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Srinu's Insectivores 10Jan04 Final

Srinu's Insectivores 10Jan04 Final

REVIEW ZOOS' PRINT JOURNAL 19(2): 1361-1371

CHECKLIST OF INSECTIVORES (MAMMALIA: ) OF SOUTH ASIA

S. Chakraborty 1, C. Srinivasulu 2*, Bhargavi Srinivasulu 2, M.S. Pradhan 3 and P.O. Nameer 4

1 Zoological Survey of , ‘M’ Block, New Alipur, Kolkata, West Bengal 700053, India. 2 Wildlife Biology Section, Department of Zoology, Osmania University, Hyderabad, Andhra Pradesh 500007, India. 3 Western Regional Station, Zoological Survey of India, Vidyanagar, Sector 29, Rawat Road, PCNTDA Post, Pune, Maharashtra 411044, India. 4 Department of Wildlife, College of Forestry, Kerala Agricultural University, Velanikkara, Thrissur, Kerala 680656, India. * Corresponding author Email: 1 [email protected]; 2 [email protected]; 3 [email protected]; 4 [email protected]

Abstract IUCN (1995), and, Roberts (1997) to gain insights on the current A checklist of 38 of insectivores belonging to taxonomic status of insectivores occurring in the region. 12 genera under four subfamilies in three families, Besides these, we also referred to numerous sources for (namely, Fischer, 1817; Fischer, 1817 information including Sclater (1891), Cabrera (1925), Lindsay and Soricidae Fischer, 1817) known to occur in South (1929), Allen (1938), Ellerman and Morrison-Scott (1951), Asia including Bangladesh, Bhutan, India, Nepal, Harrison (1958), Petter (1963), Biswas and Ghose (1970), Abe Pakistan and Sri Lanka is provided. (1971, 1932), Niethammer (1973), Chakraborty (1975, 1983), Doglov and Hoffman (1977), Jameson and Jones (1977), Mitchell Keywords (1975), Corbet (1978, 1988), Hutterer (1979), Corbet and Hill (1980, Checklist, Erinaceidae, Soricidae, Talpidae, 1986), Saha (1980), Hoffman (1986, 1987), Butler (1988), George Insectivora, Mammalia, South Asia (1988), Yates and Moore (1990), Abe et al. (1991), Frost et al. (1991), Agrawal (1993), Pradhan, et al. (1997), Nameer (1999), Nowak (1999), and Alfred et al. (2002).

Introduction This paper follows Srinivasulu and Pradhan (2003) for scientific Unlike Order Rodentia on which comprehensive works by names, synonyms and distribution information in South Asia. Ellerman (1940, 1961) exist, the Order Insectivora has not been given much attention. Details of species diversity of the Order Results Insectivora is available in Blyth (1863), Jerdon (1874), Sterndale A total of 38 species of insectivores belonging to 12 genera (1884), Blanford (1888), Lindsay (1929), Ellerman and Morrison- under four subfamilies of three families have been recorded Scott (1951), and, Corbet and Hill (1992). Very recently IUCN from South Asia. The family Erinaceidae Fischer, 1817 is (1995) has put together a comprehensive list of known represented by five species belonging to one genus, namely insectivores of Eurasia. As is applicable to all other groups, the Fitzinger, 1866. The family Talpidae Fischer, 1817 Order Insectivora too has witnessed numerous taxonomic includes two species in two genera, and the family Soricidae changes since the aforementioned literature became available. Fischer, 1817 includes 31 species in two subfamilies, namely – Keeping this in view, an attempt has been made, here, to record Soricinae Fischer, 1817 and Crocidurinae Milne-Edwards, 1872. an updated checklist of the insectivore species reported from The former subfamily includes 11 species in five genera, while this region. The checklist also provides information on their the latter includes 20 species in four genera (Table 1). synonyms, subspecies and distribution. Of this diversity, 15 species (39%) are endemic to South Asia Methods with four species being endemic to more than one country. Of The present work relies on Corbet and Hill (1992), Hutterer the seven species endemic to India four are restricted to (1993), IUCN (1995) and a list prepared by Dr. Mike Jordan. We Andaman and Nicobar Islands, while four are endemic to Sri reviewed the major works by Ellerman and Morrison-Scott (1951, Lanka (Table 2). Generically the genus Feroculus Kelaart, 1850 1953), Phillips (1980), Corbet and Hill (1991, 1992), Hutterer (1993), is endemic to South Asia, and the genus Solisorex Thomas,

Received 28 November 2003 Finally accepted 10 January 2004

February 2004 Zoos' Print Journal 19(2): 1361-1371 1361 Checklist of insectivores (Mammalia: Insectivora) of South Asia S. Chakraborty et al.

Table 1. Species diversity of Order Insectivora in South Asia Family: Erinaceidae Fischer, 1817 Subfamily: Erinaceinae Fischer, 1817 Scientific name No. of species Remarks Genus Hemiechinus Fitzinger, 1866 Erinaceidae Corbet and Hill (1992) treated this genus under subfamily Erinaceinae Hemiechinus 5 Endemics: 1 regional; 1 Indian Erinaceinae Fischer, 1817 in family Erinaceidae Fischer, 1817. Originally regarded as a subgenus of Linnaeus, 1758, Talpidae it was raised to generic status by Corbet (1978, 1988). Frost et al. (1991), IUCN (1995) and Nowak (1999) include two subgenus, 1 namely, Hemiechinus Fitzinger, 1866 and Parascaptor 1 Monotypic Trouessart, 1879. Ellerman and Morrison-Scott (1951) and Corbet Soricidae and Hill (1992) treat both these at the generic level. This genus Soricinae is widespread from northern Africa through and northern Anourosorex 1 Monotypic South Asia. This genus is represented by five species in two Chimmarogale 1 subgenera in the region. 4 1. Hemiechinus auritus (Gmelin, 1770) Soriculus 4 1770. Erinaceus auritus Gmelin, Nov. Comm. Acad. Sci. Petrop., Nectogale 1 Monotypic 14: 519. Crocidurinae Name: Long-eared 11 Endemic: 5 Indian; 1 Sri Lankan Type locality: Astrakhan, Southeastern Russia Synonyms: Erinaceus auritus Gmelin, 1770 Solisorex 1 Monotypic, Sri Lankan endemic Erinaceus megalotis Blyth, 1845 7 Endemic: 2 regional; 1 Indian; 2 Sri Subspecies: Hemiechinus auritus megalotis (Blyth, 1845) Lankan Distribution: Baluchistan in Pakistan. Feroculus 1 Monotypic, regional endemic Comments: Belongs to subgenus Hemiechinus Fitzinger, 1866. Ellerman and Morrison-Scott (1951) treated Hemiechinus megalotis Blyth, 1845 as a distinct species and included 16 subspecies of Hemiechinus auritus (Gmelin, 1770) of which 1924 is endemic to Sri Lanka. only one subspecies, now a species, pertains to South Asia. This was also followed by Roberts (1997). Niethammer (1973) The present known insectivore diversity on record from South treated Hemiechinus megalotis Blyth, 1845 as a subspecies, Asia is listed below. The synonyms provided are those followed by Corbet and Hill (1992), Hutterer (1993) and IUCN applicable for the South Asian region. We have not listed extra- (1995). limital synonyms that may be valid for some taxa elsewhere.

Table 2. Insular endemic species of Order Insectivora in South Asia

Scientific Name Distribution Remarks

Crocidurinae Crocidura andamanensis Andaman & Nicobar Islands Only from South Andaman Isle Crocidura hispida Andaman & Nicobar Islands Only from Middle Andaman Isle Crocidura jenkinsi Andaman & Nicobar Islands Only from South Andaman Isle Crocidura miya Sri Lanka Crocidura nicobarica Andaman & Nicobar Islands Only from Great Nicobar Isle Solisorex pearsoni Sri Lanka Only fron C Highlands Suncus fellowes-gordoni Sri Lanka Suncus zeylanicus Sri Lanka

1362 February 2004 Zoos' Print Journal 19(2): 1361-1371 Checklist of insectivores (Mammalia: Insectivora) of South Asia S. Chakraborty et al.

2. Hemiechinus collaris (Gray, 1830) Name: 1830. Erinaceus collaris Gray in Hardwicke, Illust. Indian Zool., Type locality: Bhawalpur, Punjab, Pakistan. I pl. 8. Synonyms: Erinaceus micropus Blyth, 1846 Name: Collared Hedgehog Paraechinus micropus (Blyth, 1846) Type locality: Doab, between Yamuna and Ganges rivers, India Hemiechinus mentalis Fitzinger, 1867 Synonyms: Erinaceus collaris Gray, 1830 Erinaceus pictus Stoliczka, 1872 Erinaceus spatangus Bennett, 1832 Paraechinus intermedius Biswas and Ghose, 1970 Erinaceus grayi Bennett, 1832 Paraechinus intermedius kutchicus Biswas and Erinaceus indicus Royle, 1833 Ghose, 1970 Erinaceus blanfordi Anderson, 1878 Subspecies: Hemiechinus micropus kutchicus (Biswas & Paraechinus hypomelas blanfordi (Anderson, 1878) Ghose, 1970) Subspecies: None. Distribution: Gujarat, Maharashtra, Punjab and Rajasthan in Distribution: Endemic to the region. Delhi, Gujarat, Himachal India; Pakistan. Pradesh, Jammu & Kashmir, Punjab, Rajasthan and Uttar Comments: Belongs to subgenus Paraechinus Trouessart, Pradesh in India; Sind in Pakistan. Maharashtra specimen in 1879. Ellerman and Morrison-Scott (1951) synonymized National Zoological Collection, Kolkata, India perhaps could Erinaceus nudiventris Horsfield, 1851 with Paraechinus be an ‘introduced’ or ‘isolate’ specimen. micropus (Blyth, 1846) with a doubtful status, a trend followed Comments: Belongs to subgenus Hemiechinus Fitzinger, by Corbet and Hill (1992). Frost et al. (1991), Hutterer (1993) 1866. Ellerman and Morrison-Scott (1951), and, Corbet (1978) and IUCN (1995) treat nudiventris as a distinct species – included this as a subspecies under Hemiechinus auritus Hemiechinus nudiventris (Horsfield, 1851) under subgenus (Gmelin, 1770). It has been accorded specific status by Corbet Paraechinus Trouessart, 1879. Biswas and Ghose (1970) and Hill (1992). The name Erinaceus blanfordi Anderson, 1878 described new species and subspecies, Paraechinus erroneously included under the Paraechinus hypomelas Brandt, intermedius and Paraechinus intermedius kutchicus. 1836 by Ellerman and Morrison-Scott (1951), and, Corbet (1988) is in fact a Hemiechinus form as pointed out by Agrawal (1973). 5. Hemiechinus nudiventris (Horsfield, 1851) Hutterer (1993) and IUCN (1995) also synonymize Erinaceus 1851. Erinaceus nudiventris Horsfield, Cat. Mamm. Mus. E. blanfordi Anderson, 1878 with this taxon. India Co., p 136. Name: Madras Hedgehog 3. Hemiechinus hypomelas (Brandt, 1836) Type locality: Madras, India. 1836. Erinaceus hypomelas Brandt, Bull. Sci. St. Petersb, I: 32. Synonyms: Erinaceus nudiventris Horsfield, 1851 Name: Brandt’s Hedgehog Paraechinus micropus nudiventris (Horsfield, 1851) Type locality: Northern Persia. Subspecies: None Synonyms: Erinaceus hypomelas Brandt, 1836 Distribution: Endemic to India. ?Andhra Pradesh, Kerala and Paraechinus hypomelas (Brandt, 1836) Tamil Nadu. Erinaceus jerdoni Anderson, 1878 Comments: Belongs to subgenus Paraechinus Trouessart, Paraechinus amir Thomas, 1918 1879. Ellerman and Morrison-Scott (1951) synonymized Subspecies: Hemiechinus hypomelas hypomelas (Brandt, 1836) Erinaceus nudiventris Horsfield, 1851 with Paraechinus Hemiechinus hypomelas jerdoni (Anderson, 1878) micropus (Blyth, 1846), a trend followed by Corbet and Hill Distribution: Pakistan. (1992). Frost et al. (1991), Hutterer (1993), and IUCN (1995) Comments: Belongs to subgenus Paraechinus Trouessart, treat it as a distinct species. Srinivasulu and Nagulu (2002) 1879. Ellerman and Morrison-Scott (1951) erroneously doubtfully report the occurrence of this species in southern synonymized Erinaceus blanfordi Anderson, 1878 with Nallamala Hills in Andhra Pradesh. Paraechinus hypomelas blanfordi (Anderson, 1878) that Agrawal (1973) rectified and placed it under Hemiechinus auritus Family: Talpidae Fischer, 1817 (Gray, 1830). The name Erinaceus jerdoni Anderson, 1878 too Subfamily: Talpinae Fischer, 1817 was synonymized with Paraechinus hypomelas blanfordi Genus Euroscaptor Miller, 1940 (Anderson, 1878) by Ellerman and Morrison-Scott (1951). South Asian Moles Corbet and Hill (1992) treat this taxon as Paraechinus hypomelas Genus Euroscaptor Miller, 1940 was considered either as a (Brandt, 1836) and list two subspecies. Frost et al. (1991), and subgenus of or synonym of Linnaeus, 1758 by earlier IUCN (1995) also list two valid subspecies from the region. authorities such as Ellerman and Morrison-Scott (1951). However, Yates and Moore (1990) established this taxon as 4. Hemiechinus micropus (Blyth, 1846) generically distinct from Talpa Linnaeus, 1758. Corbet and Hill 1846. Erinaceus micropus Blyth, J. Asiatic Soc. Bengal, 15: 70. (1992) did not list Euroscaptor Miller, 1940 as a separate genus

February 2004 Zoos' Print Journal 19(2): 1361-1371 1363 Checklist of insectivores (Mammalia: Insectivora) of South Asia S. Chakraborty et al. but included forms under Talpa Linnaeus, 1758. We follow Family: Soricidae Fischer, 1817 IUCN (1995) in recognising Euroscaptor Miller, 1940. The family Sori cidae Fischer, 1817 includes two subfamilies, namely, Soricinae Fischer, 1817 and Crocidurinae Milne- 6. Euroscaptor micrura (Hodgson, 1841) Edwards, 1872. 1841. Talpa micrurus Hodgson, Calcutta J. nat. Hist., 2: 221. Name: Himalayan Subfamily: Soricinae Fischer, 1817 Type locality: Central and northern hills of Nepal Genus Anourosorex Milne-Edwards, 1870 Synonyms: Talpa micrurus Hodgson, 1841 Mole- Talpa micrura Hodgson, 1841 This genus belongs to subfamily Soricinae Fischer, 1817. This Talpa cryptura Blyth, 1843 monospecific genus is restricted to Oriental region. Talpa macrura Hodgson, 1858 Subspecies: None. 8. Anourosorex squamipes Milne-Edwards, 1872 Distribution: Arunachal Pradesh, Assam, Sikkim and West 1872. Anourosorex squamipes Milne-Edwards, Rech. Hist. Nat. Bengal in India; Nepal. Mamm., 264. pl. 38, fig. 1, pl 38A, figs I-Ij. Comments: The genus Euroscaptor Miller, 1940 had been Name: Mole-Shrew earlier synonymized or treated as sub genus of the genus Talpa Type locality: Sichuan Prov., Moupin, China Linneaus, 1758 by various authors. Ellerman and Morrison- Synonyms: Anourosorex assamensis Anderson, 1875 Scott (1951) treated it as Talpa micrura Hodgson, 1841 also Subspecies: None for the region. followed by Corbet and Hill (1992). Yates and Moore (1990) Distribution: Arunachal Pradesh, Assam, Meghalaya and and Hutterer (1993), based on distinct characters, recognized it Mizoram in India; Bhutan. as a distinct genus. Comments: Corbet and Hill (1992) doubt the status of the mainland subspecies – Anourosorex squamipes capinas Allen, Genus Parascaptor Gill, 1875 1923, Anourosorex assamensis capito Allen, 1923; and Indian Mole Anourosorex squamipes schmidi Petter, 1963, but further Genus Parascaptor Gill, 1875 was considered either as a maintain that the insular form Anourosorex squamipes subgenus of or synonym of Talpa Linnaeus, 1758 by earlier yamashinai Kuroda, 1935 could possibly be accepeted as a authorities like Ellerman and Morrison-Scott (1951). However, distinct subspecies. IUCN (1995) retains this arrangement. later authorities treated it as generically disinct from Talpa However, Hutterer (1993) retains Anourosorex squamipes Linnaeus, 1758. See comments under Parascaptor leucura schmidi Petter, 1963 along with Anourosorex squamipes (Blyth, 1850). Monospecific genus restricted to Oriental region yamashinai Kuorda, 1935 as subspecies under this species from northeastern India to southern China (Allen, 1938). following Petter (1963) and Jameson and Jones (1977).

7. Parascaptor leucura (Blyth, 1850) Genus Chimmarogale Anderson, 1877 1850. Talpa leucura Blyth, J. Asiat. Soc. Bengal, 19: 215, pl 4, Water Shrew fig. 1. This genus belongs to subfamily Soricinae Fischer, 1817. This Name: Indian Mole genus is restricted to South, Southeast and East Asia and is Type locality: Cherrapunji in Khasi Hills, Assam (now represented by one species in the region (Hoffman, 1987). Meghalaya), India Synonyms: Talpa leucura Blyth, 1850 9. Chimmarogale himalayica (Gray, 1842) Talpa micrura leucura (Blyth, 1850) 1842. Crossopus himalayicus Gray, Ann. Mag. Nat. Hist., [1]10: 261. Subspecies: None. Name: Distribution: Meghalaya in India. Type locality: Chamba, NE Punjab (now Himachal Pradesh), India Comments: The genus Parascaptor Gill, 1875 had been Synonyms: Crossopus himalayicus Gray, 1842 earlier synonymized or treated as sub genus of the genus Talpa Subspecies: None for the region. Linneaus, 1758 by authors like Corbet and Hill (1991). Corbet Distribution: Himachal Pradesh, Jammu and Kashmir, Sikkim (1978), Abe et al. (1991) and Hutterer (1993) recognized it as a and West Bengal in India; Nepal. distinct genus. See comments under genus Euroscaptor Miller, Comments: Ellerman and Morrison-Scott (1951) and 1940. Ellerman and Morrison-Scott (1951) treated it as Talpa Harrison (1958) included Chimmarogale himalayica (Gray, micrura leucura (Blyth, 1850), also followed by Corbet and Hill 1842) as subspecies of Chimmarogale platycephala (Temminck, (1992) who remarked that it could be a separate species but 1842). Jones and Mumford (1971) did not consider doubted that it warranted generic separation. Chimmarogale platycephala (Temminck, 1842) distinct from Chimmarogale himalayica (Gray, 1842). Later, Hoffman (1987) showed it to be distinct species based on detailed taxonomic

1364 February 2004 Zoos' Print Journal 19(2): 1361-1371 Checklist of insectivores (Mammalia: Insectivora) of South Asia S. Chakraborty et al. study. Corbet and Hill (1992) and IUCN (1995) retained it as Linnaeus, 1766. Corbet and Hill (1992) following Hoffman (1987) distinct species. Hutterer (1993) misinterpreted Chimmarogale included planiceps under Sorex thibetanus Kastschenko, 1905. platycephala (Temminck, 1842) to be listed by Ellerman and The specimen described as Sorex thibetanus was found to be Morrison-Scott (1951) as subspecies of Chimmarogale those akin to Sorex minutus thus invalidating the former name himalayica (Gray, 1842). and being considered as nomen dubium (Hutterer, 1979). Hutterer (1993) and IUCN (1995) treated Sorex minutus Genus Sorex Linnaeus, 1758 Linnaeus, 1766 and Sorex planiceps Miller, 1911 as distinct Long-tailed species based on the latter’s distinctly larger cranial This genus is represented by four species belonging to measurements. subgenus Sorex Linnaeus, 1758. 13. Sorex planiceps Miller, 1911 10. Sorex bedfordiae Thomas, 1911 1911. Sorex planiceps Miller, Proc. Biol. Soc. Wash., 24: 242. 1911. Sorex bedfordiae Thomas, Abstr. Proc. Zool. Soc., Lond., Name: Kashmir Shrew 1911(90): 3. Type locality: Dachin, Khistwar, Kashmir, India Name: Lesser Stripe-backed Shrew Synonyms: None. Type locality: Omisan, Sichuan, China Subspecies: None. Synonyms: Sorex cylindricauda bedfordiae Thomas, 1911 Distribution: Jammu and Kashmir, and Pakistan-occupied- Subspecies: Sorex bedfordiae nepalensis Weigel, 1969 Kashmir in India; northern Pakistan. Distribution: Nepal. Comments: Ellerman and Morrison-Scott (1951) treated it Comments: Ellerman and Morrison-Scott (1951) treated under Sorex minutus Linnaeus, 1766, while Doglov and Hoffman Sorex bedfordiae Thomas, 1911 as synonym of Sorex (1977) and Hoffman (1987) listed it as a subspecies of Sorex cylindricauda Milne-Edwards, 1872. Corbet (1978), Hoffman thibetanus Kastschenko, 1905, also followed by Corbet and (1987), Corbet and Hill (1992), Hutterer (1993) and IUCN (1995) Hill (1992). Hutterer (1979) retained it as a distinct species from treated this species specifically different than Sorex either Sorex minutus Linnaeus, 1766 or Sorex thibetanus cylindricauda Milne-Edwards, 1872. Kastschenko, 1905 (now considered a nomen dubium) based on larger skull measurements, a trend followed by Hutterer 11. Sorex excelsus Allen, 1923 (1993) and IUCN (1995). 1923. Sorex excelsus Allen, Am. Mus. Novit., 100: 4. Name: Genus Soriculus Blyth, 1854 Type locality: Summit of Hoshan, Peitai, South of Chungtein, Brown-toothed (or Asiatic) Shrews Yunnan, China Corbet and Hill (1992) kept this genus under subfamily Soricinae Synonyms: Sorex araneus excelsus (Allen, 1923) Fischer, 1817. It is represented by four species in two subgenera Subspecies: None. – Soriculus Blyth, 1854 and Ellerman and Distribution: Nepal. Morrison-Scott (1951) in the region. Van Valen (1967) and Comments: Ellerman and Morrison-Scott (1951) treated Repenning (1967) treated the latter subgenus as a genus, while excelsus Allen, 1923 as a subspecies of Sorex araneus Linnaeus, Hoffman (1986) considered the differences between them to be 1758. Corbet (1978) treated it as a possible subspecies of Sorex too small to separate them generically. asper Thomas, 1914. Hutterer (1993) following Hoffman (1987) treated Sorex excelsus Allen, 1923 as a distinct species and 14. Soriculus caudatus (Horsfield, 1851) further remarked that the specimen from Nepal reported by 1851. Sorex caudatus Horsfield, Cat. Mamm. Mus. E. India Co.,: 135. Agrawal and Chakraborty (1971) possibly belonged to this Name: Hodgson’s Brown-toothed Shrew species. Type locality: Darjeeling, West Bengal, India Synonyms: Sorex caudatus Horsfield, 1851 12. Sorex minutus Linnaeus, 1766 Episoriculus caudatus (Horsfield, 1851) 1766. Sorex minutus Linnaeus, Syst. Nat. 12th ed., I: 73. Sorex gracilicauda Anderson, 1877 Name: Lesser Shrew Subspecies: Soriculus caudatus caudatus (Horsfield, 1851) Type locality: Bernaul, Western Siberia Soriculus caudatus soluensis Gruber, 1969 Synonyms: Sorex thibetanus Kastschenko, 1905 (nom. dub.) Distribution: Jammu and Kashmir, Sikkim, Uttranchal and Subspecies: Sorex minutus minutus Linnaeus, 1766 West Bengal in India; Nepal. Distribution: Jammu and Kashmir, and Pakistan-occupied- Comments: This species belongs to subgenus Episoriculus Kashmir in India; Nepal. Ellerman and Morrison-Scott, 1951. Van Valen (1967) and Comments: Ellerman and Morrison-Scott (1951) doubtfully Repenning (1967) treated this taxon as Episoriculus caudatus treated planiceps Miller, 1911 as subspecies of Sorex minutus (Horsfield, 1851).

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15. Soriculus leucops (Hodgson, 1855) India; Bhutan; Nepal. 1855. Sorex leucops Hodgson in Horsfield, Ann. Mag. Nat. Hist., Comments: This species belongs to subgenus Soriculus [2]16: 111. Blyth, 1854. specimens belong to Soriculus Name: Indian Long-tailed Shrew nigriscens radulus Thomas, 1922. Type locality: Nepal Synonyms: Sorex leucops Hodgson, 1855 Genus Nectogale Milne-Edwards, 1870 Episoriculus leucops (Hodgson, 1855) Web-footed (or Elegant) Water Shrew Sorex nivicola Gray, 1863 (nom. nud.) This genus belongs to subfamily Soricinae Fischer, 1817. It is Sorex minor Dobson, 1890 monospecific genus represented by a species that is restricted Sorex baileyi Thomas, 1914 to Oriental region and shows affinity to high altitude mountain Soriculus caudatus bailey (Thomas, 1914) streams (Mitchell, 1975). Soriculus gruberi Weigel, 1969 Subspecies: None. 18. Nectogale elegans (Milne-Edwards, 1870) Distribution: Assam, Manipur, Sikkim and West Bengal in 1870. Nectogale elegans Milne-Edwards, C.R. Acad. Sci. Paris, India; Nepal. 70: 341. Comments: This species belongs to subgenus Episoriculus Name: Web-footed/Elegant Water Shrew Ellerman and Morrison-Scott, 1951. Van Valen (1967) and Type locality: Moupin, Sichuan, China Repenning (1967) treated this taxon as Episoriculus leucops Synonyms: Nectogale sikhimensis de Winton and Styan, 1899 (Hodgson, 1855). Sometimes the names baileyi and gruberi Subspecies: Nectogale elegans elegans Milne-Edwards, 1870 are considered valid species (IUCN, 1995). However, according Nectogale elegans sikhimensis de Winton and to Corbet and Hill (1992) the forms representing Sorex baileyi Styan, 1899 Thomas, 1914 are confused with Soriculus macrurus Blanford, Distribution: Sikkim in India; Bhutan; Nepal. 1888. Hutterer (1993) includes Sorex baileyi as subspecies of Comments: Monospecific, distributed in the Himalaya from Soriculus leucops (Hodgson, 1855) following Hoffman (1986). Nepal to Sichuan in China. See Hoffman (1987) for comments on status of Nectogale sikhimensis deWinton and Styan, 1899 16. Soriculus macrurus Blanford, 1888 based on which Corbet and Hill (1992), Hutterer (1993) and 1888. Soriculus macrurus Blanford, Fauna Brit. India Mamm. I: 231. IUCN (1995) treated it as a distinct subspecies. Name: Arboreal Brown-toothed Shrew Type locality: Darjeeling, West Bengal, India Subfamily Crocidurinae Milne-Edwards, 1872 Synonyms: Sorex macrurus Hodgson, 1863 (nom. nud.) Genus Crocidura Wagler, 1832 Episoriculus macrurus (Blanford, 1888) White-toothed Shrews Subspecies: None This genus belongs to subfamily Crocidurinae Milne-Edwards, Distribution: Sikkim in India; Nepal. 1872. This genus is widespread ranging from Africa to Europe Comments: This species belongs to subgenus Episoriculus and South East Asia and includes many forms in four Ellerman and Morrison-Scott, 1951. Ellerman and Morrison- subgenera. It is represented by 11 species belonging to Scott (1951) synonymized it with Soriculus leucops (Hodgson, subgenus Crocidura Wagler, 1832 in the region. 1855) following Osgood (1932). Van Valen (1967) and Repenning (1967) treated this taxon as Episoriculus macrurus (Blanford, 19. Crocidura andamanensis Miller, 1902 1888). Corbet and Hill (1992), Hutterer (1993), and IUCN (1995) 1902. Crocidura andamanensis Miller, Proc. U. S. Natnl. Mus., 24: 777. treat it as distinct species following Hoffman (1986). Name: Andaman White-toothed Shrew Type locality: MacPherson Strait, South Andaman Isle, 17. Soriculus nigriscens (Gray, 1842) Andaman and Nicobar Islands, India 1842. Corsira nigriscens Gray, Ann. Mag. Nat. Hist., [Ser]10: 261. Synonyms: None. Name: Sikkim Large-clawed Shrew Subspecies: None. Type locality: Darjeeling, West Bengal, India Distribution: Endemic to India, known only from type locality. Synonyms: Corsira nigriscens Gray, 1842 Comments: According to Ellerman and Morrison-Scott Sorex aterrimus Blyth, 1842 (nom. nud.) (1951) this taxon is probably closely allied to Crocidura Sorex sikimensis Hodgson, 1849 (nom. nud.) nicobarica Miller, 1902. Subspecies: Soriculus nigriscens nigriscens (Gray, 1842) Soriculus nigriscens caurinus Hinton, 1922 20. Crocidura attenuata Milne-Edwards, 1872 Soriculus nigriscens pahari Hinton, 1922 1872. Crocidura attenuata Milne-Edwards, Rech. Hist. Nat. Soriculus nigriscens centralis Hinton, 1922 Mamm., pp. 263. Distribution: Assam, Sikkim, Uttranchal and West Bengal in Name: Grey Shrew

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Type locality: Moupin, Sichuan Province, China Sorex (Crocidura) myoides Blanford, 1875 Synonyms: Crocidura rubricosa Anderson, 1877 Subspecies: Crocidura horsfeildi horsfieldi (Tomes, 1856) Crocidura attenuata rubricosa Anderson, 1877 Crocidura horsfieldi myoides (Blanford, 1875) Crocidura kingiana Anderson, 1877 Distribution: Jammu and Kashmir in India; Nepal; Sri Lanka. Crocidura attenuata kingiana Anderson, 1877 Comments: Refer Jenkins (1976) and Jameson and Jones Subspecies: None. (1977) for further details. Variations between insular (Sri Lankan) Distribution: Assam, Jammu and Kashmir, Sikkim and and Himalayan forms require further taxonomic studies. Uttaranachal, in India; Nepal; Bhutan. Comments: Ellerman and Morrison-Scott (1951) included 24. Crocidura jenkinsi Chakraborty, 1978 four subspecies under this taxon. However, Corbet and Hill 1978. Crocidura jenkinsi Chakraborty, Bull. Zool. Surv. India, 1: 303. (1992), Hutterer (1993) and IUCN (1995) maintain that there are Name: Jenkin’s Andaman Spiny Shrew no subspecies of this taxon. Type locality: Wright Myo, South Andaman Isle, Andaman and Nicobar Islands, India 21. Crocidura gueldenstaedtii (Pallas, 1811) Synonyms: None. 1811. Sorex güldenstaedtii Pallas, Zoogr. Ross. Asiat., I: 132. pl. Subspecies: None. 9, fig. 1. Distribution: Endemic to India, known only from type locality Name: Gueldenstaedt’s White-toothed Shrew and Mount Harriet. Type locality: Near Dushet, Georgia Comments: None. Synonyms: Sorex güldenstaedtii Pallas, 1811 Crocidura russula güldenstaedtii (Pallas, 1811) 25. Crocidura leucodon (Hermann, 1780) Crocidura pullata Miller, 1911 1780. Sorex leucodon Hermann in Zimmerman, Geogr. Gesch. Crocidura russula pullata (Miller, 1911) Mensch. Vierf. Thiere, 2: 382. Subspecies: None. Name: Bicoloured White-toothed Shrew Distribution: Jammu and Kashmir, and Pakistan-occupied- Type locality: Bashkin, Strasburg, France Kashmir in India; North West Frontier Province in Pakistan. Synonyms: Sorex leucodon Hermann, 1780 Comments: Ellerman and Morrison-Scott (1951) and Jenkins Subspecies: None for the region. (1976) included Crocidura gueldenstaedtii Pallas, 1811 and Distribution: Jammu and Kashmir, and Pakistan-occupied- Crocidura pullata Miller, 1911 as subspecies of Crocidura Kashmir in India; probably also in Pakistan. russula Hermann, 1780. These have been treated as distinct Comments: Chakraborty (1983) reports its occurrence in species by Corbet and Hill (1992), Hutterer (1993), and IUCN India based on a male specimen collected in 1974 from Daksum, (1995). Furthermore, Hutterer (1993) reffering to Catzeflis et al. Jammu and Kashmir. (1985) opines that Crocidura güeldenstaedtii Pallas, 1811, Crocidura pullata Miller, 1911 and Crocidura russula Hermann, 26. Crocidura miya Phillips, 1929 1780 could be conspecific with Crocidura suaveolens Pallas, 1929. Crocidura miya Phillips, Spolia Zeylan., 15: 113. 1811. Name: Sri Lankan Long-tailed Shrew Type locality: Moolgama, Kandiyan Hills, Nilambe dt., Sri 22. Crocidura hispida Thomas, 1913 Lanka 1913. Crocidura hispida Thomas, Ann. Mag. Nat. Hist., [8]11: 468. Synonyms: None. Name: Andaman Shrew Subspecies: None. Type locality: North End, Middle Andaman Isle, Andaman and Distribution: Endemic to Sri Lanka. Nicobar Islands, India Comments: Refer Phillips (1980) for further details. Synonyms: None. Subspecies: None. 27. Crocidura nicobarica Miller, 1902 Distribution: Endemic to India, known only from Andaman 1902. Crocidura nicobarica Miller, Proc. U. S. Natnl. Mus., 24: 776. Isles, Andaman and Nicobar Islands. Name: Nicobar Shrew Comments: None. Type locality: Greater Nicobar Isle, Andaman and Nicobar Islands, India 23. Crocidura horsfieldi (Tomes, 1856) Synonyms: None. 1856. Sorex horsfieldi Tomes, Ann. Mag. Nat. Hist., [2]17: 23. Subspecies: None. Name: Horsfield’s Shrew Distribution: Endemic to India, known only from type locality. Type locality: Sri Lanka Comments: None. Synonyms: Sorex horsfieldi Tomes, 1856 Crocidura retusa Peters, 1870

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28. Crocidura pergrisea Miller, 1913 Type locality: Trichur (now Thrissur), Cochin (now Kochi) in 1913. Crocidura pergrisea Miller, Proc. Biol. Soc. Washington, 26: 113. Kerala, India Name: Pale Grey Shrew Synonyms: Crocidura dayi Dobson, 1888 Type locality: Skoro Loomba, Shigar, Baltistan, Jammu and Subspecies: None for the region Kashmir, India Distribution: Endemic to India, known only from a few Synonyms: None. localities in southern India Subspecies: None. Comments: None. Distribution: Endemic to India, known only from type locality in Pakistan-occupied-Kashmir. 32. Suncus etruscus (Savi, 1822) Comments: Ellerman and Morrison-Scott (1951) included 1822. Sorex etruscus Savi, Nuovo Giorn. De Litterati, Pisa, 1: 60. zarudnyi Ognev, 1928 as subspecies of this taxon. Hassinger Name: Savi’s Pygmy Shrew (1973) treated zarudnyi Ognev, 1928 as a distinct species, a Type locality: Pisa, Italy trend that was followed by Corbet and Hill (1992), Hutterer Synonyms: Sorex etruscus Savi, 1822 (1993) and IUCN (1995). Sorex perrotteti Duvernoy, 1842 Sorex micronyx Blyth, 1855 29. Crocidura zarudnyi Ognev, 1928 Sorex hodgsoni Blyth, 1855 1928. Crocidura zarudnyi Ognev, Mamm. E. Europe, N. Asia, I: 341. Sorex atratus Blyth, 1855 Name: Zarudny’s Shrew Ssorex nudipes Blyth, 1855 Type locality: Near Baluchistan border, eastern Iran. Pachyura assamensis Anderson, 1873 Synonyms: Crocidura pergrisea zarudnyi Ognev, 1928. Crocidura (Pachyura) nilagirica Anderson, 1877 Subspecies: None. Crocidura (Pachyura) pygmaeoides Anderson, Distribution: Baluchistan in Pakistan. 1877 Comments: Earlier named as Crocidura tatianae Ognev, Crocidura (Pachyura) travancorensis 1921, the name that was preoccupied by Crocidura nyansae Anderson, 1877 tatianae Dollman, 1915 a taxon from Africa. Thus, the current Crocidura (Pachyura) nitidofulva Anderson, name Crocidura zarudnyi Ognev, 1928 is valid. 1877 Subspecies: Suncus etruscus perrotteti (Duvernoy, 1842) Genus Solisorex Thomas, 1924 Suncus etruscus nudipes (Blyth, 1855) Sri Lankan Long-clawed Shrew Suncus etruscus micronyx (Blyth, 1855) This monospecific Sri Lankan endemic genus belongs to Suncus etruscus pygmaeoides (Anderson, 1877) subfamily Crocidurinae Milne-Edwards, 1872. Suncus etruscus nitidofulva (Anderson, 1877) Distribution: Andhra Pradesh, Assam, Bihar, Chattisgarh, 30. Solisorex pearsoni Thomas, 1924 Gujarat, Harayana, Himachal Pradesh, Jharkhand, Karnataka, 1924. Solisorex pearosni Thomas, Spolia Zeylan., 13: 94. Kerala, Maharashtra, Manipur, Orissa, Pondicherry, Punjab, Name: Pearson’s Long-clawed Shrew Rajasthan, Sikkim, Tamil Nadu, Uttaranchal, Uttar Pradesh and Type locality: Hakgala, Nuwara Eliya, Central Highlands, Sri West Bengal in India; Nepal; Bhutan; Pakistan; Sri Lanka. Lanka Comments: Ellerman and Morrison-Scott (1951) listed Synonyms: None. Suncus fellowes-gordoni Phillips, 1932 under this taxon as a Subspecies: None. subspecies, while Corbet and Hill (1991, 1992) synonymized it Distribution: Endemic to Central Highlands of Sri Lanka. with this taxon. However, Hutterer (1993) and IUCN (1995) Comments: None. treated it as distinct species. Eisenberg and McKay (1970) did not treat Podihik kura Deraniyagala, 1958 being distinct from Genus Suncus Erhenberg, 1833 Suncus etruscus (Savi, 1822), as followed by Hutterer (1993). Musk Shrews This genus belongs to subfamily Crocidurinae Milne-Edwards, 33. Suncus fellowes-gordoni Phillips, 1932 1872. Ellerman and Morrison-Scott (1951) and Harrison and 1932. Suncus fellowes-gordoni Phillips, Spolia Zeylan., 17: 124. Bates (1991) suggested that this genus was not more than a Name: Ceylon Pygmy Shrew subgenus of Crocidura Wagler, 1832. It is represented by seven Type locality: West Haputale Estate, Ohiya, C Province, Sri Lanka species in the region. Synonyms: Suncus fellowes-gordoni Phillips, 1932 Podihik kura Deraniyagala, 1958 31. Suncus dayi (Dobson, 1888) Subspecies: None. 1888. Crocidura dayi Dobson, Ann. Mag. Nat. Hist., [6]1: 428. Distribution: Endemic to Sri Lanka Name: Day’s Shrew Comments: Ellerman and Morrison-Scott (1951) treated this

1368 February 2004 Zoos' Print Journal 19(2): 1361-1371 Checklist of insectivores (Mammalia: Insectivora) of South Asia S. Chakraborty et al. taxon as a subspecies of Suncus etruscus (Savi, 1822). Corbet Crocidura (Pachyura) waldemarii Peters, 1870 (1978) and Corbet and Hill (1991, 1992) synonymized Suncus Crocidura (Pachyura) ceylanica Peters, 1870 fellowes-gordoni Phillips, 1932 and Podihik kura Deraniyagala, Crocidura (Pachyura) media Peters, 1870 1958 with Suncus etruscus (Savi, 1832). Hutterer (1993) opines Crocidura (Pachyura) fulvocinerea Anderson, 1877 that the Podihik kura is more similar to Suncus etruscus (Savi, Suncus murinus fulvocinereus (Anderson, 1877) 1822) than to Suncus fellowes-gordoni Phillips, 1932. Hutterer Crocidura (Pachyura) blythii Anderson, 1877 (1993) and IUCN (1995) treated Suncus fellowes-gordoni Phillips, Crocidura (Pachyura) sindensis Anderson, 1877 1932 as a distinct species. Crocidura (Pachyura) pealana Anderson, 1877 Crocidura (Pachyura) blanfordii Anderson, 1877 34. Suncus montanus (Kelaart, 1850) Suncus murinus blanfordii (Anderson, 1877) 1850. Sorex montanus Kelaart, J. Ceylon Branch Asiat. Soc., 2: 211. Crocidura (Pachyura) rubicunda Anderson, 1877 Name: Hill Shrew Crocidura andersoni Trouessart, 1879 Type locality: Pidurutalagala, Nuwara Eliya, Sri Lanka Crocidura beddomei Anderson, 1881 Synonyms: Sorex montanus Kelaart, 1850 Sorex beddomei (Anderson, 1881) Suncus murinus montanus (Kelaart, 1850) Subspecies: Suncus murinus murinus (Linnaeus, 1766) Sorex feruginea Kelaart, 1850 Suncus murinus caerulescens Shaw, 1800 Sorex ferugeneus Kelaart, 1850 Suncus murinus sindensis Anderson, 1877 ? Sorex niger Horsfield, 1851 Suncus murinus nemorivagus Hodgson, 1845 Suncus niger malabaricus Lindsay, 1929 Suncus murinus viridescens Blyth, 1859 Subspecies: None. Distribution: Andhra Pradesh, Assam, Bihar, Chattisgarh, Distribution: Endemic to the region. Karnataka, Kerala and Gujarat, Harayan, Himachal Pradesh, Jharkhand, Karnataka, Tamil Nadu in India; Sri Lanka. Kerala, Maharashtra, Manipur, Meghalaya, Mizoram, Orissa, Comments: Lindsay (1929) described darker forms akin to Pondicherry, Punjab, Rajasthan, Sikkim, Tamil Nadu, Tripura, this species from southern India as Suncus niger malabaricus Uttaranchal, Uttar Pradesh and West Bengal in India; Pakistan; that has been considered as possible subspecies of the Suncus Bangladesh; Nepal; probably in Bhutan. murinus (Linnaeus, 1766) by Ellerman and Morrison-Scott Comments: Ellerman and Morrison-Scott (1951) listed more (1951). However, Corbet (1992) proposed it to be a valid than 15 subspecies under this taxon, including Sri Lankan forms subspecies of Suncus montanus (Kelaart, 1850) and accepted Suncus montanus Kelaart, 1850 and Suncus zeylanicus Phillips, by Corbet and Hill, 1992, Hutterer, 1993 and IUCN, 1995. 1928 which are now treated as distinct species (Phillips, 1980; Corbet & Hill, 1992; Hutterer, 1993). Phillips (1980) treated Sorex 35. Suncus murinus (Linnaeus, 1766) murinus kandianus (Kelaart, 1852) as a valid subspecies from 1766. Sorex murinus Linnaeus, Syst. Nat. 12th ed., I: 74. Sri Lanka. This species has been synonymized with Name: House Shrew Suncus murinus (Linnaeus, 1766) by Corbet and Hill (1992). Type locality: Java, Indonesia According to Corbet and Hill (1992) Suncus murinus Synonyms: Sorex murinus Linnaeus, 1766 nemorivagus Hodgson, 1845 possibly includes Suncus murinus Sorex indicus Geoffroy, I., 1811 soccatus (Hodgson, 1845), Suncus murinus griffithi (Horsfield, Sorex sonerratii Geoffroy, I., 1827 1851), Suncus murinus saturatior (Hodgson, 1855) and Suncus Sorex giganteus Geoffroy, I., 1831 murinus tytleri (Blyth, 1859); and Suncus murinus Sorex serpentarius Geoffroy, I., 1831 viridescens Blyth, 1859 possibly includes Crocidura Sorex nemorivagus Hodgson, 1845 (Pachyura) blanfordii Anderson, 1877. Sorex soccatus Hodgson, 1845 Suncus murinus soccatus (Hodgson, 1845) 36. Suncus stoliczkanus (Anderson, 1877) Sorex griffithi Horsfield, 1851 1877. Crocidura (Pachuyra) stoliczkanus Anderson, J. Asiat. Suncus murinus griffithi (Horsfiled, 1851) Soc. Bengal, 46: 270. Sorex kandianus Kelaart, 1852 Name: Anderson’s Shrew Suncus murinus kandianus (Kelaart, 1852) Type locality: Bombay, Maharashtra, India Sorex heterodon Blyth, 1855 Synonyms: Crocidura (Pachuyra) stoliczkanus Anderson, 1877 Sorex kelaarti Blyth, 1855 Crocidura (Pachuyra) subfulva Anderson, 1877 Sorex saturatior Hodgson, 1855 Crocidura (Pachuyra) bidiana Anderson, 1877 Suncus murinus saturatior (Hodgson, 1855) Crocidura leucogenys Dobson, 1888 Sorex viridiscens Blyth, 1859 Suncus stoliczkanus leucogenys (Dobson, 1888) Sorex tytleri Blyth, 1859 Subspecies: None for the region Suncus murinus tytleri (Blyth, 1859) Distribution: Endemic to the region. Gujarat, Madhya

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Pradesh, Maharashtra, Uttar Pradesh, Rajasthan and Tamil Nadu Priya and Ms. Padma Priya, Zoo Outreach Organization, for in India; Pakistan. their help in literature survey and preparation of the final Comments: Ellerman and Morrison-Scott (1951) listed three checklist; and also acknowledge the individual Research Grant subspeceis under this taxon, that were synonymized by Corbet from CSIR, New Delhi. and Hill (1992), and Hutterer (1993). References 37. Suncus zeylanicus Phillips, 1928 Abe, H., S. Shiraishi and S. Arai (1991). A new mole from Uotsuri- 1928. Suncus zeylanicus Phillips, Spolia Zeylan., 14: 313. jima, the Ryukyu Islands. Journal of the Mammalogical Society of Name: Ceylon Jungle Shrew, Sri Lankan Shrew 15: 47-60. Type locality: Gonagama Estate, Kitulgala, Sri Lanka Abe, H. (1971). Small of Central Nepal. Journal of the Synonyms: Suncus murinus zeylanicus Phillips, 1928 Faculty of Agriculture, Hokkaido University, Japan 56: 367-423. Subspecies: None. Abe, H. (1982). Ecological distribution and faunal structure of small mammals in central Nepal. Mammalia 46: 477-503. Distribution: Endemic to Sri Lanka. Agrawal, V.C. (1973). A note on the taxonomic status of Erinaceus Comments: Ellerman and Morrison-Scott (1951) included it blanfordi Anderson and Erinaceus jerdoni Anderson (Mammalia: under Suncus murinus (Linnaeus, 1766), but it as been treated Insectivora). Journal of the Zoological Society of India 25: 158-159. as distinct species basing on major morphological differences Agrawal, V.C. and S. Chakraborty (1971). Notes on a collection of by Phillips ( 1980) and McKay (1984), a trend followed by Corbet small mammals from Nepal, with the description of a new mouse- and Hill (1992), Hutterer (1993), and IUCN (1995). hare (Lagomorpha: Ochotonidae). Proceedings of the Zoological Society of Calcutta 24: 41-46. Genus Feroculus Kelaart, 1850 Alfred, J.R.B., N.K. Sinha and S. Chakraborty (2002). Checklist Long-clawed Shrew of Mammals of India. Records of the Zoological Survey of India, Occasional Paper 199: 1-289. Zoological Survey of India, Kolkata. This genus belongs to subfamily Crocidurinae Milne-Edwards, Allen, G.M. (1938). The mammals of China and Mongolia. In: Granger, 1872. It is represented by an endemic species in the region. W. (editor). Natural History of Central Asia. Central Asiatic Expeditions of the American Museum of Natural History, New York, 38. Feroculus feroculus (Kelaart, 1850) 11, Part 1: 1-620. 1850. Sorex feroculus Kelaart, J. Ceylon Branch Asiat. Soc., 2(5): 211. Biswas, B. and R.K. Ghose (1970). Taxonomic notes on the Indian Name: Kelaart’s Long-clawed Shrew pale hedgehogs of the genus Paraechinus Trouessart, with descriptions Type locality: Central Mountains at ca.828m, Nuwara Eliya, of a new species and subspecies. Mammalia 36: 467-477. Sri Lanka Blanford, W.T. (1888). The Fauna of British India. Mammalia, Part Synonyms: Sorex feroculus Kelaart, 1850 1. Taylor and Francis, London, 250pp+xii. Blyth, E. (1863). Catalogue of the Mammalia in the Museum of Asiatic Sorex macropus Blyth, 1851 Society. Calcutta. 187+xiii pp. Sorex nuwara-ellia Kelaart, 1851 Butler, P.M. (1988). Phylogeny of the Insectivores. pp. 117-141. In: Sorex newara Wagner, 1855 M.J. Benton (editor). The Phylogeny and Classification of the Crocidura macropus Blyth, 1888 Tetrapods, 2 (Mammals). Clarendon Press, Oxford. Subspecies: None for the region. Cabrera, A. (1925). Genera mammalium: Insectivora, Galaeopithecia. Distribution: Endemic to South Asia. Kerala and Tamil Nadu Museo Nacional de Ciencias Naturales, Madrid, 232pp. in India; Sri Lanka Catzeflis, F.M., T. Maddalena, S. Hellwing, and P. Vogel (1985). Comments: For details refer Phillips (1980). Corbet and Hill Unexpected findings on the taxonomic status of East Mediterranean (1992) indicated that there is an old unconfirmed report of this Crocidura russula auct. (Mammalia, Insectivora). Zeitschrift für Säugetierkunde 50: 185-201. taxon from Palni Hills, Tamil Nadu in India. However, Pradhan Chakraborty, S. (1975). On a collection of mammals from Bhutan. et al. (1997) have confirmed its occurrence and distribution in Records of the Zoological Survey of India 68: 1-20. southern India. Chakraborty, S. (1983). Contribution to the knowledge of the mammalian fauna of Jammu and Kashmir, India. Records of the Acknowledgements Zoological Survey of India 38: 1-129. We thank Dr. J.R.B. Alfred, Director, Zoological Survey of India, Corbet, G.B. (1978). The Mammals of the Palaearctic Region: A Kolkatta and Officer-in-Charge, Zoological Survey of India, Taxonomic Review. British Museum (Natural History), London, 314pp. Corbet, G.B. (1988). The family Erinaceidae: a synthesis of its Western Regional Station, Pune for facilities; Ms. Sally Walker , phylogeny, ecology and zoogeography. Review and Mr. Sanjay Molur, Zoo Outreach Organization, Coimbatore 18: 117-172. helped us in many ways; Dr. Mike J.R. Jordan, Chester Zoo, Corbet, G.B. and J.E. Hill (1980). A World List of Mammalian England helped us with initial list. MSP wishes to thank Mr. Species. British Museum (Natural History), London, 226pp. S.S. Talmale from ZSI, WRS, Pune, for his help. CS and BS Corbet, G.B. and J.E. Hill (1986). A World List of Mammalian thank Dr. V. Nagulu and the Head, Department of Zoology, Species. Second ed. British Museum (Natural History), London, 254pp. Osmania University, Hyderabad for encouragement; Ms. Binu Corbet, G.B. and J.E. Hill (1991). A World List of Mammalian

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Species. 3rd edition. British Museum (Natural History) Publications, Department (Wildlife Wing) and Kerala Agriculture University, 90+ London, 243pp. xxv pp. Corbet, G.B. and J.E. Hill (1992). Mammals of the Indomalayan Niethammer, J. (1973). Zur Kenntnis der Igel (Erinaceidae) Region: A Systematic Review. Oxford University Press, Oxford, 488pp. Afghanistans. Zeitschrift für Säugetierkunde 38: 271-276 [Original Dolgov, V.A. and R.S. Hoffmann (1977). Tibetskaya burozubka - not seen]. Sorex thibetanus Kastchenko, 1905 (Soricidae, Mammalia). Zoological Nowak, R.M. (1999). Walker’s Mammals of the World. 6th ed. Vol. I Zhurnal 46: 1687-1692. [In Russian, Original not seen] and II. The John Hopkins University Press, Baltimore and London, Eisenberg, J.F. and G.M. McKay (1970). An annotated checklist of Vol. I: lxx+1-836 and Vol. II: xii+837-1936. the recent mammals of Ceylon with keys to the species. Ceylon Journal Osgood, W.H. (1932). 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