<<

Journal of Neuropterology 3: 9-23,2000 (2001)

Annotated List of the Neuroptera of Hispaniola, with New Faunistic Records of Some Species

D. E. PCrez-Gelabert (1) and 0. S. Flint Jr. (2) (1) 5714 Ridgway Ave., Rockville, MD 20851, U.S.A. E-mail: [email protected] (2) National Museum of Natural History, Smithsonian Institution, Washington, D. C. 20560-0105, U.S.A. E-mail: [email protected]

Keywords: Neuroptera, Ascalaphids, Lacewings, Antlions, , , New records.

SUMMARY

An annotated list and some new faunistic records of Hispaniolan Neuroptera are presented, based on literature records and specimens in the collections of the National Museum of Natural History, Washington, D. C. and the Carnegie Museum, Pittsburgh. Seventeen species are newly recorded here for the island. Fifty-two extant species within 26 genera are known to us. Additionally, three fossil species and four other genera of Neuroptera have been reported from Dominican amber. We provide synonymic, type and distribution data for each species that should enhance the comprehensive characterization of this group in Hispaniola.

INTRODUCTION

The Hispaniolan fauna of Neuroptera has been the subject of only haphazard studies by visiting collectors and entomologists. It appears that a significant number of species still remain to be discovered on the island. Systematic studies on Neotropical Neuroptera have been hindered by the unsettled taxonomic state of several families, making it difficult to be confident about the identity or status of particular taxa. Locally, lack of tradition in faunistic studies and the paucity of research collections necessary for comparative studies have also discouraged work on this group. The first species of Neuroptera reported from Hispaniola was the ascalaphid Ululodes villosus described by BEAUVOIS (1805). Two additional species were described prior to 1900 by WALKER (1853, 1860). The only faunistic study of the Neuroptera of the island was published by SMITH (1931), who was stationed near Port-au-Prince, Haiti while working for the Service Technique du DCpartement de I'Agriculture from July 1928 to March 1930. Smith reported 24 species, most collected in the area of the agricultural station at Damien, based on identifications by Nathan Banks. The recent catalogues of the neuropteran fauna of the New 10 PEREZ-GELABERT, D. E. & FLINT Jr., 0. S.

World produced by PENNY (1977) and PENNY et al. (1997) have summarized the early works on the insular fauna and provided the initial basis for this paper. The Hispaniolan fauna of Neuroptera contains at least 52 extant species placed in 26 genera and 8 families. Seventeen species, six genera and two families are newly recorded here for the extant Neuroptera fauna in the island. Three fossil species, a Hemerobiid, a Coniopterygid and a Myrmeleontid, have been described from Dominican amber. Four other taxa identified only to generic level (Leucochrysa, Coniopteryx, Hemerobius and Mantispa) and the family Ascalaphidae have also been reported in Dominican amber. This list is intended as a useful tool to address more effectively the remaining taxonomic problems involving Hispaniolan neuropterans, encourage their collection and to aid in the discovery of new taxa. We report new materials only for the previously unrecorded genera or species that could be identified with certainty in the collections of the National Museum of Natural History, Washington, D. C. and the Carnegie Museum of Natural History, Pittsburgh. We list all relevant citations of Neuroptera species for the island of Hispaniola. Type data includes kind of type, sex, deposition and type locality. Finally the known general distribution of each species is noted. Institutions and collections holding the types of these species are coded in this paper using the following abbreviations: A.M.N.H. (American Museum of Natural History), New York, U.S.A.; B.M.N.H. (Natural History Museum), London, England; C.A.A.S. (California Academy of Sciences), San Francisco, California, U.S.A.; C.M.N.H. (Carnegie Museum of Natural History), Pittsburgh, Pennsylvania, U.S.A.; H.M.B. (Museum fur Naturkunde der Humboldt Universitat), Berlin, Germany; I.E.S.H. (Instituto de Ecologia y Sistemitica), La Habana, Cuba; I.R.S.N.B. (Institut Royal des Sciences Naturelles de Belgique), Brussels, Belgium; J.W.C. (Personal collection of Jim Work), Oregon, U.S.A.; M.C.Z. (Museum of Comparative Zoology), Cambridge, Massachussetts, U.S.A.; M.N.H.N.P. (MusCum National d7Histoire Naturelle), Paris, France; N.M.N.H. (National Museum of Natural History), Washington, D. C., U.S.A.; and Z.S.M.C. (Zoologische Staatssamlung), Munich, Germany.

LIST OF SPECIES

Order NEUROPTERA Family ASCALAPHIDAE Subfamily ASCALAPHINAE Genus ULULODES Currie, 1899 Ululodes banksi Weele, 1908 UluEodes banksi: Smith, 1931: 819. Ululodes banksi: NavAs, 1932: 34. Type: Syntypes 2 1 9, B.M.N.H. Type locality: Jamaica: Rae-Town. Distribution: Jamaica, Haiti. Journal of Neuropterology 3: 9-23, 2000 (2001) 11

Ululodes sanctidomingi Weele, 1908 Ululodes sancti-domingi Weele, 1908: 115, fig. 74. Ululodes sancti-dominici: NavBs, 1913: 30. Ululodes macleayana venezolensis: Smith, 1931: 818, fig. 19 [erroneous identification]. Ululodes sanctidomingi: Smith, 1931 : 8 19. Ululodes sancti-domingi: Banks, 1938b: 179. Type: Syntypes 2 99, B.M.N.H. and I.R.S.N.B. Type locality: "San Domingo"and "Haiti". Distribution: Hispaniola.

Ululodes smithi Banks, 1938 Ululodes subvertens: Smith, 1931 : 8 18 [erroneous identification]. Ululodes smithi Banks, 1938b: 179. Type: Holotype 9, M.C.Z. no. 23192. Type locality: Haiti: Port-au-Prince. Distribution: Haiti.

Ululodes villosus (Beauvois, 1805) Ascalaphus villosus Beauvois, 1805: 86, pl. 7. Ululodes villosa: Weele, 1908: 118. Ululodes villosa: NavBs, 1913: 30. Ululodes villosus: Banks, 1938b: 178. Type: Holotype Q, B.M.N.H. Type locality: "Saint Domingue". Distribution: Hispaniola. Comments: Aggregations of this species on low lying vegetation were first reported by NEEDHAM (1941). They seem to be a common occurrence in the Dominican Republic.

Family CHRYSOPIDAE Subfamily APOCHRYSINAE Genus LAINIUS NavBs, 1913 Lainius constellatus NavBs, 1913 New generic record for Hispaniola! Type: 3 syntypes, sex unknown, B.M.N.H. Type locality: Guatemala: Cerro Zunil. Distribution: Guatemala, Dominican Republic. Specimens: 1 individual from DOMINICAN REPUBLIC: Pedernales Prov., 26 km N. Cabo Rojo, 18O06' N, 71°38' W, 730 m, VI1.16.1992, C. Young, R. Davidson, S. Thompson, J. Rawlins, mesic deciduous forest with scattered pines. C.M.N.H. Determined by N. Penny in 1999.

Subfamily CHRY SOPINAE Genus CERAEOCHRYSA Adams, 1982 Ceraeochrysa claveri (NavBs, 1911) Chrysopa haitiensis: Smith, 1931 : 806. Ceraeochrysa claveri: Adams, 1982: 72. Ceraeochrysa claveri: Adams & Penny, 1985: 444. Type: Lectotype d, M.N.H.N.P. Type locality: Colombia. Distribution: Colombia, Cuba, Brazil, Mexico, Haiti. 12 PEREZ-GELABERT,D. E. & FLINT Jr., 0. S.

Comments: Smith's Haitian described the larva of this species as a trash carrier.

Ceraeochrysa cincta (Schneider, 1851) New record for Hispaniola! Type: Type, sex unknown, H.M.B. Type locality: Brazil. Distribution: Argentina to Mexico, Cuba, Galapagos, United States (Florida), Dominican Republic. Specimens: 1 6' from DOMINICAN REPUBLIC: , VI.3-4.1969, Flint & Gbmez, N.M.N.H. Determined by P. A. Adams in 1993.

Ceraeochrysa cubana (Hagen, 1861) Chrysopa cubana Hagen, 1861: 215. Chrysopa freemani: Smith, 1931: 81 1. Ceraeochrysa cubana: Adams, 1982: 72. Ceraeochrysa cubana: Adams & Penny, 1985: 445, figs. 135-139, pl. 17. Type: Neotype 0, M.C.Z. Neotype locality: Cuba: Soledad, near Cienfuegos. Distribution: Cuba, Haiti, Jamaica, Barbados, Florida, Mexico, Guyana, Brazil.

Ceraeochrysa valida (Banks, 1895) Chrysopa lioni: NavBs, 1927b: 54. Chrysopa lioni: Smith, 1931: 812 [no specimens were studied]. Chrysopa damiensis: Smith, 1931: 803, pl. 2, figs. 1, 11. Chrysopa wolcotti: Smith, 1931: 8 10. Ceraeochrysa valida: Adams, 1982: 73. Type: Syntypes 2 QQ, M.C.Z. Type locality: Mexico: Baja California. Distribution: Haiti, Puerto Rico, Mexico, Ecuador, Brazil, Guyana, Guatemala, United States (Florida and Texas). Comments: The larva of this species was described by SMITH (1931) when he studied C. damiensis.

Genus CHRYSOPA Leach, 1815 Chrysopa domingensis Banks, 1941 Chrysopa domingensis Banks, 1941a: 394. Type: Holotype 9, M.C.Z. no. 25289. Type locality: Dominican Republic: Constanza. Distribution: Dominican Republic. Comments: BROOKS & BARNARD (1990) did not list this species. It is the only one of the Hispaniolan species originally described in the genus Chrysopa still remaining in this genus. Further study will probably place it within one of the other five genera of Chrysopidae cited for Hispaniola.

Genus CHRYSOPERLA Steinmann, 1964 Chrysoperla dozieri (Smith, 1931) Chrysopa dozieri Smith, 1931 : 808, fig. 4. Chrysoperla dozieri: Brooks, 1994: 166, figs. 196-202. Type: Holotype d, M.C.Z. Type locality: Haiti: Port- au-Prince. Distribution: Haiti. Journal of Neuropterology 3: 9-23, 2000 (2001) 13

Chrysoperla exotera (NavBs, 1913) Chrysopa estradai NavBs, 1924: 334. Chrysopa estradai: Smith, 1931: 809. Clzrysopa exterior: Smith, 1931: 804, fig. 2. Chrysopa exotera: Brooks, 1994: 167. Type: Holotype 9, B.M.N.H. Type locality: Guatemala: Duenas, Capetillo. Distribution: Guatemala, Cuba, Mexico, Costa Rica, Haiti.

Chrysoperla externa (Hagen, 1861 ) New record for Hispaniola! Type: Neotype 9, M.C.Z. Neotype locality: United States: Winter Park, Florida. Distribution: Argentina, Mexico, Bahamas, Barbados, Grand Cayman, Trinidad, United States (Florida), Dominican Republic. Material examined: 2 BB 1 9 from DOMINICAN REPUBLIC: Pedernales Prov., roadside, 33 km N. Pedernales, 1100 m, 18O13.2' N, 71°45.4' W, 111.19.1999, 0. S. Flint, Jr., N.M.N.H. 1 0 from Santo Domingo, Urb. El Portal, 111.29.1999, 0. S. Flint, Jr., N.M.N.H. Determined by 0. S. Flint, Jr. in 1999.

Chrysoperla harrisii (Fitch, 1855) New record for Hispaniola! Type: Holotype B,M.C.Z. Type locality: United States: New York. Distribution: Canada, United States, Mexico, Dominican Republic. Specimens: 1 B from DOMINICAN REPUBLIC: La Vega Prov., Hotel Montaiia, ca. 520 m, 10 km NE Jarabacoa, V.28.1973, Don & Mignon Davis, N.M.N.H. 1 0 1 9 from , Constanza, 1164 m, Hotel Nueva Suiza, V.29.1973, Don & Mignon Davis, N.M.N.H. Determined by P. A. Adams in 1992.

Chrysoperla rufilabris (Burmeister, 1839) New record for Hispaniola! Type: Syntypes, sexes unknown, destroyed. Type locality: United States and Mexico. Distribution: United States, Canada, Mexico, Bermuda, Dominican Republic. Specimens: 1 individual from DOMINICAN REPUBLIC: Pedernales Prov., 30 km N. Cabo Rojo, 18O07' N, 71'39' W, 1070 m, VII.23-24.1990, C. Young, J. E. Rawlins, S. Thompson. C.M.N.H. 2 individuals from Pedernales Prov., 1 km E. Cabo Rojo, 17O55' N, 71°39' W, 10 m, VII.30.1990, J. E. Rawlins, C. W. Young, S. A. Thompson. C.M.N.H. 1 individual from Independencia Prov., 4 km S. Los Pinos, Loma de Vientos, 1S035' N, 71°46' W, 455 m, VII.23.1992, R. Davidson, J. Rawlins, S. Thompson, C. Young. Semiarid deciduous forest with pastures. C.M.N.H. All determined by N. Penny in 1999.

Genus CHRYSOPODES NavBs, 1913 Chrysopodes (Neosuarius) collaris (Schneider, 1851) Chrysopa thoracica: Walker, 1853: 243. Chrysopa thoracica: Smith, 1931 : 807, fig. 3. Chrysopodes (Neosuarius) collaris: Adams & Penny, 1985: 436, figs. 14 PEREZ-GELABEKT,D. E. & FLINT Jr., 0. S.

91-97, pl. 13. Type: Holotype d, H.M.B. Type locality: Puerto Rico. Distribution: Puerto Rico, Dominican Republic, United States (Texas and Florida), Mexico, Colombia, Venezuela, Guyana.

Genus EREMOCHRYSA Banks, 1903 Eremochrysa sp. New generic record for Hispaniola! Material examined: 4 individuals without abdomen from DOMINICAN REPUBLIC: Dunas de Las Calderas, Bani, VIII.23.1966, E. de J. Marcano F., in Cassia angustifolia, N.M.N.H. Determined by 0. S. Flint, Jr.

Genus LEUCOCHRYSA MacLachlan, 1868 Leucochrysa (Leucochrysa) christophei Banks, 1938 Leucochrysa christophei Banks, 1938a: 292. Type: Holotype 9, M.C.Z no. 22679. Type locality: Desbarriere, Haiti: Mt. La Hotte. Distribution: Haiti. Comments: Known only from the type locality.

Leucochrysa (Leucochrysa) haitiensis Smith, 1931 Leucochrysa haitiensis Smith, 1931: 812, fig. 5, 8, 20. Type: Holotype 0, M.C.Z. Type locality: [Haiti]. Distribution: Haiti.

Leucochrysa (Leucochrysa) insularis (Walker, 1853) New record for Hispaniola! Type: Holotype d, B.M.N.H. Type locality: Jamaica. Distribution: Eastern United States, Puerto Rico, Cuba, Jamaica, Dominican Republic. Material examined: 1 0 from DOMINICAN REPUBLIC: La Vega prov., La Palma, 12 km E. of El Rio, VI.2-13.1969, Flint & G6mez. 1 d 1 Q from Convento, 12 km S. of Constanza, VI.6-13.1969, Flint & G6mez. 1 0 from Pedernales Prov., streams & falls, 19 km N. Pedernales, 230 m, 18O09.2' N, 71'44.8' W, 111.19.1999, 0. S. Flint, Jr. 1 6 from Pedernales Prov., Rio Mulito, 21 km N. Pedernales, 270 m, 18'09.3' N, 71°45.6' W, 111.20.1999, 0. S. Flint, Jr. 1 cT Barahona Prov., San Rafael, 8.3 km S. Bahoruco, 30 m, 18'01.9' N, 71°08.4' W, 111.22.1999, 0. S. Flint, Jr. All determined by 0. S. Flint, Jr. in 1999 and deposited at N.M.N.H.

Leucochrysa (Nodita) ocampina Banks, 1941 Nodita ocampina Banks, 1941a: 395. Type: Holotype, sex unknown, M.C.Z no. 25290. Type locality: Dominican Republic: Monte Diego de Ocampo. Distribution: Dominican Republic. Comments: This species was overlooked by BROOKS & BARNARD (1990). In their generic revision they proposed Nodita as a subgenus of Leucochrysa. This would have produced the combination used above. Journal of Neuropterology 3: 9-23,2000 (2001)

?-Leucochrysasp. Leucochrysa: Poinar & Poinar, 1999: 203. Fossil in Dominican amber. Genus MELEOMA Fitch, 1855 Meleoma sp. New generic record for Hispaniola! Specimens: 1 individual from DOMINICAN REPUBLIC: Independencia Prov., 3 km ESE of El Aguacate, north slope Sierra de Bahoruco, 1980 m, 18'18' N, 71°42' W, IX.28-29.1991, J. Rawlins, S. Thompson, C. Young, R. Davidson, Dense clound forest. C.M.N.H. 1 specimen from Independencia Prov., Sierra de , near crest, 5.5 km NNW Angel Feliz, 18O41' N, 7 1'47' W, 1750 m, VII.21-22.1992, J. Rawlins, S. Thompson, C. Young, R. Davidson, dense cloud forest. C.M.N.H. Determined by N. Penny in 1999.

Genus PLESIOCHRYSA Adams, 1982 Plesiochrysa brasiliensis (Schneider, 1851) Chrysopa cubana: NavBs, 1921 : 120 [preoccupied name]. Chrysopa antillana: Smith, 1931: 809. Type: Holotype 9, H.M.B. Type locality: Brazil. Distribution: Cuba, Colombia, Brazil, Dominican Republic, Costa Rica, Mexico, United States (Florida).

Plesiochrysa elongata (NavBs, 1913) New record for Hispaniola! Type: Holotype d', Z.S.M.C. Type locality: Jamaica. Distribution: Jamaica, Guatemala, Peru, Guadeloupe, Costa Rica, Honduras, Brazil, Dominican Republic. Specimens: 6 individuals from DOMINICAN REPUBLIC: Pedernales Prov., 5 Km NE Los Arroyos, 18O15' N, 7 1°45' W, 1680 m, VII. 17-18.1990, C. Young, J. E. Rawlins, S. Thompson. C.M.N.H. 4 individuals from Pedernales Prov., 30 km N. Cabo Rojo, 18O07' N, 71'39' W, 1070 m, VII.23-24.1990, J. E. Rawlins, C. Young, S. Thompson. C.M.N.H. Determined by N. Penny in 1999.

Family CONIOPTERYGIDAE Subfamily ALEUROPTERYGINAE Genus NEOCONIS Enderlein, 1929 Neoconis cubana (Banks, 1938) Neoconis cubana: Meinander, 1990: 28. Type: Holotype 0, M.C.Z. Type locality: Cuba: Oriente. Distribution: Cuba, Dominican Republic.

Genus SPZLOCONIS Enderlein, 1907 tSpiloconis glaesaria Meinander, 1998 Spiloconis glaesaria Meinander, 1998: 33, figs. 1-4. Type: Holotype 9, A.M.N.H. DR-10-59. Fossil in Dominican amber. 16 PEREZ-GELABERT, D. E. & FLINT Jr., 0. S.

Comments: MEINANDER (1998) indicates that recent species of Spiloconis are found in south and east Asia, Australia and the Fiji islands. The finding of this genus in Dominican amber indicates that it used to have a more extensive distribution.

Subfamily CONIOPTERYGINAE Genus CONZOPTERYX Curtis, 1834 Coniopteryx haitiensis Smith, 1931 Coniopteryx haitiensis Smith, 1931 : 8 19, figs. 7a, 7b. Coniopteryx haitiensis: Meinander, 1972: 198. Coniopteryx haitiensis: Meinander, 1986: 41. Type: Holotype Q, M.C.Z. Type locality: Haiti: Damien. Distribution: Haiti. Comments: Being based on a female, the species is currently unrecognizable. There are questionable records from Cuba and Jamaica.

Coniopteryx (Scotoconiopteryx)fumata Enderlein, 1907 Coniopteryx (Scotoconiopteryx) fumata: Meinander, 1990: 48. Type: Holotype 0, lost? Type locality: Brazil: Santa Catharina. Distribution: Dominican Republic, Colombia, Venezuela, Brazil. lconiopteryx sp. Coniopteryx: Meinander, 1998: 35, fig. 5. Specimen 1, 9 A.M.N.H. DR-10-4 and specimen 2, d A.M.N.H. DR-10-15. Fossils in Dominican amber.

Genus SEMZDA LZS Enderlein, 1905 Semidalis marginalis (Banks, 1930) Semidalis marginalis: Meinander, 1990: 78. Type: Holotype 0, M.C.Z. Type locality: Cuba: Sierra Maestra. Distribution: Cuba, Dominican Republic.

Family DILARIDAE Subfamily NALLACHIINAE Genus NALLACHZUS NavBs, 1909 Nallachius pulchellus (Banks, 1938) New family and generic record for Hispaniola! Type: Lectotype 0, M.C.Z. No. 22681. Type locality: Cuba: Central Soledad. Distribution: Cuba, United States, Dominican Republic. Material examined: 1 d from DOMINICAN REPUBLIC: Dajab6n province, 13 km S. , ca. 400 m, V.20-22.1973, Don & Mignon Davis, N.M.N.H. 1 d from , 2 km S. DuvergC, 10 m, 18'22.0' N, 7 1°31.4' W, 111.25.1999, 0. S. Flint, Jr., N.M.N.H. 4 00 from Independencia province, Rio Guayabal, 4.5 km N. Postrer Rio, 150 m, 18'34.7' N, 71'37.7' W, 111.25.1999, 0. S. Flint, Jr., N.M.N.H. Determined by 0. S. Flint, Jr. in 1999. Journal of Neuropterology 3: 9-23, 2000 (2001)

Family HEMEROBIIDAE Subfamily HEMEROBIINAE Genus HEMEROBZUS Linnaeus, 1758 Hemerobius antigonus Banks, 1941 Hemerobius antigonus Banks, 1941a: 396, fig. 3. Hemerobius antigonus: Monserrat, 1996: 426, figs. 21-23, 114-115, 163, 188 (map). Type: Holotype 0, M.C.Z. no. 25409. Type locality: Dominican Republic: Valle Nuevo. Distribution: Dominican Republic, Haiti. Comments: MONSERRAT (1996) included in this species one Dominican specimen that Banks had designated a paratype of Hemerobius domingensis. Hemerobius darlingtoni Banks, 1938 Hemerobius darlingtoni Banks, 1938a: 290, fig. 4. Hemerobius darlingtoni: Monserrat, 1996: 435, figs. 36-38, 120-121, 168, 189 (map). Type: Holotype 0, M.C.Z. no. 22121. Type locality: Jamaica: Blue Mt. Peak. Distribution: Jamaica, Dominican Republic, Cuba. Comments: MONSERRAT (1996) records this species for Dominican Republic from 1 9 reported by BANKS (1941a) as paratype 25409 of Hemerobius antigonus. Hemerobius domingensis Banks, 1941 Hemerobius domingensis Banks, 1941a: 395, figs. 4, 7. Hemerobius domingensis: Monserrat, 1996: 462, figs. 80-83, 150- 15 1, 184, 191 (map). Type: Holotype 0, M.C.Z. no. 25408. Type locality: Dominican Republic: Loma Rucilla. Distribution: Dominican Republic, Cuba. Hemerobius gaitoi Monserrat, 1996 Hemerobius gaitoi Monserrat, 1996: 476, figs. 84-87, 140-141, 181, 192 (map). Type: Holotype 0, C.A.A.S. Type locality: Costa Rica: Puntarenas. Distribution: Costa Rica, Brazil, Venezuela, Guatemala, Ecuador, Mexico, Dominican Republic. j-Hemerobius sp. Hemerobius: Poinar & Poinar, 1999: 130, fig. 130. Fossil in Dominican amber. Comments: This same specimen without the generic identification was previously depicted by POINAR (1992) as fig. 71.

Subfamily MEGALOMINAE Genus MEGALOMUS Rambur, 1842 Megalomus minor Banks, 1905 Megalomus lioni: NavBs, 1927a: 26. Megalomus lioni: Smith, 1931: 801 [no specimens were studied]. Megalomus minor: Monserrat, 1997a: 152, figs. 3, 45- 18 PEREZ-GELABERT, D. E. & FLINT Jr., 0. S.

49, 106. Type: Lectotype 0, M.C.Z. Type locality: United States: Nevada. Distribution: Haiti, Cuba, Mustique Island, St. Vincent, Costa Rica, Honduras, Mexico, Panama, Dominican Republic, Trinidad, Venezuela, Colombia, Peru, Bolivia, United States.

Subfamily MICROMINAE Genus MICROMUS Rambur, 1842 Micromus posticus (Walker, 1853) Micromus posticus: Monserrat, 1998: 144. Type: Syntypes dQ, B.M.N.H. Type locality: United States: Georgia. Distribution: United States, Mexico, Dominican Republic.

Micromus subanticus (Walker, 1853) Micromus haitiensis Smith, 1931: 800, pl. 2, fig. 10. Menutus haitiensis: NavBs, 1932: 35. Micromus subanticus: MacLeod & Stange, 1981:2. Micromus subanticus: Monserrat, 1993510. Micromus subanticus: Monserrat, 1998: 144. Type: Holotype d, B.M.N.H. Type locality: United States: East Florida. Distribution: United States, Mexico, Costa Rica, Dominican Republic.

Micromus sp. This represents a new species of Micromus. Specimens: 2 dd 2 QQ from DOMINICAN REPUBLIC: La Vega Prov., 4 krn E of La CiCnaga de Manabao, 3050 ft., 19'04'47" N 70'49'29" W, IV.19.2000, T. J. Henry and R. E. Woodruff, black lights. N.M.N.H. Determined by J. D. Oswald in 2000.

Genus NUSALALA NavBs, 1913 Nusalala cubana (Hagen, 1886) Nusalala damiensis Smith, 1931: 800, pl. 2, fig. 12. Nusalala cubana: Monserrat (in lit.), confirms synonymy. Type: Holotype B, M.C.Z. Type locality: Cuba. Distribution: Cuba, Haiti, Dominican Republic, Puerto Rico.

Nusalala n. sp. This represents a species being described by Monserrat (in lit.) Material examined: 4 QQ from DOMINICAN REPUBLIC: La Vega Prov., 11.5 km S. Constanza, I km N. El Convento, 14 10 m, 18'5 1.7' N, 70'41.0' W, 111.27.1999, 0. S. Flint, Jr., N.M.N.H. 1 0 from La Vega, 15 krn SW jct. 1 & Rt. 12, above , 1200 m, 19'00'36" N, 70'33'35" W, IV.18.2000, T. J. Henry, R. E. Woodruff. NMNH. 1 Q from La Vega, Rt. 12, 20 km SW jct. 1, above Bonao, 1030 m, 19'00'37" N, 70'33'34'' W, IV.18.2000, T. J. Henry, R. E. Woodruff. NMNH. 1 d from La Vega prov., 4 km E La Ciknaga de Manabao, Journal of Neuropterology 3: 9-23,2000 (2001) 19

925 m, 19O04'47" N, 70°49'29" W, IV.19.2000, T. J. Henry, R. E. Woodruff. NMNH. Determined by 0. S. Flint, Jr. in 2000. Subfamily NOTOBIELLINAE Genus NOTIOBIELLA Banks, 1909 Notiobiella israeli (Alayo, 1968) Notiobiella israeli: Monserrat, 1998: 140. Type: Holotype d,I.E.S.H. Type locality: Cuba: Marianao. Distribution: Cuba, Dominican Republic. no ti obi el la thaumasta Oswald, 1999(2000) Notiobiella: Poinar & Poinar, 1999: 203. Notiobiella thaumasta Oswald, 1999(2000): 298, figs. 1-5. Type: Holotype 0,A.M.N.H. DR-14-1124. Fossil in Dominican amber. Subfamily SYMPHEROBIINAE Genus SYMPHEROBIUS Banks, 1904 Sympherobius amiculus (Fitch, 1 856) New generic record for Hispaniola! Type: Lectotype 9,M.C.Z. Type locality: New York, Washington Co., Salem. Distribution: Eastern United States, Canada, Dominican Republic. Material examined: 1 0 2 QQ from DOMINICAN REPUBLIC: La Vega Prov., 4 km E of La Cienaga de Manabao, 3050 ft., 19'04'47" N 70'49'29" W, IV. 19.2000, T. J. Henry and R. E. Woodruff, black lights. N.M.N.H. Determined by J. D. Oswald in 2000. Family MANTISPIDAE Subfamily MANTISPINAE Genus MANTISPA Illiger, 1798 Mantispa sayi Banks, 1897 New generic record for Hispaniola! Type: Lectotype d,M.C.Z. Type locality: Brazos Co., Texas. Distribution: United States, south to Panama, Bahamas, Cuba, Puerto Rico, Hispaniola. Material examined: 1 Q from DOMINICAN REPUBLIC: Azua prov., 8 krn NE Padre Las Casas, Rio Las Cuevas, 18O46' N 70°53' W, 580 m, VIII.1990, J. Rawlins, S. Thompson. C.M.N.H. 1 Q from Barahona prov., 5 krn SE Polo, slopes of Loma La Torre, 18O03' N, 71°16' W, 980 m, VII.18.1992, disturbed forest with coffee, C. Young, R. Davidson, S. Thompson, J. Rawlins, C.M.N.H. Both determined by 0. S. Flint, Jr. in 1999. Mantispa viridis (Walker, 1853) New record for Hispaniola! Type: Holotype 0, B.M.N.H. Type locality: E. Florida. Distribution: Eastern and Southern United States, south to Costa Rica, Venezuela, Cuba and Dominican Republic. 20 PEREZ-GELABERT, D. E. & FLINT Jr.. 0. S.

Material examined: 1 d from DOMINICAN REPUBLIC, Azua, East side of crest Sierra Martin Garcia, 7 km WNW Barrero, 1S021' N 70°58' W, 860 m, VII.25-26.1992. C. Young, R. Davidson, S. Thompson, J. Rawlins. C.M.N.H. Determined by 0. S. Flint, Jr. in 1999.

?Mantispa sp. Mantispa: Poinar & Poinar, 1999: 130, fig. 129. Fossil in Dominican amber. Species of Mantispa pulchella group. Collecting data: 1 d from "R. DOMINICANA, 14 km W. Puerto Plata, V.10- 11.1985, E. Giesbert coll., F.S.C.A." Comments: This is an officially undescribed species recorded in the unpublished dissertation of HOFFMAN (1992).

Family MY RMELEONTIDAE Comments: POINAR & STANGE (1996) state that 12 species of antlions are known from Hispaniola. Most of these represent undescribed species (STANGE, pers. comm.). One fossil adult specimen of Myrmeleontidae in Dominican amber was depicted by WU [1997]: 104, photo 90.

Subfamily ACANTHACLISINAE Genus VELLA NavBs, 1913 Vella fallax haitiensis Smith, 1931 Vella fallax haitiensis Smith, 1931: 815, pl. 11, fig. 16. Vella haitiensis: Navds, 1932: 33. Type: Holotype 9, M.C.Z. Type locality: Haiti: Jacmel. Distribution: Haiti.

Subfamily MYRMELEONTINAE Genus ABATOLEON Banks, 1924 Abatoleon indiges (Walker, 1860) Myrmeleon indiges Walker, 1860: 189. Austroleon indiges: Smith, 1931 : 8 16. Brachynemurus indiges: Stange, 1965: 120, figs. 65, 66, 234. Type: Holotype 0, B.M.N.H. Type locality: "Haiti". Distribution: Cuba, Dominican Republic, Haiti, United States.

Genus EREMOLEON Banks, 1901 Eremoleon cewerai (Navds, 1921 ) Glenurus cewerai: Smith, 1931 : 8 14, fig. 13. Type: Holotype 9, M.C.Z. Type locality: Cuba: Habana, Santiago de las Vegas. Distribution: Cuba, Haiti.

Eremoleon cerverinus (Navds, 1921) New record for Hispaniola! Type: Holotype 9, M.C.Z. Type locality: Cuba: Habana, Rio Almendares. Distribution: Cuba, Haiti. Journal of Neuropterology 3: 9-23, 2000 (2001) 2 1

Specimens: 1 Q from HAITI, A. Curtiss, N.M.N.H. Determined by L. A. Stange in 1961.

Genus MYRMELEON Linnaeus, 1767 Myrmeleon insertus Hagen, 1861 Myrmeleorz insertus Hagen, 1861 : 233. Myrmeleon insertus: Smith, 1931 : 817, fig. 15. Type: Two syntypes, sex unknown, M.C.Z. Type localities: Cuba and Haiti: Port-au-Prince. Distribution: Cuba, Haiti, United States (Florida).

Genus PORRERUS NavBs, 1913 "yorrerus dominicanus Poinar & Stange, 1996 Porrerus dominicanus Poinar & Stange, 1996: 384, figs. 1-5, 6-8. Type: Holotype 9, J.W.C. Fossil in Dominican amber. Comments: The genus Porrerus is found extant from Venezuela to Paraguay.

Genus PSAMMOLEON Banks, 1899 Psammoleon bistictus (Hagen, 1861) Psammoleon bistictus: Smith, 1931: 815, fig. 14. Psammoleon bistictus: Navds, 1932: 34. Type: Holotype, sex unknown, H.M.B. Type locality: Cuba. Distribution: Cuba, Hispaniola, Puerto Rico, United States (Florida).

Psanzmoleon imbellis (Banks, 1941) Puren imbellis Banks, 1941b: 102. Type: Holotype 9, M.C.Z. no. 22520. Type locality: Haiti: Port-au-Prince. Distribution: Haiti.

Psammoleon minor Banks, 1927 New record for Hispaniola! Type: Lectotype 9, M.C.Z. no. 15746. Type locality: Florida: Dry Tortugas, Loggerhead Key. Distribution: Cuba, Puerto Rico, United States (Florida). Specimens: 1 individual from DOMINICAN REPUBLIC: Pedernales Prov., 8 km N. Cabo Rojo, 30 m, VII.18.1987, J. Rawlins, R. Davidson. C.M.N.H. Determined by L. Stange.

ACKNOWLEDGMENTS

Our appreciation to John D. Oswald (Texas A & M University) and Raymond GagnC (Systematic Entomology Lab, U. S. Department of Agriculture, Washington, D. C.) for multiple corrections and suggestions that improved this paper. Lionel Stange (Florida State Collection of Arthropods, Gainesville) provided important information and John Rawlins (Carnegie Museum of Natural 22 PEREZ-GELABERT,D. E. & FLINT Jr., 0. S.

History, Pittsburgh) gave us information and loaned specimens. We are also indebted to unknown reviewers for their valuable help.

REFERENCES

ADAMS, P. A,, 1982. Ceraeochrysa, a new genus of Chrysopinae (Neuroptera) (Studies in New World Chrysopidae, part 11). Neuroptera International I1 (2): 69-75. ADAMS, P. A. & PENNY, N. D., 1985. Neuroptera of the Amazon basin. Part 1la. Introduction and Chrysopini. Acta Amazonica 15 (3-4): 41 3-479. ALAYO D. P., 1968. Los Neurbpteros de Cuba. Poeyana, Serie B, No. 2: 1-127. BANKS, N., 1938a. New West Indian neuropteroid insects. Revista de Entomologia 9: 285-304. BANKS, N., 1938b. Antillean Ascalaphidae. Journal of Agriculture of the University of Puerto Rico 22: 177-180. BANKS, N., 1941a. New neuropteroid insects from the Antilles. Memorias de la Sociedad Cubana de Historia Natural 15: 385-402. BANKS, N., 1941b. Three new species of Myrmeleonidae. Psyche 48: 101-104. BEAUVOIS, A. M. F. J. P., 1805-1821. Insectes Recuellis en Afrique et en Amerique dans les Royaumes d'Oware a Saint-Domingue et dans les Etats-Unis pendant les Annees 1786-1797. Paris ,276 pp. BROOKS, S. J., 1994., A taxonomic review of the common green lacewing genus Chrysoperla (Neuroptera: Chrysopidae). Bulletin of the Natural History Museum, Entomology Series 63: 137-210. BROOKS, S. J. & BARNARD, P. C., 1990. The green lacewings of the world. A generic review (Neuroptera: Chrysopidae). Bulletin of the British Museum (Natural History) Entomology Series 59: 117-286. HAGEN, H., 1861. Synopsis of the Neuroptera of North America. Smithsonian Miscellaneous Collections 4 (1): xx + 347 pp. HOFFMAN, K. M., 1992. Systematics of the Mantispinae (Neuroptera: Mantispidae) of North, Central and South America. PhD Thesis, Clemson University. MACLEOD, E. G. & STANGE, L. A,, 1981. The brown lacewings of Florida (Neuroptera: Hemerobiidae). Florida Department of Agriculture and Consumer Services, Division of Plant and Industry, Entomology Circular No. 227: 1-4. MEINANDER, M., 1972. A revision of the family Coniopterygidae (Planipennia). Acta Zoologicu Fennica 136: 1-357. MEINANDER, M., 1986. The Coniopterygidae of America (Neuroptera). In: GEPP, J., ASPOCK, H. and HOLZEL, H., Recent Research in Neuropterology, Graz, 31-44. MEINANDER, M., 1990. The Coniopterygidae (Neuroptera, Planipennia), A check-list of the species of the world, descriptions of new species and other new data. Acta Zoologica Fennica 189: 1-95. MEINANDER, M., 1998. Coniopterygidae (Neuroptera) in amber from the Dominican Republic. Journal of Neuropterology 1: 33-36. MONSERRAT, V. J., 1993. New data on some species of the genus Micromus Rambur, 1842. Annuli del Museo Civico di Storia Naturale "Giacomo Dorian 89: 477-516. MONSERRAT, V. J., 1996. A revision of the Latin American species of the genus Hemerobius (Neuroptera, Hemerobiidae). Fragmenta Entomologica 27: 399-523. MONSERRAT, V. J., 1997. Revision del gdnero Megalomus de Latinoamdrica (Neuroptera, Hemerobiidae). Fragmenta Entomologica 29: 123-306. MONSERRAT, V. J., 1998. Nuevos datos sobre 10s hemerdbidos de AmCrica (Neuroptera: Hemerobiidae). Journal of Neuropterology 1: 109- 153. NAVAS, L., 1913. Sinopsis de 10s Ascalifidos (Ins. Neur.). Arxius de l'lnstitut de Ciencias, Barcelona 1 (3): 99 pp. Journal of Neuropterology 3: 9-23, 2000 (2001) 23

NAVAS, L., 1921. Insectos americanos nuevos o criticos. Broteria , Serie Zoologia 19: 113-124. NAVAS, L., 1924. Algunos insectos de Cuba. Revista de la Real Academia de Ciencias Exactas, Ftsicas y Naturales de Madrid 21: 333-340. NAVAS, L., 1927a. Insectos del Museo de Paris (4a Serie). Broteria, Serie Zoologica 24: 5-33. NAVAS, L., 1927b. Veinticinco formas nuevas de insectos. Boletin de la Sociedad Ibe'rica de Ctencias Naturules 26: 48-75. NAVAS, L., 1932. Neur6pteros de Haiti. Boletin de la Sociedad Entomologica de EspaAa 15: 33- 37. NEEDHAM, J. G., 1941. A note concerning aggregations of Ululodes villosa Beauvois. Psyche 48: 112. OSWALD, J. D., 1999 (2000). The brown lacewing genus Notiobiella (Neuroptera: Hemerobiidae) from Dominican amber. Journal of the New York E~~tomologicalSociety 107: 297-303. PENNY, N. D., 1977. Lista de Megaloptera, Neuroptera e Raphidioptera do MCxico, AmCrica Central, ilhas Caraibas e America do Sul. Acta Amazonica 7 (Suplemento): 1-61. PENNY, N. D., ADAMS, P. A. & STANGE, L.A., 1997. Species catalog of the Neuroptera, Megaloptera, and Raphidioptera of America North of Mexico. Proceedings of the California Academy qf Sciences 50 (3): 39-114. POINAR, G. O., JR., 1992. Life in Amber. Stanford University Press, Stanford, California. 350 pp. POINAR, G. O., JR. & POINAR, R., 1999. The Amber Forest - A Reconstruction of a Vanished World. Princeton University Press, Princeton, New Jersey. 239 pp. POINAR, G. O., JR. & STANGE, L. A. ,1996. A new antlion from Dominican amber (Neuroptera: Myrmeleontidae). Experientia 52: 383-386. SMITH, R. C., 1931. The Neuroptera of Haiti, West Indies. Annals of the Entomological Society of America 24: 798-821. STANGE, L. A., 1965. Revision of the Brachynemurini of North America (Neuroptera: Myrmeleontidae). Ph. D. Thesis, University of California, Davis. 396 p. WALKER, F., 1853. Catalogue of the specimens of Neuropterous insects in the collection of the British Museum, part I1 (Sialides - Nemopterides). British Museum of Natural History, London. 193-476. WALKER, F., 1860. Characters of undescribed Neuroptera in the collection of W. W. Saunders, Esq., F. R. S. & c. Transactions of the Entomological Society of Lolldon 5: 176-199. WEELE, H. W. VAN DER., 1908. Ascalaphiden. Monographisch Bearbeitet. Collections Zoologiques du Baron Edm. de Selys Lonchamps. Fasc. 8: 326 pp. WU, R. J. C., [1997]. Secrets of a Lost World - Dominican Amber and its Inclusions. Privately published. Santo Domingo. 222 pp.

Received: 5.1V.2000 Accepted: 4.VII.2000 Bibliography of the Neuropterida

Bibliography of the Neuropterida Reference number (r#): 9972

Reference Citation: Pérez-Gelabert, D. E.; Flint, O. S., jr. 2000 [2001.04.30]. Annotated list of the Neuroptera of Hispaniola, with new faunistic records of some species. Journal of Neuropterology 3:9-23.

Copyrights: Any/all applicable copyrights reside with, and are reserved by, the publisher(s), the author(s) and/or other entities as allowed by law. No copyrights belong to the Bibliography of the Neuropterida. Work made available through the Bibliography of the Neuropterida with permission(s) obtained, or with copyrights believed to be expired.

Notes:

File: File produced for the Bibliography of the Neuropterida (BotN) component of the Global Lacewing Digital Library (GLDL) Project, 2008.