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我mmonoidsinja腿n 一stratigralphic Dlistribution and Bib且iogra］Phy一

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我mmonoidsinja腿n 一stratigralphic Dlistribution and Bib且iogra］Phy一 Bulletin of the Geological Survey of Japan，vol．51（ll），p． 559－613 2000 ， ：Notes an且Comments Da流abase of’the Cretaceous 我mmonoidsinJa腿n 一stratigralphic dlistribution and bib且iogra］phy一 Seiichi TosHIMITsul and Hiromichi HIRANo2 Seiichi TosHIMITsu and Hiromichi HIRANo（2000）Database of the Cretaceous ammonoids in Japan －stratigraphic distribution and bibliography一．Bκ1乙 G60乙 Sπ鰍ゆ硯v・1．51（ll），P．559－613，1 fig，19 tables． Abstract：A database with a bibliography on the Japanese Cretaceous ammonoids has been made by the survey of various literature（Tables2－19）．This database consists of a list of species，based on higher taxonomy of Wright8砲乙（1996），with their stratigraphic distribution data and a bibliog－ raphy on them．As a result，it clarified that790species are detected from the Japanese Cretaceous deposits，and the species－diversity of the Cretaceous ammonoids in Japan changed cyclically， although the diversity of ammonoids in・the Early Cretaceous is much lower than the Late Cretaceous．Diversity changes are controlled by environmental changes，and therefore this database will be useful not only for stratigraphical and biochronological studies but also for paleoenviron－ mental study of the Cretaceous Period． genera and65subgenera were recognised in Arke11α α乙（1967），while684genera and l53subgenera were 1． Introduction recognized in Wright6厩孟（1996）． Ammonoids are one of the most common fossil On the other hand，Matsumoto （1942e，1943， animals in the Late Paleozoic and Mesozoic Eras．A 1954a）1isted many ammonoid species，obtained from great many species of ammonoids have been reported Japanese Cretaceous deposits， by ：regional groups． until now，and it is said that the mmber of described Recently，Tanabeαα乙（2000）made a database of the ammonoid species in the world is greater than5，000， Cretaceous ammonoid specimens （about l500 in－ or more than IO，000．Moreover，many unknown species dividuals，including about200type specimens）from will be found from now onward．Ammonoids are Japan and Sakhalin housed in the University Museum we11－known to be useful for biostratigraphical zona－ of the University of Tokyo．However，a catalogue of tion and correlation，because of theiHemarkable evolu． all Japanese Cretaceous ammonoids was not made until tion and wide distribution．Besides，they are known to now．Therefore，we have surveyed records of all have been influenced by environmental changes，for Cretaceous ammonoid species from Japan from litera－ which the term bioseismometer was intruduced by ture in geology，stratigraphy and paleontology。As a House（1989）．We can expect to clarify the Cretaceous result，we made a list comprising about800am． environmental changes recorded in Japan and the monoid species with data of their stratigraphic distribu． Northwestem Pacific through the study of am－ tions（Tables2－19）． monoids（Hirano8オα乙，1999，2000）． A catalogue containing all Paleozoic and Mesozoic 2． Database of the Japanese Cret＆ceous ammonoid genera of the world，with their diagnosis， amm㎝oiαS was written by Arke11αα乙（1957）．However，there This database is made up of a list of all Japanese is no catalogue containing all ammonoid species of the Cretaceous ammonoid species with their stratigraphic Paleozoic and Mesozoic of the world．After Arke11α distribution data（Tables2－19）and a bibliography on α乙 （1957），many new genera were added through them．We follow Wright6オα乙（1996）for higher many studies on ammonoids．Wrightαα乙（1996） taxonomy in this（1atabase．Stratigraphic subdivision of revised only the Cretaceous ammonoid genera．As far the Japanese Cretaceous System is based on Matsu－ as the Cretaceous ammonoids are concemed，466 moto（1978c）and Matsumoto6渉α乙（1982）for the Lower Cretaceous，and Toshimitsuαα乙（1995a，b） Geological Museum，GSJ Department of Earth Sciences，School of Education， Keywords：database，Ammonoidea， Cretaceous， Japan， Waseda University，Tokyo，169－8050Japan stratigraphic disnribution，bibliography 一559一 Bκllεオ初げオhεGεoZog歪oαl S銘7η鯉げ」αメ》αη，Vb乙 51， ！〉α11， 2000 for the Upper Cretaceous．However，the subdivisions α脚no歪pointed out by Matsumoto （1980）， of the Lower Aptian，Upper Aptian and upper part of ‘‘y罐oッα耀068郷乞5h乞乏α測α！’by Maeda（1993），and Lower Campanian Substages are modified in this Po4y少砂oん06εrα5 メ》5εz6409αz61扉n麗ηz by Okamoto and paper．．For example，K3al and K3a20f the Lower Shibata （1997）．However，we treated the related Aptian are united into one substage（K3al－2） （Fig． species to them as independent species in this da．tabase。 1），because・fthedifficulty・fbi・stratigraphically There is also a problem on sexual dimorphism be－ higher resolution on this substage at the present．As tween the following two genera， M18肱舵5 and a result，the Japanese Cretaceous are subdivided into ／1nα〆》α6hツ4Z56z6～ discussed by Cobban and Kenne（iy 31substagesinthispaper（Fig。1）． （1993）．We treated these two genera as indep6ndent Our term“species”mean not only paleontologically from each other in this paper．On the other hand， described ones but also undescribed ones such as the biostratigrαphic problem w＆s recently pointed out ‘‘sp．nov．” and “aff．”．Both are reported in geology， by Naruse 6オα乙 （2000）。They reported that G側4 stratigraphy and paleontology journals and re1αted ぴ66rα5 hα刀zαn8ゑα6n36 and 。翫6hこy4乞56z45 ノγ6∬μ05μ5， magazines．Among these species，some species have whose ranges were previously regarded as late problems related to taxonomy and biostratigraphy。 Mαastrichtian，are early Maastrichtian species based The taxonomic problem is of synonyms on Sオol舵勉宛 on the stratigraphical study of the Cretaceous of South 」＆panese Stages＆Substages sc田e u． l l l l l l ： ： ： ： Up． K6b2 iii l ： ： ： ： L 1 ： 1 ： ： Maastrichtian u． l l l l ： Low。 K6b1 1 ： 1 ： 1 ： ： ： ： L ： ： ： ： ： ；＝…脳二…iii l ： u． l l l K6a4 I l Up。 1 ： ： 『 1 ： l L K6a3 I I ： Campanian ： ： u。 K6a1－2iii ・欄培 1 1 1 ！ 1 ： Low． 1 ： ＝＝i＝iGi＝i＝≡ ： L K5b2 ： の38題。δお鼠P ．，；3：鵬：i Santor直an K5b！ 9．＝＝…＝…：i；i ：＝i＝…c…；…＝… Up K5a2 ≡………ili…i…… …………………………… i Coniacian Mid。 …無……≡… ・掘：…：… 融1：1＝1 K5a1 ： 1『 Low． ＝：…：…‘…＝…＝≡ ： 1 礁韓：OAE3 繍…＝ 1……i…嫁 ii…iil…1…1… Up K4』b3 Turonian Mid。 K4b2 ……i…iii…i…i ・紳…＝…＝… ：：…＝1‘1：…＝1 LDW。 K4b1 騰 ……………臨 ： ： l ： ： u． K4a6 l 『 1 噸韓：OAE2 蕪華 Up． M． K4a5 ： 1 ： ： L K4a4 ： ； Mi（L K4a3 ii轟垂i 難ilil ［ ： u． …liiiiiiiii K4a2 －＝：’＝1＝＝＝＝＝ 1 ： Low， 1 K4a1 』．：：≡‘…＝…：… ．：：…＝…＝…：…： 蕪錨i ： L I I l Up。 K3b3 1…………i…………… 隅… 灘箋 一・雨：噸嚇：OAEld i Albian Mid． 1 ： ： ： 畢08題ohQお30■ K3b2 鴫ll l l l 鴫ρAElc …iiiiliiiii I ： ： ： Low． K3b1 磯： ： ： ： 1 ：磯：OAElb ＝＝i＝i‘i：i＝i ： ： ： Up K3a3－5ili 1 Aptian ： ： l l l Low． K3a1－2iii ： ： ： 1 囎： ： ： ： 囎：OAEla Up． …罫 i ： ： ： Barremian K2b ： ： ： i…i…ili…i…i ： ： ： ： Low。 ： ： ： ： ：：1：：：〕 Hauterivian K2a ： ： ： 1 ： ： ： ： ： ： ： Valan●nlan Klb ： ： ： ： Bcrriasian Kla ： 、 1 1 1 0 20 40 60 80 100 number ofspecies Fig，1．Changes of number of ammonoid species during the Cretaceous in Japan，Subdivision of substages are modified from Matsumoto（1978c），Matsumotoαα孟（1982）and Toshimitsu6厩乙（1995a，b）． 一560一 D伽伽6げオh6伽鰭・％5㈱脚n・跳乞n鋤齪（乃5h伽オ5観n4H加n・） Sakhalin by Maeda8厩乙（1995：0ral presentation）． A．K，Moore，R．C．，Schindewolf，0．H．，Sy1． Moreover，the lower biostratigraphic resolution of the vester－Bradley，P．C．and Wright，C．W．（1957） Lower Cretaceous System in Japan is another prob－ 丁紹αオ魏 on 乞n泥π66昭オ8勿160n6010gy，」％π五． 1em．Future studies are required to solve these prob． Mollz6s6α 4r G砂hαloρ04¢∠肋z刀zono弼6α GeoL lems． Soc。America，Inc．＆Univ．Kansas，490p． This database can clarify the diversity of the Cobban，W．A．and Kemedy，W．J．（1993）The Japanese Cretaceous ammonoids，which consist of four UpPer Cretaceous dimorphic pachydiscid am－ suborders，that is，Phyllocemtina，Lytoceratina，Am－ moniteハ46n％撚in the Westem Interior of the monitina and Ancyloceratina．The total number of United States．U S G80乙S祝澱，P7犀拓ρ．， species is790（Table l）．As preliminarily mentioned no。1533，1．14，14 pls． byHiran・6厩乙（1999，2000），speciesdiversity Hirano，H．，Toshimitsu，S．，Matsumoto，T。and （＝number of species）of the Japanese Cretaceous am， Takahashi，K．（1999）Bioevents and paleoen－ monoids is higher in the Lower Barremian，Upper Ap． vironmental changes in Mid－Cretaceous。Eo∬痂 tian to Lower Albian，Upper Albian to Lowermost 侮1α6・nよS・‘」砂αnク，no．66，47－49．［in Cenomanian，Middle to Upper Turonian，Middle Jαpanese］ Coniancian and Santonian Substages （Fig． 1）． Hirano，H．，Toshimitsu，S．，Matsumoto，T．and Contrastly，the minima occur in the Lower Aptian， Takahashi，K．（2000） Changes in Cret＆ceous Middle Albian，Upper Cenomanian，and Lower ammonoid diversity and marine environments of Coniacian Substages．As suggested by・Hirano66α乙 the Japanese Islands．Zn Okada，H．and Mateer， （1999，2000），the timing of these declines in species N．J．，eds．，α6嬬60％56π吻on耀n劣∫n∠45毎 diversity are comparatively harmonious with those of Developments in P＆1eontology and Stratig－ the Cretaceous Anoxic Events（OAE），except for the raphy，no．17，Elsevier，Amsterdam，145．154． Upper Albian Substage at the time of OAEld．Thus， Maeda，H．，Shigeta，Y．，Okamoto，T．，Hasegawa，T．， this database could be the useful source of informa． Kase，T．，Futakami，M．and Uemura．，K． tion to analyse the environmental changes an（玉 res－ （1995）Stratigraphy and fossil assemblages of ponse of the ammonoids． the Upper Cretaceous in the Makarov area， Sakhalin．ノ16鉱 1995／17zn。ハ48ε凌 ん伽80nム Soo Acknowle通gements：We express our sincere thanks J4ノ》αn，no．92，ll9．［in Japanese］ to Emeritus Professor Tatsuro Matsumoto of Kyushu Tanabe，K．，Ito，Y．，Moriya，K．and Sasaki，T．’ University for his valuable suggestions and com－ （2000） Database of Cretaceous ammonite ments．We wish to thank Dr．Fumihisa Kawabe of specimens registered in the Dep＆rtment of His－ W＆seda University，and Dr．Satoshi Nakae and Mr． torical Geology and Paleontology of the Univer－ Naotomo Kaneko
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