Of the United States of America

Proceedings OF THE National Academy of Sciences OF THE UNITED STATES OF AMERICA

Officers FRANK PRESS, President of the JAMES D. EBERT, Vice President Academy BRYCE CRAWFORD, JR., Home Secretary WALTER A. ROSENBLITH, Foreign Secretary ELKAN R. BLOUT, Treasurer Editorial Board MAXINE F. SINGER, Chairman of the GORDON A. BAYM EDWARD E. DAVID, JR. HERBERT E. SCARF Proceedings PAUL BERG STUART A. KORNFELD ROBERT L. SINSHEIMER WILLIAM F. BRACE DANIEL E. KOSHLAND, JR. SOLOMON H. SNYDER RONALD BRESLOW PETER D. LAX THOMAS A. WALDMANN MARY-DELL CHILTON DANIEL NATHANS FRANK H. WESTHEIMER PETER H. RAVEN Managing Editor: FRANCES R. ZWANZIG Senior Associate Editor: GARY T. COCKS Associate Editor: CAY BUTLER Associate Editor: JOHN M. MALLOY Associate Editor: JANET L. MORGAN Associate Editor: T. PEARSON Associate Editor: DOROTHY P. SMITH Associate Editor: COLENE WALDEN Assistant Managing Editor: JOANNE D'AMICO Senior Production Editor: LYNN A. WASSYNG Production Editors: BARBARA A. BACON, RUTH E. CROSSGROVE, SCOTT C. HERMAN, PEGGY LEONARD, MARGARET M. MADELEINE, JANET L. OVERTON, KATHLEEN RUBY, DON C. TIPPMAN, DEBORAH I. WEINER Administrative Assistants: DELORES BANKS, BEAULAH EDWARDS Manuscript Coordinators: CYNDY MATHEWS, JACQUELINE PERRY Circulation: JULIA LITTLE, VIRGINIA TREADWAY Editorial correspondence: PROCEEDINGS OF THE NATIONAL ACADEMY OF SCIENCES, 2101 Constitution Avenue, Washing- ton, DC 20418. Business correspondence: Circulation Office of the PROCEEDINGS, National Academy of Sciences, 2101 Constitution Ave- nue, Washington, DC 20418. Information for Contributors: See issue Number 1, January 1986. Copyright: The National Academy of Sciences has copyrighted this journal as a collective work and does not own copy- rights for individual articles. Requests for permission to reproduce parts of individual articles or for reprints of individual articles should be addressed to the authors. Microforms of complete volumes are available to regular subscribers only and may be obtained from University Microfilms, Xerox Corporation, Ann Arbor, MI 48103. Subscriptions: All correspondence concerning subscriptions should be addressed to the Circulation Office ofthe PROCEED- INGS. Subscriptions are entered on a calendar year basis only. For 1986, subscription rates are as follows-in the United States: student/postdoctoral, $65; personal, $185; institutional, $215; elsewhere: student/postdoctoral, $100; personal, $220; institutional, $250. Subscribers are requested to notify the Circulation Office ofthe PROCEEDINGS 6 weeks in advance of any change of address; also the local postmaster. The Academy is not responsible for nonreceipt of issues because of an improper address unless a change of address is on file. The notice of address change should list both the old and new addresses. Claims for replacement copies will not be honored more than 60 days after the issue date for domestic subscribers and not more than 90 days after the issue date for foreign subscribers. Back Issues: Volumes 78-82, January 1981 and thereafter, are available from the Circulation Office of the PROCEEDINGS. The price of a single issue is $21.00 for Volume 78 or $11.00 for Volumes 79-83. Second class postage paid at Washington, DC, and at additional mailing offices. PRINTED IN THE USA PROCEEDINGS OF THE NATIONAL ACADEMY OF SCIENCES OF THE UNITED STATES OF AMERICA (ISSN-0027-8424) is pub- lished semimonthly by THE NATIONAL ACADEMY OF SCIENCES, 2101 Constitution Avenue, Washington, DC 20418. C 1986 by THE NATIONAL ACADEMY OF SCIENCES OF THE UNITED STATES OF AMERICA. POSTMASTER: Send address changes to: PROCEEDINGS OF THE NATIONAL ACADEMY OF SCIENCES OF THE UNITED STATES OF AMERICA, 2101 Constitution Ave., Washington, DC 20418. ProceedIngs OF THE National Academy of Sciences OF THE UNITED STATES OF AMERICA

June 1986 Volume 83, Number 11 pp. 35694108 Table of Contents

AUTHOR INDEX ix

Physical Sciences

CHEMISTRY A revised set of values of single-bond radii derived from the observed Linus Pauling and Barclay Kamb 3569 interatomic distances in metals by correction for bond number and resonance energy Photoinduced oxidation of a water-soluble manganese(III) porphyrin Anthony C. Maliyackel, John W. Otvos, 3572 Larry 0. Spreer, and Melvin Calvin

MATHEMATICS Generalized random fields related to self-intersections of the Brownian motion E. B. Dynkin 3575

PHYSICS Thomas-Fermi theory for atomic systems Robert G. Parr and Swapan K. Gosh 3577

Biological Sciences

APPLIED BIOLOGY Meiosis in sesquidiploid hybrids of Lycopersicon esculentum and Solanum Charles M. Rick, Joseph W. DeVerna, 3580 lycopersicoides Roger T. Chetelat, and M. Allen Stevens

i AUTHOR INDEX, JUNE 1986

Alpers, D. H. 3781 Croce, C. M. 3934 Harik, S. I. 4067 Alpers, J. D. 3982 Cusanovich, M. A. 3693 Hartig, P. R. 4086 Argraves, W. S. 3766 Dafni, N. 3619 Hassell, J. R. 3761 Arlinghaus, R. B. 3629 Dalton, D. A. 3811 Hasstedt, S. 3909 Ar-Rushdi, A. 3934 Daly, J. W. 4089 Hawke, D. H. 4081 Baraban, J. M. 4053 Danciger, E. 3619 Hays, J. B. 3885 Bardin, C. W. 3589 Davidson, E. H. 3875 Hedge, A.-M. 3629 Barnard, M. B. 3738 DeAk, F. 3766 Heinemann, S. 4076 Barstad, L. 4021 Deftos, L. J. 4104 Heinzmann, C. 3929 Beard, P. 4016 DeMars, R. 3909 Henikoff, S. 3919 Begovich, A. B. 3594 Deom,C.M. 3771 Herlyn, M. 3728 Bellavite, P. 4002 De Souza, E. B. 4053 Hiken, J. F. 3649 Ben-Bassat, I. 4012 DeVerna, J. W. 3580 Hoffman, B. J. 4086 Bergstrom, S. 3890 Dobson, D. E. 3786 Honess, R. W. 3604 Berk, P. D. 3584 Doege, K. 3761 Hood, L. 3958 Berman, J. 3713 Douer, D. 4012 Hoover, R. G. 3982 Bernanke, J. 3844 Driscoll, C. J. 3870 Hopkins, N. K. 3703 Bernatzky, R. 3880 Duvall, E. J. 3939 Hosey, M. M. 3733 Bernstein, Y. 3619 Dynkin, E. B. 3575 Hsiao, H.-Y. 3743 Berton, G. 4002 Earl, P. L. 3659 Hsieh, T.-s. 3664 Binder, L. I. 4040 Eaves, C. J. 3851 Huang, S.-Y. 3738 Bleskan, J. 3919 Edelstein, S. J. 37% Huber, R. 3816 Blocker, H. 3746 Edsall, J. T. 37% Huebers, H. 3723 Blumenthal, T. 3865 Eisen, A. Z. 3756 Huebner, K. 3934 Bodemer, W. 3604 Ellingson, J. S. 3669 Hunt, C. R. 3786 Boger, J. 3972 Emanuel, B. S. 3934 Huo, L. 3654 Bohr, V. A. 3830 Eppenberger, H. M. 3816 Hwu, H. R. 3875 Boime, I. 3781 Ermine, A. 3914 Idzerda, R. L. 3723 Bonner, T. I. 3934 Erwin, C. R. 3599 Ignatius, M. J. 4108 Borsotto, M. 3733 Estess, P. 3594 Ikeda, R. A. 3614 Bosselman, R. A. 4035 Evans, H. J. 3811 Inkster, M. 3649 Botchan, M. R. 3609 Evans, M.G. 4099 Isobe, M. 3934 Boulet, A. M. 3599 Ezekowitz, R. A. B. 4002 Itakura, K. 4081 Brambl, R. 3644 Faber, S. 4016 Itoh, H. 3776 Braun, M. J. 4007 Felsted, R. L. 3847 Itoh, N. 4081 Breslauer, K. J. 3746 Finch, C. A. 3723 Iwai, S. 3776 Britten, R. J. 3875 Finke, U. 3844 Jacobson, K. A. 4089 Brotcorne-Lannoye, A. 3904 Fleckenstein, B. 3604 Jen, G. C. 3895 Brown, M. S. 3679 Frank, R. 3746 Joachim, C. L. 4044 Brunger, A. T. 3801 Freed, J. H. 3%8 Johnson, E. M., Jr. 4094 Bryan, P. 3743 Gallo, R. C. 4007 Jones, E. V. 3659 Buus, S. 3968 Galuska, S. 3972 Jones, P. P. 3594 Caffrey, M. 3693 Garcea, R. L. 3624 Kalckar, H. M. 3684 Calvin, M. 3572 Garman, R. D. 3987 Kalyanaraman, B. 3708 Cameron, K. R. 3604 Garrard, W. T. 3738 Kamb, B. 3569 Cameron, P. 3972 Gebicke-Harter, P. J. 4108 Kaplan, D. R. 3624 Cantley, L. 3624 Geever, R. F. 3944 Kaploon, A. 4012 Carswell, E. A. 3949 Geller, R. 3929 Karplus, M. 3801 Case, M. E. 3944 Ghosh, S. K. 3577 Katada, T. 3776 Cashmore, A. R. 3880 Gilden, R. V. 4007 Kato, Y. 4049 Caterson, B. 3761 Giles, N.H. 3944 Katz, J. J. 3791 Caton, A. J. 3771 Gillette, R. 4058 Kawasaki, H. 3776 Chen, T. T. 3698 Glowacki, J. 4104 Kaziro, Y. 3776 Chetelat, R. T. 3580 Goetinck, P. F. 3766 Keith, G. 3914 Chilton, M.-D. 3895 Goldberg, G. 3756 Khoury, S. 4026 Chin, J. 3972 Goldman, A. E. 3839 Kirk, K. L. 4089 Chin, W. W. 4026 Goldman, D. 4076 Kirk, T. K. 3708 Chou, J. Y. 3781 Goldman, R. D. 3839 Kiss, I. 3766 Chui, D. H. K. 3851 Goldsborough, M. D. 3934 Knott, T. J. 3929 Chung, S. W. 3851 Goldstein, J. L. 3679 Ko, J.-L. 3987 Clark, H. B. 4094 Gonda, M. A. 4007 Koeffler, H. P. 4035 Clark, S. 4049 Gordon, S. 4002 Konopka, J. B. 4049 Clark, S. D. 3756 Gorman, R. R. 3703 Koo, M. 3594 Clarkson, B. 4049 Gottesman, M. M. 3847 Koomey, J. M. 3890 Clements, J. E. 4007 Grey, H. M. 3968 Koprowski, H. 3728 Cleveland, J. L. 3934 Griffin, C. A. 3934 Kosik, K. S. 4044 Clore, G. M. 3801 Gronenborn, A. M. 3801 Kozak, C. 3934 Cohn, M. 3634 Groner, Y. 3619 Kozasa, T. 3776 Colon, S. 3968 Grossman, A. R. 3924 Kristensen, T. 3%3 Conley, P. B. 3924 Hammel, K. E. 3708 Kronberger, A. 3756 Conn, P. J. 4086 Hampikian, G. 3766 Kujiraoka, T. 4063 Connor, W. E. 4021 Hanawalt, P. C. 3830 Landefeld, T. D. 4026 Cornwell, M. M. 3847 Hanson, L. K. 3693 Lautenberger, J. A. 3698 Cox, K. A. 4104 Hanus, F. J. 3811 Laver-Rudich, Z. 3619 Crawford, G. D. 4081 Haranaka, K. 3949 Lawrence, J. C., Jr. 3649 Crespi, H. L. 3634 Hards, R. 3919 Lazdunski, M. 3733

ix AUTHOR INDEX, JUNE 1986

Leach, R. 3909 Papas, T. S. 3698 Scott, C. W. 3649 Lee, M.P. 3664 Papavasiliou, S. S. 4026 Scott, J. 3929 Lehrman, M. A. 3679 Parr, R. G. 3577 Seeholzer, S. H. 3634 Lemaux, P. G. 3924 Pascoe, G. A. 3811 Sekiguchi, M. 3997 Limjuco, G. 3972 Pastan, I. 3847 Selkoe, D. J. 4044 Lin, D. S. 4021 Patrick, J. 4076 Serra, M. C. 4002 Lomax, T. L. 3924 Patterson, D. 3919 Shastri, N. 3958 London, J. A. 4058 Paul, S. M. 4071 Shimizu, K. 3997 Lovett, P. S. 3939 Pauling, L. 3569 Shinnick, T. M. 3751 Luck, S. 4021 Pedersen, N. 3718 Shively, J. E. 4081 Lusis, A. J. 3929 Pederson, T. 3718 Shooter, E. M. 4108 Lusky, M. 3609 Pesold-Hurt, B. 3820 Shyu, A.-B. 3865 McDevitt, H. 0. 3594 Peterson, S. W. 3684 Silen, W. 3844 McKinnon, R. D. 3751 Pichersky, E. 3880 Simmons, D. 4076 McKnight, G. S. 3723 Pierres, M. 3958 Singh, B. 3629 McLaughlin, J. 4049 Pimpinelli, S. 3900 Skene, J. H. P. 4108 MacLeod, C. L. 3825 Pizer, L. I. 4016 Skolnick, P. 4071 Maenhaut-Michel, G. 3904 Pledger, W. J. 3834 Slemmon, J. R. 4081 Magun, B. E. 3825 Plesofsky-Vig, N. 3644 Smith, C. 3968 Mahley, R. W. 4108 Poch, 0. 3914 Snyder, S. H. 4053 Malissen, B. 3958 Pollock, N. J. 4040 Sparkes, M. C. 3929 Maliyackel, A. C. 3572 Poulos, T. 3743 Sparkes, R. S. 3929 Marchal, G. 3977 Prendergast, J. S. 3949 Sparks, K. J. 3766 Marky, L. A. 3746 Price, D. H. 3674 Sparrow, J. T. 3629 Marshall, J. C. 4026 Putkey, J. A. 3634 Spiegelman, B. M. 3786 Martin, K. 3654 Pyper, J. M. 4007 Spreer, L. 0. 3572 Marty, A. 4099 Quill, S. G. 3743 Steinert, P. M. 3839 Marzluff, W. F. 3674 Radbruch, A. 3954 Stevens, M. A. 3580 Matsui, S.-i. 3738 Raff, R. A. 3865 Stockdale, F. E. 3860 Matsukawa, S. 3997 Rajewsky, K. 3954 Strauss, A. W. 3781 Maul, G. 3839 Rakowicz-Szulczynska, E. M. 3728 Stremmel, W. 3584 Mayrand, S. H. 3718 Ramot, B. 4012 Strife, A. 4049 Means, A. R. 3634 Randall, R. E. 3604 Strohmeyer, G. 3584 Mendelsohn, J. 3825 Ranyard, J. 4035 Sugimura, T. 3997 Meyer, T. E. 3693 Rapp, U. 3934 Sunada, H. 3825 Michalski, T. J. 3791 Raptis, L. 3624 Sussman, F. 3806 Miles, C. 3968 Raulet, D. H. 3987 Sutcliffe, J. G. 3751 Miller, C. 4035 Reed, J. C. 3982 Suzdak, P. D. 4071 Miller, J. B. 3860 Reisner, Y. 4012 Suzuki, K. 3776 Miller, R. L. 3589 Rhodes, J. A. 3844 Swanson, J. 3890 Milon, G. 3977 Richardson, C. C. 3614 Swanson, L. W. 4076 Min, H. Y. 3786 Rick, C. M. 3580 Tachibana, C. Y. 3713 Mirra, S. S. 4040 Ripoll, P. 3900 Tack, B. F. 3963 Mohandas, T. 3929 Risau, W. 3855 Taira, M. 3997 Mori, M. 3997 Ro, J. H.-S. 3786 Tam, J. P. 3844 Morita, E. 4081 Robb, R. J. 3992 Taniuchi, M. 4094 Morris, G. F. 3674 Robbins, K. 3890 Tanksley, S. D. 3880 Moser, H. 3816 Roberts, E. 4081 Taraschi, T. F. 3669 Moss, B. 3659 Roberts, J. W. 3875 Tarpley, W. G. 3703 Muller, W. 3954 Roberts, T. M. 3624 Terada, M. 3997 Mumby, M. C. 3649 Rodeck, U. 3728 Teter, M. 3756 Murayama, T. 3944 Rose, S. M. 3738 Tezon, J. 3589 Murphy, R. A. 3844 Rosen, H. 4002 Tollin, G. 3693 Nagarajan, L. 3934 Rossi, F. 4002 Tordo, N. 3914 Nagata, S. 3776 Rougeon, F. 3914 Tye, B.-K. 3713 Naguib, F. 3919 Rubin, E. 3669 Ui, M. 3776 Nakamura, S. 3776 Russell, D. W. 3679 Ukena, D. 4089 Nash, D. 3919 Russell, S. A. 3811 Ullrey, D. B. 3684 Neer, A. 3619 Rutter, W. J. 3599 Van Beneden, R. J. 3698 Neuringer, M. 4021 Safa, A.R. 3847 van Loon, A. P. G. M. 3820 Nicolson, M. A. 4035 Saito, T. 4063 Vulpe, C. D. 3987 Niller, H.-H. 3604 Sakamoto, H. 3997 Wallimann, T. 3816 Nitta, M. 4049 Salvaterra, P. M. 4081 Warshel, A. 3806 Nolan, J. M. 3664 Sanders-Bush, E. 4086 Watson, D. K. 3698 Nowell, P. C. 3982 Satomi, N. 3949 Wazuddin, M. 3870 Ohtsuka, E. 3776 Saunders, M. 3844 Wegmann, G. 3816 O'Keefe, E. J. 3834 Scarborough, G. A. 3688 Weisgraber, K. H. 4108 Okumoto, D. S. 3830 Schaffhausen, B. 3624 Wen, D. 3639 Olashaw, N. E. 3834 Schatz, G. 3820 White, R. 3909 Old, L. J. 3949 Schilling, J. W. 4108 Whitman, M. 3624 Osborne, W. R. A. 4030 Schleif, R. F. 3654 Wilcox, K. W. 4016 O'Sullivan, J. 4104 Schlesinger, M. J. 3639 Wilhelm, S. M. 3756 Otvos, J. W. 3572 Schmidt, J. A. 3972 Wilkie, D. 4104 Ovitt, C. E. 3781 Schulze, I. T. 3771 Williamson, B. D. 3949 Pallas, D. 3624 Schwartz, E. 4012 Williamson, R. 4081 Pantoliano, M. W. 3743 Schwartz, R. D. 4071 Wilson, C. 3684

x AUTHOR INDEX, JUNE 1986

Windle, B. E. 3885 Worcester, D. L. 3791 Yang, H.-Y. 3839 Witte, O. N. 4049 Worley, P. F. 4053 Yoshida, T. 3997 Wong, P. M. C. 3851 Wu, A. 3669 Zimmerman, R. 3669 Wong-Staal, F. 4007 Wyckoff, E. 3664 Zmeili, S. 4026 Wood, J. G. 4040 Xu, M. 3738 Woolford, J. F. 3919 Yamada, Y. 3761