Puget Sound, Washington, With

WASHU- T- 72-005 c. 2

IeCUI.@+II>CG Cm V a Gra.~t D="-posttory

CHECKLIST OF PUGET SOUND FISHES by Allan C. Del acy, Bruce S. Miller, and Steven F. Borton

WSG 72 3 July 1972

DIVISION OF MARINE RESOURC.ES UNIVERSITY OF WASHINGTON 98IO5

Prepared under the National science Foundation SEA GRANT PROGRAM

ACKNOWLEDGMENTS Wewant to makespecial acknowledgment of thework put into theearly stagesof this checklistby R. L. Dryfoosand R. B. Grinols. Wewould also like to thankthe following individuals for thelog books they made available to us: T, S. Englishand C. M.Halstead University of WashingtonDepartment of Oceanography!;G.S. DiDonato, A. E. Millikan,R. M. Buckley, and F ~ Haw WashingtonDepartment of Fisheries!; R. W. Tyler, E. 0. Salo,and Q.J, Stober University of Washington Fisheries Research Institute!; W. L. High andC. R. Hitz NationalMarine Fisheries Service!; R. I. Matsudaand R. S. Domenawski Municipality of MetropolitanSeattle!; R. VanCleve Universityof WashingtonCollege of Fisheries!;R. B. Grinols Peninsula College!; and D. M. DesVoignes Port of Seattle!. Supportfor this projectwas from the Washington SeaGrant Program, NationalOceanic and Atmospheric Administration grant NC-1-72. CONTENTS

1 Introduet ion

3 Synopsis of Families 4 Checklist of Puget Sound Fishes

21 Bibliography

40 Index to Common Names 123o

122 Figure 1. The 12 sampling areas used to group reports of fish occurrence in Puget Sound. CHECKLIST of PUGET SOUNDFISHES*

Theaccompanying list of fishesconstitutes a compilationof dataon Puget Soundfish speciesdistribu ion up to 1971. It is compiledfrom the literature; fromunpublished catalogued entries in tneUniversity of Washingtoncollection of fishes; andfrom unpublished log recordsfrom the Universityof Washington Collegeof Fisheries,University of WashingtonDepartment of Oceanography, Universityof WashingtonFriday Harbor Laboratories, Washington State Department of Fisheries,National Marine Fisheries Service, Municipality af Metropolitan Seattle,and other less extensive sources. We have tried to avoidany duplica- tion of reportsand have made the general nature of this list conservative,since wehave attempted ta avoid any reports of questionablevalidity. A totalof 211 speciesof fish havebeen recorded in this checklist. In this list the symbol+ meansthat a specieshas been reported from one to ninetimes in the indicatedarea, the symbol 0+ indicates 10 ta 49reports, and thesymbol ~ means that 50or moreoccurrences have been reported. A blankspace meansthat we haveno record of occurrencein the particular area. In theuse of names,we have followed the standardset by theAmerican FisheriesSociety 970!; but in thosecases where the American Fisheries Society doesnot list a name,wehave used as our reference, Clemens andWilby 961!. Analphabetical listing of thefamilies with their page locations is on page 3, andan index to commonnames begins on page 40, Wehave also included a bibliography page 21! with this checklist, since published records are often of special interest. ThePuget Saund region Figure 1!, for faunisticpurposes, is defined as an arealimited on the west by a linedrawn northward along longitude 123o28' from EdizHook, near Port Angeles, Washington, to the U.S.-Canada international bound- aryline, and to the northwest andnorth by the U.S.-Canada international boundary line. Thefurther division of thePuget Sound region into 12 areas is basedin parton oceanographic conditions butalso reflects an arbitrary and expedient definitionof areas,with same attention to theintensity of thecollecting

efforts. ContributionNo. 371, College of Fisheries,University of Washington. A wideassortment of gearhas been been employed in the collectionof fish includedin this list beachseines, bottom trawls, gillnets, purse seines, traps, spearfishing,hook-and-line, poisoning, etc.!. It hasnot beenpossible to presentmeaningfully the relative samplingeffort in the various areas. Lower GeorgiaStrait, theinnter Strait of Juande Fuca, Admiralty Inlet, andColvos Passagehave been sampled less intensively than other areas. We recognize that for certainspecies e.g. the cutthroat trout, galenclarki! thefrequency oi reports probablyunderesti~ates the true frequencyof that speciesin certain areas,but for otherspecies e.g. theblackfin sculpin, Malacocottus kincaidi! wefeel the frequencyof reportsis of particularvalue and have r therefore added this roughlyquantitative aspect to ourlist, Theauthors would like to emhasize that the welcomean additionalinformation that readersma be re aredto provide. periodic updating of this list is contemplated. Thi.slist is basedon notebooks at the University of WashingtonCollege of Fisheriescontaining maps and data source sheets for eachspecies of PugetSound fish. In thenotebooks there is a mapshowing the distribution of reportsfor eachspecies. There is alsoa datasource sheet for eachspecies giving the exactsource of eachreport e.g. museumcatalog number, published record, log bookdate, etc.!. Thesource may be consultedfor detailedinformation on a particularspecies e.g. exactlocation of capture,date of capture,depth of capture,type of gearused, etc.!. Thenotebooks are availableat the Collegeof Fisheries for perusal by peoplehaving a needfor such data. SYNOPSIS OF FAMILIES

Page Family Page Family Acipenseridae sturgeons! Myctophidae lanternfishes! Agonidae poachers! Nemichthyidae snipe eels! Alepisauridae lancetfishes! Ophidiidae brotulas! Alopiidae thresher sharks! Osmeridae smelts! Paralepidae barracudinas! 12 Ammodytidae sand lances! Percichthyidae temperate basses! 12 Anarhichadidae wolffishes! Petromyzontidae lampreys! 15 Anoplopomatidae sablef ishes!

10 Bathymasteridae ronquils! Pholidae gunnels! Batrachoididae toadfishes! 19 Pleuronectidae righteye flounders! Ptili,chthyidae quillfishes! 19 Bothidae lefteye flounders! 12

10 Bramidae pomfrets! Ragidae skates! Carcharhinidae requiem sharks! Salmonidae salmons and trouts! Chimaeridae chimaeras! 10 Sciaenidae drums! Clupeidae herrings! Scomberesocidae sauries! 13 Scombridae mackerels and tunas! 15 Cottidae sculpins! Scorpaenidae rockfishes! 12 Cryptacanthodidae wrymouths! 13 Scyliorhinidae cat sharks! 18 Cyclopteridae lumpfishes and snailfishes! 12 Scytalinidae graveldivers! 10 Sphyraenidae barracudas! 9 Cyprinodontidae killif ishes! Squalidae dogfish sharks! 10 Embiotocidae surf perches! Engraulidae anchovies! Squatinidae angel sharks! Gadidae codfishes! Stichaeidae pricklebacks! Gasterosteidae sticklebacks! 13 S tromateidae but ter f ishes ! Gobiesocidae clingf ishes! Syngnathidae pipef i shee! 12 Gobiidae gobies! Torpedinidae electric rays! Trachipteridae ribbonfishes! 15 Hexagrammidae greenlings! Hexanchidae cow sharks! 13 Trichiuridae cutlassfishes! Icosteidae ragfishes! 10 Tr ichodontidae sand f ishes! Lamnidae mackerel sharks! Zoarcidae eelpouts!

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Ã Dg O 4 C l 4J 0 0 M CO EA il II ii + % ~ 20 BI BLIOGRAP HY Acenay Puntanilla, S. P. 1942.A newgenus and three new species of digenetic trematodesfrom Puget Sound fishes. M. S. Thesis. Univ. Washington. Allen,G. H. 1956. Migration,distribution, and movement of Puget Sound silver

salmon. Ph. D. Thesis. Univ. Washington. Allen,G. H., A. C. DeLacy,and D, W.Gotshall. 1960. Quantitive sampling of marinefishes a problemin fish behaviorand fishing gear, p. 448-511. In Wastedisposal in the marineenvironment. Pergamon Press. AlsoUniv. WashingtonColl, Fish. Contrib. 70.1 AmericanFisheries Society. 1970. A list of commonand scientific namesof fishesfrom the United States and Canada. 3rd ed. Am.Fish. Soc. Spec.

Publ. 6. Washington, D. C. 150 p, Batts,B. S. 1960.Further occurrence of the California pompano, Palometa simillima Ayres!in PugetSound, Washington. Copeia 1960 !:146-147. Batts,B. S. 1961.Intertidal fishes as food of thecommon garter snake.

Copeia 1961 !:350-351. Bayliff,W. H. 1953.Marked salmon returns in 1952,p. 33-38.In Puget Soundinvestigations. Washington State Dep. Fish. Prog. Rep. Dec. 1952-

Feb. 1953 ' Bayliff,W. H. 1959.Notes on the taxonomy anddistribution ofcertain zoarcid fishesin the northeasternPacific. Copeia1959 !:78-80. Bean,B. A. 1899Notes on the capture of rarefishes. Proc. U. S. Nat. Mus.

21:639-640. Bean,T. H. 1884.Notes on some fishes collected by James C.Swan in Washington Territory,including a new species of Macrurus.Proc. U. S. Nat.Mus.

6: 362-364.

21 Bean,T. H., andB. A. Bean.1897. Fishes collected at Beringand Copper Islandsby NicolaiA. Grebnitskiand Leonhard Stejneger. Proc. U. S. Net~

Mus. 19:237-251.

Bonham,K. 1954. Food of the dogfish, Washington State Dep. Fish. Fisheries Res. Pap. 1!:25-36 ' Bonbon,R,, andp, R.Olson. 1996. The whlee sea bass, ~Cnosclon nobi.lis. in Puget Sound. Copeia 1956 ! -64-65. Burke,CD V. 1930. Revisionof the fishesof the familyLiparidae. U, S. Nat. Mus Bull. 150:i-xii, 1-204 ' Carter,W. R., III. 1965. Racialvariations of the arrowgaby, Clevelandia ios Jordanand Gilbert! 1882 in PugetSound and on the coast of Washington State. M. S. Thesis. Univ. Washington. Chapman,W. M., andA. C. DeLacy.1933. Noteson the fishes of the State of Washington. Copeia 1933 !:102-103. Chapman,W.M., M.Katz, and D. W.Erickson. 1941. The races of herringin the Stateof Washington.Washington State Dep. Fish. Biol. Rep.38A. 35 p. Chew,K. K. 1970.Preliminary survey of macrofaunain Anderson Cove. A report preparedfor Boise CascadeProperties, Inc. 43 p. Chuinard,R. G. 1966. Thenatural history andpathology of epidermal

OrcasIsland, Washington. M. S. Thesis. Univ. Oregon. Clemens,W. A., and G. V. Wilby, 1961. Fishes of the Pacific Coastof Canada. Fish. Res. Board Can. Bull. 68. 443 p. Cobb,J; N. 1930. Pacific salmonfisheries. Bur. Fish. Rep.U. S. Comm.Fish. 1930. p. 409-704. Also Bur. Fish. Doc. 1092!. Collins, J. W. 1892. Reporton the fisheries of the Pacific Coastof the UnitedStates. U. STComm. Fish andFisheries Rep. Comm. 1888. p. 3-269.

22 Cooney,R. T. 1967.Diel differencesin trawlcatches of somedemersal fishes.

M. S. Thesis. Univ. Washington. Crawford,D. R. 1927. Recordsof rare fishes from the NorthPacific during

1925. Copeia 1927 60!:182-184. Dayton,L. M. 1969.Morphological and physiological studies of thethyroid of the ratfish, H drola us colliei, M. S. Thesis.Univ. Washington. DeLacy,A. C., andW. M, Chapman, 1935. Notes on some elasmobranchs of Puget Sound,with descriptionsof their eggcases. Copeia1935 !:63-67. DeLacy,A. C., andT. S. English.1954. Variations in beachseine samples causedby net lengthand repeated hauls. Ecology35!:18-20. DeLacy,A. C., andF. Neave.1947. Migrationof pink salmonin southern British Columbiaand Washington in 1945. Fish. Res.Board Can. Bull. 74, Also WashingtonState Dep.Fish. Biol. Rep.47A. 11 p.! DiDonato,G. 1969. The1968 Washington trawl landings by Pacific Marine FisheriesCommission and state statistical area. WashingtonState Dep.

Fish. Res. Div. Groundfish Data Rep. Ser. 5. 43 p. el-Sayed,S. Z. 1959.Population dynamite of English sole ~Farohtvs vetulus, Girard!in PugetSound, Washington, with special reference to theproblems of sampling. Ph. D. Thesis. Univ, Washington. English,T. S. 1967.Preliminary assessment of the English sole in Port Gardner,Washington. J. WaterPollut. ControlFed. 39 8!:1337-1350. Evermann,B. W., and E. L. Goldsborough.1907, The fishes of Alaska.Bull.

U. S. Bur. Fish. 26: 219-360. Evermann,B. W., and S. E. Meek.1898. A reportupon salmon investigations in the ColumbiaRiver Basin and elsewhere on the PacificCoast in 1896. Bull.

U. S . F ish. Comm. 17: 15-84.

23 Fisherman's News. 1963a. Sound gillnets takebonito, Seattie, Washington.

19 !: 8. Fisherman'sNews, 1963b. Unusual catch. Seattle,Washington. 197!:7 ~ Fisherman's News. 1963c. Bonito. Seattie, Washington.199!: 8. Fitzgerald, J. W. 1953. PugetSound sport catch, chinooksand silvers 1949-1952by areas,p. 46. ZnPuget Sound investigations. Washington State Dep. Fish. Prog. Rep. Dec. 1952-Feb. 1953. Fowler,H. W. 1912, Noteson salmonoidand related fishes. Proc Acad.' Nat.

Sci. Philadelphia 63:551-571. Garman,S. 1908. NewPlagiostomia and Chismopnea. Harvard Coll. Bull. Mus.

Comp, Zool. 51 9!:251-256. Garman,S. 1913. ThePlagiostomia sharks, skates, andrays!. HarvardColl. Mem. Mus. Comp. Zool. 36:1-528.

fromPuget Sound and Alaska. Proc'U. S. Nat. Nus. 11:554' Gilbert, CDH. 1895. Theichthyological collections of the steamer"Albatross" duringthe years1890 and 1891. U, S. Comm.Fish and Fisheries Rep. Comm. 1893. p. 393-476. Gilbert, C. H. 1904. Noteson fishes from the Pacific Coastof NorthAmerica. Proc. California Acad. Sci. Ser. 3, 3:255-271. Gilbert, C. H., andJ. C. Thompson.1905. Noteson the fishesof PugetSound. Proc. U. S. Nat. Mus. 28:973-987 ' Gill, T. 1863a. Descriptions of newspecies of Alepidosauroidae. Proc. Acad.

Nat. Sci Philadelphia 14:127-132. Gill, T. 1863b.On the classificationof the familiesand genera of Squallof California. Proc. Acad. Nat, Sci. Philadelphia 14:483-501.

24 Gill, T. 1865. Secondcontribution to the selachologyof California. Proc.

Acad. Nat. Sci. Philadelphia 16:147-151. Girard,C. 1856a.Characteristics of somecartilaginous fishes of the Pacific Coastof North America. Proc, Aced.Nat. Sci. Philadelphia7:196-197. Girard,C. 1856b. Noticeupon the viviparousfishes inhabiting the Pacific Coast of North America, with an enumerationof the species observed. Proc,

Acad. Nat. Sci. Philadelphia 7:318-323. Girard,C. 1857a.Contributions to the ichthyologyof the westerncoast of the United States, from specimensin the Museumof the SmithsonianInstitution.

Proc' Acad. Nat. Sci. Philadelphia 8:131-137. Girard, C. 1857b. Noticeupon the speciesof the genusSalmo, of authors, observed chiefly in Oregonand California. Proc. Aced. Nat. Sci.

Philadelphia 8:217-220 ' Girard,C. 1858. Noticeupon new genera and new species of marineand fresh- water fishes fromwestern North America. Proc. Aced. Nat. Sci.

Philadelphia 9:200-202. Girard,C. 1859 Fishes,Part IV, p. 1-400. In Reportsof explorationsand surveys,to ascertainthe most practicable and economical route for a railroad from the Mississippi River to the Pacific Ocean. 33rd Congr~

2nd Session House Executive Doc. 91 110!, Good,H. V. 1940. A studyof anepithelial tumorof ~Parohr s vetulus.

M. S. Thesis. Univ. Washington. Greenfield,D. W., andR. 3. Grinols. 1965. C rinodon varie atus on the OlympicPeninsula, Washington. Copeia1965 .!:115-116.

25 Greenwood,M. R. 1957. Bottomtrawling exploration in the Strait of Juande Fuca February to March1956. Comm.Fish. Rev. 19!:1-10. Gunther,A. 1860.Catalogue of thefishes in theBritish Museum. Printed by order of the Trustees, London2. 548 p. Gunther,A. 1862.Catalogue of thefishes in theBritish Museum.Printed by order of the Trustees, I ondon4. 534 p. Gunther,A. 1864.Catalogue of the fishesin theBritish Museum.Printed by order of the Trustees, London 5. 455 p. Gunther,A. 1866.Catalogue of the fishes in theBritish Museum. Printed by order of the Trustees, London6. 368 p. Halkett, A. 1913. Checklist of the fishes of the Dominionof Canadaand Newfoundland. King's Printer, Ottawa. 138 p. Haletead,B. W. 1950.The capture of a wattledeel pout,~L codea palearie, from Fidalgo Island, Washington,and the distributional record of this

species. Copeia 1950 !:317. Hanson,D. 1967. Cardiovasculardynamics and aspects of gasexchange in Chondrichthyes. Ph, D. Thesis. Univ. Washington. Harris, J. A. 1965. A comparativestudy af a newbottom-sampler with a Van Veengrab and an otter trawl. M. S. Thesis. Univ. Washington. Hart, J. F. 1935. Cestodafrom fishes of PugetSound. Ph. D. Thesis. Univ.

Washington. Haw,F., andR. M. Buckley. 1971. Saltwaterfishing in Washington. Stanley N. Jones, Seattle. 192 p. Heath,HE 1910. Theassociation of a fish with a hydroid. Biol. Bull. 19!:73-78.

26 High,W. L. 1966.Recent captures of theCalifornia pompano Palometa simillima! and the sandfish Trichodontrichodon! in PugetSound. WashingtonState Dep. Fish, Fisheries Res. Pap. 2!:53-54. Holland,G. A. 1969. Age,growth and mortality of racesof Englishsole ~perphr s vetulus!in PugstSound, Washington. Pac. Har, Fish. Comm.

Bull. 7: 35-50. Horn,A. D. 1932. Anecological survey of the marinelife of LongTree Point, Skagit Bay, Puget Sound. M. S. Thesis. Univ. Washington. Hubbs,C. LE 1921. Thelatitudinal variation in the numberof vertical fin

Hubbs,C, L. 1926. Descriptionsof newgenera of cottoid fishes relatedto Artedius. Univ. Michigan Mus. Zool. Occas. Pap. 170:1-18. Hubbs,C. L. 1928. A check-listof the marinefishes of Oregonand Washington.

J. Pan-Pac. Res, inst. 3!:9-16. Hubbs,C. L., andL. P. Schultz. 1933. Descriptionsof twonew American species referable to the rockfish genusSebastodes, with notes on related species. Univ. WashingtonPubl. Biol. 2!:15-44.

nerha!. Ph. P. Thesis. Wniv. Washington. JamisongD. W. 1966. Comparativeanatomy and histology of the olfactory apparatusof somecottid. fishes. M. S. Thesis.Univ. Washington. Jamison,D. W. 1970.Design of a long-termbenthic, marine biological, pollution- monitoringprogram evaluation of selectedhabitats, variables, and

communities. Ph. D. Thesis, Univ. Washington.

27 Jensen,H. M. 1952. 1952Puget Sound reef net taggingat BushPoint, p. 72-84. In PugetSound investigations. Washington State Dep, Fish. Prog.Rep.

March-June 1953 ' Johnson,D. R. 1952.Gillnet test drifts in PortGardner and Port Susan, p. 3-4. In PugetSound investigations. Washington State Dep. Fish Prog. Rcpt July-Nov. 1952. Johnson,M.S. 1970.Lactate dehydrogenase variation in thecrested blenny, M. S. Thesis. Univ. Washington. Jordan,D. S. 1884.Fishes, p. 169-682,In G.B. Goode,The fisheries and fishery industriesof the UnitedStates. Sect.I. U~ STComm. Fish and FisheriesU. S. GovernmentPrinting Office, Washington. Jordan,D. S. 1885. Themountain or salmontrout of Oregon.Bull. U. S. Fish

Comm. 5:310. Jordan,D. S. 1887. Thefisheries of the Pacific Coast,p. 591-629. In G. B. Goode,The fisheries andfishery industries of the UnitedStates. Sect.ZI. U. S. Comm.Fish and Fisheries. U. S. GovernmentPrinting Office, Washington. Jordan,D. S., andC. H. Gilbert. 1881a.Description of a newspecies of

3:409-410. Jordan,D. S., andC. H. Gilbert. 188lb. Descriptionsof twonew species of

Sound. Proc. U. S. Nat. Mus. 3:276-280. Jordan, D. S., and C. H. Gilbert. 188lc. List of the fishes of the Pacific Coastof the United States, with a table showingthe distribution of the

species. Proc. U. S. Nat. Mus. 3:452-458.

28 Jordan,D. SsH and C. H. Gilbert. 1881d.Observations on the sal~onof the

Pacif ic. Am. Nat. 15:177-186 ~ Jordan,D. Ssyand C. H. Gilbert. 1882.Notes on the fishesof thePacific Coastof the United States. Proc. U. S. Nat. Mus. 4:29-70. Jordan,D. Ssgand P. L. Jouy. 1882,Check list of duplicatesof fishesfrom the Pacific Coastof NorthAmerica, distributed by the Smithsonian Institution in behalf of the UnitedStates National Museum. Proc. U. S.

Nat. Mus. 4:1-18. Jordan,D. Sspand E. C. Starks. 1896. Thefishes of PugetSound, Proc.

California Acad. Sci. Ser. 2, 5:785-855. Katz,M. 1942.The herring races of Washington,with a noteon the fecundity of the Seal Rockpopulation. M. S. Thesis. Univ. Washington. Kelly,T. K. 1946.The commercial fisheries of Washington.Ph. D. Thesis.

Univ. Washington. Kendall,A. WsgJr, 1966,Sampling juvenile fishes on some sandy beaches of PugetSound, Washington. M. S. Thesis.Univ, Washington. Kilambi,R, V. 1965.Heterogeneity among three spawning populations of the surf smelt,~gomasus pretiosus Girard! in thegrate of Washington.Ph. D.

Thes is. Univ. Washington. Hilambi,R. V.. andA, C. Dehacy.1967. Heterogeneity of surf smelt,H~omesus Hretfosus Girard!, in thestate of Washington,asjudged by incidence of larval Anisakis Nematoda!. J. Fish. Res.Board Can. 24 !:629-633. Kilambi,R. Vsg F. M.Utter, and A. C.DeLacy. 1965. Differentiation of spawn- ingpopulations of the surf smelt ~H omesus pretiosus Girard! by serologi- cal methods.J. Mar. Biol. Assoc.India 7!:364-368. Also Univ. WashingtonColl. Fish. Contrib. 230.! Kincaid,T. 1919.An annotated liat of PugetSound fishes. WashingtonState Dep.Fish. F. M. LambornPublic Printer, Olympia.51 p, Kohlruss,F. J. 1933, Studieson the morphologyof somefish trematodesfrom PugetSound. M. STThesis' Univ. Washington. Leach,G. C. 1925.Artificial propagationof shad. Bur. Pish. Rep. U. S. Comm. Fish. 1924.p. 459-486. Also Bur. Fish. Doc.981.! Leong,G. 1967.Facundity of surf-smelt,~g omasus gretlosus Glrard!, in the state of Washington.M. S. Thesis. Univ. Washington. Lloyd,L. C. 1936. Somedigenetic trematodes from Puget Sound fish. Ph.D. Thesis. Univ. Washington Lord, J. K. 1866. Thenaturalist in VancouverIsland andBritish Columbia. RichardBentley, London. 2 vol. 358p,; 375figs. McArngG.Eeg R. G.Chuinerd, B. S. Miller, R. E. Brooks,and S. R.Wellings. 1968.Pathology of skin tumorsfound on English sole and starry flounder fromPuget Sound, Washington. J. Natl. CancerInst. 41:229-242. MacPhee,Ceg and W. A. Clemens.1962. Fishes of theSan Juan Island Archipelago, Washington. Northwest Sci. 36!'-27-38. Meek,S. E. 1899. Noteson a collection of cold-bloodedvertebrates from the Olympic Mountains' Field ColumbianMus ~ Zool. Series Publ. 31

12!:225-232. Menasveta,D. 1956. Migration and fishing mortality of Englishsole ~Farohr s vetulus! in SaratogaPassage, and adjacent waters. M. S. Thesis. Univ.

Washington. Miles, W.R. 1918. Experimentson the behaviorof somePuget Sound shore fishes Blenniidae!. Publ. PugetSound Biol. Stat. 27!:79-94.

30 Miller, B. S. 1965.Food and feeding studies on the adults of twospecies of melanostictus! pleuronectids stellatus and in EastSound, Orcas Island Washington!,H. S. Thesis.Univ. Washington. Hiller, B. S. 1967.Stomach contents of adultstarry flounder and sand sole in EastSound, Orcas Island, Washington. J. Fish.Res. Board Can. 242!:2515-

2526. Miller, B. S, 1969.Life historyaspects of normaland tumor-bearing flathead sole in EastSound, Orcas Island Washington!.Ph. D. Thesis.Univ.

Washington. Miller, B. S. 1970.The food of flatheadsole Hi o lossoideselassodon! in EastSound, Orcas Island, Washington. J, Fish,Res. Board Can. 27 9!:1661-

1665. Miller, B. S. 1971a,Seahurst County Park fish survey,18 p. In A. J. Kohnet al., Anecological survey of SeahurstCounty Park. Report submi.tted to King CountyDesign Commission in June, 1971. Miller,B. S. 197lb. Inshorefish surveyof DumasBay Park, King County, Washington,20p. In D. R.Paulson et al., Anecological survey of Dumas BayCounty Park. Report submitted to King County Design Commission in

August, 1971. Miller, B. S., andD. W. Greenfield. 1965. A juvenilesix-gilled shark Hexanchuscorinus! from the San Juan Islands, Washington. J. Fish. Res.

Board Can. 22!:857-859. Miller,B. S., andS. R. Wellings. 1971. Epizootiology of tumors on flathead sole Hi o lossoideselassodon! in East Sound, Orcas Island Washington!.

Trans. Am. Fish. Soc. 100!:247-266.

31 Miller,D. M., and G. D. Stauffer. 1967. Study of themigration and spawning distribution of the runsof chinookand coho in the Green-DuwamishRiver syste~in the fall of 1965. Univ. WashingtonColl. Fish. FisheriesRes. Inst. Circ. 67-4. 21 p ~ Norman,J. R. 1933.Notes on the flat-fishes Heterosomata!IV. A synopsisof thegenera of thesubfamily Pleuronectinae. Annu. Mag. Nat, Hist. ser.10, 9:214-222. Norman,J. R. 1934.A systematicmonograph of the flatfishes Heterosomata!, vol. 1. Psettodidae,Bothidae, Pleuronectidae, Brit. Mus.Nat. Hist. i-viii, 459 p,; 317 figs. O'Malley, H., and W. H. Rich. 1920. Migration of adult sockeye salmon in Puget Soundand Fraser River. Bur.Fish. Rep. U. S. Comm.Fish. 1918 Append. VIII. 38 p. Also Bur. Fish. Doc. 873.! Penis,M. R. 1932. Anepithelial tumorof ~Farohr s vetulus. M. S. Thesis. Univ. Washington. Parr, A. E. 1926. Investigationson the Cyclopterini. BergensMus, Aarb. 1924-1925. Naturvidensk. raekk nr. 7:1-31. Patten,B. G., D. T. Rodman,andK. D.Waldron. 1965. The Pacific bonito, Sarda chiliensis Cuvier!,in PugetSound, Washington. California Fish and Game 51!:298-299. Peden,A. E. 1966.Reexamination of two species in thestichaeid genus,

Powers,E. B. 1921. Experimentsand observations on the behaviorof marine fishes towardthe hydrogen-ionconcentration of the seawaterin relation to their migratorymovements and habitat. Publ. PugetSound Biol. Stat. 3 7!: 1-22. Horbuscha!in southern British Columbiaand Washington in 1943. Fish. Res. BoardCan. Bull. 66. AlsoWashington State Dep. Fish. Biol. Rep.44A. 23 p. ! Pruter,A. T.9 andR. VanCleve. 1954. A preliminarystudy of thepopulation of Englishsole, ~Parohr s vetulus!,in HolmesHarbor, Washington. Washington State Dep. Fish. Fisheries Res. Pap. l!:3-18. Rathbun,R. 1894. Summaryof the fishery investigationsconducted in the north Pacific Oceanand Bering Seafrom July 1, 1888, to July 1, 1892, by the U. S. Fish Commissionsteamer "Albatross." Bull. U. S. Fish. Comm.12:127-

201. Rathbun,R. 1900. A reviewof the fisheries in the contiguouswaters of the State of Washingtonand British Columbia.U. S. Comm.Fish andFisheries

Rep. Comm. 1899. p. 251-350. Read,L. Jtg Jr. 1969. Aspectsof nitrogenmetabolism and osmoregulation in lowervertebrates with emphasison the class Chondrichthyes.Ph. D. Thesis.

Univ. Washington. Richardson,J. 1836. Fishes,Part 3, 24pls. In FaunaBoreali-Americana; or the zoologyof the northernparts of British Americacontaining descriptions of the ob!ectsof naturalhistory collectedon the late northernland expeditions,under command of Sir JohnFranklin, R. N. RichardBentley,

London. Rosenblatt,R. H. 1964. A newgunnel, Pholis clemensi, from the coastof western North America. J. Fish. Res. Board Can. 21!:933-939. Rosenblatt,R. H., andD. Wilkie. 1963. A redescriptionof the rare cottid fish, Artedlus~mean i, new to thefauna of BritishColumbia. 3. Fish.Ras. Board

Can. 20 !: 1505-1511,

33 Rounsefell,G. As! andE. H. Dahlgren.1933. Tagging experiments onthe Pacific herring. ~Cluea ~allasii. 3. du ConseilS!:371-334. Salo,E, 0. 1969.Final report for theperiod June I, 1965-Sept.30, 1968! estuarineecology research project, Univ,Washington Coll. Fish. Fisheries

Res. Inst. Schaefer,M. B. 1936.Contribution to the life historyof thesurf smelt ~B oeesusBretiosus! in pugetSound. washington State Bep. Fish. Biol. Rep. 35B. 45 p. Schultz,L. P. 1930.Miscellaneous observations onfishes of Washington. Copeia 1930 !-'137 140 ' Schultz,L. P. I935. Thespecies of salmonand trout in thenorthwestern United States. Proc. 5th Pac. Sci. Congr, 1933. p. 3772-3782.

Schultz,L. P., andWE M. Chapman. 1934. A newOsmerid fish, dilatus, fromPuget Sound. Annu. Mag. Nat. Hist, ser. 10, 13:67-78. Schultz,L. Ps!and A. C.DeLacy. 1935-36. Fishes of theAmerican northwest, a catalogueof the fishes of Washingtonand Oregon, with distributional recordsand a bibliography.J. Pan-Pac.Res. Inst. five parts in five issues! 1935,10 !: 365-380;1936, II I!: 63-78! ll !: 127-142,11! 211- 226,Il!:275-290. Publishedaspart of Mid-Pac.Mag. 48!, 49 l!, 49!, 49!, and 49! respectively ~! Schultz,L. P., andH. A. Hanson.1935. Salmonoidgame fishes in the national forestsof northwesternUnited States. U. S. Dep.Agric. For. Serve! Seattle. 26 p. Also WashingtonSportsman. July, 1935!. Schultz,L. Pe! J. L. Hart, andF. J. Gunderson.1932. Newrecords of marine west coast fishes. Copeia 1932 !:65-68.

34 Schultz,L. Pe9 andA. D.Welander. 1935. A reviewof the cods of thenorth- easternPacific with comparativenotes on related species. Copeia 1935

!: 127-139. Shelford,V. E., A. 0. Weese,L, A. Rice,D. I. Rasmussen,andA. Maclean. 1935.General survey of thecommunities, p. 251-332. In Somemarine biotic communitiesof the Pacific Coast of NorthAmerica. Ecol. Monogr. 5. Shimada,B. M. 1948.Records of lantern fish in PugetSound. Copeia 1948

!:227. Sivalingam,S. 1953.Age and growth of Taeniotocalatezalis ~ M.ST Thesis.

Univ. Washington. Sjolseth,D. E. 1969.Studies of juvenilesalmon in the Nooksack River system andBellingham Bay. M. S. Thesis.Univ. Washington. Slipp,J. W.,and A. C.DeLacy. 1952. On the distribution and habits of the wattledeal-pout, ~L codes palearis Cop.eia1932 !:201-203. Smiley,C.W. 1885.Notes upon fish and the fisheries. Bull. U. S. Fish Comm.

5-'337-352. Smith,R. T. 1936.Report on the Puget Sound otter trawl investigations. WashingtonStateDep. Fish. Biol. Rep. 36B. 61 p. AlsoM, S. Thesis. Univ.

Washington. 1937.! Smith,R. T. 1937.Observations onthe shrimp fishery in PugetSound. WashingtonState Dep. Fish. Biol. Rep.36D. 19 p. Smoker,W.A. 1954.A preliminaryreview of salmonfishing trends on inner PugetSound. Washington State Dep. Fish. Res. Bull. 2. 55p. Sommani,p. 1969. Growth anddevelopment of sandsole post larvae psettich ~ths melanostictus!. M. S. Thesis.Univ. Washington.

35 Starks,E. C. 1896a.Description of a newgenus and species of cottoidfish fromPuget Sound. Proc. Acad. Nat. Sci. Philadelphia47:410-412. Starks,E, C. 1896b.List of fishescollected at PortLudlow, Washington. Proc. California Acad. Sci. ser. 2, 6:549-562. Starks,E. C. 1911.Results of anichthyological survey about the San Juan Islands, Washington.Ann. Carn. Mus. 7:162 213. Starks,E. C., and W. F. Thompson. 1911. A reviewof the flounders belonging to the genusPleuronichth s. Proc. U. S. Nat. Mus.38:277-287 ' Steindachner,F. 1877.Ichthyologische Beitrage V!. Sitz. Akad.Wiss. Wien

1876 74!:1-190, Steindachner,F. 1879.Ichthyologische Beitrage VII!. Sitz. Akad.Wiss. Wien

1878 78!:377-400. Stober,Q. J., andE. 0~ Salo. l970a. Biologicalstudies of theKiket Island nuclear-powersite. Annualreport for the periodSeptember 1, 1969- August31, 1970. Univ.Washington Coll. Fish. FisheriesRes. Inst. Stober,Q. J., andE. 0. Salo. 1970b.Biological studies of the Kiket Island nuclear-powersite. Reportto the WashingtonState Dep. Fish. of 1970 fish andshellfish coll. Univ.Washington Coll. Fish. FisheriesRes. Inst. Suckley,G. 1860.Report upon the fishescollected on the survey,p. 307-368.In Reportsof explorations and surveys, to ascertain the most practicable and economical route for a railroad from the Mississippi River to the Pacific Ocean.36th Congr.1st SessionHouse Executive Doc. 56 12!. Suckley, G. 1862a. Descriptions of several new species of Salmonidaefrom the northwest coast of America. Ann. Lyc. Nat. Hist. NewYork 7:1-10, Suckley,G. 1862b. Noticesof certain newspecies of NorthAmerican Salmonidae, chiefly in the collection of the NorthwestBoundary Commission. Ann. Lyc.

Net. Hist. New York 7:306-313.

36 Suckley,G. 1874.On the North American species of salmonand trout. U. S. Comm.Fish andFisheries Rep. Comm.1872 and 1873. p. 91-160,

lected in PugetSound, Washington. M. S. Thesis,Univ. Washington. Swan,J. G, 1881.The eulachon or candle-fishof thenorthwest coast. Proc.

U, S. Nat. Mus. 3:257-264. Swan,J. G. 1883,Shad in PugetSound. Bull. U, S. FishComm. 2, 1882:152. Swan,J. G. 1887.Codfish in thePacific. Bull. U. S. FishComm. 6:131. Tanner,Z. L. 1894.Report upon the investigations of the U. S. FishCommission steamer"Albatross" for the year endingJune 30, 1892. U. S. Comm. Fish and Fisheries Rep. Comm.1892. p- 1-64. Thompson,W.F., andN. L. Freeman.1930. History of thePacific halibut fishery. Int. Fish Comm.Rep. 5. 61 p. Ting,R. Y.-M. 1965. Ecology of demersal animals; problems in sampling. Ph. D.

Thesis. Univ. Washington. Townsend,L. D. 1935.The variation in themeristic characters of certain

Pacific flounders. M. S. Thesis. Univ. Washington. Tyler,R. W. 1964.Distribution and migration of young salmon in Bellingham Bay, Washington.Univ. Washington Coll. Fish.Fisheries Res. Inst. Circ.212.

26 p. VanCleve, R., andA. T. Pruter. 1956.Problems of sampling a Puget Sound popu- lerion oEniglishsole, ~Parahr s versine. Cons.Inr. Explor.Mer Rspp.

P. -V. 140 !: 87 93. Walker,E. T. 1953.Records of uncommon fishes from Puget Sound. Copeia 1953

!:239. WashingtonState Dep. Fish. 1967.1967 Annual Report. State Printing Plant,

Olympia.

37 WashingtonState Dep. Fish. 1969. 1969Annual Reports State Printing Plant,

Olympia.

armatus. Proc. Montana Acad. Sci. 29:63-71. Welander,A. D., andD. L. Alverson. 1954. Newand little knownfishes of the EasternPacific. WashingtonState Dep. Fish. FisheriesRes. Pap. 1!:1-8. Williams,g. W. 1999.The fishery for herring ~Cluea pallasit! onPuget Sound.Washington State Dep.Fish. FisheriesRes. Pap, 2!:5-30. Wismer, N. M~ 9 and J. H. Swanson. 1935. A study of the communitiesof a restricted area of soft bottom in San Juan Channel, p ~ 333-354. In Some marine biotic communitiesof the Pacific Coast of North America. Ecol.

Monogr. 5. Yap-Chiongoo,J. V. 1941. Hyyomssuspretiosus: its developmentand early life history. Ph. D. Thesis. Univ. Washington. Zebold, S. L. 1970. Inter- and intraspecific comparisonsof the diel distribu- tions and the food and feeding habits of five species of demersalfishes fromDuwamish Head, Puget Sound, Washington. M.S. Thesis.Univ. Washington.

38 INDEX TO COMMONNAMES For easein locating Puget Soundfishes by their commonnames, the following inde~ is included. Readersare remindedof the synopsisof scientific familynames appearing on page 3 of this checklist.

39 INDEX TO COMMONNAMES

17 allig ator f ish, smooth Poacher! Americans'had Herring! Calif ornia headlightf ish 67 Lanternfish! Anchovies 6 California skate 5 Angel Sharks 14 canary rockfish 12 arrow goby 7 capelin Smelt! 19 arrowtooth flounder 5 Cat Sharks 6 Chimaeras 14 China rockfish 7 chinook salmon 13 chub mackerel Barracudas 107 7 chum salmon Barracudinas 8 Clingfishes basking shark 5 9 20 Basses, Temperate C-O sole Righteye Flounder! ll cockscombs Prickleback! 12 bay goby 8 Codfishes 9 bay pipefish 7 coho salmon 19 Bering snailfish 15 18 bigeye poacher combfish, longspine Greenling! 13 copper rockfish big skate 5 Cow Sharks 69 blackbelly9 eelpout 11 black eelpout crescent gunnel 13 Cutlassfishes 12 blackeye goby 7 cutthroat trout 18 blackfin poacher 16 blackfin sculpin ll black prickleback 14 black rockfish 6 black skate 13 darkblotched rockfish 18 blacktail snailfish 17 darter sculpin 18 blacktip poacher ll daubedshanny Prickleback! 118 bluebarred prickleback blue lanternfish decorated warbonnet Prickleback! ll57 Dogfish Sharks 145 blue rockfish blue shark Dolly Varden Salmon, Trout! 20 18 bluespotted poacher Dover sole Righteye Flounder! 10 Drums 14 bocaccio Rockfish! 16 dusky sculpin 15 bonehead sculpin 12 dwarf wrymouth bonito, Pacific Mackerel, Tuna! 138 Brotulas5 brown cat shark 16 brown Irish lord Sculpin! 13 brown rockfish 9 Eelpouts 16 buffalo sculpin 6 Eels, Snipe 13 Butterfishes 6 Electric Rays 19 butter sole Righteye Floun der! 17 cabezon Sculpin! 207 English sole Righteye Flounder! eulachon Smelt! 16 calico sculpin

40 Lampreys 19 flathead sole Righteye Flounder! 4 Lancetfishes 19 Flounders, Lefteye 8 8Lanternfishes 19 Flounders, Righteye 17 fluffy sculpin 19 Lefteye Flounders 18 fourhorn poacher 15 lingcod Greenling! 13 frostfish Cutlassfish! 11 longfin gunnel longfin sculpin 1676longfin 8 smelt longnose lancetfish longnose skate 15 longspine combfish Greenling! 12 giant wrymouth 18 Lumpfishes, Snailfishes 12 Gobies 18 lumpsucker, Pacific spiny 12 Graveldivers 17 gray starsnout Poacher! 16 great sculpin 15 Greenlings greenstriped rockfish 136 Mackerel Sharks green sturgeon 5 13 Mackerels, Tunas 17 grunt sculpin 17 manacled sculpin 11 Gunnels marbled snailfish 188 midshipman, plainfin Toadfish! 20 Molas 16 mosshead sculpin ll mossheadwarbonnet Prickleback! 8 hake, Pacific Codfish! 19 halibut, Pacific Righteye Flounder! headlightfish, California Lanternfish! northern anchovy 6 Herrings 78 northern8 clingfish 11 high cockscomb Pri.ckleback! nor them lampfish Lanternfish! 10 northern ronquil 16 northern sculpin 17 northern spearnose Poacher! 16 Irish lord, brown Sculpin! 16 Irish lord, red Sculpin!

13 ocean perch, Pacific Rockfish! 20 ocean sunfish Mola!

15 kelp greenling 10 kelp perch 9 Killifishes 9 king-of-the-salmon Ribbonfish! 5 Pacific angel shark 10 Pacific barracuda 13 Pacific bonito Mackerel, Tuna! 8 Pacific cod 8 lampfish, northern Lanternfish! 6 Paci,fic electric ray

41 8 Pacific hake codfish! 5 Requiem Sharks 19 Pacific halibut Righteye Flounder! 19 rex sole Righteye Flounder! 6 Pacific herring ribbed sculpin 4 Pacific lamprey 177 ribbon barracudina 13 Pacific ocean perch Rockfish! 9 Ribbonfishes 10 Pacific pomfret ll ribbon prickleback 13 Pacific pompano Butterfish! 18 ribbon snailfish 19 Pacific sanddab Lefteye Flounder! 19 Righteye Flounders 10 Pacific sandfish 4 river lamprey 12 Pacific sand lance 13 Rockfishes Pacific sardine Herring! 6 9 15 rock greenling Pacific saury 18 rockhead Poacher! 5 Pacific sleeper shark 11 rock prickleback 18 Pacific spiny lumpsucker l9 rock sole Righteye Flounder! 16 Pacific staghorn sculpin 12 rockweed gunnel 8 Pacific tomcod 10 Ronquils 15 padded sculpin 15 rosylip sculpin pai.nted greenling 159 14 rosy rockfish pallid eelpout 15 roughback sculpin 11 penpoint gunnel 13 rougheye rockfish 19 petrale sole Righteye Flounder! 17 roughspine sculpin 107 pile perch pink salmon 9 8Pipefishes plainfin midshipman Toadfish! 17 Poachers 15 Sablef ishes 8 pollock, walleye Codfish! ll saddleback gunnel 10 Pomfrets 17 saddleback sculpin 13 pompano, Pacific Butterfish! sailf in sculpin ll Pricklebacks 165 salmon7 shark 14 Puget Sound rockfish Salmons, Trouts 15 Puget Sound sculpin 10 Sandfishes 18 pygmy poacher 12 Sand Lances 206 sandsole Righteye Flounder! sardine, Pacific Herring! 9 Sauries 15 scalyhead sculpin 14 quillback rockf ish 15 Sculpins 12 Quillfishes l0 seabass, white Drum! 10 seaperch, striped 5 6sevengill shark shad, American ll5 shanny, daubed Prickleback! 137 Ragfishes Sharks rainbow trout 14 sharpchin rockfish 6 ratfish Chimaera! 159 sharpnose sculpin 13 redbanded rockfish sheepshead minnow 8 red brotula 109 shiner perch ll red gunnel shortfin eelpout 16 red Irish lord Sculpin! 14 shortspine thornyhead 14 redstripe rockfish 19 showy snailfish

42 S continued!

9 Temperate Basses 15 silverspotted sculpin 14 thornyhead,shortspine Rockfish! silver surfperch threadfin sculpin 105 sixgill shark 16 6 threadfish Snipe Eel! 6 Skates 9 threespine stickleback ll slender cockscomb Prickleback! 5 Thresher Shark 19 slender sole Righteye Flounder! 17 tidepool sculpin 17 slim sculpin tidepool snailfish slipskin snailfish 18 197 14 tiger rockfish Smelts 8 Toadfishes 17 smooth alligatorfish Poacher! 8 tomcod, Pacific 15 smoothhead sculpin 7 Trouts, Salmons 18 Snailfishes, Lumpfishes 18 tubenose poacher snake prickleback tube-snout Stickleback! ll6 Snipe7 Eels 9 sockeye salmon 16 soft sculpin speckledsanddab Lefteye Flounder! 195 spiny dogfish 14 vermilion rockf ish 17 spinycheek starsnout Poacher! 16 spinyhead sculpin 13 splitnose rockfish 16 spotfin sculpin 18 spotted snailfish 8 walleye pollock 16 staghorn sculpin, Pacific starry flounder RighteyeFlounder! Warbonnets Prickleback! wattled eelpout 206 starry skate ll9 7 starsnouts Poacher! whitebait smelt whitebarred prickleback 179 Sticklebacks9 11 striped bass TemperateBass! 10 white seabass Drum! 10 white seaperch 10 striped seaperch 15 whitespotted greenling 14 stripetail rockfish sturgeon poacher 6 white sturgeon lj6 wolf-eel Sturgeons 12 12 Wolffishes 20 sunfish, ocean Mola! Wrymouths 107 Surfperches 12 surf smelt

14 yelloweye rockf ish 14 yellowtail rockfish 17 tadpole sculpin 19 tadpole snailfish