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Flying Dromaeosaurs
A winged, but flightless, Deinonychus by Stephen A. Czerkas FLYING DROMAEOSAURS Stephen A. Czerkas, Dianshuang Zhang, Jinglu Li and Yinxian Li The Dinosaur Museum, 754 South 200 West, Blanding, Utah 84511, USA; Liaoning Provincial Bureau of Land Resources Management, Liaoning Fossil Administration Office, and Liaoning Museum of Paleontology, Left of Nanshan Park, Beipiao, Liaoning Province 122100, People’s Republic of China. The Dinosaur Museum © 2002 Abstract Dromaeosaurs have been regarded as theropod dinosaurs that were among the closest avian ancestors which were strictly terrestrial having not yet evolved the ability to fly. Consequently, phylogenetic analyses have resulted in the claims of birds having evolved from “the ground up” within a dinosaurian ancestry. Though widely accepted, the relationship between birds and dinosaurs has remained highly controversial and disputed by advocates of birds as having been derived from an arboreal, non-dinosaurian type of archosaur. The cladistical interpretation of the dinosaur/bird relationship hinges upon the presumption of the dromaeosaurs inability to fly. Recent discoveries of dromaeosaurs have revealed impressions of feathers and avian characters in the skeleton that nearly equal and even surpass that of Archaeopteryx. Yet despite this, the ability to fly has been discounted due to the shorter length of the forelimbs. Described below are two such dromaeosaurs, but preserved with impressions of primary flight feathers extending from the manus which demonstrate an undeniable correlation towards the ability to fly. This compelling evidence refutes the popular interpretation of birds evolving from dinosaurs by revealing that dromaeosaurs were already birds and not the non-avian theropod dinosaurs as previously believed. -
Anchiornis and Scansoriopterygidae
SpringerBriefs in Earth System Sciences SpringerBriefs South America and the Southern Hemisphere Series Editors Gerrit Lohmann Lawrence A. Mysak Justus Notholt Jorge Rabassa Vikram Unnithan For further volumes: http://www.springer.com/series/10032 Federico L. Agnolín · Fernando E. Novas Avian Ancestors A Review of the Phylogenetic Relationships of the Theropods Unenlagiidae, Microraptoria, Anchiornis and Scansoriopterygidae 1 3 Federico L. Agnolín “Félix de Azara”, Departamento de Ciencias Naturales Fundación de Historia Natural, CEBBAD, Universidad Maimónides Buenos Aires Argentina Fernando E. Novas CONICET, Museo Argentino de Ciencias Naturales “Bernardino Rivadavia” Buenos Aires Argentina ISSN 2191-589X ISSN 2191-5903 (electronic) ISBN 978-94-007-5636-6 ISBN 978-94-007-5637-3 (eBook) DOI 10.1007/978-94-007-5637-3 Springer Dordrecht Heidelberg New York London Library of Congress Control Number: 2012953463 © The Author(s) 2013 This work is subject to copyright. All rights are reserved by the Publisher, whether the whole or part of the material is concerned, specifically the rights of translation, reprinting, reuse of illustrations, recitation, broadcasting, reproduction on microfilms or in any other physical way, and transmission or information storage and retrieval, electronic adaptation, computer software, or by similar or dissimilar methodology now known or hereafter developed. Exempted from this legal reservation are brief excerpts in connection with reviews or scholarly analysis or material supplied specifically for the purpose of being entered and executed on a computer system, for exclusive use by the purchaser of the work. Duplication of this publication or parts thereof is permitted only under the provisions of the Copyright Law of the Publisher’s location, in its current version, and permission for use must always be obtained from Springer. -
Birds of Bharatpur – Check List
BIRDS OF BHARATPUR – CHECK LIST Family PHASIANIDAE: Pheasants, Partridges, Quail Check List BLACK FRANCOLIN GREY FRANCOLIN COMMON QUAIL RAIN QUAIL JUNGLE BUSH QUAIL YELLOW-LEGGED BUTTON QUAIL BARRED BUTTON QUAIL PAINTED SPURFOWL INDIAN PEAFOWL Family ANATIDAE: Ducks, Geese, Swans GREATER WHITE-FRONTED GOOSE GREYLAG GOOSE BAR-HEADED GOOSE LWSSER WHISTLING-DUCK RUDDY SHELDUCK COMMON SHELDUCK COMB DUCK COTTON PYGMY GOOSE MARBLED DUCK GADWALL FALCATED DUCK EURASIAN WIGEON MALLARD SPOT-BILLED DUCK COMMON TEAL GARGANEY NORTHERN PINTAIL NORTHERN SHOVELER RED-CRESTED POCHARD COMMON POCHARD FERRUGINOUS POCHARD TUFTED DUCK BAIKAL TEAL GREATER SCAUP BAER’S POCHARD Family PICIDAE: Woodpeckers EURASIAN WRYNECK BROWN-CAPPED PYGMY WOODPECKER YELLOW-CROWNED WOODPECKER BLACK-RUMPED FLAMBACK Family CAPITONIDAE: Barbets BROWN-HEADED BARBET COPPERSMITH BARBET Family UPUPIDAE: Hoopoes COMMON HOOPOE Family BUCEROTIDAE: Hornbills INDAIN GREY HORNBILL Family CORACIIDAE: Rollers or Blue Jays EUROPEAN ROLLER INDIAN ROLLER Family ALCEDINIDAE: Kingfisher COMMON KINGFISHER STORK-BILLED KINGFISHER WHITE-THROATED KINGFISHER BLACK-CAPPED KINGFISHER PIED KINGFISHER Family MEROPIDAE: Bee-eaters GREEN BEE-EATER BLUE-CHEEKED BEE-EATER BLUE-TAILED BEE-EATER Family CUCULIDAE: Cuckoos, Crow-pheasants PIED CUCKOO CHESTNUT-WINGED CUCKOO COMMON HAWK CUCKOO INDIAN CUCKOO EURASIAN CUCKOO GREY-BELLIED CUCKOO PLAINTIVE CUCKOO DRONGO CUCKOO ASIAN KOEL SIRKEER MALKOHA GREATER COUCAL LESSER COUCAL Family PSITTACIDAS: Parrots ROSE-RINGED PARAKEET PLUM-HEADED PARKEET Family APODIDAE: -
LETTER Doi:10.1038/Nature14423
LETTER doi:10.1038/nature14423 A bizarre Jurassic maniraptoran theropod with preserved evidence of membranous wings Xing Xu1,2*, Xiaoting Zheng1,3*, Corwin Sullivan2, Xiaoli Wang1, Lida Xing4, Yan Wang1, Xiaomei Zhang3, Jingmai K. O’Connor2, Fucheng Zhang2 & Yanhong Pan5 The wings of birds and their closest theropod relatives share a ratios are 1.16 and 1.08, respectively, compared to 0.96 and 0.78 in uniform fundamental architecture, with pinnate flight feathers Epidendrosaurus and 0.79 and 0.66 in Epidexipteryx), an extremely as the key component1–3. Here we report a new scansoriopterygid short humeral deltopectoral crest, and a long rod-like bone articu- theropod, Yi qi gen. et sp. nov., based on a new specimen from the lating with the wrist. Middle–Upper Jurassic period Tiaojishan Formation of Hebei Key osteological features are as follows. STM 31-2 (Fig. 1) is inferred Province, China4. Yi is nested phylogenetically among winged ther- to be an adult on the basis of the closed neurocentral sutures of the opods but has large stiff filamentous feathers of an unusual type on visible vertebrae, although this is not a universal criterion for maturity both the forelimb and hindlimb. However, the filamentous feath- across archosaurian taxa12. Its body mass is estimated to be approxi- ers of Yi resemble pinnate feathers in bearing morphologically mately 380 g, using an empirical equation13. diverse melanosomes5. Most surprisingly, Yi has a long rod-like The skull and mandible are similar to those of other scansoriopter- bone extending from each wrist, and patches of membranous tissue ygids, and to a lesser degree to those of oviraptorosaurs and some basal preserved between the rod-like bones and the manual digits. -
Oldest Known Dinosaurian Nesting Site and Reproductive Biology of the Early Jurassic Sauropodomorph Massospondylus
Oldest known dinosaurian nesting site and reproductive biology of the Early Jurassic sauropodomorph Massospondylus Robert R. Reisza,1, David C. Evansb, Eric M. Robertsc, Hans-Dieter Suesd, and Adam M. Yatese aDepartment of Biology, University of Toronto Mississauga, Mississauga, ON L5L 1C6, Canada; bRoyal Ontario Museum, Toronto, ON M5S 2C6, Canada; cSchool of Earth and Environmental Sciences, James Cook University, Townsville, 4811 QLD, Australia; dDepartment of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013; and eBernard Price Institute for Palaeontological Research, University of the Witwatersrand, Wits 2050, Johannesburg, South Africa Edited by Steven M. Stanley, University of Hawaii, Honolulu, HI, and approved December 16, 2011 (received for review June 10, 2011) The extensive Early Jurassic continental strata of southern Africa clutches and recognition of the earliest known dinosaurian nesting have yielded an exceptional record of dinosaurs that includes complex at the Rooidraai locality. scores of partial to complete skeletons of the sauropodomorph Massospondylus, ranging from embryos to large adults. In 1976 an Results incomplete egg clutch including in ovo embryos of this dinosaur, Our work at the Rooidraai locality has yielded multiple in situ the oldest known example in the fossil record, was collected from clutches of eggs as well as fragmentary eggshell and bones, all from a road-cut talus, but its exact provenance was uncertain. An exca- a 2-m-thick interval of muddy siltstone 25 m from the top of the vation program at the site started in 2006 has yielded multiple in Lower Jurassic Upper Elliot Formation (“Stormberg Group,” situ egg clutches, documenting the oldest known dinosaurian Karoo Supergroup) (11). -
A Phylogeny and Timescale for Marsupial Evolution Based on Sequences for Five Nuclear Genes
J Mammal Evol DOI 10.1007/s10914-007-9062-6 ORIGINAL PAPER A Phylogeny and Timescale for Marsupial Evolution Based on Sequences for Five Nuclear Genes Robert W. Meredith & Michael Westerman & Judd A. Case & Mark S. Springer # Springer Science + Business Media, LLC 2007 Abstract Even though marsupials are taxonomically less diverse than placentals, they exhibit comparable morphological and ecological diversity. However, much of their fossil record is thought to be missing, particularly for the Australasian groups. The more than 330 living species of marsupials are grouped into three American (Didelphimorphia, Microbiotheria, and Paucituberculata) and four Australasian (Dasyuromorphia, Diprotodontia, Notoryctemorphia, and Peramelemorphia) orders. Interordinal relationships have been investigated using a wide range of methods that have often yielded contradictory results. Much of the controversy has focused on the placement of Dromiciops gliroides (Microbiotheria). Studies either support a sister-taxon relationship to a monophyletic Australasian clade or a nested position within the Australasian radiation. Familial relationships within the Diprotodontia have also proved difficult to resolve. Here, we examine higher-level marsupial relationships using a nuclear multigene molecular data set representing all living orders. Protein-coding portions of ApoB, BRCA1, IRBP, Rag1, and vWF were analyzed using maximum parsimony, maximum likelihood, and Bayesian methods. Two different Bayesian relaxed molecular clock methods were employed to construct a timescale for marsupial evolution and estimate the unrepresented basal branch length (UBBL). Maximum likelihood and Bayesian results suggest that the root of the marsupial tree is between Didelphimorphia and all other marsupials. All methods provide strong support for the monophyly of Australidelphia. Within Australidelphia, Dromiciops is the sister-taxon to a monophyletic Australasian clade. -
71St Annual Meeting Society of Vertebrate Paleontology Paris Las Vegas Las Vegas, Nevada, USA November 2 – 5, 2011 SESSION CONCURRENT SESSION CONCURRENT
ISSN 1937-2809 online Journal of Supplement to the November 2011 Vertebrate Paleontology Vertebrate Society of Vertebrate Paleontology Society of Vertebrate 71st Annual Meeting Paleontology Society of Vertebrate Las Vegas Paris Nevada, USA Las Vegas, November 2 – 5, 2011 Program and Abstracts Society of Vertebrate Paleontology 71st Annual Meeting Program and Abstracts COMMITTEE MEETING ROOM POSTER SESSION/ CONCURRENT CONCURRENT SESSION EXHIBITS SESSION COMMITTEE MEETING ROOMS AUCTION EVENT REGISTRATION, CONCURRENT MERCHANDISE SESSION LOUNGE, EDUCATION & OUTREACH SPEAKER READY COMMITTEE MEETING POSTER SESSION ROOM ROOM SOCIETY OF VERTEBRATE PALEONTOLOGY ABSTRACTS OF PAPERS SEVENTY-FIRST ANNUAL MEETING PARIS LAS VEGAS HOTEL LAS VEGAS, NV, USA NOVEMBER 2–5, 2011 HOST COMMITTEE Stephen Rowland, Co-Chair; Aubrey Bonde, Co-Chair; Joshua Bonde; David Elliott; Lee Hall; Jerry Harris; Andrew Milner; Eric Roberts EXECUTIVE COMMITTEE Philip Currie, President; Blaire Van Valkenburgh, Past President; Catherine Forster, Vice President; Christopher Bell, Secretary; Ted Vlamis, Treasurer; Julia Clarke, Member at Large; Kristina Curry Rogers, Member at Large; Lars Werdelin, Member at Large SYMPOSIUM CONVENORS Roger B.J. Benson, Richard J. Butler, Nadia B. Fröbisch, Hans C.E. Larsson, Mark A. Loewen, Philip D. Mannion, Jim I. Mead, Eric M. Roberts, Scott D. Sampson, Eric D. Scott, Kathleen Springer PROGRAM COMMITTEE Jonathan Bloch, Co-Chair; Anjali Goswami, Co-Chair; Jason Anderson; Paul Barrett; Brian Beatty; Kerin Claeson; Kristina Curry Rogers; Ted Daeschler; David Evans; David Fox; Nadia B. Fröbisch; Christian Kammerer; Johannes Müller; Emily Rayfield; William Sanders; Bruce Shockey; Mary Silcox; Michelle Stocker; Rebecca Terry November 2011—PROGRAM AND ABSTRACTS 1 Members and Friends of the Society of Vertebrate Paleontology, The Host Committee cordially welcomes you to the 71st Annual Meeting of the Society of Vertebrate Paleontology in Las Vegas. -
Supplementary Material
Pterocles alchata (Pin-tailed Sandgrouse) European Red List of Birds Supplementary Material The European Union (EU27) Red List assessments were based principally on the official data reported by EU Member States to the European Commission under Article 12 of the Birds Directive in 2013-14. For the European Red List assessments, similar data were sourced from BirdLife Partners and other collaborating experts in other European countries and territories. For more information, see BirdLife International (2015). Contents Reported national population sizes and trends p. 2 Trend maps of reported national population data p. 3 Sources of reported national population data p. 5 Species factsheet bibliography p. 6 Recommended citation BirdLife International (2015) European Red List of Birds. Luxembourg: Office for Official Publications of the European Communities. Further information http://www.birdlife.org/datazone/info/euroredlist http://www.birdlife.org/europe-and-central-asia/european-red-list-birds-0 http://www.iucnredlist.org/initiatives/europe http://ec.europa.eu/environment/nature/conservation/species/redlist/ Data requests and feedback To request access to these data in electronic format, provide new information, correct any errors or provide feedback, please email [email protected]. THE IUCN RED LIST OF THREATENED SPECIES™ BirdLife International (2015) European Red List of Birds Pterocles alchata (Pin-tailed Sandgrouse) Table 1. Reported national breeding population size and trends in Europe1. Country (or Population estimate Short-term -
Transport of Water by Adult Sandgrouse to Their Young Tom J
THE CONDOR VOLUME69 JULY-AUGUST,1967 NUMBER4 TRANSPORT OF WATER BY ADULT SANDGROUSE TO THEIR YOUNG TOM J. CADE and GORDONL. MACLEAN In 1896 the English aviculturist Meade-Waldo published an astonishing and seemingly incredible account of how the males of sandgrouse that he successfully bred in captivity carried water to their young in their breast feathers. To quote from his original report: As soon as the young were out of the nest (when twelve hours old) a very curious habit developed itself in the male. He would rub his breast violently up and down on the ground, a motion quite distinct from dusting, and when all awry he would get into his drinking water and saturate the feathers of the under parts. When soaked he would go through the motions of flying away, nodding his head, etc. Then, remembering his family were close by, would run up to the hen, make a demonstration, when the young would run out, get under him, and suckthe water from his breast. This is no doubt the way that water is conveyed to the young when far out on waterless plains. The young . are very independent, eating hard seed and weeds from the first, and roosting independently of their parents at ten days old (Meade-Waldo, 1896). See also Meade- Waldo (1921). Despite the fact that .Meade-Waldo (1897 ; 1921) observed 61 broods from three different species of sandgrouse hatched in his aviaries between 189.5 and l915, and soon received confirmation from another breeder for two species (St. Quintin, 1905), and despite the fact that field naturalists and native hunters have frequently observed wild male sandgrouse wetting their breast feathers at water holes in the way described (Meade-Waldo, 1906; Buxton, 1923; Heim de Balsac, 1936; Hoesch, 1955), the idea that the young do receive water in this exceptional way has met with a great deal of scepticism (Archer and Godman, 1937; Meinertzhagen, 1954, 1964; Hiie and Etchkcopar, 1957; Schmidt-Nielsen, 1964). -
Re-Evaluation of the Haarlem Archaeopteryx and the Radiation of Maniraptoran Theropod Dinosaurs Christian Foth1,3 and Oliver W
Foth and Rauhut BMC Evolutionary Biology (2017) 17:236 DOI 10.1186/s12862-017-1076-y RESEARCH ARTICLE Open Access Re-evaluation of the Haarlem Archaeopteryx and the radiation of maniraptoran theropod dinosaurs Christian Foth1,3 and Oliver W. M. Rauhut2* Abstract Background: Archaeopteryx is an iconic fossil that has long been pivotal for our understanding of the origin of birds. Remains of this important taxon have only been found in the Late Jurassic lithographic limestones of Bavaria, Germany. Twelve skeletal specimens are reported so far. Archaeopteryx was long the only pre-Cretaceous paravian theropod known, but recent discoveries from the Tiaojishan Formation, China, yielded a remarkable diversity of this clade, including the possibly oldest and most basal known clade of avialan, here named Anchiornithidae. However, Archaeopteryx remains the only Jurassic paravian theropod based on diagnostic material reported outside China. Results: Re-examination of the incomplete Haarlem Archaeopteryx specimen did not find any diagnostic features of this genus. In contrast, the specimen markedly differs in proportions from other Archaeopteryx specimens and shares two distinct characters with anchiornithids. Phylogenetic analysis confirms it as the first anchiornithid recorded outside the Tiaojushan Formation of China, for which the new generic name Ostromia is proposed here. Conclusions: In combination with a biogeographic analysis of coelurosaurian theropods and palaeogeographic and stratigraphic data, our results indicate an explosive radiation of maniraptoran coelurosaurs probably in isolation in eastern Asia in the late Middle Jurassic and a rapid, at least Laurasian dispersal of the different subclades in the Late Jurassic. Small body size and, possibly, a multiple origin of flight capabilities enhanced dispersal capabilities of paravian theropods and might thus have been crucial for their evolutionary success. -
A Model Based on Curvatures of Extant Avian Ungual Bones
Inferring lifestyle for Aves and Theropoda: A model based on curvatures of extant avian ungual bones A thesis submitted to the University of Manchester for the degree of Master of Science by Research in the Faculty of Science & Engineering 2019 Savannah E. Cobb School of Earth and Environmental Sciences Contents List of Figures.........................................................................................................................4-5 List of Tables..............................................................................................................................6 List of Abbreviations..............................................................................................................7-8 Abstract......................................................................................................................................9 Declaration...............................................................................................................................10 Copyright Statement...............................................................................................................11 Acknowledgements..................................................................................................................12 1 Literature Review........................................................................................................13 1.1 Avians, avialans, and theropod dinosaurs..........................................................13 1.2 Comparative study and claws............................................................................18 -
The Water-Holding Mechanism of Sandgrouse Feathers by A
'. Exp. BM. (1972), 56, 195-200 ith 2 text-figures Printed in Great Britain THE WATER-HOLDING MECHANISM OF SANDGROUSE FEATHERS BY A. M. RIJKE Department of Materials Science, University of Virginia, Charlottesville, Virginia 22901 {Received 15 June 1971) INTRODUCTION The detailed studies of Cade & Maclean (1967) on the mechanism of water transport by the sandgrouse Pterocles have contributed important evidence to the long-standing controversy regarding the manner in which the young obtain their drinking water. The field observations of these authors have shown conclusively that the male of the species soaks his abdominal feathers by squatting down in water-holes, usually during the early hours. He then speeds off to the nesting site where the young gather around his exposed abdomen and obtain their water by stripping his belly feathers with their beaks. In addition to this behaviour it has been noted that the abdominal feathers of the male, and to a lesser extent those of the female, show a number of structural charac- teristics which render the uptake of water particularly effective. The distal fifths of these feathers are very similar to the feathers of other parts of the sandgrouse body in that they show the conventional structural array of barbs and barbules with their characteristic water-repelling properties. The structural parameter (r+d)jr for both male and female rates between 5-5 and 6-2 with slightly higher values within this range for the dorsal side. This result is in line with similar data on other terrestrial birds, indicating an effective water repellency without the necessity of preventing water penetration (Rijke, 1970).