Isla Del Coco National Park Species Lists*

Isla del Coco National Park Species Lists* Updated 22 July 2013

Fishes

SUPERCLASS PISCES CLASS CHONDRICHTHYES SUBCLASS ELASMOBRANCHII

ORDER Squaliformes Echinorhinidae — Bramble sharks Echinorhinus cookei Pietschmann, 1928

ORDER Orectolobiformes Rhincodontidae — Whale shark Rhincodon typus Smith, 1828

ORDER Lamniformes Alopiidae — Thresher sharks Alopias vulpinus (Bonnaterre, 1788) Odontaspididae - Sand tiger sharks Odontaspis ferox (Risso, 1810)

ORDER Carcharhiniformes Carcharhinidae — Requiem sharks Carcharhinus albimarginatus (Rüppell, 1837) Carcharhinus falciformis (Müller & Henle, 1839) Carcharhinus galapagensis (Snodgrass & Heller, 1905) Carcharhinus limbatus (Müller & Henle, 1839) Galeocerdo cuvier (Peron & LeSueur in: LeSueur, 1822) Rhizoprionodon longurio (Jordan & Gilbert, 1882) Triaenodon obesus (Rüppell, 1837) Sphyrnidae — Hammerhead sharks Sphyrna lewini (Griffith & Smith, 1834) Sphyrna mokarran (Rüppel, 1837)

ORDER Rajiformes Rhinobatidae — Guitarfishes Rhinobatos planiceps Garman, 1880

ORDER Myliobatiformes Dasyatidae — Whiptail stingrays Taeniura meyeni Müller & Henle, 1841 Myliobatidae — Eagle rays Aetobatus narinari (Euphrasen, 1790) Rhinopteridae — Cow-nosed rays Rhinoptera steindachneri Evermann & Jenkins, 1892 Mobulidae — Mantas, devil rays and mobulas Manta birostris (Walbaum, 1792) Mobula munkiana Notarbartolo-di-Sciara, 1987 Mobula tarapacana (Philippi, 1893)

CLASS ACTINOPTERYGII

ORDER Anguilliformes Myrocongridae — Conger-morays Myroconger nigrodentatus Castle and Bearez, 1995 Muraenidae — Morays Anarchias galapagensis (Seale, 1940) Echidna nebulosa (Ahl, 1789) Echidna nocturna (Cope, 1872) Enchelycore octaviana (Myers & Wade, 1941) Gymnomuraena zebra (Shaw in: Shaw & Nodder, 1797) Gymnothorax angusticeps (Hildebrand and Barton, 1949) Gymnothorax buroensis (Bleeker, 1857) Gymnothorax castaneus (Jordan & Gilbert, 1883) Gymnothorax dovii (Günther, 1870) Gymnothorax flavimarginatus (Rüppell, 1830) Gymnothorax javanicus (Bleeker, 1859) Gymnothorax meleagris (Shaw in: Shaw & Nodder, 1795) Gymnothorax panamensis (Steindachner, 1876) Gymnothorax pictus (Ahl, 1789) Muraena argus (Steindachner, 1870) Muraena lentiginosa Jenyns, 1842 Scuticaria tigrina (Lesson, 1828) Uropterygius macrocephalus (Bleeker, 1865) Uropterygius versutus Bussing, 1991 Congridae — Conger and garden eels Ariosoma gilberti (Ogilby, 1898) Heteroconger klausewitzi (Eibl-Eibesfeldt & Köster, 1983) Paraconger californiensis Kanazawa, 1961 Ophichthidae — Snake eels Bascanichthys bascanoides Osburn & Nichols, 1916 Callechelys eristigma McCosker & Rosenblatt, 1972 Gordichthys cf. combibus Lavenberg, ms Herpetoichthys fossatus (Myers & Wade, 1941) Ichthyapus selachops (Jordan & Gilbert, 1882) Myrichthys aspetocheiros McCosker & Rosenblatt, 1993 Myrichthys tigrinus Girard, 1859 Ophichthus rugifer Jordan & Bollman, 1890 Paraletharchus pacificus (Osburn & Nichols, 1916) Quassiremus evionthas (Jordan & Bollman, 1890) Quassiremus nothochir (Gilbert, 1890)

ORDER Gonorynchiformes Chanidae — Milkfishes Chanos chanos (Forsskål, 1775)

ORDER Aulopiformes Aulopidae — Flagfins Aulopus bajacali Parin & Kotlyar, 1984 Chlorophthalmidae — Greeneyes Chlorophthalmus mento Garman, 1899 Synodontidae — Lizardfishes Synodus lacertinus Gilbert, 1890

ORDER Ophidiiformes Ophidiidae — Cusk-eels Brotula ordwayi Hildebrand & Barton, 1949 Otophidium indefatigabile Jordan & Bollman, 1890 Petrotyx hopkinsi Heller & Snodgrass, 1903 Carapidae — Pearlfishes Carapus dubius (Putnam, 1874) Echiodon exsilium Rosenblatt, 1961 Encheliophis vermicularis Müller, 1842 Bythitidae — Viviparous brotulas ∑Ogilbia cocoensis Möller, Schwarzhans & Nielsen, 2005

ORDER Gadiformes Moridae - Codlings (Moras) Physiculus sp.

ORDER Lophiiformes Lophiidae — Goosefishes Lophiodes caulinaris (Garman, 1899) Antennariidae — Frogfishes Antennarius avalonis Jordan & Starks, 1907 Antennarius coccineus (Lesson, 1831) Antennarius commerson (Latreille, 1804) Antennarius sanguineus Gill, 1863 Antennatus strigatus (Gill, 1863) Ogcocephalidae — Batfishes ∑Ogcocephalus porrectus Garman, 1899 Zalieutes elater (Jordan & Gilbert, 1882)

ORDER Mugiliformes Mugilidae — Mullets Agonostomus monticola (Bancroft, 1834) Chaenomugil proboscideus (Günther, 1861) Mugil curema Valenciennes in: Cuvier & Valenciennes, 1836

ORDER Atheriniformes Atherinopsidae — New World silversides Atherinella eriarcha Jordan & Gilbert, 1882 Melanorhinus cyanellus (Meek & Hildebrand, 1923)

ORDER Beloniformes Exocoetidae — Flyingfishes Exocoetus monocirrhus Richardson, 1846 Exocoetus volitans Linnaeus, 1758 Hirundichthys speculiger (Valenciennes, 1847) Prognichthys sealei Abe, 1965 Prognichthys tringa Breder, 1928 Hemiramphidae — Halfbeaks Euleptorhamphus viridis (Van Hasselt, 1823) Hyporhamphus rosae (Jordan & Gilbert, 1880) Oxyporhamphus m. micropterus (Valenciennes in: Cuvier & Valenciennes, 1847)

Belonidae — Needlefishes Platybelone argalus pterura (Osburn & Nichols, 1916) Strongylura exilis (Girard, 1854) Tylosurus acus melanotus (Bleeker, 1851) Tylosurus crocodilus fodiator Jordan & Gilbert, 1882

ORDER Ateleopodiformes Ateleopodidae — Jelly nose fishes Guentherus altivela Osório, 1917

ORDER Beryciformes Holocentridae — Soldierfishes and squirrelfishes Myripristis berndti Jordan & Evermann, 1903 Myripristis leiognathus Valenciennes, 1846 Plectrypops lima (Valenciennes in: Cuvier & Valenciennes, 1831) Sargocentron suborbitalis (Gill, 1863)

ORDER Gasterosteiformes Aulostomidae — Trumpetfishes Aulostomus chinensis (Linnaeus, 1766) Fistulariidae — Cornetfishes Fistularia commersonii Rüppell, 1838 Syngnathidae — Pipefishes and seahorses Bryx veleronis Herald, 1940 Doryrhamphus excisus excisus Kaup, 1856 Hippocampus ingens Girard, 1858

ORDER Scorpaeniformes Scorpaenidae — Scorpionfishes Pontinus clemensi Fitch, 1955 Pontinus strigatus Heller & Snodgrass, 1903 Scorpaena afuerae Hildebrand, 1946 ∑Scorpaena cocosensis Motomura, 2004 Scorpaena histrio Jenyns, 1840 Scorpaena mystes Jordan & Starks in: Jordan, 1895 Scorpaena russula Jordan & Bollman, 1890 DEEP Scorpaenodes rubrivinctus Poss, McCosker & Baldwin, 2010 Scorpaenodes xyris (Jordan & Gilbert, 1882) Triglidae — Searobins Bellator farrago Richards & McCosker, 1998

Peristediidae — Armored searobins ∑Peristedion nesium Bussing, 2010

ORDER Perciformes Serranidae — Groupers and sea basses Alphestes immaculatus Breder, 1936 Cephalopholis panamensis (Steindachner, 1877) Dermatolepis dermatolepis (Boulenger, 1895) Epinephelus cifuentesi Lavenberg & Grove in: Heemstra & Randall, 1993 Epinephelus itajara (Lichtenstein, 1822) Epinephelus labriformis (Jenyns, 1840) Epinephelus mystacinus (Poey, 1852) Epinephelus niphobles Gilbert & Starks in: Gilbert, 1897 Liopropoma fasciatum Bussing, 1980 Mycteroperca olfax (Jenyns, 1840) Paranthias colonus (Valenciennes, 1846) Pronotogrammus eos Gilbert, 1890 Pronotogrammus multifasciatus Gill, 1863 Pseudogramma thaumasium (Gilbert in: Jordan & Evermann, 1900) Rypticus bicolor Valenciennes, 1846 Serranus aequidens Gilbert, 1890 ∑Serranus tico Meisler & Lavenberg in: Allen & Robertson, 1998 Kuhliidae — Flagtails Kuhlia mugil (Forster in: Bloch & Schneider, 1801) Priacanthidae — Bigeyes Cookeolus japonicus (Cuvier in: Cuvier & Valenciennes, 1829) Heteropriacanthus cruentatus (Lacépède, 1801) Pristigenys serrula (Gilbert, 1891) Apogonidae — Cardinalfishes Apogon atradorsatus Heller & Snodgrass, 1903 Apogon dovii Günther, 1861 Malacanthidae — Tilefishes Caulolatilus affinis Gill, 1865 Caulolatilus hubbsi Dooley, 1978 Malacanthus brevirostris Guichenot, 1848 Echeneidae — Suckerfishes, remoras Echeneis naucrates Linnaeus, 1758 Remora australis (Bennett, 1840) Remora brachyptera (Lowe, 1839) Remora osteochir (Cuvier, 1829) Remora remora (Linnaeus, 1758) Remorina albescens (Temminck & Schegel, 1850) Carangidae — Jacks and pompanos Alectis ciliaris (Bloch, 1787) Carangoides orthogrammus (Jordan & Gilbert, 1882) Caranx caballus Günther, 1868 Caranx caninus Günther, 1867 Caranx lugubris Poey, 1860 Caranx melampygus Cuvier in: Cuvier & Valenciennes, 1833 Caranx sexfasciatus Quoy & Gaimard, 1825 Decapterus macarellus (Cuvier in: Cuvier & Valenciennes, 1833) Elagatis bipinnulata (Quoy & Gaimard, 1825) Gnathanodon speciosus (Forsskål, 1775) Naucrates ductor (Linnaeus, 1758) Selar crumenophthalmus (Bloch, 1793) Seriola rivoliana Valenciennes in: Cuvier & Valenciennes, 1833 Trachinotus rhodopus Gill, 1863 Trachinotus stilbe (Jordan & McGregor, 1898) Uraspis helvola (Forster, 1801) Coryphaenidae — Dolphinfishes Coryphaena equiselis Linnaeus, 1758 Coryphaena hippurus Linnaeus, 1758 Lutjanidae — Snappers Aphareus furca (Lacépède, 1801) Hoplopagrus guntheri Gill, 1862 Lutjanus aratus (Günther, 1864) Lutjanus argentiventris (Peters, 1869) Lutjanus guttatus (Steindachner, 1869) Lutjanus inermis (Peters, 1869) Lutjanus jordani (Gilbert in: Jordan & Evermann, 1898) Lutjanus novemfasciatus Gill, 1862 Lutjanus viridis (Valenciennes, 1846) Gerreidae — Mojarras Eucinostomus currani Zahuranec in: Yáñez-Arancibia, 1980 Lobotidae — Tripletails Lobotes pacificus Gilbert, 1898 Haemulidae — Grunts Anisotremus caesius (Jordan & Gilbert, 1882) Anisotremus interruptus (Gill, 1862) Anisotremus taeniatus Gill, 1861 Sciaenidae — Drums Odontoscion eurymesops (Heller & Snodgrass, 1903) Umbrina xanti Gill, 1862 Mullidae — Goatfishes Mulloidichthys dentatus (Gill, 1862) Kyphosidae — Sea chubs Kyphosus elegans (Peters, 1869) Sectator ocyurus (Jordan & Gilbert, 1882) Chaetodontidae — Butterflyfishes Chaetodon humeralis Günther, 1860 Chaetodon lunula (Lacépède, 1803) Johnrandallia nigrirostris (Gill, 1862) Prognathodes carlhubbsi (Nablant, 1995) Pomacanthidae — Angelfishes Holacanthus passer Valenciennes, 1846 Pomacanthus zonipectus (Gill, 1862) Pomacentridae — Damselfishes Abudefduf concolor (Gill, 1862) Abudefduf troschelii (Gill, 1862) Chromis alta Greenfield & Woods, 1980 Chromis atrilobata Gill, 1862 Microspathodon bairdii (Gill, 1862) Microspathodon dorsalis (Gill, 1862) Stegastes acapulcoensis (Fowler, 1944) Stegastes arcifrons (Heller & Snodgrass, 1903) Stegastes flavilatus (Gill, 1862) Stegastes beebei (Nichols, 1924) Cirrhitidae — Hawkfishes Cirrhitichthys oxycephalus (Bleeker, 1855) Cirrhitus rivulatus (Valenciennes, 1846) Oxycirrhites typus Bleeker, 1857 Sphyraenidae — Barracudas Sphyraena idiastes Heller & Snodgrass, 1903 Polynemidae — Threadfins Polydactylus approximans (Lay & Bennett, 1839) Labridae — Wrasses Bodianus diplotaenia (Gill, 1862) Decodon melasma Gomon, 1974 Halichoeres adustus (Gilbert, 1890) ∑Halichoeres discolor Bussing, 1983 Halichoeres dispilus (Günther, 1864) Halichoeres nicholsi (Jordan & Gilbert, 1882) Halichoeres notospilus (Günther, 1864) ∑Halichoeres salmofasciatus Allen & Robertson, 2002 Iniistius pavo (Valenciennes in: Cuvier & Valenciennes, 1840) Novaculichthys taeniourus (Lacépède, 1801) Polylepion cruentum Gomon, 1977 Stethojulis bandanensis (Bleeker, 1851) Thalassoma grammaticum Gilbert, 1890 Thalassoma lucasanum (Gill, 1862) Thalassoma purpureum (Forsskål, 1775) Xyrichtys victori Wellington, 1992 Scaridae — Parrotfishes Scarus compressus (Osburn & Nichols, 1916) Scarus ghobban Forsskål, 1775 Scarus perrico Jordan & Gilbert, 1882 Scarus rubroviolaceus Bleeker, 1847 Opistognathidae — Jawfishes Opistognathus panamaensis Allen & Robertson, 1991 Tripterygiidae — Triplefins ∑Axoclinus cocoensis Bussing, 1991 Chaenopsidae — Tube blennies ∑Acanthemblemaria atrata Hastings & Robertson, 1999 Chaenopsis new species Coralliozetus boehlkei Stephens, 1963 Coralliozetus springeri Stephens & Johnson in: Stephens, Hobson & Johnson, 1966 Emblemaria nivipes Jordan & Gilbert, 1883 Dactyloscopidae — Sand stargazers Dactyloscopus pectoralis fallax Dawson, 1975 ∑Gillellus chathamensis Dawson, 1977 Labrisomidae — Labrisomid blennies Dialommus fuscus Gilbert, 1891 Paraclinus mexicanus (Gilbert, 1904)

Blenniidae — Combtooth blennies Entomacrodus chiostictus (Jordan & Gilbert, 1882) Hypsoblennius brevipinnis (Günther, 1861) Ophioblennius steindachneri Jordan & Evermann, 1898 Plagiotremus azaleus (Jordan & Bollman, 1890) Gobiesocidae — Clingfishes Arcos rhodospilus (Günther, 1864) Gobiesox adustus Jordan & Gilbert, 1882 ∑Gobiesox fulvus Meek, 1907 ∑Gobiesox woodsi (Schultz, 1944) ∑Tomicodon vermiculatus Briggs, 1955 Callionymidae — Dragonets Synchiropus atrilabiatus (Garman, 1899) Eleotridae — Sleepers Eleotris picta Kner, 1863 ∑Eleotris tubularis Heller & Snodgrass, 1903 Gobiidae — Gobies Bathygobius ramosus longipinnis Ginsburg, 1947 ∑Chriolepis atrimelum Bussing, 1997 ∑Chriolepis dialepta Bussing, 1990 Coryphopterus urospilus Ginsburg, 1938 Elacatinus nesiotes Bussing, 1990 Gobulus hancocki Ginsburg, 1938 ∑Lythrypnus alphigena Bussing, 1990 ∑Lythrypnus cobalus Bussing, 1990 ∑Lythrypnus lavenbergi Bussing, 1990 Lythrypnus rhizophora (Heller & Snodgrass, 1903) ∑Sicydium cocoensis (Heller & Snodgrass, 1903) Microdesmidae — Wormfishes Clarkichthys bilineatus (Clarke, 1936) Acanthuridae — Surgeonfishes Acanthurus nigricans (Linnaeus, 1758) Acanthurus triostegus (Linnaeus, 1758) Acanthurus xanthopterus Valenciennes in: Cuvier & Valenciennes, 1835 Ctenochaetus marginatus (Valenciennes in: Cuvier & Valenciennes, 1835) Naso annulatus (Quoy & Gaimard, 1825) vagrant Naso hexacanthus (Bleeker, 1855) vagrant Prionurus laticlavius (Valenciennes, 1846) Zanclidae — Moorish idol Zanclus cornutus (Linnaeus, 1758) Gempylidae — Snake mackerels Gempylus serpens Cuvier, 1829 Scombridae — Mackerels and tunas Acanthocybium solandri (Cuvier in: Cuvier & Valenciennes, 1832) Auxis rochei eudorax Collette & Aadland, 1996 Auxis thazard brachydorax Collette & Aadland, 1996 Euthynnus lineatus Kishinouye, 1920 Katsuwonus pelamis (Linnaeus, 1758) Sarda orientalis (Temminck & Schlegel, 1844) Scomber japonicus Houttuyn, 1782 Scomberomorus sierra Jordan & Starks in: Jordan, 1895 Thunnus albacares (Bonnaterre, 1788 ) Thunnus obesus (Lowe, 1839) Istiophoridae — Billfishes Istiophorus platypterus (Shaw in: Shaw & Nodder, 1792) Kajikia audax (Philippi, 1887) Makaira nigricans Lacépède, 1802

ORDER Pleuronectiformes Bothidae — Lefteye flounders Bothus leopardinus (Günther, 1862) Bothus mancus (Broussonet, 1782) Paralichthyidae — Sand flounders Hippoglossina bollmani Gilbert, 1890 Syacium latifrons (Jordan & Gilbert, 1882) ∑Syacium maculiferum (Garman, 1899) Soleidae — Soles Aseraggodes herrei Seale, 1940 Cynoglossidae — Tonguefishes Symphurus varius Garman, 1899

ORDER Tetraodontiformes Balistidae — Triggerfishes Balistes polylepis Steindachner, 1876 Canthidermis maculata (Bloch, 1786) Melichthys niger (Bloch, 1786) Melichthys vidua (Richardson, 1845) Pseudobalistes naufragium (Jordan & Starks in: Jordan 1895) Sufflamen verres (Gilbert & Starks, 1904) Xanthichthys caeruleolineatus Randall, Matsura & Zama, 1978 Xanthichthys mento (Jordan & Gilbert, 1882) Monacanthidae — Filefishes Aluterus monoceros (Linnaeus, 1758) Aluterus scriptus (Osbeck, 1765) Cantherhines dumerilii (Hollard, 1854) Ostraciidae — Boxfishes Ostracion meleagris Shaw in: Shaw & Nodder, 1796 Tetraodontidae — Puffers Arothron hispidus (Linnaeus, 1758) Arothron meleagris (Lacépède, 1798) Canthigaster punctatissima (Günther, 1870) Sphoeroides angusticeps (Jenyns, 1842) Sphoeroides lobatus (Steindachner, 1870) Diodontidae — Porcupinefishes Chilomycterus reticulatus (Linnaeus, 1758) Diodon holocanthus Linnaeus, 1758 Diodon hystrix Linnaeus, 1758

Σ indicates endemic to Isla del Coco DEEP indicates found very deep