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Title a Revisional Study of the Spider Family Thomisidae (Arachnida A revisional study of the spider family Thomisidae (Arachnida, Title Araneae) of Japan( Dissertation_全文 ) Author(s) Ono, Hirotsugu Citation 京都大学 Issue Date 1988-01-23 URL https://doi.org/10.14989/doctor.r6388 Right Type Thesis or Dissertation Textversion author Kyoto University 学位 請 論 文 (主 論 文) 小 野 族 嗣 1灘 灘灘 灘轟 1 . Thomisidae aus Japan. I. Das Genus Tmarus Simon (Arachnida: Araneae). Acta arachnol., 27 (spec. no.): 61-84 (1977). 2 . Thomisidae aus Japan. II. Das Genus Oxytate L.Koch 1878 (Arachnida: Araneae). Senckenb. biol., 58: 245-251 (1978). 3 . Thomisidae aus dem Nepal-Himalaya. I. Das Genus Xysticus C.L.Koch 1835 (Arachnida: Araneae). Senckenb. biol., 59: 267-288 (1978). 4 . Thomisidae aus dem Nepal-Himalaya. II. Das Genus Lysiteles Simon 1895 (Arachnida: Araneae). Senckenb. biol., 60: 91-108 (1979). 5 . Fossile Spinnen aus miozanen Sedimenten des Randecker Maars in SW- Deutschland (Arachnida: Araneae). Jh. Ges. Naturkde. Wurttemberg, 134: 133-141 (1979). (W.Schawaller t 4E1t) 6 . Thomisidae aus Japan. III. Das Genus Lysiteles Simon 1895 (Arachnida: Araneae). Senckenb. biol., 60: 203-217 (1980). 7 . Thomisidae aus dem Nepal-Himalaya. III. Das Genus Stiphropus Gerstaecker 1873, mit Revision der asiatischen Arten (Arachnida: Araneae). Senckenb. biol., 61: 57-76 (1980). 8 . Erstnachweis einer Krabbenspinne (Thomisidae) in dominikanischem Bernstein (Stuttgarter Bernsteinsammlung: Arachnida, Araneae). Stuttgart. Beitr. Naturk., B, (73): 1-13 (1981). 9 . Revision japanischer Spinnen. I. Synonymieeiniger Arten der Familien Theridiidae, Araneidae, Tetragnathidae and Agelenidae (Arachnida: Araneae). Acta arachnol., 30: 1-7 (1981). 10 . Verwandtschaft von Tetrablemma phulchoki Lehtinen 1981 (Araneae: Tetrablemmidae). Senckenb. biol., 62: 349-353 (1982). (1) 11 • Zodariidae aus dem Nepal-Himalaya. I. Neue Arten der Gattung Storena Walckenaer 1805 (Arachnida: Araneae). Senckenb. biol., 63: 211- 217 (1983). 12 Eine neue japanische Synaema-Art (Araneae, Thomisidae). Acta arachnol., 31: 59-63 (1983). 13 • The Thomisidae of Japan. IV. Boliscus Thorell, 1891 (Arachnida, Araneae), a genus new to the Japanese fauna. Bull. natn. Sci. Mus., Tokyo, (A), 10: 63-71 (1984). 14 • Revision einiger Arten der Familie Thomisidae (Arachnida, Araneae) aus Japan. Bull. natn. Sci. Mus., Tokyo, (A), 11: 19-39 (1985). 15 • The Thomisidae of Japan. V. Monaeses Thorell, 1869, and its new junior synonym, Mecostrabus Simon, 1903 (Arachnida, Araneae). Bull. natn. Sci. Mus., Tokyo, (A), 11: 91-97 (1985). 16 • Eine neue Art der Gattung Misumenops F.O.Pickard-Cambridge, 1900, aus Japan (Araneae: Thomisidae). Proc. Jpn. Soc. syst. 'tool., Tokyo, (31): 14-19 (1985). 17 • A new spider of the group of Clubiona corticalis (Araneae, Clubionidae) found in Japan. Ent. Pap. pres. Kurosawa, Tokyo, pp.19-25 (1986). 18 • A new Sino-Japanese species of the genus Cupa (Araneae, Thomisidae) from the coastal areas of the East China Sea. Bull. natn. Sci. Mus., Tokyo, (A), 12: 25-29 (1986). (D.-X. Song k) 19 Little-known Japanese spider, Clubiona zilla (Araneae, Clubionidae)-- Representative of a new and peculiar species-group. Bull. natn. Sci. Mus., Tokyo, (A), 12: 117-121 (1986). 20 • Spiders of the families Clubionidae, Gnaphosidae and Thomisidae from the Noto Peninsula and the southern part of Ishikawa Prefecture, Japan. Mem. natn. Sci. Mus., Tokyo, (19): 167-174 (1986). (2) 21 • Twonew species of the family Zodariidae from Saudi Arabia (Arachnida: Araneae). Fauna Saudi Arabia, 8: 6-11 (1986). (R.Jocque ti 1l n) 22 • A new Japanese castianeirine genus (Araneae, Clubionidae) with presumptive prototype of salticoid eyes. Bull. natn. Sci. Mus., Tokyo, (A), 13: 13-19 (1987). (3) A Revisional Study of the Spider Family Thomisidae (Arachnida, Araneae) of Japan By Hirotsuau Ono Department of Zoology, National Science Museum, Tokyo Contents Introduction ............................................. • • • 1 Acknowledgement ..................................... • • • • 2 A Brief History of the Study in Japanese Spiders ......... • • 4 Material and Methods ..................................... • • 10 Type Depository of the Thomisid Species Recorded from Japan • • • • • . 12 Terminology .............................................. • • • 17 Abbreviations ....................................... • • 20 Taxonomy: Family Thomisidae Sundevall, 1833 ............. • • • • 21 Subfamily Stephanopinae 0. Pickard-Cambridge, 1871 .. • • • • • 27 Genus Cupa Strand, 1906 ......................... • • • • • 30 Genus Phrynarachne Thorell, 1869 ................ • 38 Subfamily Dietinae Simon, 1895 ...................... • • • • • • 50 Genus Oxytate L. Koch, 1878 ..................... • • Genus Loxobates Thorell, 1877 ................... Genus Alcimochthes Simon, 1885 .................. • Subfamily Thomisinae Sundevall, 1833 ................ • Genus Tmarus Simon, 1875 ........................ • Genus Monaeses Thorell, 1869 .................... • Genus Coriarachne Thorell, 1870 ................. • • Genus Bassaniana Strand, 1928 ................... • • Genus Xysticus C.L. Koch, 1835 .................. Genus Oxyptila Simon, 1864 ...................... • • • -i- • Genus Lysiteles Simon, 1895 ............. • • • • • • • • • 220 Genus Synaema Simon, 1864 ............... • • • • • • • • 241 Genus Takachihoa Ono, 1985 .............. • • • • • • • • 258 Genus Misumenops F.O. Pickard-Cambridge, 1900 ... • • • • • • • • . 264 Genus Diaea Thorell, 1869 ............... • • • • • • • • • • 289 Genus Heriaeus Simon, 1875 .............. • • • • • • • • • 295 Genus Misumena Latreille, 1804 .......... • • • • • • • • 303 Genus Pistius Simon, 1875 ............... • • • • • • 311 Genus Runcinia Simon, 1875 .............. • • • • • • • • 319 Genus Thomisus Walckenaer, 1805 ......... • • • • • • 332 Subfamily Bominae Simon, 1886 ............... • • • • • • • • • 351 Genus Boliscus Thorell, 1891 ............ • • • • • • • • • • 353 Species Incertae Sedis ...................... • • • •• • •• • • • • • • • • • • • • 359 Doubtful Records, Misplaced and Misidentified Species • • • • • • • • • 361 Nomina Nuda ................................. • • • • • • • • • • • 365 List of the Taxa Described in this Paper .... • • • • • • •• • •• • •• ^• • • • • • • • • • 366 Higher Classification and Phylogeny .............. • • • • • • • • • • 370 Zoogeographical Notes ............................ • • • • ^• • • • • • . 389 References ....................................... • • . • • • •. • • • • . • . 399 Explanation of Figures ........................... 447 -ii- Introduction The purpose of this study is to present the natural history of thomisid spiders in Japan by investigating through comparative morphology, taxonomy and partly zoogeography. In order to pursue this subject, it was necessary first to read through literature referring to the thomisid spiders not only from Japan and her adjacent regions but from all parts of the world, because very little has been made generally on the family Thomisidae in the sense of modern biological sciences. Since L. Koch (1878c) described the first thomisid spider from Japan, more than 70 specific and subspecific names excluding philodromids have been reported from this country (details are given in a later chapter). However, some records were based on doubtful identification or inappropriate naming; some species were placed in wrong taxonomic positions and many have never been recognized since their original descriptions. Re-examination of the original specimens of most species described from Japan and her adjacent countries became inevitable. I spent several years for studying spiders at the Zoological Institute of the University of Mainz in Germany. During my stay in Europe, I visited other institutions as well, and communicated with curators of many museums for examining those old collections on loan. Other than the type specimens, general and comparative material used in this study consists of the specimens personally collected by myself from various places in Japan since 1966 and those -1- obtained from or loaned from several institutions, friends and colleagues. Total number of these specimens may exceed 3,000. A large quantity of them are deposited in the collection of the Department of Zoology, National Science Museum (Nat. Hist.), Tokyo, where I have prepared this paper. Based upon the preliminary research of records hitherto published and actual examination of material, each taxon of the family Thomisidae in Japan will be systematically revised and diagnosed in this paper, that is, 4 subfamilies, 22 genera and 53 species including 6 new ones. Then, the peculiarity of the thomisid fauna of Japan is delineated and the relationship of the Japanese fauna will be discussed in comparison with the world fauna, especially Eurasian, to which attaches the Japanese one. The relationship among subfamilies of the Thomisidae will be phylogenetically analysed. Acknowledgement I wish to express my sincere thanks to Professor Dr. Toshitaka Hidaka, Zoological Institute, Kyoto University, for giving me the opportunity to present this thesis, to Dr. Shun-Ichi Ueno, Department of Zoology, National Science Museum (Nat. Hist.), Tokyo, for his generous support in the planning and realization of this study and for revising the manuscript of this paper, to Professor Dr. Jochen Martens, Institut fur Zoologie, Johannes Gutenberg-Universitat, Mainz, -2- for his guidance in various ways, and to Professor Dr. Takeo Yaginuma, Biological Laboratory, Ohtemon-Gakuin University, Osaka, for giving me useful advice in arachnology
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