesia Stuhlmann, 1x88 Genus Fabaeformiscandona Krstic, 1972 Strandesia obliqua (Brady, 1868)"' Fabaeformiscandona acuminara (Fischcr, 1854)K Sub-family Cypridinae Baird, 1845 Fabaeformiscandona balaronica (Daday, 1894)''' Genus Cypris 0. F. Miiller, 1776 Fabaeformiscandona caudata (Kaufmann, 1900)"" Cypris bispinosa Lucas, 1849" Fabaeformiscandona fabella (Nuchterlein, 1969) Cypris pubera 0. F. Miiller, 1776p''R Fabaeformiscandonafabaeformis (Fischer, 1851)''"R Sub-family Dolerocypridinac Triebcl, 1961 Fabaeformiscandona fragilis (Hartwig, 1898)" Genus Dolerocypris Kaufmann, 1900 Fabaeformiscandona hyalina (Brady & Robertson, Dolerocyprisfasciata (0.F. Miiller. 1776)R 1870)' Sub-family Eucypridinae Bronshtein, 1947 Fabaeformiscandona protzi (Hartwig, 1898)'' Genus Eucypris (Vivra, 1891) Fabaeformiscandona siliquosa (Brady, 1910)' Eucypris anglica Fox, 1967' Genus Pseudocandona Kaufmann, 1900 Eucypris crassa (0.F. Miillcr, 1785)" Pseudocandona albicans (Brady, 1 864)"'11' Eucypris dulcifrons Diebel & Pietrzeniuk, 1969' Pseudocandona breuili (Paris. I 920)'' Eucypris elliprica (Baird, 1846)R
62 British freshwater ostracods
Eucypris cf. gemella Bodina, 1961 nian Museum of Natural History, Skopje), Dr D. .1. Horne Eucypris heinrichi Diebcl & Pietrzcniuk, 1978"" (University of Greenwich, London), Dr P. A. Henderson Eucypris lilljeborgi (G. W. Miiller. 1900)"' (National Power Marine Biology Labs, Fawley) and Dr C. Eucypris ornafa (0.F. Muller, 1776)K Meisch (National Museum of Natural History, Luxembourg) Eucypris pigra (Fischer. 1851)""" provided valuable discussions and advice. Dr P. Marmonier Eucypris uirens (Jurine, 1820)PFiR Genus Prionocypris Brady & Norman, 1896 (University of Savoy) and Dr J. E. Robinson (University Prionocypris serrafa (Norman, 186 1 College, London) are thanked for comparative material. Genus Tonnacypris Diebel & Pietrzeniuk, 1975 Special thanks to Dr K. Martens (KBIN, Brussels) who Tonnacypris lufaria (Komch. 1838)"' kindly read through an earlier version of this manuscript, Genus Trajancypris Martens. 1989 made many useful comments and criticisms, and also Trajancypris clauafa (Baird. 1838)"" provided a preprint of his study of Purnlirnnocythere. ? Trujancypris serrafa ((3.W. Muller, 1900)R Thanks also to our many colleagues for their ready help, Sulis-family Herpetocypridinae Kaufmann. 1900 advice, and assistance. Genus lferpefocypris Brady & Norman, 1889 This study was funded by a SERC research grant to J. G. Herpefocypris breuicaudafa Brady & Norman. 1889Pi'R Evans. Herpefocypris cheureuxi Sars, 18Y6PHR ? Herpefocypris ehringsdorfensis Diebel & Wolfschliiger, Manuscript received August 1992 1975' Manuscript accepted May 1993 Herpetocypris repfans (Elaird. 1X35)PHR Gcnus Psychrodromus Danielopol & McKenzie, 1977 REFERENCES Psychrodromus oliuaceus (Brady & Norman. 1 889)'"' Bowman, T. E. & Abele. L. G. 1982. Classification of the Recent Psychrodromus roberfsoni (Brady & Robcrtson, 1889)' Crustacea. In Abele, L. G. (Ed.), The biology of Crusruceu. volume 1. Systemnrics, the fossil record and cwolurion, 1-27. Genus Stenocypria W. IMiiIler. 1901 G. Academic Press, New York. Sfeno,cypriafischeri (Li Ill e borg, 1 883) 'I Brady, G. S. 1910. A revision of the British species of ostracod Subfamily Scottinae Bronshtein, 1947 Crustacea belonging to the subfamilies Candoninae and Genus Scoffia Brady & Norman. 1889 Herpetocypridinae, (with a note on a parasitic worm, by Miss Scoftia browniana (Jones. 1850)' M.V. Lcbour, MSc.). Proceedings of the Zoological Societ-y of Scoftia pseudobrowniantz Kempf. I971 "'' London. London. 1910 1Y4-220. pls 19-30. Scoftia tumida Kcmpf, 197 1 " Brady, G. S. & Norman, A. M. 1889. A monograph of the marine Subfamily Cyprinotinae Bronshtein. 1947 and freshwater Ostracoda of the North Atlantic and of North-western Europe. Section Podocopa. Scienr$c Trunsuc- Genus Heterocypris Claus, 1892 I. tion.r of the Roynl Dublin Socic,rv. Dublin, 4 63-270, pls 8-23. Heferocypris incongruens (Ramdohr, 1808)HR Brady, G. S. & Robertson. D. 1869. Notes on a week's dredging in Heferocypris salina (Brady, 1868)"HR the west of Ireland. Annn1.y nnd Mngazine of Nuturul History Subfamily Isocypridinae Rome, 1965 London, Ser. 4.3: 353-374, pls 18-22. Genus lsocypris G. W. Muller. 1908 Broodbakkcr, N. W. 1983. The genus Srrundrsiu and other Isocypris beauchampi (Paris. 1920)K Cypricercini (Crustacca, Ostracoda) in the West Indies. Part 1. Sub-family Cypridopsinae Kaufmann, 1900 Taxonomy. Bijdragen tot de Dierkunde, Leiden, 53: 3277368. Genus Cypridopsis Brady, 1867 Carbonel, P., Colin, J. P., Danielopol, D. L.. Loffler, H. & Cypridopsis harfwigi G. W. Miiller, 190OPHK Neustrueva. 1. 1988. Palcoccology of limnic ostracodes: a review Cypridopsis lusafica Schaffer, 1943K of some major topics. Pulaeogeography. Polrroclimcrtolog.v, Pulueoecology, Amsterdam, 62: 413-461. Cypridopsis obesa Brady & Robertson, 186YK Carbonnel, G. 1965. Sur un nouveau genre (Pnralimnorvtkrre) et Cypridopsis uidua (0.F. Muller, 1776)PHR une nouvelle espt-ce (P. bouleigensis) d'ostracodcs du Tortonien. Genus Plesiocypridopsis (Romc, 1965) Archives des Sciences, GenPve, Geneva, 18: 146-150. pls 1-2. Plesiocypridopsis newtoni (Brady & Robertson, 1870)i'R Catt, J. A. & Penny, L. F. 1962. The Pleistocene deposits of Genus Sarscypridopsis McKcnzic , 1977 Holdcrncss, East Yorkshire. Proceedings of the Yorkshirr Sarscypridopsis aculeafa (Costa. 1847)HK Geologicul Society. Leeds, 35: 375-420. Genus Cauernocypris Hartmann. 1964 Colin, J. P. & Danielopol, D. L. 1980. Sur la morphologic. la Cauernocypris subrerranea (Wolf, I ~20)"~ systgmatique, la biogiographie et I'kvolution des ostracodes Genus Potamocypris Brady. 1870 Timiriaseviinae (Limnocytheridae). PnlPobiologie conrinentule, Montpellier, 11: 1-51, pls l,-16. Poramocypris arcuata (Sars, 1 903)""' Danielopol. D. L. 1978. Uber Herkunft und Morphiogie der Potamocypris fallar Fox, 1967HR Susswasser-hypogaischen Candoninae (Crustacea, Ostracoda). Potarnocypris fulua (Brady, 1868)'"K Sitzunpberichte rler Osterreichixhen Akudemie cler Pofanzocypris pallida Alm, 1914K Wissenschuften. Mathemcrtisch-naturwissenschrrftliche Klusse. Pofanzocypris similis G. W. Miiller, 1912"R Abwilung 1, Vienna, 187: 1-162. Potanzocypris smaragdina (Vivra. 1891)HK Danielopol, D. L. 1982. Nouvelles don& sur Ies Candoninac Pofanzocypris uariegafa (Brady & Norman. 1889)"' (Ostracoda) hypogis de Roumanie et Yugoslavie. Bulletin rlu Pofanrocypris uillosa (Jurine, 1820)PHR Museum nuriorial rl'Histoire natiirelle tie Paris. (Secr. A)., Paris, 4 Pofanzocypris zschokkei (Kaufmann, 1900)PHR 369-396. Danielopol, D. L. & McKenzie, K. G. 1977. Psychrorlromiis gen. n. (Crustacea, Ostracoda), with redescription of the cypridid genera ACKNOWLIEDGEMENTS Prionocypris and 1Iyodromu.s. Zoologica Scriptu. Stockholm, 6: Our thanks to Dr S. Halsey (Crustacea Section, BMNH) for 301-322. Danielopol. D. L., Martens, K. & Casale, L. M. 1989. Revision of allowing our examination of type material from the the genus Leucocythere Kaufmann, 1892 (Crustacea, Ostracoda, Museum's collections. Professor T. K. Petkovski (Macedo- Limnocytheridae), with the description of a new species. Bulletin
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van her koninklijk Belgisch lnstituut voor Natuurwetenschappen, History Society of Northumbria, Newcastle-upon-Tyne, 55: Riologie, Brussels, 59 63-94. 147-148. Dc Deckker, P. & Forester, R. M. 1988. The use of ostracods to Keen, D. H., Jones, R. L. & Robinson, J. E. 1984. A late Devensian reconstruct continental palaeoenvironmcntal records. In De and early Flandrian fauna and flora from Kildale, north-east Deckker, P., Colin, J. P. & Peypouquet, J. P. (Eds), Ostracoda in Yorkshire. Proceedings of the Yorkshire Geological Society, the Earth Sciences, 175-199 Elsevier Science Publishers, Leeds, 44: 385-397. -, Amsterdam. Kempf, E. K. 1971. Okologie, Taxonomie und Verbreitung der Delorme, L. D. 1970. Freshwater ostracodes of Canada. Part 4. nichtmarincn Ostracoden-Gattung Scottia im Quartar von Families Ilyocyprididae, Notodromadidae, Darwinulidae, Cyther- Europe. Eiszeitalrer und Gegenwurt, Ohringen, 22: 43-63. ideidae and Entocytheridae. Cunadian Journal Zoology, Ottowa, Kempf, E. 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