Great Basin Naturalist

Volume 53 Number 3 Article 1

9-27-1993

1993 update, checklist of the vascular of Bryce Canyon National Park, Utah

John R. Spence Manzanita BioStudies, Page,

Hayle Buchanan Weber State University, Ogden, Utah

Follow this and additional works at: https://scholarsarchive.byu.edu/gbn

Recommended Citation Spence, John R. and Buchanan, Hayle (1993) "1993 update, checklist of the vascular plants of Bryce Canyon National Park, Utah," Great Basin Naturalist: Vol. 53 : No. 3 , Article 1. Available at: https://scholarsarchive.byu.edu/gbn/vol53/iss3/1

This Article is brought to you for free and open access by the Western North American Naturalist Publications at BYU ScholarsArchive. It has been accepted for inclusion in Great Basin Naturalist by an authorized editor of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. The Great Basin Naturalist PUBLISHED AT PROVO, UTAH, BY BRIGHAM YOUNG UNIVERSITY

ISSN 0017-3614

VOLUME 53 SEPTEMBER 1993 No.3

Great Basin Natualist 53(3), pp. 207~221

1993 UPDATE, CHECKLIST OF THE VASCULAR PLANTS OF BRYCE CANYON NATIONALPARK, UTAH

John R. Spence1 and Rayle Buchanan2

ABSTRACT.-An updated checklist of the vascular flora of Bryce Canyon National Park is presented, including 458 species arranged in 232 genera and 73 families. Four species have been added to the park list based on 1991 fieldwork: Arabis perennans, Arceuthobium divaricatum, Gentiana parryi, and Haplopappus acauUs. Recent changes in nomencla­ ture have been included, and several species identifications have been corrected. The six largest families are, in decreasing size, Asteraceace, Pocaceae, , Brassicaceae, Scrophulariaceae, and Rosaceae. Common names and an index are provided.

Key words: Bryce Canyon National Park, flora, vascular plants.

Recent fieldwork in Bryce Canyon Nation­ for the ferns, as fern phylogeny has become al Park, along with nomenclatural changes in increasingly stahle over the last few years, and the Utah flora, has neccesitated an updated well-estahlished classifications currently exist. version of the flora of Bryce Canyon National We follow Weber (1987), who has provided an Park, south central Utah. The last puhlished analysis of the new classification for regional version of this species list is now 12 years old ferns. Only two genera occur at Bryce Canyon, (Buchanan and Grayhosch 1981). We offer this Cheilanthes, in the Sinopteridaceae, and revised list hoping it will be useful to Pteridium, in the Hypolepidaceae, We also researchers currently engaged in vegetation follow the now well-established classification and floristic work in and around the park. for the Liliaceae and its allies, which has Nomenclature in general follows the Inter­ become generally accepted worldwide mountain Flora (Cronquist et al, 1972--1990), (Dahlgren et al. 1985). Hence, in addition to except for the substitution of the more recent the Liliaceae, we have members of the treatment of the Poaceae in Arnow (1987). For Agavaceae, Alliaceae, Asparagaceae, Calo­ those families not yet treated, Welsh et al. chortaceae, Convallariaceae, and Melanthi­ (1987) is used. We differ from the above vol­ aceae in the park flora. Although we feel that umes in the delimitation of some families. In the alternative classifications of Weber (1987) particular, we cannot accept the outmoded have much to offer, we think it premature to use ofthe Polypodiaceae as a "catch-all" group utilize his system in its entirety, as this would

lManzanita BioSludies, Box 833, Page, Arizona 86040. 2Depaltmenl ofBotany, Weher State University, Ogden, Utah 84408.

207 208 GREAT BASIN NATURAlJST [Volume 53 neccessitate numerous nomenclatural innova­ nized alphabetically by family, genus, and tions for Utah taxa. Hence, only three genera species. are accepted that differ from the treatments in Table 2 includes a list of species that have A Utah Flora and Intermountain Flora: Bis­ been found around the boundaries of Bryce torta (Polygonum in parI), Ipomopsis (Cilia in Canyon National Park but are not yet known part), and Pseudocymopterus (CynwptenUi in from within the park. FUture fieldwork is like­ part). We utilize these because they bave been ly to lind some of these species in the park. widely accepted in North American floristic work. Finally, we note that there are funda­ CHECKLIST OF THE VASCULAR PLANTS OF mental differences in nomenclature at the BRYCE CANYON ATIONAL PARK species level for some planIs at Bryce Canyon when Welsh et al. (1987) and Weber (1987) ACRIV.CRAE are compared; e.g., in Arabis, Euplwrhia., Sol­ 1. Ace,- glabrum Ton', Rocky Mountai.n maple i

TABLE 1. Changes in species identifications in, and deletions hum, the 1981 list of the flora of Bryce Canyon National Park. 1 = outside range, misidentification, deleted from list; 2 = inside range, misidentification, retained in list. Taxa previously reported CurrcDtjuslification and slatus Arertatia nuttaLlii Pax (1) no specimen. outside range Campanula t"otundifolia L. (2) C. parryi Gray Erigeron simplex Gr(.'t;:oc (2) E. ecmpQsUus Pursh Genl.ianeU(~ tenella Rottb. (2) G. tortuosa oones) J. Gillett Lmnalilm~ nuttallii (Gmy) Machr. (2) l. parryi (Wats.) Mach.. Phlox condetlSata (Cr.lY) E. Nels. (2) P. puLvinWn (Wherry) Cnmq. P. dijJusa Bcnth. (2) r. hoodu Richards P. tumulosa Wheny (2) P. puloiMW (Wheccy) Crunq. Physaria newberryi Gray (1) no spocimen, olltside range Polemonil.lm tM'cosum Nut!. (1) no specimen; outside range ScirplJ$ americanus Pers. (2) S. pungem Vahl Senecio cymb(darioide~' Duck (2) S. hurtianus Heller SoluUl/lo uccidentafu (Nutt.) T. & C. (2) S. sparsiflom Gray 'fanacetwll ntdtallii T. & G. (2) SphaeromeriIJ cop;ww NUll. Tludypodiumsagittatum (Nutt.) Endl. (1) no specimen, outsi.de range 1993] VASCULAR PLANTS OF BRYCE CANYON 209

TABLE 2. List of species reported from near the boundary of Bryce Canyon National Park, but not yet collected from within the park.

ASCLEPIADACEAE JUNCACEAE Asclepias cryptoceras Wats. funGus mertensianw Bong.

ASTEMCEAE LAMIACEAE Artemisiajtlifolia Torr. Mentha arvensis L. Cirsium clavatum (Jones) Petrak C. wheeled (Gray) Petrak LrNACEAE Crepis atrabarba Heller !.inum australe Heller Erigeron aphan(J{;tis (Gray) Greene E. londwphyUus Hook OIl.CHIDACEAE Hap[opappus lancevlatus Hook. Hahenaria hyperborea (L.) R.Br. H. parryi Gray Hymenoxys depressa ('1'. & G.) Welsh & Reveal PINACEAE Senecio canus Hook. Abieslasiocarpa (Hook.) Nutt. Solidago nana Nutt. Picea engelmannii Engelm. Stephanorrwria exigul/. Nutt. POACEAE BRASSICACEAE Lolium perenne L. Arabis drummondii Gray Lepidium lasiocarpum Nutt. POLEMONIACEAE Lesquerella wardii Wats. Leptodactylon caepitosum Nutt. Thlaspi flwnumum L. POLYGONACEAE CAPRIFOLlACEAE Eriogonum a.retioides Barneby Sambucus mcemosa L. RANUNCULACEAE CYPERACEAE Caltha leptosepala DC. Carex egglestonii Mack. C. peta8ata Dewey ROSACEAE C. simulata Mack. Amelanchier alnifolia Nutt. lvesia kingii Wats. EPHEDRACEAE Ephedra viridis SALICACEAE Salix boothii Darn FABACEAE Astragalus Imtiginoslls Hook. SCHOPIlUUHlACEAE A. wardii Gray Cordylanthus wrightii Gray Trifolium kingii Wats. Pedicularis groenlandica Retz. Penstemon laevis Pennell GROSSULARIACEAE P. procerus Graham Ribes monngenum McClat. SOLANACEAE HYPERlCACEAE Datura stramunium L. Hypericumfornwsum H.B.K. Nicutiana attenuata Wats.

ImDACEAE SPAIl.GANIACEAE Sisyrinchium idalwense Biekn. Sparganium eurycarpum Engelm.

VrOLACEAE Viola nephrophylla Greene

14. L. parryi (Wats.) Machr. Parry hiscuitroot 18. A. spee.-wsa Torr. showy or common 15. Pseudocymopterus montanus milkweed (Gray) Coult. & Rose mountain parsley ASPARAGACEAE APQCYNACEAE 19. Asparagus officinalis L. asparagus 16. Apocynum androsaemifolium L. spreading dogbane ASTERACEAE ASCLEPIADACEAE 20. Achillea lanulosa Nutt. yarrow 17. Asclepias asperula (Dene.) 21. Agoseris aurantiam (Hook.) Woodson spider milkweed Greene orange agoseris 210 GREAT BASIN NATURALIST [Volume 53

vax. aurantiaca 64. E. eatonii Gray Eaton fleabane Vat. purpurea (Cray) Greene 65. E. fiageU.ns Gray trailing fleabane 22. Agoseri8 glaoco (Pursh) 66. E. pumilus Nutt. low fleabane D. Dietr. mountain dandelion ssp. concin.twides Cronq. vaT. paroij/ora (NUlt.) Rydb. 67. Eo ",-=(LindJ.) DC. Oregon fleabane 23. Antennaria dimmpha (Nutt.) 68. E. ur.sinus D.C. Eaton bear fleahme T. &G. low pussytoes 69. E. vagus Payson wandering fleabane 24. A mkruphylls Rydb. rosy pussytoes 70. Grindelia squarrosa (Pursh) 25. A. paroifolia Nutt. small-leaf pussytoes Dunal curlycup gumweed. 26. A. rosulnt<; I\ydb. woolly pussytoes 71. Gutierrezia sarothrae 27. Arnica chamissonis Less. Chamisso arnica (Pursh) Britt. & Rusby broom snakeweed 28. A. cordifolia Hook heartleaf arnica 72. Haplopappus acaulis (Nutt.) 29. Artemisia carnpestris L. field sagewort Gray stemless goldenweed 30. A cctrTUthii Wood ex Carruth Carruth sagewort 73. H. arrrwrioide.s (Nutt.) Gray thrifty goJdenweed 31. A drocuncu/u$ L. tarragon 74. H. scopul.orum Oone,) 32. A frigida Willd. fringed sagewort Blake Rocky Mountain 33. A ludoviciana Nutt. foothill sagebrush goldenweed 34. A nova A Nels. black sagebrush 75. H. zionis L.e. Anderson Cedar Breaks 35. A pygmaea Gray pigmy sagebrush goldenweed 35. A tridenWa Null. big sagebrush 76. Heliantiudla unifl«a 37. Aster chUensis Nees pacific aster (Nutt.) T. & G. little sunflower 38. A eatonii (Gray) HoweU Eaton aster 77. HeUan.t-hw 01IIUA4S L. common sunflower 39. A glaocedes Blake bJueleaf asler 78. Heliomeris trnJ.tiflora Nutt. showy goldeneye 40. A occidentali.s (NUll.) Torr. 79. Heterotheca jonra var. arizonicwn (Nutl) T. & C. showy rushpin1c: var. nid.ulwn Oones) Welsh 89. Machaeranthera canescena 52. C. caIcareum Oones) Woot. (Pu"h) Cray hoary lansyaster & StandI. limestone thistle 90. M. grin

102. Stephanomeria spinosa 135. D. sophia (L.) Webb flixweed (Nutt.) Tomb thorn wirclettucc 136. Draha suhalpina Goodman 103. S. tenuifolia (Torr.) Hall slenderleafwirelettuce & Hitchc. breaks draba 104. Taraxacum ufficinale 137. Erysimum asperum (NutL) Weber common dandelion DC. western wallflower 105. Tetradymia canescens DC. Gray horsebrush 138. E. inconspicu.u.m (S. Wats.) 106. Townsendia exscapa MacMill small wallflower (Richards) Porter stemless townsendia l.39, Lepidium densiflorum 107.1'. incana Nutt. hoary townsendia Schrad. prairie pepperweed lOR. T. minima Eastw. least townsendia 140. L. montanutn Nutt. mountain peppcl'weed 109. Tragopogon dubius Scap. yellow salsifY 141.l.Rsquerella interrnedia 110. Verbesina encelioides (S. Wats.) Heller Watson bladderpod (Cav.) Benth. & Hook. golden crownheard 142. L. kingii S. Wats. Kin!!: hladderpod val'. parvifolia (Maguire BERBEHllJACEAE & Holmgren) Welsh & 111. Mahoniafremontii (Torr.) Heveal Fedde Fremont mahonia 143. L rubicundula Rullins breaks bladderpod 112. M. repens (Lind!.) G. Don Oregon grape 144. Malcolmia africana (L.) R. Br. African mustard 145. Physaria chambersii Rollins Chamber twinpod BETULACEAE 146. P. newherryi Gray Newberry twinpod 113. Betula occidentulis Hook. western water birch 147. Schoencrambe Unijolia (Nutt.) Greene schocncrambe BORAGINACEAE 114. Cryptantha abata LM. 148, Slanleya pinnata (Pursh) prince's plume Johnston low cryptanth Britton lIS. C. baked (Greene) Payson Baker cryptanth 149. Streptunthus cordatus Nlltt. twistflower 116. C. confertifolia (Greene) 150. Thelypodium sagittatum Payson golden cryptanth (Nutt.) EndL arrowleaf thclytxxly 117. G.fendled (Gray) Greene sand dune cryptanth var. ovalifolium (Rydb.) Welsh & Reveal 118. C. jlava (Nels.) Payson yeIlow cl)'ptanth field pennycress 119. C. fulvocanescens (Gray) 1.51. Thlaspi arllcnse L. Payson plateau cryptanth 120. C. humilus (Greene) CACTACEAE Payson roundspike clyptanth 152. Echinocereus 121. C. jamesii (Torr.) Payson James clyptanth triglochidiatus Engelm. claret cup 122. C. ochroleuca Higgins yellowish clyptanth 153. Opunlia erinacea Engelm. 123. C. setosissima (Gray) & Bigel. grizzlybear pric:klypcar Payson bristly cryptanth 154. Pediacactus simpsonii 124. Lappula occidentalis (Engelm.) Britt. & Rose Simpson pediocactus (Wats.) Greene western stickseed 1,5,5. Sclerocactus whipplei 12,5. Lithospennun incisUin (Engelmm. & Bigcl.) Lehm. showy stickseed Britt. & Rose fishhook cactus 126. L multiflorum Ton. southwestern stickseed 127. Mel"tensia arizonica CALOCHORTACEAF: Greene aspen bluebell 156. Calvchortus nuftallii T. & G. sego Jjly

BHASSICACEAE CA:>.1PAI\ULACEAE 128. Arabis demissa Greene roekcress 157. Campanuw parryi Gray Parry bellflower 129. A. holboelii Bornem. Holboell rockcress var.fendleri Wats. CAPPARACEAE var. holboelii 158. Cleome lutea Hook. yellow beeplant 130. A. pel'ennans Wats. common rockcress 131. Capsella bursa-pastoris CAPRIFOLIACEAE (L.) Medic shepherd's purse 159. Sambucus cerulea Raf. blue elderberry 132. Caulanthu.s crassicaulis 160. Symphvricarpos oreophilus (Torr.) S. Wats. thickstem wildcabbage Gray mountain snowberry 133. Conringia orientalis (L.) Dumort hare's ear mustard CAHYOPHYI.LACEAF. 134. Descurainia richm-dsonii 161. Arenaria fendleri A. Gray Fcndlcr·sandwort (Sweet) O.E. Schulz Richardson 162. Lychnis drumrnondii tansymustard (Hook.) S. Wats. Drummond catchfly 212 GREAT BASIN NATURALIST [Volume 53

163. Paronychia sessiliflora 193. C. ,wnophyUa Wilil. narrowleaf sedge Nutt. creeping nailwort 194. Eleocharis palustris (L.) 164. Silene petersonii Maguire Peterson campion R. & S. common spikerush vaT. minor C.L. Hitchc. 195. E. paucifbw (Light!.) & Maguire Link spikerush 165. SteUariajamesiana Torr. James chickweed 196. E. rosteUata (Torr.) Torr. beaked spikerush 166. S. longipes Goldie longstalk chickweed 197. Scirpus pungens Pers. common threesquare bulrush CELASTRACEAE 167. Paxistirna myrsinites EUEAGNACEAE (Pursh) Raf. mountain lover 198. Slwpherdia argentea (Pursh) Nutt. silver buffalobeny CHENOPODJACEAE 199. S. canadensw (L.) Nut!. russet buffaloberry 168. Atriplex canescens 200. S. roturzdifoUa Parry roundleafbuffaloberry (Pursh) Nutt. fourwing saltbush 169. A confertifolia (Torr. & EQUISETACEAE Frem.) Wats. shadsca1e 20 1. Equisetum aroense L. field horsetail 170. A gardneri (Maq.) D. Dietr. Gardner saltbush 202. E. laevigatum A. Br. smooth horsetail var. tridentata (Kuntze) MacBr. ERICACEAE 171. Chenopodium atrovirens 203. Arctostaphylos patula Rydb. mountain goosefoot Greene greenleaf manzanita 172. G.frerrwntii Wats. Fremont goosefoot 204. A. uva-ursi (L.) Spreng. bearberry var. incanum Wats. 173. C. leptophyUum Nutt. narrowleaf goosefoot EUPHORBlACEAE 174. Eurotia lan.ata (Pursh) Maq. winterfat 205. Euphorbia brachycera 175. Salsola iberica Sennen Engelm. shorthorn spurge &Pau Russian thistle 206. E. fendleri T. & G. Fendler spurge 176. Sarcobatus vermieulatus (Hook.) Torr. greasewood FABACEAE 207. Astragalus agrestis G. Don field milkvetch CONVALLARIACEAE 208. A. amphioxys Gray crescent milkvetch 177. Smilacina steUata (L.) 209. A. argophyllus T. & G. silverleaf milkvetch Desf. starry false Solomon seal 210. A. bWulcai= (Hook.) G

228. Melilotus (~fficinalis (L.) LA;-"lIACEAE Pall. yellow sweetclover 256. Dracoceplwlum parviflorum 229. Oxytrupis dejlexa (Pall.) Nutt. American dragonhead DC. hangpod locoweed 257. Hedeoma drummondii vaT. serieea T. & G. Benth. mock pennyroyal 230. 0. ureophila Gray mountain locoweed 258. Marruhium vulgare L. hoarhuund 231. Pediomelwn pariense 259. Monardella odoratissima (\Vebb & Atwood) Benth. cloverhead horsemint J. Grimes Paria breadroot 260. Prunella vulgaris L. selfheal 232. Trifolium repens L. \vhite 233. Vicia americana 'Villd. American vetch LJUACEAE 261. Fritillaria atropurpurea FAGACEAE NutL leopard lily 234. Quercus gamhelii Nun. Gumbel oak 262. LeucuCTinum muntanmn Nutt. ex Gray star lily

FUMAlUACEAE 23.'5. Corydalis ([urea Willd. golden corydalis LINACEA!': 263. Linum anstatwn Engelm. broom flax L. GENTIANACEAE 264. kingii Wats. King flax 236. Frasera speciosa Griseb. elkweed 265. L. lewisii Pursh blue flax 237. Gentiana affinis Griseb. Rocky Mountain gentian 1(lASACEAE 266. Mentzelia multiflora 238. G. pmTlji Engelm. Parry gentian (Nutt.) Cray desert blazing star 239. Gentianella heterusepala (Engclm.) Holub Engelmann gentian MAl.VACEAE 240. G. tortuosa (Jones) 267. Sphaeralcea coccinea J. Gillett Jones gentian (Pursh) Rydb. scarlet globemallow 241. Gentianopsis detonsa 268. S. pamifolia A. Nels. small-flowered (Rattb.) Ma fringed gentian gl()l}l;rrlall()\v

GEHANIACEAE MEL.-'..NTI-IIACEAE 242. Geranium caespitosum 269. Zigadenus elegans Pursh mountain dealh Jame:; small-leafgeranium camas 24:). G. dchardsOIlii Fisch. & Trautv. Richardson geranium NYCTACINACEAE 270. Ahroniafragrans Hook. fragrant sand verbena CHOSSULAlUACEAE 271. A. nana'Vats. low sand verbena 244. Ribes aw"eum Pursh golden currant

245. Rihes cereum Dougl. wax or squaw currant ONAGRACEAE 272. Calylophus lavandulifoUus HYPOLEPlDACEAE (T. & G.) Raven lavender evening 246. Pteridiwn aquilinum pnmrose (L.) Kubn bracken fern 273. Epilobium angustifolium L. fireweed IHlDACEAE 274. E. ciliatum Rat n()rthern willuwherb 247. Iris missouriensis NulL Missouri iris 275. Ga.yophytum 248. Sisyrinchiurn dernissum ranwsissimum T. & G. branchy Greene blue-eyed grass groundsmoke 276. G. mcemosum T. & G. kitchenweed JUNCACEAE 277. Oenothera caespitosa Nutt. tufted evening 249. Juncus arcticus Willd. wiregrass primrose 250.]. bufonius L. toad rush 278. O. coronopifi)lia T. & G. motstalk evening 251.]. ensifoliw; Wikstr. swordleafrush pnmrose 252.]. longistylis Torr. longstyle rush 279. O. flava (A. Nels.) Garrett yellow evening 253. J. torreyi Coville Torrey rush pnmrosc 280. O. howardii (A. Nels.) JUKCAcrNACEAE W.L. Wagner bronze evening 2.54. Trig70chin maritima L. maritime pnmrosc arrowgrass 281. 0. longissima Rydb. Bridges evening 255. T. palustrus L. marsh arrowgrass primrose 214 GREAT BASIN NATURALIST [Volume 53

282. 0 paUida Lind!. pale evening 319. E. smithii (Rydb.) Gould \vestern \Vlleatgrass pnmrose 320. E. spicatus (Pursh) Gould bluebunch wheatgrass ORCHlDACI\AE 321. E. trachycaulis (Link) Gould 283. Corallorhiza maculata Raf. spotted coralroot ex Shinners slender wheatgrass 284. C. striata Lind!. striped coralroot 322. Festuca octojlora Walter six weeks fescue 285. Epipactis gigantea Doug!. 323. F. ovina L. sheep fescue ex Hook. giant helleborine 324. F. pratensis Hudson meadow fescue 286. lIahenaria dilatata 325. Glyceria striata (Lam.) (Pursh) Hook. white bog orchid A.S. Hitchc. fowl mannagrass 287. H. sparstflora S. Wats. Watson bog orchid 326. Hilariajamesii (Torr.) Benth. galleta 327. Hordeum brachyantherum OROBANCHACEAE Nevski meadow barley 288. Orobanche fasciculata 328. H. jubatum L. foxtail barley Nutt. clustered broomrape 329. Koeleria macrantha (Ledeb) Schultes Junegrass PrNACEAE ,330, Muhlenberp,ia andina (Nutt.) 289. Abies concolor (Corel. & A.S, IIitchc, foxtail muhly Glend.) Lind!. white fir 331. M. asperifolia (Necs & 290. Picea pungens Engelm. blue spruce Meyer) L.n. Parodi scratchgrass 291. Pinus edulis Engelm. two-needle pinyon 332. M, montana (Nutt,) 292. P.flexilis James limber pine A.S. Ilitchc. mountain muhly 293. P. longaelJa D. K. Bailey bristlecone pine 333. M. pungens Thurber sandhill muhly 294. P. ponderosa Laws. ponderosa pine 334, M. riclwrdsonis (Trin.) Rydb. mat muhly 295, Pseudotsuga menziesii 335. Oryzopsis micrantha (Trin. (Mirb.) Franco Douglas fir & Rupr.) Thurber littleseed ricegrass 336. Phleum pratense L, timothy PLANTACINACEAE 337. Poa compressa L. Canada bluegrass 296. Plantago eriopoda Torr. woolly-foot plaintain 338. P. fendleriana (Steudel) 297. P. lanceolata L. English plaintain Vasey muttongrass 339. P. pmtensv,' L Kentucky bluegrass POACEAE 340. P. rejlexa Vasey & Scribn, nodding bluegrass 298. Agropyron cristatum 341. P, secunda Pres!' Sandberg bluegrass (L.) Gaertn. crested wheatgrass 342. Puccinellia nuttalliana 299. Agrostis stolonifera L. carpet bentgrass (Schultes) A.S. Hitchc. Nuttall alkaligrass 300. Aristida purpurea NutL purple threeawn 343. Schizachyrium scoparium 301. Bouteloua gracilis (Michx.) Nash little bluestem (H.B.K.) Lag. ex Steudc1 blue grama 344. Sporobolus airoides (Torr.) 302. Brumus anumalus Rupr. Torr. alkali sacaton ex Fourn. nodding brame 345. S. cryptandrus (Torr.) Gray sand dropsecd 303. B. carinatus H. & A. mountain brame 346. Stipa cornata Trin. & Rupr. needle-and-thread 304. B. ciliatus L. fringed brame 347. S. hymenoides R. & S. Indian ricegrass 305. B. inermis Leysser smooth brame 348, S. lettermanii Vasey Letterman 306. B.japonicus Thunb. needlegrass ex Murray meadow bramc 349. S. nelsonii Scribn. Nelson ncedlegrass 307. B. tectorurn L. cheatgrass 350. S. pinetOl'um Jones pinewoods 308. Calamagrostis scopulorum needlegrass M.E. Jones Jones reedgrass 309. C. stricta (Timm) Koeler slimstem reedgrass POI.EMONIACEAE 310. Dactylis glornerata L. orchard grass 351. Ipomopsis aggregata (Pursh) 311. Deschampsia caespitosa V. Crant scarlet gilia, skyrocket (L.) Beauv. tufted hairgrass 352. I. congesta (Hook.) V. Grant ballhead gilia 312. Elymus canadensis L. Canada wildrye 353. Leptodactylon pungens (TorL) 313. E. elynwides (Raf.) Swezey squirrel tail Nutt, prickly phlox 314. E. glaucu~ Buckley hlue wildrye 354. Linanthastrum nuttallii (Gray) 315. E. junceus Fisch. Russian wildrye Ewan Nuttall flaxflower 316. E. lanceolatus (Scribn. 355. Phlox austromontana Cov. desert phlox & Sm.) Gould thickspike wheatgrass 356. P. hoodii Richards, carpet phlox 317. E. salinus Jones Salina wildryc 357. P, longifolia NutL longleafphlox 318. E. scribneri (Vasey) Jones spreading whcatgrass 358. P, muscoides NutL moss phlox 1993J VASCULAR PLANTS OF BRYCE CANYON 215

359. P. pulvinata (Wherry) Cronq. cushion phlox ROSACEAE 360. Polenwnium caeruleum L. western Jacobs 392. Amel.anchier utahensis ladder Koehne Utah serviceberry 393. Cercocarpus ledifolius POLYGONACEAE Null. curlleafmountain 361. Bistorta vivipam (L.) S. Gray Amcricml bistort mahogany 362. Eriogonum alatum Torr. winged buckwheat 394. C. montanus Raf alder leaf mountain 363. E. corumbosum Benth. COlymb hucbvheat mahogany 364. E. microthecum Nutt slender buckwheat 395. Fallugia pamdoxa (D. Don) var.}oliosum (T. & G.) Endl. Apache plume Reveal 396. Fragaria virginiana 365. E. panguicense (M.E. Jonc~) Duchesne mountain strawberry Reveal Panguitch buckwheat 397. IIolodiscus dumoslls 366. E. racemosum Nutt. redroot buckwheat (I-look.) Heller hush ocean spray 367. E. subreniforme S. Wats. Stokes buckwheat .'398. Ivesia sabulosa (M.E. Jones) 368. E. umhellatum Torr. sulphur buckwheat Keek Sevier ivesia var. subaridum Stokes 399. Peraphyllum ramosissimum 369. Polygonum aviculare L. knotweed NuH. squaw-apple 370. P. douglasii Greene Douglas knotweed 400. l'otentilla anserina L. silverwced cinquefoil var.johnstonii Munz 401. P. concimw Richards pretty cinquefoil 371. P. ramosissimwn Michx. bushy knotweed 402. P. crinita Gray bearded cinquefoil 372. Rumex crispus L. curly dock 403. P. fntticosa L. shrubby cinquefoil 373. R. salicifolius Wcinm. beach dock 404. P. gmcilis Doug\. slender cinquefoil var. triangulivalis Danser. val'. pulcherrima (Lehm.) Fern. 405. P. p/atlensis Nutt. cinquefoil PRIMULACEAE 406. Prunus virginiana L. chokecherry 374. Androsace septentrionalis L. pygmy-flower 407. Purshia me-rieana (D. Don) rockjasmiJle Welsh 375. Dodecatheon pulchellurn cliffrose 408. p. lridentata (Pursh) DC. bitterbrush (Raf.) Merrill pretty shooting star 409. Rosa nutkana Pres!' Nutka rose 410. R. woodsii Lind!. Woods rose PYROLACEAE 411. Rubus Maeus L. raspberry 376. Pymla asarifolia Michx. liverleafwintergreen val'. saehalinensis (levI.) 377. P. oirens Schweigg. green wintergreen Focke 37R. Pterospora andromedea Nutt. pinedrops HUBJACEAE 412. Galium trijiJurn L. small bedstraw RANl;NCULACEAE 413. Kelloggia galioides Torr. kelloggia 379. Aconitum columbianum

Nutt. monkshood SALICACEAE 380. Anemone multi{1da PoiI'. globeflower 414. Populus angustifolia James narrowleaf 381. Aquilegia caerulea Jamcs Colorado columbine cottonwood 382. A. scopulorum Tides. rock columbine 415. P.jremontii S. Wats. Fremont 383. Clematis columbiana (Nutt.) cottonwood T. & G. Rocky Mountain clcmatis 416. P. tremuloides Michx. quaking aspen 384. C. ligusticifoUa Nutt. white virgin's bower 417. Salixbebbiana Sarg. Bchb willow 385. Delphinium andersonii Gray Anderson larkspur 418. S. drummondiana Barratt Drummond willow vaL scaposum (Greene) 419. S. exigua Nutt. narrowleafwillow Welsh 420. S. geyeriana Andcrss. Geyer willow 386. RanunCIJlU8 cymbalaria 421. S. lasiandra Benth. \\'}lipl,ISil willo\\' Pursh marsh buttercup vaL caudata (Nutt.) Sudw. 387. R. j1ammula L. spean",ort 388. R. inarnoenus Greene drab buttercup SANTALACEAE 389. Thalictrumfendleri Engelm. Fendler meadowruc 422. Comandra urnbellata (L.) Nutt. bastard toadflax RUAMNACEAE 390. Ceanothusfendleri Gray Fendler mountain SAXIFHAGACEAE lilac 423. Litlwphmgma tenella 391. C. marlinii M.E. Jones Utah mountain lilac Nutt. slender woodland star 216 GREAT BASIN NATURALIST [Volume 53

424. Pamassia palustri:s L. northern grass­ 455. A divaricatum Engelm. pinyon pine of-Parnassus dwarfmistletoe 456. A douglasii Engelm. Douglas dwarf SCROPI-IULARIACEAE mistletoe 425. Castilleja chromosa A. Nels. desert paintbrush 457, A vaginatum (Willd.) Presl. southwestern 426. C. linariifolia Benth. Wyoming paintbrush dwarf mistletoe 427. C. revealii N. Holmgren Bryce Canyon 458, Phoradendronjuniperinum paintbmsh Gray juniper mistletoe 428. Cordylanthw; kingii S. Wats. King birdsbeak 429. C rarviflorus (Ferris) ACKNOWLEDGMENTS Wiggim smallflower birdsbeak 430. Mimulus guttatus DC. common monkeyflower We would like to thank the superintendent 431. M. rubellus Gray monkeyflower of Bryce Canyon National Park for providing 432. Orthocarpus Iuteus Nutt. yellow owlclover housing, collecting permits, and the use of 433. 0. purpureo-albus Gray purple-white equipment and office space, and the Bryce owlclover 434. Pedicularis centranthera Canyon Natural History Association for finan­ Gray dwarf lousewort cial support. Therese Johnson and Charlie 435. Penstemon bracteatus Schelz provided us with the oppurtunity to Keck Platy penstemon work on the flora, for which we are grateful. 436. P. caespitosus Nutt. ex Linda Allen and Leila Schultz helped confirm Gray mat penstemon critical collections and helped check species 437. P. comarrhenus Gray dusty penstemon locations. 438. P. eatonii Gray firecracker penstemon 439. P. leiophyllus Pennell Markagunt penstemon 440. P. rostriflorus Kellogg beaked penstemon LITERATURE CITED 441. P. rydbergii A. Nels. Rydberg penstemon 442. P. stnctus Benth. Rocky Mountain ALBEE, B. ]., L. M. SCHULTZ, AND S. GOODRICH. 1988. penstemon Atlas of the vascular plants of Utah. Salt Lake City, 443. P. utahensis Eastw. Utah penstemon Utah Museum of Natural History Occasional Publi­ 444. Verbascum thapsus L. flannel mullein cations No.7. 445. Veronica americana Schw. American brooklime AHNOW, L. A. 1987. Gramineae. In .. S. L. \Ve!sh, N. D. Atwood, L. C. Higgins, and S. Goodrich, A Utah SINOPTERIDACEAE flora. Great Basin Naturalist Memoirs 9: 1-894. 446. Cheilanthes feei Moore slender lipfern BUCHANAN, H, AND R. GRAYBOSCH. 1981. Revised check­ list of the vascular plants of Bryce Canyon National SOLANACEAE Park, Utah. Great Basin Naturalist 41: 109-120. 447. Solanum triflorum Nutt. cutleaf nightshade CRONQVIST, A., A. H. HOL.MGREN, N. H. HOLMGREN, J. L. REVEAL, AND P. K. HOLMGRE:-l. 1972-1990. Inter­ TAMAIUCACEAE mountain flora. Vok 1, 3B, 4, 6. Hafner Publishing, 448. Tamarix ramosissima Columbia University Press, and New York Botanic Ledeb. salt cedar, tamarisk Gardens, New York. DAHLGREN, R., H. T. CLIFFORD, AND P. E. YEO. 1985. VALERIANACEAE The families of the monocotyledons. A comparative 449. Valeriana acutiloba Rydb. cordilleran valerian study. Springer-Verlag, Berlin. 450. V. edulis Nutt. taprooted valerian HALLSTEN, G. P., AND D. W. ROBERTS. 1988. Additions to the vascular flora of Bryce Canyon National Park, VEHBE;-JACEAE Utah. Great Basin Naturalist 48: 352. 451. Verbena bracteaw Lag. SPEl\'CE, J. R. 1992. An inventory and monitoring program & Rodr. prostrate vervain of the vegetation in the breaks, Bryce Canyon National Park, Utah. Final report to the National VIOLACEAE Park Service, Bryce Canyon National Park. Man­ zanita BioStudies, Torrey, Utah. 452. Viola adunca J.E. Smith blue violet WEBER, \'\T. A. 1987. Colorado flora: Western Slope. Col­ 453. V. purpurea Kellogg pine violet orado Associated University Press, Boulder. WELSH, S. L., N. D. ATWOOD, L. C. HIGGINS, AND S. VrSCACEAE GOODRICH. A Utah flora. Great Basin Naturalist 454. Arceuthobium cyanocarpum Memoirs 9: 1-894. Coult. & Rose limber pine dwarf mistletoe 1993J VASCULAR PLANTS OF BRYCE CANYON 217

INDEX TO GENERA AND COMMON NAMES

Abajo Heabane 60 Bebb willow 417 bulrush 197 Abies 289 bedstraw 412 bush ocean spray 397 Abronw 270, 271 lx:cplant 158 bushy knotweed 371 Ace,. 1, 2 bellflower 157 buttercup 386. 388 Achillea 20 belltgraSS 2!)C) Aconitum 379 lJerula 9 Ca[amagroslis 308, 309 African mustard 144 Belllla 113 Calochortus 156 Agoseris 21, 22 big sagehrush 36 Culylophus 272 Agropyron 298 bigtooth maple 2 Campanula 157 AgrostiJ 299 bindweed 178 c£lmpion 164 alderleaf mountain mahogany 394 hirch 113 C£lnada hll1egra~s 337 alfalfa 227 birdsbcak 428, 429 Canada wildrye 312 alkali sacaton 344 biscuitroot 13, 14 CupseUa 131 Allium 5, 6 Bistorta 361 Cu,'ex 183-193 Amanmthus 7 billerbru.c;h 408 carpet benlgmss 299 Amelanchier 392 black creeper sedge 190 carpet phlox 356 American bistort 361 black groundsel 93 Carruth sage\vort 30 American brooklime 445 black medic 226 Castilleja 425-427 American dragonhead 256 black sagebmsh 34 calchfly 162 American vetch 233 bladde.-pod 141-143 CuuLanthw; 132 Anderson larkspur 385 blazing star 266 Ceonothus 390, 391 Androsace 374 bluebell 127 Cedar Breaks guldenweed 75 Anemone 380 blucbunch whentl:.rnlss 320 Cenf(lUreCl 43 AtltCTInanCJ 23-26 blue elderherry 159 Cercocar"u.~ 393 Apache plume 395 blue-eyed grass 248 Chuenactis 44 Apocynum 16 blue flax 26.5 Chamber twtnpod L45 Aquilegia 381, 382 hlue grama 301 Chamisso arnica 27 Arabis 128-130 bluegrass 337, 339-341 chefltgrass 307 Arceuthobium 454-457 blueJeaf aster 39 CMikmthes 446 Arctostdphylos 203, 204 blue lettuce 86 Chenopodium 171-173 Arenaria 161 blue spruce 290 chicl.-weed 165 Aristida 300 bluestem 343 chokecherry 406 Arizona thistle 51 blue viold 452 Ch,'ysutlwmnus 45-50 Arnica 27. 2B blue wildrye 314 Cieuta 10 arrowgrass 254. 255 bog orchid 286, 287 cinquefoil 400-405 armwleafbalsamroot 41 Boute!ouu 30 1 Cin'ium 51--55 Artemisia 29--36 bracken fem 246 claret cup 152 Asclepias 17, 18 branchy groundsmoke 275 Clerrwtis 383, 384 Asparagus 19 breadroot 231 Cleome 1s..~ aspen bluebell 127 breaks bbdde.-pod 143 c1ifTrose 407 Aster 37-40 breaks dmha L36 clover 232 Astragalus 207-219 Brickelliu 42 c10verhead horsemint 259 Atrlplex 168-170 brickcllbush 42 clustered hroomrape 288 Bridges evening primrose 281 coast goldenrod 98 Baker cryptanth 115 bristleconc pine 293 Colorado columbine 381 hallhead gilia 3.52 bristly clyptanth 123 Colorado rubber 83 Balsumorhiza 41 Bromns 302-307 columbine 381, 382 barberry III brome 302-306 Cumwulra 422 barley 327, 328 bronze evening primrose 280 common dandeliun 104 bastard toadHax 422 brooklime 445 common juniper 180 beach dock 373 broom flax 263 common milkweed IS beaked penstemon 440 broom groundsel 97 common monkeyflower 430 beaked sedge 192 broomrape 288 cummon rockcress 130 beaked spikerush 196 broom snakcweed 71 common spikerush 194 bearberry 204 BryL't: Canyon pnintbmsh 427 common sunflower 77 bearded cinqucfoil402 buckwheat 362-368 Cunringia 133 bear flealr..:me 6B buU thistle 55 Convolvulus 178 218 GREAT BASIN NATURALIST [Volume 53

CO'alLx-hlza 283, 2lI4 Epilobium 273, 274 Clyceria 325 coralroot 283, 284 Epipaa;. 2B5 globeOower 380 Cordilleran valerian 449 Equisetum 201. 202 globemalJow 267, 268 CO'clylantlws 428, 429 Erigeron 60--69 goldenaster 79, 80 Comus 179 Eriog01ltlm 362-368 golden corydalis 235 Corvdnlis 235 Erysimum 137. 138 golden crownbeard 110 Corymb buckwheat 363 Euplwrbia 205, 206 golden CT)'Ptanth 116 cottonwood 414, 415 Eurotia 174 golden currant 244 creeping nailwort 163 evening primrose 272, 277-282 goldeneye 78 Crepis 57-59 goldenrod 92, 98-100 crescent milkvetch 208 Fallugia 395 golden sedge 184 crested wheatbfra.ss 298 Fendler meadowrue 389 goldenweed 72-75 crownbeard 110 Fendler mountain lilac 390 goosefoot 171-173 Cryptantha 114-123 Fendler sandwort 161 Gray hawksbeard 57 curlleafmountain mahogany 393 Fendler spurge 206 Gray horsebrush 105 curlycup gumwecd 70 femleaf fleabane 62 grass-of-Pamassus 424 curl)' dock 372 Festuca 322-324 great bladdet}' milkvetch 217 currant 244, 245 fewflowered goldenrod 100 great rushy mill.-vetch 216 cushion phlox 359 field bindweed 178 greasewood 176 cutleafnightshade 447 field horsetail 201 greenleafmanzanita 203 cutJeafwaterparsnip 9 field mill..-vetch 207 green sedge 189 Cyrnopterus 11 6eld pennycress 151 green wintergreen 377 field sagewort 29 Grindelia 70 Dacty!is 310 R, 289, 295 grizzJybear ptickJypear 153 dandelion 22, 104 firecracker penstemon 438 groundsel 93-97 death C"d.mas 269 fireweed 273 ground smoke 275 Delphinium 385 fishhook cactus ]55 gumweed 70 Deschampsia 311 flannel mullein 444 gumweed tansyaster 90 Descurainia 134, 135 flax 263-265 Cutierrezia 71 desert blazing star 266 f1axflower 354 desert paintbl"U,':ih 425 fleabane 60-69 Hobenaria 286, 287 desert phlox 355 flixweed 135 hairgrass 311 dock 372, 373 foothill sagebnlsh 33 Hall milkvetch 213 DodecatheOtl 375 fourwing saltbush ]68 hangpod locoweed 229 dogbane 16 fowl mannagrass 325 Haplopappus 72-75 dogwood 179 foxtail barley 328 hare's ear mustard 133 Douglas dustymaiden 44 foxtail muhly 330 Harriman yucca 4 Douglas dwarf mistletoe 456 Fragaria 396 Hart groundsel 94 Douglas fir 295 fragrant sand verbena 270 hawksbeard 57-59 Douglas knotweed 370 Frasera 236 heartleafarnica 28 Douglas sedge 186 Fremont barberry 111 HedeOl'n(J 257 Dralm 136 Fremont cottonwood 415 Hedysarum 220 drab buttercup 388 Fremont goosefoot 172 Helianthella 76 Dracocephalum 256 fringed brome 304 Heliattthus 77 dr'.tgonhead 256 fringed sagebrosh 32 Heliomeds 78 Drummond catchfly 162 Fritillaria 261 helleborine 285 Drummond willow 418 Heterotheca 79, 80 dwat{ lousewort 434 gallet.326 Hilaria 326 dwad mistletoe 454-457 Galium 412 hoary fleabane 61 dwarf rabbithrush 45 Gambel oak 234 hoary goldenaster 80 Gardner saltbush 170 hoary tansyaster 89 Eaton aster 38 GfJyaphytum 275. 276 hoary townsendia 107 Eaton fleabane 64 gentian 237-241 Holboell Tockcress 129 &:hin{)(;eretl$ 152 Gentiana 237. 238 Holodiscus 397 elderberry 159 Gen...."lla 239, 240 Hordeum 327, 328 El.acha'" 194-196 GentUmopsi.l 241 horehound 258 elkweed 236 Geranium 242, 243 horsebrush 105 Elymus 312-;321 Geyer willow 420 horsemint 259 Engelmann gentian 239 giant helleborine 285 horsetail 201, 20'2 English plantain 297 gi~a 351, 352 Hymenopappus 81 1993J VASCULAR PLA:ITS OF BRYCE CANYON 219

Hymenoxys 82 Lomatium 13 muhly 330-334 longleafphlox 357 mullein 444 Indian ricegrass 347 wtufi 222 muttongrass 338 Ipomopsw 351, 352 lousewort 434 Iris 247 low cryptanth 114 l1ailwort 163 Iva 84 low fleabane 66 narrowleafcottonwood 414 lvesia 398 low pussytoes 23 narrowJeaf goosefoot 173 low .sand \'erbena 271 narrowleaf sedge 193 Jacobs ladder 360 Lu,rinus 223--225 narrowleaf willow 419 James chicJ...weed 165 Lyclm~ 162 narrowleafyucca 3 James cryptanth 121 Lygoo.mlia 88 Nebraska sedge 188 Jones gentian 240 needle-and-thread grass 346 Jones goldenaster 79 Machoerantltera 89-91 needle grass 348-350 Jones reedgrass 308 Mahon'" III, 112 Nelson needlegrass 349 JUrn:wl 249-253 Malcolm'" 144 Nevada: onion 6 junegrass 329 mannagrass 325 Newberry twinpod 146 Juniperus 18D-182 manzanita 203 New Mexico thistle 53 juniper mistletoe 458 maple 1, 2 nightshade 447 maritime arrowgJ.-a5s 254 nodding bluegrass 340 Kelloggia 413 Markagunt penslemon 439 nodding brome 302 kentrophyta 215 Marrumum 258 northem grnss-of-Pamassus 424 Kentucky bluegrass 339 marsh arrowgrass 255 northern m-eetvetch 220 King birdsbeak 428 marsh buttercup 386 northern willowherb 274 King bladderpod 142 mat muhJy 334 Nutka rose 409 King flax 264 mat penstemon 436 Nuttall all:aligrass 342 King lupine 224 meadow barley 327 Nuttall Oaxflower 354 kitchenweed 276 meadow brome 306 knapweed 43 meadow fescue 324 oak 234 knotweed 369-371 meadow gentian 241 ocean spray 397 Koeleria 329 meadow hawksbeard 59 OerlOthera 277-282 meadowrue 389 onion 5, 6 wctuca 85, 86 Medicago 226, 227 Opuntia 153 Loppula 124 Melilotus 228 orange agoseris 21 larkspur 385 Mentze/ia 266 orchard grass 310 Lathy",-, 221 Merlensia 127 Oregon fleabane ff7 lavender evening primrose 272 milkvetch 207-214, 216-219 Oregon grape 112 least biscuitroot 13 miu...... veed 17, 18 Orobanche 2BB least townseDdia 108 Mimulm 430. 431 Orthocarpus 432, 433 leoparo lily 261 Missouri his 247 Oryzopsis 335 LRpidium 139, 140 mistletoe 454-458 ow1clover 432, 433 LRptodactylon 353 mock pennyroyal 257 Oxytropis 229, 230 Lesquerella 141-143 Monardella 259 Letterman needlegrass 348 monkeyflower 430, 431 Paxistima 167 lettuce 85, 86 monkshood 379 Pacific aster 37 Let/celene 87 moss phlox 358 paintbrush 425--427 Leucocrimnn 262 mountain brame 303 pale evening primrose 282 Ligustiwm 12 mountain chess 306 pale sedge 185 lily 261, 262 mountain dandelion 22 Panguitch buckwheat 365 limber pine 292 mountain deathcamas 269 Paria breadroot 231 limber pine dwarf mistletoe 454 mountain goosefoot 171 parsley 15 limestone thistle 52 mountain lilac 390, 391 Parnas$ia 424 LinantJu<.trwn 354 mountain locoweed 230 Paronychia 163 Unum 263-265 mountain lover 167 Parry bellflower 157 Lithophragma 423 mountain muhly 332 Parry hiscuitroot 14 Uthospermum 125, 126 mountain parsley 15 Parry gentian 238 little bluestem 343 mountain pepperweed 140 Parry goldenrod 99 littleseed ricegrass 335 mountain rabbHbrush 50 Parry rabbitbrush 48 little sun(lower 76 mountain snowberry 160 pedicularis 434 liverleaf wintergreen 376 mountain strawberry 396 PedioctJ(;tus 154 locoweed 229, 230 MuhlenbergiA 330-334 Pe&omelum 231 220 GREAT BASIN NATURALIST [Volume 53 pennycrcss 151 Ranunculus 386-388 Sclerocac'tus 155 pennyroyal 257 raspberry 411 seratchgrass 331 Pensfemon 435-443 red-osier dogwood 179 sedge 183-193 pepperweed 139, 140 redrootbuck\vheat366 sego lily 1.56 Peraphyllum 399 reedgrass .'308, .'309 selfhea1260 Peterson campion 164 Rhus 8 Senecio 93-97 Petradoria 92 Rihes 244, 245 serviceberry 392 Phleum 336 rieegrass 335, 347 Sevier ivesia 398 Phlox 355-359 Richardson geranium 243 shadscaJe 169 Phoradendron 458 Richardson tansymustard 134 sheep fescue 323 Physaria 145, 146 rock columbine 382 ShephL~(Ua 198-200 Picca 290 rockeress 128-130 shepherd's purse 131 pigmy sagebrush 35 rock goldenrod 92 shooting star 375 pinedrops 378 rockjasmine 374 shorthorn spurge 205 pine violet 453 Rocky Mountain clematis 383 shov',y goldeneye 78 pinewoods necdlcg:rass 350 Rocky Mountain gentian 237 showy milkweed 18 Pinus 291-294 Rocky Mountain goldenwced 74 showy rush pink 88 pinyon 291 Rocky Mountain juniper 182 showy stoneseed 125 pinyon pine dwarf mistletoe 455 Rocky Mountain maple 1 shrubby cinquefoil 403 Plantago 296, 297 Rocky Mountain penstemon 442 Silene 164 plantain 296, 297 rootstalk evening primrose 278 silky lupine 22.5 plateau cryptanth 119 Rosa 409, 410 silver buffaloberry 198 Platy penstcmon 435 rose heath 87 silverleaf milkvetch 209 Paa 337-341 Ross sedge 191 silvcrweed cinquefoil 400 Po[emonium 360 rosy pussytoes 24 Simpson pedioeadus 154 Po[ygonum 369-371 TOundleaf buflaloherry 200 Sisyrinchium 248 ponderosa pine 294 roundspike cryptanth 120 sixweeks fescue 322 Populus 414-416 rubherplant 83 skyrocket 351 Poteutilla 400--405 rubber rabbitbrush 47 slenderbeak sedge 183 poverty sumpweed 84 Rubus 411 slender buckwheat 364 prairie pepperweed 139 Rumex 372, 373 slender cinquefoil 404 pretty cinquefoil 401 rush 250-253 slender Iipfern 446 pretty shooting star 375 rush pink 88 slender wheatgrass 321 prickly lettllce 85 russet huffaloberry 199 slender wirelettuec 103 prickly pear 153 Russian knapweed 43 slender woodland star 423 prickly phlox 353 Russian thistle 175 slimstem reedgrass 309 prince's plume 148 Hussian wildrye 315 small bedstraw 412 prostrate pigweed 7 Rydberg penstemon 441 srnallflower birdsbeak 429 prostrate vervain 451 small-flowered globemallow 268 Prunella 260 sagebrush 33-36 small-leafgeranium 242 Primus 406 sagewort 29-30, 32 small-leafpussytoes 2.5 Pseudocymopterus 15 Salina wildrye 317 small wallflower 138 Pseudo'tsuga 295 Salix 417--421 smallwing sedge 187 Pteridium 246 Salsola 175 Smilacina 177 Pterospora 378 salsify 109 smooth brame 305 Puccinellia 342 saltbush 168, 170 smooth horsetail 202 pulse milkvetch 219 salt cedar 448 snakeweed 71 purple peavine 221 SambucuR 1.59 snowberry 160 purple springparsley 11 Sandberg bluegrass 341 Solanum 447 purple threeawn 300 sand dropseed 345 Solidago 98-100 purple-white owlclover 433 sand dune cryptanth 117 southern Iigusticum 12 Purshia 407, 408 sandhill muhly 333 southwestern dwarf mistletoe 457 pussytoes 23-26 sand verbena 270, 271 southwestern stoneseed 126 pygmy-flower rockjasmine 374 sandwort 161 squaw apple 399 Pyrola 376, 377 Sarcobatus 176 squawbush 8 searlet gilia 351 squaw currant 245 quaking aspen 416 scarlet globemallow 267 squirrel tail 313 Quercus 234 Scirpus 197 Sphaeralcea 267, 268 Schizachyrium 343 Sphaerumeria 101 rabbitbrush 4,5--50 Schoencrambe 147 spider milkweed 17 1993] VASCULAR PLANTS OF BRYCE CANYON 221 spikerush 194-196 tufted evening primrose 277 woolly pussytoes 26 Sporobolus 344, 345 tufted hairgrass 311 Wyoming paintbrush 426 spotted coralroot 283 twinpod 145, 146 spreading dogbane 16 twistflower 149 yarrow 20 spreading fleabane 63 two-grooved milkvetch 210 yellow beeplant 158 spreading rabbitbrush 46 two-needle pinyon 291 yellow clyptanth 118 spearwort 387 yellow evening primrose 279 spurge 205, 206 Uintah groundsel 96 yellowish cryptanth 122 Stanleya 148 Utah juniper 181 yellow owlclover 432 star lily 262 Utah mountain lilac 391 yellow salsify 109 starry false Solomon seall77 Utah penstemon 443 yellow sweetclover 228 Stellari{l 165, 166 Utah serviceberry 392 Ymca 3, 4 Stephanomeria 102, 103 Utah trefoil 222 stemless goldenweed 72 Zigadenus 269 stemless townsendia 106 Valeriana 449, 450 stemless woollybase 82 Vasey rabbitbrush 49 stickseed 124 Verhascum 444 Received 29 janua1'y 1992 Slipa 346-350 Verbena 451 Accepted 8 May 1993 Stokes buckwheat 367 Verhesina 110 stoneseed 125, 126 Veronica 445 strawberry 396 vervain 451 Streptanthus 149 vetch 233 striped coralroot 284 Vida 233 sulphur buckwheat 368 Viola 452, 453 sumpweed84 sunflower 76, 77 wallflower 137, 138 sweetvetch 220 wandering fleabane 69 Symphoricarpos 160 water hirch 113 swordleaf rush 251 water groundsel 95 water hemlock 10 tamarisk 448 waterparsnip 9 Tamarix 448 Watson bladderpod 141 tansyaster 89-91 Watson bog orchid 287 tansymustard 134 wavyleafthistle 54 tapertip onion 5 wax currant 245 taprooted valerian 450 weedy milkvetch 218 Taraxcu;um 104 western aster 40 tarragon 31 western hawksbeard 58 Tetradymia 105 western Jacohs ladder 360 Thalictrum 389 western stickseed 124 Thelypodium 150 western wallflower 137 thelypody 150 western water birch 113 thickspike wheatgrass 316 western wheatgrass 319 thistle 51-55 wheatgrass 298, 316, 318-321 Thlnspi 151 whiplash willow 421 thorn wirelettuce 102 white bog orchid 286 threeawn 300 white clover 232 threesquare bulrush 197 white fir 289 timber milkvetch 212 white virgin's bower 384 timothy 336 wildcabbage 132 thickstem wildcabbage 132 wildrye 312, 314, 315, 317 thrifty goldenweed 73 willow 417--421 toad rush 250 willowherb 274 Torrey milkvetch 211 winterfat 174 Torrey rush 253 winged buckwheat 362 Townsendia 106-108 wintergreen 376, 377 Tragopogon 109 wiregrass 249 trailing flebane 65 wirelettuce 102, 103 Trifolium 232 Woods rose 410 Triglochin 254, 255 woolly-foot plantain 296