Volume 8 – 20 June 2014 A newsletter for the Pyraloidea fans
Editorial Collecting is a primordial component will be the regulation on access to genetic of our research and more important than resources, the “ABS Regulation” in short Dear Pyraloidea fans, ever in the current sorry state of nature (see http://register.consilium.europa.eu/doc/ conservation. Thus, I was glad to read a srv?l=EN&f=PE%20131%202013%20INIT), Welcome to the 8th edition of our news- letter in support of collecting in Science which was adopted by the Parliament and letter. I hope that you had or are having a recently (see http://www.sciencemag.org/ Council of the European Union and several good collecting season. Here in Switzerland content/344/6186.toc#Letters). It seems to countries. As its Article 1 reads, “this Regula- we were experiencing our first hot spell just be more complex now to go out collecting, tion establishes rules governing compliance recently and although I saw lots of moths at borrow and loan specimens, obtain funding with access and benefit-sharing for genetic light, I wasn’t very lucky with pyraloids. for collections maintenance and improve- resources and traditional knowledge associ- ment, and publish taxonomy, than it was ated with genetic resources in accordance in the past. One more possible obstacle with the provisions of the Nagoya Protocol on Access to Genetic Resources and the Fair and Equitable Sharing of Benefits Aris- ing from their Utilization to the Convention on Biological Diversity.” When ratified by at least 50 states, the Nagoya Protocol will take effect, and this is scheduled to happen in 2015. I assume that the Nagoya Protocol is
This issue was made possible with the help of Todd Gilligan, John Hawking, James Hayden, Sangmi Lee, Houhun Li, Richard Mally, Shankar Murthy, Hideshi Naka, Matthias Nuss, Harsimran Singh, Alma Solis, and Stephen Sutton.
The logo of The Pyraloid Planet was created by Florence Marteau of the Muséum d’histoire naturelle, Geneva, Switzerland, and the layout of this issue was made by Corinne Charvet of the A new genus and species of Midilinae from the Serra Bonita Reserve, Bahía, Brazil, currently being same institution. described by Vitor Becker. Photo by B. Landry.
1 not against collecting with proper authoriza- increased by 688 (+ 311 synonyms) to a total The larval morphology contained apo- tions, but it will probably impact on our work. of 14,823 (+ 5,949 synonyms). Nearly 4,800 morphic characters which supported taxo- For example, article 28 mentions that “a sys- changes were made by five contributors to nomic placement of many species. The tem of registered collections within the Union GlobIZ pages. These counts well demon- larval characters provided diagnostic apo- should be put in place through the establish- strate the increasing efforts made to improve morphic characters which distinguished 20 ment of a voluntary register of collections the quality of the data in contrast to the sole previously misplaced species and confirmed to be maintained by the Commission. Such addition of new records to the database. the generic placement of 11 species. The a system would ensure that collections structure of the forecoxae was found to be included in the register effectively apply In 2011, for Zootaxa’s special issue on a major diagnostic character state which measures restricting the supply of samples animal biodiversity, we provided a count of allowed generic placement of species. This of genetic resources to third persons with 15,579 pyraloid species (Nieukerken et al., character was strongly supported by other documentation providing evidence of legal 2011) for which we calculated the number of morphological characters (breathing appa- access, and ensure the establishment of species per subfamily using GlobIZ as well ratus structure and arrangement, head mutually agreed terms, where required.” as LepIndex. Already in 2012, we announced direction and mandible form) and ecological More in years to come… that the calculated number for crambines parameters (habit, habitat, food type and was exceeding the previous year’s count. case or retreat type). On a personal note my trip to Brazil Since then, the count slightly exceeds that at the end of last year was great as I was for other subfamilies too by a few species, or The CO1 gene inferred the association able to sample several new habitats, thanks several dozens, as in Spilomelinae. Richard of 31 larvae with their respective adults, to the help and guidance of Vitor Becker. I Mally, who is busy with the latter taxon, says with the larval taxonomy supported by diag- have made no plans to organize a real col- that it is not the end of the line. Let’s look for nostic morphological characters. The CAD, lecting expedition this year. Instead I hope the progress during the next twelve months, CAD+CO1 and CO1 gene trees inferred to finish some manuscripts and visit the generic and specific relationships, and phy- and as usual to conclude this news item: Natural History Museum, London. Given the logeny, which were supported by the tra- Everybody is welcome to take part in GlobIZ! lack of technical assistance I am faced with ditional morphological descriptions of the and the extra administrative duties that fell adults. on me since early last year, I must target Matthias Nuss fewer taxa in the field and take extra care in The larval morphology, larval ecology planning my work time. In this context, I am and molecular sequencing greatly increased really VERY much grateful to those of you the quantity of taxonomic information and who responded positively to Théo Léger’s Systematics and character states to provide an informed sys- request last year (see PP 7: 4–5) for Cram- tematic placement of Australian acentropine binae and Scopariinae specimens in ethanol Ecology of the species. The knowledge gained will allow a for our phylogeny project with Mathias Nuss. Australian Aquatic more critical evaluation of the world Acentro- Please don’t forget us if you haven’t sent us pinae fauna and the use of these techniques material yet and planned to do so! We are Moths, Acentropinae will help resolve some of the taxonomic still missing many taxa. impediments currently experienced in the systematics of the Australasian and Oriental I thank the contributors to this issue, The Aquatic Moths, Acentropinae (Cram- faunas. which I hope you will find entertaining, or at bidae) of Australia were revised by the least interesting! The references cited in the traditional morphological taxonomic tech- John Hawking texts can be found in GlobIZ (www.pyraloi- niques of wing patterns, wing venation and dea.org), unless otherwise stated. As usual genitalia of the males, which were comple- please don’t hesitate to recruit new pyral- mented by adding larval morphology, larval An illustrated guide oid fans by disseminating this issue widely ecology and molecular sequencing. The or otherwise, send any change of address revision recognised 17 genera, with 54 spe- to the Thyridoid & before the next issue, and feel free to let me cies, of which seven genera and 10 species know if you would like to edit the next issue. were undescribed. Forty one species were Pyraloid Moths of consistent with 10 described genera, Any- Borneo Cheers, draula, Elophila , Eoophyla, Margarosticha, Bernard Landry Nymphicula, Paracataclysta, Paracymoriza, Parapoynx, Strepsinoma and Tetrernia, The macro-moths of Borneo are now well and 11 species were consistent with seven known, thanks to the 30 years of sustained GlobIZ News 2014 undescribed genera. Three genera, Hyle- effort of Jeremy Holloway with the support bates, Nyctiplanes and Neoshoenobia were of Henry Barlow through Southdene Sdn inconsistent with the Acentropinae and were Bhd. Among the ‘micros’ of Borneo the Pyra- During the last 12 months, the num- removed. Taxonomic descriptions, plus taxo- loidea have long stood out as a group crying ber of valid species included in the Global nomic keys of the adults were given for the for attention. They are abundant and highly Information System on Pyraloidea (GlobIZ) genera and species. diverse, collections are available for study
2 The Pyraloid Planet 8 - 2014 and much preliminary work has been done In 2009 we realized that the medium of stage of preparation for uploading as Phase to make sense of the generic structure and the printed page was unsuitable for an illus- 2 of the website development. Extensive species groups. However, with a likely mini- trated guide of a group with a rapidly evol data from specimen labels and the literature mum of 2500 species, at least 50 person- ving taxonomy and burgeoning number of has been collected by TMW, and some 4,500 years would be needed to revise genera and described species and identifiable morpho- images have been prepared by Southdene. name species, and time is pressing. types, and that a website, with all the flexi The challenge now is to distil these data and bility it provided for use and updating, was images into uploads to the website and find a With this in mind, in 2004 Henry Barlow the best way forward. provisional sub-family and generic structure initiated the production of ‘An Illustrated to provide a framework. Guide to the Pyraloid Moths of Borneo’ We are now happy to introduce to you, the members of the Pyraloid Planet (PP) col- to be published in book form, as a bridge laborative network (or collective for short), to the eventual production of taxonomic the first phase of the website, comprising monographs. Note that the concept was 295 described species and morphotypes (and remains) of an illustrated guide with no at or near the species level (designated attempt to formally name species or revise as ‘sp.1’, ‘sp.2’, etc.). For us the PP is an genera. He recruited Terry Whitaker and Ste- extremely valuable organisation to act as a phen Sutton to assist. We added some 300 sounding board and hopefully to produce a species of thyridoids to this project because team to eventually take over the develop- they were ‘joined at the hip’ with pyraloids ment of the site. in the earlier literature and make a group of more convenient size to work up before We currently have some 2500 species/ entering the formidably speciose pyraloid morphotypes of pyraloids in Borneo (in the Dysodia sp.14, an unnamed specimen from jungle. approximate ratio of 55:45) in an advanced Sabah, Mt. Kinabalu N.P., Poring Hot Springs.
Screenshot extract from the ‘Getting Started’ section of the website, showing the structure of a species ‘page’.
The Pyraloid Planet 8 - 2014 3 The site can be found at http://www. loids hasn’t yet been decided, but we will museum collections. Many of the morpho- pyralidsofborneo.org/index.php?index. It is certainly have the individual species ‘pages’ types are unnamed series set aside by the built on a software platform prepared by Oli- and 80+ Plates with 30 or so species imaged late Michael Shaffer and Eugene Munroe ver Georgi in Germany. Lim Kooi Fong, the on each and the facility to click through to the in the collections of the Natural History owner of Biovis Informatics Sdn Bhd in Kuala individual species or morphotype. Museum in London, as detailed in LepIndex. Lumpur is using this software under licence Their work has contributed heavily to the to produce a range of websites. This pyraloid The pyraloids to be uploaded will be foundations for this website. one is the prototype for a series of illustrated treated in the same way as the thyridoids guide websites cataloguing the biodiversity already on the site, with one difference. For The PP Collective has the expertise and of Malaysia. These in turn will be the source the thyridoids we were able to include spe- membership size to make major additions to for downloadable ‘AppBooks’ to provide cies/morphotypes not found in Borneo but the data we need to ‘fill out’ this site. Please research tools for identifying plant and ani- recorded, as it were, from both sides of the use the Feedback form (last Section of mal groups when out of range of broadband island. Due to the large number of pyraloids the site Menu) for contributions and thereby e.g. in the 750K ha Sabah Foundation forest we will not be able to do this in the next phase make the development of this site a PP Col- concession of central Sabah, NE Borneo, of uploading (it will probably take 4 years to lective activity. a major biodiversity hotspot. Typically an upload our existing list of 2500 species/mor- ‘AppBook’ will consist of images of species, photypes). Unless/until significant further Stephen Sutton, Terry Whitaker & names and minimal text, with links to the funding materialises we are constrained. Henry Barlow parent website from which fuller details can be downloaded when in broadband range Apart from the site being developed by (available in Malaysian towns and research Biovis Informatics for Southdene publishing News from… stations with direct satellite links). company for the macro moths of Borneo, we are not aware of other websites for identify- The basic unit of the pyralidsofborneo site ing SE Asian moths, or indeed of any site James Hayden is a ‘page’ for each species, with a tab for anywhere with the same approach as ours. ‘Taxonomy’, ‘Description’, ‘Distribution’ and We hope very much that others will follow Like weeds, diseases, and giant snakes, more (see image of the screenshot from the and be able to use the same software and pyraloids invade Florida regularly. For exam- ‘Getting Started’ section of the site). There design. ple, I have been tracking Nacoleia “charesa- are empty tabs for such data as DNA barcod- lis” (Walker) (a widespread Palaeotropical ing and future uploading of genitalia images. Hardly any of the species/morphotypes species, possibly a complex). It is moving The tabs for ‘Distribution & Habitat’ and ‘Life on our site have been named or imaged on fast: since 2012, it has appeared on the History & Pest Status’ have absolutely mini- the internet before and are only accessible southeast and southwest coasts of Florida, mal content reflecting our minuscule amount to the few people with clearance to study and very recently near Orlando in the state’s of knowledge of these aspects. A major pur- pose of the site is to provide an opportunity for the PP Collective and others to fill these gaps.
On each ‘page’ there is a large, magnifi- able image of the species, and a thumbnail. A long introduction section covers a range of subjects such as ‘Material Examined’ and ‘LepIndex’ (to which there is a hotlink). There is a Checklist and some 800 literature refer- ences which are being added to all the time. A Gazetteer covering all the collection sites mentioned on the site will prove useful for all of the South and South-east Asian region.
Special effort has been made to maxim- ise ease of use. For the thyridoids, individual species/morphotype pages can be reached by clicking on the image of the species on one or other of the 10 Plates or you can call up a ‘genus gallery’ with an image of a fairly typical species of each genus. You can then click on this and get images of all the species in that genus. Further aids are being devel- oped. How we will access the 2,500 pyra- A male of the Mexican Rice Borer, Eoreuma loftini (Dyar).
4 The Pyraloid Planet 8 - 2014 center. Adults fly to sugar bait and foul- North American Moth Photographers’ Group, southeast Asia feeding on ferns, a new spe- smelling Tephritidae traps, and I suspect that we would get somewhere. In the meantime, cies of Diatraea from the U.S., and new mor- the larvae are in stems of banana plants. I please ask if you want a photo of anything. phological keys to the identity of Diatraea depend greatly on the expertise of all my species. I plan to visit the Natural History international colleagues to figure out these Museum in London in August, 2014 to solve adventive species. some of the problems uncovered during the Alma Solis type pyraloid database project. Unfortunately The most serious introduction, and for it is not up on the web yet, but I am more me the most time-consuming, is the haim- As of October 1, 2013 I stepped back into than willing to share type database informa- bachiine Eoreuma loftini (Dyar), the Mexican my research from being Research Leader for tion with anyone with a research project (just Rice Borer. A polyphagous stem-borer in the Systematic Entomology Laboratory. In send me an e-mail). I have also been busy Poaceae, it is probably the worst insect pest addition, my colleagues of many years, Mike working on the writing of my next 5-year pro- of sugarcane in North America, and Florida Pogue and John Brown, retired in 2014. This ject; I will be proposing to finish the Costa has the largest cane industry within the transition period has been easy and I am Rican pyraloid project with D. Janzen and W. U.S. It has been in Texas since 1980, but its located at the National Museum of Natural Hallwachs. appearance in 2012 in north-central Florida History in Washington, DC, all the time now, surprised me. Survey by FDACS-DPI and but I have also lost some of my technical Last fall I was invited by Annette Aiello to University of Florida has delimited the popu- support. So Mark Metz and I worked hard to present two talks at the Smithsonian Tropi- lation’s extent. Since the larvae are hard to finish some projects, primarily in the genus cal Research Institute in Panama entitled control, I plan for long-term monitoring as it Diatraea and the pyraloid type database “Chasing Snout Moths: A Woman’s Quest for spreads southward. Meanwhile, I struggle of the National Museum of Natural History, Discovery in the Neotropics” in Panama City, with haimbachiine systematics. The North Smithsonian Institution. He was able to get and a BAMBI seminar on Barro Colorado American genera were revised by Capps many of the genitalia dissections that he Island entitled “Snout Moth Feeding and (1965) and Klots (1970), but they are a sam- did on slides, primarily those of type speci- Habits (Lepidoptera: Pyraloidea).” ple of a poorly understood global tribe. mens. I have managed to get some tempo- rary support to continue this with non-type In case you missed it, I submitted a blog The online ID Tool “Microlepidoptera on specimens. on sloth moths on the NMNH Department of Solanaceae” was published in September Entomology Blog fueled by a recent paper by 2013 at
I have volunteers databasing the pyra- loids of the late Dale Habeck. He was a University of Florida professor who raised Lepidoptera from the 1960s through the 1990s. There are many new host records and larval specimens. The long-term plan is to publish summaries of the records for each family group; Drs. Deborah Matthews and Charles Covell have done the Geometridae. Because aquatic weeds are a major problem in Florida, Prof. Habeck’s Acentropinae are a major focus.
I continue to make diagnostic dissection slides for the FSCA/MGCL. Time allowing, I want to put photographs of them on line. In the spirit of Pierce and Metcalfe, North Ame rica needs more images of pyraloid genitalia to serve regulatory identification and supple- ment revisionary works. The fauna’s large size and incomplete study would obviously require a piecemeal approach, but if regional collections could pool carefully identified Volunteers Nick Silverson (left) and Alvira Hasan working on incorporating the Ron Leuschner collection slides of important taxa on a site such as the into the main USNM Pyraloidea collection.
The Pyraloid Planet 8 - 2014 5 of the Leuschner donation of over 11,000 University. As a graduate student of Dr. Kyu- at Colorado State University (CSU) focused pyraloids. I needed to utilize the space for Tek Park, she received her M.Sc. degree with primarily on the systematics and identifica- the Pyraloidea of Costa Rica project (we also her thesis entitled “Systematics of Subfamily tion of economically important tortricids. received the reared material of Janzen and Gelechiinae in Korea.” Sangmi received her Hallwachs from the Guanacaste Conserva- Ph.D. under Dr. Richard L. Brown at Missis- Todd currently works as a Research Sci- tion Area in Costa Rica) and I had a number sippi State University with her dissertation entist in the Department of Bioagricultural of students available to work on this incor- entitled “Systematics of Holarctic genera of Sciences and Pest Management at CSU. poration. I also worked on material that has Teleiodini (Lepidoptera: Gelechiidae).” The majority of his research involves pro- been slowly donated by Kelly Richers and ducing morphological and molecular identi- Bo Sullivan. My office also had a variety of Sangmi has specialized on Gelechiidae fication resources for invasive Lepidoptera material that accumulated over the years for the past 15 years, and has published in conjunction with the USDA-APHIS-PPQ- and needed to be incorporated into the main 25 scientific papers and 6 non-refereed S&T Identification Technology Program collection. identification aids on Gelechiidae and other (ITP) located in Fort Collins. Todd has microlepidoptera. She also made many authored or coauthored more than 20 peer- presentations at regional, national, and reviewed publications, including a book on Houhun Li international meetings. Since 2002 she has olethreutine moths. He maintains a website curated and identified gelechiids in many dedicated to tortricids and has performed The Fauna of China chapter on Pyralidae, collections in North America, becoming one extensive field work across North America, Phycitinae by Houhun Li, Yindang Ren and of the foremost experts in this difficult group. Europe, Australia, and Africa. He received Hongxia Liu, has been completed with the Sangmi developed the most comprehensive numerous awards for his entomological support of the National Natural Science website on Gelechiidae which includes a work, the most notable including CSU’s Foundation of China. A total of 503 species global framework for phylogenetics and clas- inaugural University Distinguished Profes- in 116 genera and three tribes are described sification of Gelechioidea. She contributed sors Scholarship (2012), the Entomological in detail, including new species, new educational videos on collecting and dissect- Society of America’s John Henry Comstock synonyms, and new combinations. Images ing microlepidoptera that are available on Award (2011), and the USDA-APHIS-PPQ of the venation for each genus and of the YouTube. Sangmi is the Collection Manager Deputy Administrator’s Safeguarding Award habitus and genitalia for each species are of the Hasbrouck Insect Collection at Arizona (2011). Todd currently serves as President of provided along with distribution maps. The State University, since 2012, and she serves the Lepidopterists’ Society, and he is web- manuscript is now with the editor. as a referee to the Moth Photographs Group master for the Foundation’s website. (MPG) site for identifications of gelechiids. Everyone is invited to visit the Founda- The Wedge Entomological Todd was born and raised in a small town tion’s website (http://www.wedgefounda- in northern Ohio. He became interested in tion.org). Other board members include: Research Foundation Lepidoptera at an early age, thanks to his John Brown, Oliver Dominick, Larry Gall, father, who was a high school chemistry Don Lafontaine, Ron Hodges, Eric Metzler, Ron Hodges, President of the Wedge teacher. Summers would involve rearing sat- Jackie Miller, Paul Opler, Kelly Richers, and Entomological Research Foundation, urniid moths and traveling the state collecting David Wagner. The Board welcomes com- announced the addition of two new mem- butterflies, moths, and other insects. Todd munication from all who are interested in the bers of the Foundation’s Board of Directors. joined the Ohio Lepidopterists at around age Foundation’s activities. Dr. Sangmi Lee and Dr. Todd Gilligan both 10, and was very active in that organization accepted appointments to the board. The for the next 20 years. He began collecting For further information, contact Eric term of each member is 5 years with the micro moths in the early 1990’s during the Metzler, [email protected]. option of renewal. height of the Ohio Survey of Lepidoptera, and eventually became interested in moths Ron, a founding member, tracks his in the family Tortricidae, which remain his involvement in the Foundation back to late specialty. His undergraduate education 1960s. Ron said “What began as a larval began at Ohio Northern University and con- Book review organization eclosed into a mature force in tinued at Ohio State University (OSU) where the study of moths of the World. Todd and he earned a bachelor’s degree in entomol- Sangmi will provide continuity for many ogy. His original plans to attend graduate František Slamka 2013. Pyraloidea (Lepi- years to come.” school were postponed when his wife moved doptera) of Europe, Volume 3: Pyraustinae from Sydney, Australia to Ohio, and he spent & Spilomelinae. Identification - Distribution - Sangmi Lee was born and raised in South the next eight years in the field of computer Habitat - Biology. - Bratislava, 357 pp. - ISBN Korea. She started learning about insects systems administration. Deciding that study- 978-80-969052-8-7. when she was a young girl who chased ing moths was better than being employed, grasshoppers so she could eat their fried he left the computer world and returned to In his third volume of “Pyraloidea of legs. Her fascination with moths led her to OSU to obtain a M.Sc. in entomology. In Europe”, published in October 2013, Fran- concentrate on microlepidoptera during her 2007 he moved with his wife and two dogs to tisek Slamka focuses on Pyraustinae and undergraduate program at Kangwon Nat’l Colorado to pursue a Ph.D. His dissertation Spilomelinae, two of the most diverse groups
6 The Pyraloid Planet 8 - 2014 among snout moths. This fact is mirrored in cal boundaries of Europe, but are found in dopagyda without investigating the type the extent of the book: with a total of 357 North Africa and the Near East. Although species of the presumably related genera pages it is by far the most voluminous of the information on these species is undoubt- Pagyda Walker, 1859 and Paliga Moore, series. As in Volume 2, it is written in English edly interesting, their inclusion “because of 1886. This is in opposition to the attempts only, therefore reaching a broad readership. their similarity to European species” (p. 7) towards nomenclatorial stability, one of the The introduction includes a short outline is beyond the scope of this book, impairing prime objectives of the International Code on relevant literature, a list of nomenclato- its overall consistency and comprehensibil- on Zoological Nomenclature. In addition, rial changes, illustrated basic terminology ity. Besides, other European species have Slamka proposes to transfer Pionea acutan- of genitalia, a passage on the basic biol- similar non-European species that are gulata Swinhoe, 1901, Paliga contractalis ogy of Spilomelinae and Pyraustinae, and a not incorporated in the book, so Slamka’s Warren, 1896, and Botys aureolalis Lederer, paragraph on the species distribution maps. selection of non-European species remains 1863 to Pseudopagyda even though Bän- The distribution maps differentiate between unclear. A number of infrequently inter- ziger states that the latter two species “prob- six statuses: 1) confirmed for the country/ cepted tropical or subtropical (pest) species, ably each belong to a different genus [than to state, 2) confirmed for the region of country/ such as Glyphodes stolalis Guenée, 1854, his proposed combination with Microstega]” state, 3) not recently (since 1980) confirmed, Maruca vitrata (Fabricius, 1787), and Zebro- (Bänziger, 1995: p. 267). Ultimately, most of 4) occurrence questionable, 5) migrant, 6) nia phenice (Stoll in Cramer & Stoll, 1782) the 188 valid genus group names of Pyraus- introduced. The introduction also contains a complement the information on the autoch- tinae are in need of revision, and with Pseu- short paragraph on DNA analysis for taxo- thonous and established fauna of Europe. dopagyda, another genus name now needs nomic and phylogenetic purposes. to be taken into consideration. These kinds Slamka establishes an extensive number of descriptions are of no additional value to The book is well-manufactured: the of nomenclatorial changes: 40 new syn- the field of systematics. robust high gloss hardcover encloses the onyms, 7 new or revised statuses, and 8 solidly bound pages printed on strong paper. new generic combinations. Furthermore, he Altogether, the seemingly random inclu- The book’s size (17 x 23.6 cm) is the same describes 5 species, 3 subspecies, 1 subge- sion of non-European species and the incon- as in volumes 1 and 2, but it is significantly nus and 1 genus as new to science. How- sequent and non-reflective descriptions of thicker (3 cm). All illustrations are sharp ever, only one of the five newly described new names make this book unnecessarily and clear and the colour plates display the species, Udea uralica Slamka, 2013, is dis- inconsistent. Nonetheless, I find it to be a body and right wings of well-set, evenly tributed within the European region. very valuable book for the identification of illuminated adults with naturally appear- adults of European spilomelines and pyraus- ing colours. The depicted genitalia are well The many problems that still await revi- tines. It is only just handy enough to be taken cleaned and of appropriate contrast, and sion even for the well-studied European along on field trips. The numerous images important diagnostic characters, e.g. cornuti fauna become well apparent when looking depicting the variation in the maculation of in the phallus, are magnified and without the at Metasia: One new subgenus, two new adults are of great value and the high-quality vesica in the background. In the majority of species, one new subspecies, and eleven genitalia illustrations allow identification cases multiple adults and genitalia are illus- new synonyms are among the changes even of difficult groups through dissection. trated for each species, giving the reader an established (yet the new subgenus and both idea of the variability within the respective new species do not occur in Europe). Apart species. The text part on description, biology from the two new species, Slamka mentions Richard Mally and distribution of the species is consist- four potentially new species in the section ently two-columned as in volumes 1 and 2, “Unplaced Metasia spp.” on page 121. One with the square distribution maps spanning may wonder why he didn’t describe them as one column in breadth. All species are con- well, because these and the new species secutively numbered and the illustrations fol- Metasia arenbergeri and M. rebeli are all First Red List of low the same numbering system, therefore based on only one single specimen each. In German Pyraloidea greatly easing the location of the respective my opinion, a careful, comprehensive revi- adult and genitalia images on the plates. sion of this complex genus would have been The 110 grey-scale images in the text part, more useful, instead. In 2012, the first and official Red List of illustrating interspecific differences of closely German Pyraloidea was published. Long related species, are numbered consecutively Slamka also describes a new genus been neglected from the list of flagship throughout the text, not matching the spe- of Pyraustinae: Pseudopagyda Slamka, species in nature conservation, Pyraloidea cies numbers, but they are always arranged 2013. According to Bänziger (1995) the have been the subject of an important along with the species under discussion, so correct generic placement of homoculorum increase in knowledge during the last their association is straightforward. Bänziger, 1995, the type species of the new three decades. This holds especially true genus, is not clear. Despite the comment for Central Europe, where numerous Of the 220 species treated in this issue, that a “comprehensive revision of all related publications are now available for species 98 species in 17 genera are Pyraustinae and genera is needed to clarify their status” identification. A growing number of collectors 122 species in 34 genera are Spilomelinae. (Bänziger 1995: p. 267) (a comment which who had traditionally restricted themselves Almost 20% (40 species) of these 220 spe- Slamka even cites in his book (p. 30)!), he to Macroheterocera and Papilionoidea, also cies are not distributed within the geographi- decides to describe the new genus Pseu- take attention now to these exciting moths.
The Pyraloid Planet 8 - 2014 7 Ostrinia quadripunctalis (Denis & Schiffermüller, 1775), a congener of the European Corn Borer, is critically endangered in Germany. It is only known from two spots, has been recorded at one of these spots only by the sighting of one moth, and the larval host plant is unknown. Photo: Peter Buchner.
As a result, our knowledge on the frequency have died since. Certainly, we need more planned systematic collections and studies of occurrence of species has increased young experts for field work, research, and on the Pyraustinae and Spilomelinae fauna and allowed to prepare the first Red List of the conservation of pyraloids. of far flung forests of this vast country. My German Pyraloidea. The method applied approach is basic alpha taxonomy, based on basically requires that one is able to judge A pdf of the Red List is available upon wing venation and genitalia. I find Pyraloid the current frequency of occurrence and request. Planet a great platform to interact with compare it to long term (about 100 years) and eminent scholars working throughout the short term (about 20 years) developments. Matthias Nuss globe; it is a noble interactive initiative for The data were edited and analyzed via an students and teachers as well. With an open automated spreadsheet, provided by the heart I welcome critiques and suggestions Federal Agency for Nature Conservation. by fellow scholars, so kindly contact me at As a result, 273 pyraloid species are [email protected]. regarded as permanently occurring in Budding pyraloid Germany, of which 18 species are exotics. specialists Harsimran Singh Among the 255 autochthonous species, 44.7% are on the Red List, seven of them being extinct and 14 critically endangered. I am doing my doctoral thesis on the These proportions are more or less similar taxonomy of Pyraustinae and Spilomelinae to proportions known from other groups of of the Western Ghats and North-East organisms. Unfortunately, our knowledge is India. Through this research initiative I am more limited the rarer the species are. This addressing questions regarding the species holds especially true for some of the extinct composition of both of these biodiversity hot and critically endangered species. For some spots. I am focusing on species complexes of them, we do not even know the larval food as I believe that many species are hidden plants. In this context, the Red List points to under so-called widespread species. I the necessity of research in order to acquire emphasize biological species (recognized by the knowledge needed for the conservation reproductive isolation) based on genital and of species. In ten years, the next edition of other morphological characters. the Red List will have to be prepared. How Western Ghats and North-East India much will be achieved until then, in theory are two ‘lost worlds’, i.e. some of the as well as in practice? And, will we be able last, least-explored places on Earth. My to prepare the next list? The first Red List of PhD supervisor, Prof. (Dr.) Jagbir Singh, German Pyraloidea was made possible by a Head of the Department of Zoology and team of 26 experts across Germany. When Environmental Sciences, did his PhD thesis the list was published, half of them were on Pyraustinae of North-East India. After a already (much) over 60 years old and two gap of three decades my guide and I have
8 The Pyraloid Planet 8 - 2014 Introducing Life history and food plants I found Nacoleia commixta and Piletocera sodalis feeding on various leaf molds or I aim to provide an exhaustive analysis of detritus. These species are easily reared My name is NAKA Hideshi, an entomolo- the female sex pheromones of Pyraustinae with the widely-used artificial diet Insecta gist studying moth sex pheromones, mating and Spilomelinae species. To this end I have LFS (Nosan Co.). behaviour and pest control. I studied mainly collected female adults and obtained eggs female sex pheromones and mating behav- of various species, for which sometimes the Last September I collected many rolled iour of Sesiidae species (e.g., Naka et al., food plant was unknown. leaf shelters of Broussonetia kazinoki 2013). (Moraceae). I expected that Glyphodes For example, the beautiful Goniorhyn- spp. would emerge from these shelters, but I started to work on Pyraustinae and Spi- chus butyrosus is common in Japan, but surprisingly, Syllepte taiwanalis, which was lomelinae two years ago and to identify the no reports are available on its food plant. I thought to feed on Ampelopsis glandulosa, female sex pheromones of some species. thought that they may feed on Rubiaceae or emerged in the rearing cages. Though there are many research papers on Caprifoliaceae because the closely related the female sex pheromones of Pyraustinae G. exemplaris feeds on the Rubiaceae and Spilomelinae species, pheromone lures Paederia scandens. At first I tried to find the impregnated with synthetic pheromones can- larvae of G. butyrosus on various Rubiaceae Portable cases of Paratalanta pandalis not attract as many males as virgin females. and Caprifoliaceae, but I failed to find any. larvae One day I obtained two G. butyrosus females Just lately, our research was published in near my laboratory. When I put a Paederia Presently I am rearing Japanese Para- the Journal of Chemical Ecology. Omphisa scandens leaf into a cage with the females, talanta pandalis (synonym: Microstega jes- anastomosalis is a serious pest of sweet they laid many eggs on the leaf. Hatched sica) with the artificial diets mentioned above potato in tropical or subtropical Asia-Pacific larvae fed on this plant, and finally I easily using Artemisia and basil leaf powder. Last regions. Wakamura et al. (2010) already obtained many adults. Of course I already September I collected a female and it laid identified the female sex pheromone of identified their female sex pheromone many egg clusters. At first I thought that they this species including a novel compound components, and now I prepare some field fed on woody bamboos because a related E10,E14-16:Ald, but the attractiveness of attraction tests using synthetic pheromones. species, Demobotys pervulgalis, feeds on the synthesized pheromone did not effec- various bamboos. But the emerged cater- tively attract male moths in the field. My Likewise I found food plants of the follow- pillars did not eat bamboo leaves. Finally I collaborators and myself newly identified ing species: Nomis albopedalis, Circobotys found that they feed on various Asteraceae Z3,Z6,Z9-23:H as a female sex pheromone aurealis and C. nycterina which feed on and Lamiaceae. This species is highly inter- component from crude extract of female various woody bamboos, and Pyrausta uni- esting in that larvae make portable cases sex pheromone glands, and the pheromone punctata on Isodon trichocarpus. Besides, from leaves like bagworm moths (Psychi- lure containing this compound and already- reported compounds well attracted male moths (Yan et al., 2014).
Mass rearing by artificial diets
To identify the female sex pheromone of various Pyraustinae and Spilomelinae spe- cies, we regularly breed over 15 species using artificial diets prepared from dry leaf powder of their food plants that were quickly dried with silica gel and a widely-used pow- der for insect artificial diets (Insecta F-II, Nosan Co.).
Unfortunately, we have not succeeded in breeding vitis-feeding species such as Herpetogramma luctuosale or Syllepte pal- lidinotalis with artificial diets. However, some closely related species (e.g., H. submar- ginale, which feeds on various Asteraceae) were easily reared with the artificial diet.
The portable case of the larva of Paratalanta pandalis (Hübner, 1825).
The Pyraloid Planet 8 - 2014 9 dae). Larvae cut off a leaf into a disk, roll it Ando T (2013) Identification of the sex pheromone of the sweet potato vine borer into a tube and close both ends. They never pheromone secreted by Synanthedon tenuis moth: identification and field evalation ofa feed on the portable case itself. When larvae (Lepidoptera: Sesiidae). Appl. Entomol. Zool. tricosatriene. J. Chem. Ecol. in press. are mature, they close the portable case and 48(1): 27-33. pupate in it. Recently I realized that UKmoths (http://ukmoths.org.uk) already described Wakamura S, Ohno S, Arakaki N, Kohama T, that the European Bordered Pearl also feed Haraguchi D, Yasui H (2010) Identification on Lamiaceae and makes a portable case. I and field activity of the sex pheromone “Membership” list would be grateful to know of other portable- component of the sweetpotato vine borer case-making Pyraustinae species. moth Omphisa anastomosalis (Lepidoptera: David Agassiz Crambidae) Appl. Entomol. Zool. 45(4): The Garden House, Stafford Place References 635-640. Weston-super-Mare BS23 2QZ UNITED KINGDOM Naka H, Suzuki T, Watarai T, Horie Y, Mochizuki Yan Q, Vang LV, Khanh CNQ, Naka H, Ando T e-mail: [email protected]; F, Mochizuki A, Tsuchida K, Arita Y, (2014) Reexamination of the female sex [email protected] Stacey Anderson Entomology Technician NAQS - AQIS Darwin PO Box 37846, Winnellie, NT 0821 AUSTRALIA e-mail: [email protected]
J. E. F. Asselbergs Neerland 20 4614 GD Bergen-op-Zoom NETHERLANDS e-mail: [email protected]
Yang Seup Bae Incheon University Incheon, KOREA e-mail: [email protected]
George J. Balogh 6275 Liteolier Street Portage, Michigan 49024-2394 U.S.A. e-mail: [email protected]
Hans Bänziger Department of Entomology Faculty of Agriculture Chiang Mai University Chiang Mai 50200 THAILAND e-mail: [email protected]
Alejandro Barro Dpto Biología Animal y Humana Facultad de Biología Universidad de La Habana Calle 25 # 455 entre J e I Vedado CP 10400 La Habana CUBA e-mail: [email protected]
Graziano Bassi Via San Martino 25 I-10051 Avigliana (TO), Italy e-mail: [email protected]
Male and female of Paratalanta pandalis (Hübner, 1825).
10 The Pyraloid Planet 8 - 2014 Xicui Du College of Plant Protection Request for identification Southwest University Beibei District, Chongqing 400716 Pyralidae larva feeding on green seeds of pongamia pod from India. Photo by M. CHINA Shankar Murthy. If you can help to identify the species, please contact S. Murthy. e-mail: [email protected]
Marc Epstein Senior Insect Biosystematist, Lepidoptera Plant Pest Diagnostic Branch California Dept. of Food & Agriculture 3294 Meadowview Rd Sacramento, CA 95832-1448 U.S.A. e-mail: [email protected]
Guillermo Fernandez Institut Cavanilles de Biodiversitat i Biologia Evolutiva C/ Catedrático José Beltrán nº2 46980 Paterna SPAIN e-mail: [email protected]
Clifford D. Ferris 5405 Bill Nye Avenue, R.R. #3 Laramie, WY 82070 U.S.A. e-mail: [email protected]
Reinhard Gaedike Florusstraße 5 53225 Bonn Franziska Bauer Fuqiang Chen GERMANY Sektion Lepidoptera Institute of Zoology e-mail: [email protected] Museum für Tierkunde Chinese Academy of Sciences Senckenberg Naturhistorische Sammlungen 1, Beichen West Road, Chaoyang District Barry Goater Dresden Beijing 100101, P. R. CHINA 27 Hiltingbury Road “The Ridge” Königsbrücker Landstraße 159 e-mail: [email protected] GB-SO53 5SR Chandlers Ford (Hampshire) 01109 Dresden UNITED KINGDOM GERMANY José Clavijo e-mail: [email protected] e-mail: [email protected] Museo del Instituto de Zoología Agrícola Facultad de Agronomía Kurt Grimm Vitor O. Becker Universidad Central de Venezuela Fruthwilerstrasse 65d Reserva Serra Bonita Apartado 4579, C.P. 2101-A 8272 Ermatingen P.O. Box 001 Maracay (Aragua) SWITZERLAND 45880-970 Camacan VENEZUELA e-mail: [email protected] BRAZIL e-mail: [email protected] e-mail: [email protected] Chris Grinter Alexandre-Pierre Cotty Denver Museum of Nature & Science Rebecca Bennik Avenue Maurice Troillet 160 2001 Colorado Blvd. New Zealand Arthropod Collection (NZAC) 1950 Sion Denver, Colorado 80205 Landcare Research SWITZERLAND U.S.A. Private Bag 92170 e-mail: [email protected] e-mail: [email protected] Auckland 1142 NEW ZEALAND Willy De Prins Christian Guillermet e-mail: [email protected] Dorpstraat 401B 11 Ruelle des Amandiers 3061 Leefdaal Garbejaire 108 Richard L. Brown Belgium 06560 Valbonne Sophia Antipolis Mississippi Entomological Museum e-mail: [email protected] FRANCE Mississippi State, MS 39762 e-mail: [email protected] U.S.A. Julian P. Donahue Email: [email protected] Natural History Museum of John Hawking Los Angeles County Murray Darling Freshwater Research Centre Alain Cama 900 Exposition Boulevard La Trobe University 24, rue des Parfaits Los Angeles University Drive 37140 La-Chapelle-sur-Loire California 90007-4057 PO Box 991 FRANCE U.S.A. Wodonga, Vic 3690 e-mail : [email protected] e-mail: [email protected] AUSTRALIA e-mail: [email protected]
The Pyraloid Planet 8 - 2014 11 James E. Hayden Peter Huemer Jean-François Landry Curator of Lepidoptera Tiroler Landesmuseum Naturwissenschaften Agriculture and Agri-Food Canada Florida State Collection of Arthropods Feldstrasse 11a Central Experimental Farm, Neatby Bldg. FDACS, Division of Plant Industry 6020 Innsbruck 960 Carling Avenue P.O. Box 147100 AUSTRIA Ottawa (Ontario) Gainesville, FL 32614-7100 e-mail: [email protected] K1A 0C6, CANADA U.S.A. e-mail: [email protected] e-mail: [email protected] Utsugi Jinbo Department of Zoology Théo Léger Bob Heckford National Museum of Nature and Science Muséum d’histoire naturelle 67 Newnham Road Amakubo 4-1-1 Tsukuba-shi Route de Malagnou 1 GB-PL7 4AW Plympton (Plymouth), S. Devon Ibaraki, 305-0005 1208 Genève UNITED KINGDOM JAPAN SWITZERLAND e-mail: [email protected] e-mail: [email protected] e-mail: [email protected]
John B. Heppner Ole Karsholt Patrice Leraut McGuire Center for Lepidoptera and Biodiversity Zoologisk Museum Muséum national d’histoire naturelle Florida Museum of Natural History Universitetsparken 15 45, rue de Buffon University of Florida 2100 København Ø 75005 Paris P. O. Box 112710 DENMARK FRANCE Gainesville, FL 32611-2710 e-mail: [email protected] e-mail: [email protected] U.S.A. e-mail: [email protected] Gareth Edward King Houhun Li Departamento de Biología (Zoología) College of Life Sciences, Nankai University Alvaro Herrera Villalobos Universidad Autónoma de Madrid Tianjin 300071 Enlace con Investigadores C/. Darwin, 2, 28049 Cantoblanco (Madrid) CHINA UEA de Vertebrados, INBio SPAIN e-mail: [email protected] Apdo Postal 22-3100 e-mail: [email protected] Santo Domingo, Heredia Weichun Li COSTA RICA Valentina Kirpichnikova Department of Plant Protection e-mail: [email protected] Mountain-Taiga Station College of Agronomy Far Eastern Branch of Russian Academy of Jiangxi Agricultural University Robert J. B. Hoare Sciences Nanchang Economic & Technological New Zealand Arthropod Collection (NZAC) 692533 Gornotajozhnoe, Ussuri region Development Area Landcare Research RUSSIA Nanchang, Jiangxi Province 330045 Private Bag 92170 e-mail: [email protected] CHINA Auckland 1142 e-mail: [email protected] NEW ZEALAND Rajesh Kumar e-mail: [email protected] Central Muga Eri Research & Training Institute Jiayu Liu Central Silk Board College of Life Sciences, Nankai University Ronald W. Hodges Ministry of Textiles, Govt. of India Tianjin 300071 85253Ridgetop Drive P.O. Ladoigarh CHINA Eugene, Oregon 97405-9535 785 700 Jorhat, Assam e-mail: [email protected] U.S.A. INDIA e-mail: [email protected] e-mail: [email protected], Jean-Michel Maes [email protected] Museo Entomologica Terence Hollingworth AP 527, Leon 6, impasse Chopin Gregor Kunert NICARAGUA 31700 Blagnac Kunert Business Software e-mail : [email protected], jmmaes@yahoo. FRANCE Deutscher Platz 5c com e-mail: [email protected] 04103 Leipzig GERMANY Koen Maes Martin Honey e-mail: [email protected] AgroBioSys Intl. Department of Entomology Kleine Smetledestraat 192 Natural History Museum Hiroshi Kuroko Cromwell Road 9230 Wetteren London SW7 5BD JAPAN BELGIUM UNITED KINGDOM e-mail: [email protected] e-mail: [email protected], kvmaes@ e-mail: [email protected] telenet.be Bernard Landry Marianne Horak Muséum d‘histoire naturelle Richard Mally Australian National Insect Collection Route de Malagnou 1 University Museum of Bergen GPO Box 1700, Canberra, ACT, 2601 1208 Genève Natural History Collections AUSTRALIA SWITZERLAND Realfagbygget, Allégaten 41 e-mail: [email protected] e-mail: [email protected] 5007 Bergen NORWAY Robin Howard e-mail: [email protected] Las Descargues 46210 Gorses FRANCE email: [email protected]
12 The Pyraloid Planet 8 - 2014 Edda Martinez Valentin Nidergas Amanda Roe Mississippi Entomological Museum Le Castelnau N°4 Systematics and Evolution Box 9775 Maltaverne CW315 Biological Sciences Bldg Mississippi State, MS 39762 73390 Châteauneuf University of Alberta U.S.A. FRANCE Edmonton, Alberta T6G 2E9 e-mail: [email protected] e-mail : [email protected] CANADA e-mail: [email protected] Eric Metzler Matthias Nuss P.O. Box 45 Staatliche Naturhistorische Sammlungen Daniel Rubinoff Alamogordo, New Mexico 88311-0045 Museum für Tierkunde 310 Gilmore Hall U.S.A. Königsbrücker Landstr. 159 Dept. of Entomology e-mail: [email protected] 01109 Dresden University of Hawaii GERMANY 3050 Maile Way, Honolulu Wolfram Mey e-mail: [email protected] Hawaii 96822-2231 Museum für Naturkunde U.S.A. Humboldt-Universität Eivind Palm e-mail: [email protected] Invalidenstr. 43 Sejerslevvej 14 10115 Berlin Sdr. Sejerselv Michael Sabourin GERMANY 6280 Højer 630 Beaver Meadow Rd. e-mail: [email protected] DENMARK Marshfield, VT 05658 e-mail: [email protected] USA Joël Minet e-mail: [email protected] Muséum national d’histoire naturelle Eugenie Phillips 45, rue de Buffon COSTA RICA Akio Sasaki 75005 Paris e-mail: [email protected] 11-5, Onoba 5 FRANCE Akita City e-mail: [email protected] Colin W. Plant Akita Pref., 010-1424 14 West Road, JAPAN Andrew Mitchell Bishops Stortford e-mail: [email protected] Agricultural Scientific Collections Unit, OAI Hertfordshire NSW Department of Primary Industries CM23 3QP Brian Scholtens Forest Rd ENGLAND Biology Department Orange NSW 2800 e-mail: [email protected] College of Charleston AUSTRALIA 66 College Street e-mail: [email protected] Jerry A. Powell Charleston, South Carolina 29424-0011 Essig Museum of Entomology U.S.A. Charlie Mitter 201 Wellman Hall e-mail: [email protected] Department of Entomology University of California 4112 Plant Sciences Building Berkeley, CA 94720 Rob Schouten University of Maryland U.S.A. Museon, Dept. of Natural History College Park, Maryland 20742 e-mail: [email protected] Stadhouderslaan 41 U.S.A. 2517 HV Den Haag e-mail: [email protected] Mu-jie Qi NETHERLANDS Department of Life Sciences e-mail: [email protected] Shankar Murthy Animal Diversity Laboratory Department of Agricultural Entomology University of Incheon Christian H. Schulze UAS, Raichur Incheon, 406-772 Department für Populationsökologie, IECB / College of Agriculture KOREA Universität Wien Bheemarayanagudi - 585 287 e-mail: [email protected] Althanstr. 14 INDIA 1090 Wien e-mail: [email protected] Rahul Ranjan AUSTRIA Zoological Survey of India e-mail: [email protected] Hideshi Naka Gangetic Plains Regional Center Associate Professor, PhD Sector-8, Bahadurpur Housing Colony Andreas Segerer Lab. Applied Entomology Patna-8000 026,Bihar Zoologische Staatssammlung München Fac. Agriculture, Tottori Univ. INDIA Münchhausenstr. 21 Koyama-Minami 4-101 Tottori e-mail: [email protected] 81247 München Tottori, 680-8553 GERMANY JAPAN Yingdang Ren phone: +49-89-8107-151; FAX: +49-89-8107-300 e-mail: [email protected] Institute of Plant Protection e-mail: [email protected] Henan Academy of Agricultural Science Herb H. Neunzig Zhengzhou 450002 Jay Shaffer Department of Entomology CHINA Department of Biology-3E1 North Carolina State University e-mail: [email protected] George Mason University Raleigh, North Carolina, 27695-7613 4400 University Drive U.S.A. Fairfax, Virginia 22030-4444 e-mail: [email protected] U.S.A. e-mail: [email protected]
The Pyraloid Planet 8 - 2014 13 P. R. Shashank Hari Sutrisno David L. Wagner Scientist (Agricultural Entomology) LIPI - The Indonesian Institute of Sciences Department of Ecology and Evolutionary Biology Division of Entomology Zoological Division University of Connecticut Indian Agricultural Research Institute Research Center for Biology Storrs, CT 06269 New Delhi - 110 012 PO Box 25, Cibinong 16911, Bogor U.S.A. INDIA INDONESIA e-mail: [email protected] e-mail: [email protected] e-mail: [email protected] Terry Whitaker Ayuna A. Shodotova Stephen Sutton 4 Crowtrees, Low Bentham Institute of General and Experimental Biology Borneo Books Lancaster LA2 7EE Siberian Branch of the Russian Academy of PO Box 13908 UNITED KINGDOM Sciences 88845 Kota Kinabalu, Sabah e-mail: [email protected] Sakhyanovoi Street 6, Ulan-Ude, 670047 MALAYSIA RUSSIA e-mail: [email protected] Chunsheng Wu e-mail: [email protected] Institute of Zoology Kevin Tuck Chinese Academy of Sciences Thomas J. Simonsen Department of Entomology Beichen West Road, Chaoyang District Department of Entomology Natural History Museum Beijing 100101 The Natural History Museum Cromwell Road P. R. CHINA Cromwell Road, London London SW7 5BD e-mail: [email protected] SW7 5BD UNITED KINGDOM UNITED KINGDOM e-mail: [email protected] Hiroshi Yamanaka e-mail: [email protected] 4-18, Eiraku-cho Peter Ustjuzhanin Toyama City Harsimran Singh (Dino) Engelsa Str., 23, app. 106, Toyama Pref. # 177 Charan Bagh,Patiala Novosibirsk 63005 930-0853 JAPAN Punjab 147001 RUSSIA e-mail: [email protected] INDIA e-mail: [email protected] e-mail: [email protected] Zhaofu Yang Erik van Nieukerken Canadian Centre for DNA Barcoding Frantisek Slamka Naturalis Biodiversity Center Biodiversity Institute of Ontario Racianska 61 Dep. of Terrestrial Zoology University of Guelph 83102 Bratislava PO Box 9517 Guelph, ON, N1G 2W1 SLOVAQUIA 2300 RA Leiden CANADA e-mail: [email protected] NETHERLANDS e-mail: [email protected] e-mail: [email protected] M. Alma Solis Shen-Horn Yen SEL, USDA, Smithsonian Institution Héctor Vargas Department of Biological Sciences P.O. Box 37012 Facultad de Agronomía National Sun Yat-Sen University National Museum Natural History Universidad de Tarapacá Kaohsiung 804 E-517, MRC 168, Washington CASILLA 6D TAIWAN DC 20013-7012 Arica, CHILE e-mail: [email protected] U.S.A. e-mail: [email protected] e-mail: [email protected] Ping You Francesca Vegliante Institute of Zoology, Shaanxi Normal University Wolfgang Speidel Staatliche Naturhistorische Sammlungen, Xi’an 710062 Museum Witt Museum für Tierkunde CHINA Tengstr. 33 Königsbrücker Landstr. 159 e-mail: [email protected] 80796 München 01109 Dresden GERMANY GERMANY Dandan Zhang e-mail: [email protected] e-mail: [email protected] Institute of Entomology, Sun Yat-sen University Guangzhou, Guangdong 510275 Felix Sperling CHINA Department of Biological Sciences e-mail: [email protected] University of Alberta Edmonton, Alberta T6G 2E9 CANADA e-mail: [email protected]
14 The Pyraloid Planet 8 - 2014