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Supplementary Table 1. List of 82 Bacillus species covering 477 strains used in this study: strain

numbers and number of profiles.

Species name Strain numbers (number of profiles) Bacillus alcalophilus LMG 7120T (4) Bacillus amyloliquefaciens LMG 9814T (7); LMG 12325 (1); LMG 12329 (1); LMG 12385 (1) Bacillus aquimaris LMG 23073T (4) Bacillus arvi LMG 22165T (6); R-16994 (1) Bacillus atrophaeus LMG 16797T (1); LMG 8198 (9); LMG 8199 (5); LMG 17795 (4); LMG 17796 (2) Bacillus axarquiensis LMG 22476T (5) Bacillus azotoformans LMG 9581T (2); LMG 15443 (1) Bacillus badius LMG 7122T (4); LMG 12332 (1); LMG 18004 (1); LMG 18005 (1); LMG 18006 (1) Bacillus barbaricus LMG 23067 (5) Bacillus bataviensis LMG 21833T (7); LMG 21832 (1); R-15415 (1); R-15454 (2); R-16296 (1); R-16308 (1);

R-16321 (1); R-16324 (1); R-16325 (2); R-16336 (1); R-17019 (1) Bacillus bogoriensis LMG 22234T (8) Bacillus carboniphilus LMG 18001T (6) Bacillus cereus LMG 6923T (7); LMG 2098 (1); LMG 6910 (1); LMG 6924 (1); LMG 8221 (9); LMG 8396

(2); LMG 9676 (1); LMG 12235 (1); LMG 12236 (1); LMG 12237 (1); LMG 12334 (1);

LMG 12335 (3); LMG 12365 (9); LMG 13569 (4); LMG 14742 (4); LMG 17612 (1); LMG

18241 (2); LMG 18365 (2); LMG 18698 (1); LMG 22728 (1); LMG 22729 (1); LMG

22730 (1); LMG 22731 (1); LMG 22732 (1); LMG 22733 (1); R-20144 (1) Bacillus circulans 1 1) LMG 13261T (5); LMG 13271 (1); LMG 16568 (1); LMG 16628 (1); LMG 16629 (2)

2) LMG 6927 (1); LMG 12238 (1); LMG 12343 (2); LMG 13265 (1); LMG 13266 (3);

LMG 13267 (2); LMG 13268 (1); LMG 13270 (1); LMG 13272 (1); LMG 13273 (1); LMG

13274 (1); LMG 14421 (1); LMG 14422 (1); LMG 14423 (1); LMG 14424 (1); LMG

14636 (1); LMG 16560 (1); LMG 16561 (1); LMG 16564 (1); LMG 16565 (1); LMG

16567 (1); LMG 16570 (1); LMG 16583 (1); LMG 16585 (1); LMG 16593 (1); LMG

16594 (1); LMG 16595 (2); LMG 16624 (2); LMG 16626 (1); LMG 16627 (1); LMG

16630 (1); LMG 16634 (1); LMG 16635 (1); LMG 16637 (1); LMG 16638 (1); LMG

16640 (1); LMG 16641 (1); LMG 16645 (1); LMG 16646 (1); LMG 16647 (1); LMG

16649 (1); LMG 16650 (1); LMG 16693 (1); LMG 17441 (1); LMG 17442 (1); LMG

17444 (3); LMG 17447 (2); LMG 17449 (1); LMG 17464 (1); LMG 17465 (1); LMG

17469 (1); LMG 17470 (1); LMG 17472 (2); LMG 17473 (2); LMG 17486 (1); LMG

18014 (2); B0949 (1); R-36277 (1); R-36278 (1); R-36279 (1); R-36280 (1); R-36281 (1);

R-36282 (1); R-36283 (1); R-36284 (1) Bacillus clausii LMG 17945T (6); LMG 18518 (1) Bacillus coagulans LMG 6326T (11); LMG 12345 (2); LMG 12346 (2); LMG 12398 (2); LMG 12399 (1);

LMG 12400 (2); LMG 12401 (2); LMG 12402 (2); LMG 17451 (1); LMG 17452 (1); LMG

17453 (1); LMG 17456 (1); LMG 17457 (1); LMG 17474 (3); LMG 17475 (1); LMG

17476 (1); LMG 17477 (1); LMG 17478 (1); LMG 7376 (2) Bacillus cohnii LMG 16678T (8) Bacillus decolorationis LMG 19507T (4); R-5454 (1) Bacillus drentensis LMG 21831T (26); LMG 21830 (1); R-15416 (1); R-15427 (1); R-15445 (1); R-16310 (3);

R-16313 (1); R-16328 (2); R-16338 (1); R-16986 (1) Bacillus endophyticus LMG 21715T (5) Bacillus farraginis LMG 22081T (1); R-7343 (1); R-7148 (1) Bacillus firmus LMG 7125T (5); LMG 12241 (1); LMG 12242 (1); LMG 12243 (1); LMG 12352 (1); R-

15586 (1) Bacillus flexus LMG 11155T (6) Bacillus fordii LMG 22142 (3) Bacillus fortis LMG 22079T (1); LMG 22141 (2); R-7163 (1) Bacillus fumarioli R-10919 (1); R-13595 (1); R-13623 (1); R-13624 (1); R-13860 (1); R-13992 (1); R-14704

(1); R-14705 (1); R-14711 (1); R-16404 (1); R-19905 (1); R-19906 (1); R-19910 (1); R-

20285 (1); R-20287 (1); R-20342 (1); R-20444 (1) Bacillus funiculus LMG 22472T (5) Bacillus fusiformis LMG 9816T (6); LMG 17347 (2); B0661 (1) Bacillus galactosidilyticus LMG 17892T (5); LMG 12353 (2); LMG 12396 (3); R-15577 (1); R-16004 (1) Bacillus gelatini LMG 21880T (3); R-13476 (2); R-13565 (2); R-13588 (1); R-13635 (1); R-13810 (1); R-

13864 (1); R-13975 (2) Bacillus gibsonii LMG 17949T (7) Bacillus halmapalus LMG 17950T (6) Bacillus halophilus LMG 17942T (3) Bacillus horikoshii LMG 17946T (7); Bacillus horti LMG 18497T (5) Bacillus humi LMG 22167T (4); R-17036 (1) Bacillus indicus LMG 22858T (6) Bacillus insolitus LMG 17757T (3); LMG 17153 (3); B0433 (1) Bacillus jeotgali LMG 21653T (6) Bacillus laevolacticus LMG 6923T (9) Bacillus lentus 2 LMG 16798T (9); LMG 12354 (2); LMG 12359 (1); LMG 21649 (6); LMG 21758 (1); R-

36285 (1) Bacillus licheniformis LMG12363T (8); LMG 6934 (1); LMG 7558 (5); LMG 7559 (1); LMG 7560 (4); LMG

7561 (2); LMG 7562 (3); LMG 7626 (1); LMG 7627 (3); LMG 7628 (1); LMG 7629 (1);

LMG 7630 (1); LMG 7631 (3); LMG 7632 (2); LMG 7633 (1); LMG 7634 (1); LMG 7635

(1); LMG 7636 (1); LMG 7637 (2); LMG 12245 (1); LMG 12246 (1); LMG 12247 (1);

LMG 12248 (1); LMG 12360 (1); LMG 12361 (1); LMG 12362 (1); LMG 17334 (1); LMG

17337 (1); LMG 17339 (1); LMG 17340 (1); LMG 17649 (1); LMG 17651 (1); LMG

17652 (1); LMG 17653 (1); LMG 17654 (1); LMG 17655 (1); LMG 17656 (1); LMG

17657 (1); LMG 17658 (1); LMG 17659 (1); LMG 17660 (1); LMG 17661 (1); LMG

17662 (1); LMG 17663 (1); LMG 18685 (1); R-1210 (1); R-6452 (1); R-6646 (1); R-6979

(1); R-7199 (1); R-7478 (1); R-21381 (2); R-21382 (2); R-21383 (2); R-21384 (2); R-21385

(2) Bacillus luciferensis LMG 18422T (4); LMG 21400 (1); R-11670 (1); R-14109 (1); R-14110 (1); R-14111 (1) Bacillus malacitensis LMG 22477T (5) Bacillus marisflavi LMG 23072T (5) Bacillus megaterium LMG 7127T (7); LMG 3585 (1); LMG 11162 (1); LMG 12249 (1); LMG 12250 (1); LMG

12252 (1); LMG 12253 (1); LMG 12254 (1); LMG 12255 (1); LMG 12408 (1); LMG

12409 (1); LMG 18670 (1); LMG 18686 (1); LMG 18687 (1); LMG 18688 (1); LMG

18705 (1); LMG 18710 (1); LMG 18714 (1); LMG 23147 (2) Bacillus mojavensis LMG 17797T (4); LMG 17798 (4); R-28501 (1); R-28502 (1); R-28503 (1); R-28504 (1) Bacillus muralis LMG 20238T (4); R-8204 (1); R-8210 (1); R-8251 (1) Bacillus mycoides LMG 7128T (7); LMG 12256 (1) Bacillus neidei LMG 22737 (5) Bacillus niacini LMG 16677T (5) Bacillus novalis LMG 21837T (10); LMG 21836 (2); R-15418 (1); R-15446 (2); R-15450 (2); R-15453 (2);

R-16295 (1); R-16297 (1); R-16309 (1); R-16340 (1); R-16342 (1); R-16345 (2); R-16347

(1) Bacillus okuhidensis LMG 22468T (3) Bacillus oleronius LMG 17952T (2); LMG 17882 (1); LMG 17884 (1); LMG 17887 (1) Bacillus patagoniensis LMG 23070T (4) Bacillus pseudalcaliphilus LMG 17951T (6) Bacillus pseudofirmus LMG 17944T (5) Bacillus pseudomycoides LMG 18993T (6) Bacillus psychrodurans LMG 23063T (5) Bacillus LMG 9580T (6) psychrosaccharolyticus Bacillus psychrotolerans LMG 23062T (6) Bacillus pumilus LMG 18928T (17); LMG 3455 (3); LMG 8196 (3); LMG 8942 (2); LMG 10642 (3); LMG

12257 (1); LMG 12258 (2); LMG 12259 (3); LMG 12372 (1); LMG 12374 (1); LMG

12375 (1); LMG 12376 (4); LMG 12377 (1); LMG 18517 (9); LMG 18658 (1); LMG

18676 (1); B0296 (1); R-5334 (1); R-10579 (1); R-36286 (1); R-36308 (2); R-36309 (3); R-

36310 (6); R-36311 (2) Bacillus pycnus LMG 21634T (5) Bacillus ruris LMG 22866T (3) Bacillus shackletonii LMG 18435T (3); R-11667 (1); R-14112 (1); R-14113 (1); R-14114 (1) Bacillus silvestris LMG 18991T (6) Bacillus simplex LMG 11160T (8); LMG 12364 (2); LMG 17634 (1); LMG 17636 (1); LMG 17643 (1);

LMG 18473 (2); LMG 18508 (1); LMG 19489 (1); B0405 (1); R-5269 (1); R-5275 (1); R-

5282 (1); R-5307 (1); R-8191 (1); R-8193 (1); R-8202 (1); R-8207 (1); R-8208 (1); R-8214

(1); R-8215 (4); R-8216 (3); R-8218 (2); R-8220 (1); R-8225 (1); R-8231 (1); R-8234 (1);

R-8253 (1); R-8254 (1); R-15936 (1); R-15943 (1) Bacillus siralis LMG 22467T (6) Bacillus smithii LMG 12526T (3); LMG 6327 (2) Bacillus soli LMG 21838T (5); LMG 21839 (1); R-16301 (1); R-16307 (1) Bacillus sonorensis LMG 21636T (4) Bacillus sphaericus LMG 7134T (13); LMG 18663 (1) Bacillus sporothermodurans LMG 17668T (13); LMG 17895 (1); LMG 17896 (1); LMG 17897 (1); LMG 18460 (1);

LMG 18461 (1); LMG 18462 (1); LMG 18463 (1); LMG 18464 (1); LMG 18465 (1); LMG

18466 (1); R-1952 (2) Bacillus subterraneus LMG 23065 (5) Bacillus subtilis LMG 12260 (1); LMG 12261 (2); LMG 12262 (1); LMG 12263 (4); LMG 12264 (1); LMG

12417 (1); LMG 13579 (1); LMG 17723 (1); LMG 17725 (6); LMG 18271 (4); LMG 2099

(5); LMG 3589 (1); LMG 3590 (1)

subsp. spizizenii: LMG 19156T (1); LMG 19155 (1); LMG 19545 (4); LMG 8197 (15)

subsp. subtilis: LMG 7135T (13); LMG 19154 (1) Bacillus thermoamylovorans LMG 18084T (8) Bacillus thermoantarcticus LMG 23032T (4) Bacillus thuringiensis LMG 7138T (6); LMG 12265 (1); LMG 12266 (1); LMG 12267 (2); LMG 12268 (1); LMG

12269 (1) Bacillus vallismortis LMG 18725T (5); LMG 17799 (2); LMG 17800 (4); R-28507 (1); R-28553 (1) Bacillus velezensis LMG 22478T (5) Bacillus vireti LMG 21834T (5); R-15428 (1); R-15441 (1) Bacillus weihenstephanensis LMG 18989T (6) 1 Bacillus circulans is a very heterogeneous group of strains. The strains reported in point 1 are considered to be Bacillus circulans. The strains reported in point 2 are assigned to Bacillus circulans but further revision is needed.

2 The strain LMG 21649 is annotated as Bacillus lentus and as Bacillus halodurans by different culture collections (http://www.straininfo.net).

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