Early Cretaceous) Ammonites from the Aralar Mountains, Basque-Cantabrian Basin, Northern Spain

Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

The Early Aptian (Early Cretaceous) ammonites from the Aralar Mountains, Basque-Cantabrian Basin, Northern Spain

1 Seyed Naser RAISOSSADAT

Abstract: The Aralar Mountains located in northern Spain exposes a 983-m-thick succession of sediments of Early Aptian age. The lithological succession evolves from lutites, marls, and calcarenites of the Errenaga Formation to rudist micritic limestones of the Sarastarri Formation, and finally marls, lutites, and sandstones of the Lareo Formation. Based on ammonite assemblage faunas, the Deshayesites oglanlensis, D. weissi, D. deshayesi, and Dufrenoyia furcata biozones have been identified. A transition between the deshayesi and furcata zones with the co-occurrence of the ammonite genera Deshayesites and Dufrenoyia is described in the Aralar succession and is currently unique. The ammonites are described here and correlations are made with other Tethyan regions. Key Words: Ammonite; Aptian; biozonation; Basque Country; Spain; Tethys.

Citation: RAISOSSADAT S.N. (2011).- The Early Aptian (Early Cretaceous) ammonites from the Aralar Mountains, Basque-Cantabrian Basin, Northern Spain.- Carnets de Géologie / Notebooks on Geology, Brest, Memoir 2011/02 (CG2011_M02), p. 163-201. Résumé : Les ammonites de l'Aptien inférieur (Crétacé inférieur) des Monts Aralar, Bassin basco-cantabrique, Espagne septentrionale.- Le massif d'Aralar, situé dans le nord de l'Espagne, montre sur 983 m d'épaisseur une succession de sédiments d'âge Aptien inférieur. La succession litho- logique débute par les lutites, puis les marnes et les calcarénites de la Formation Errenaga, auxquelles succèdent les calcaires micritiques à rudistes de la Formation Sarastarri, et enfin les marnes, puis les lutites et les grès de la Formation Lareo. Basées sur des assemblages d'ammonites, les biozones à Deshayesites oglanlensis, D. weissi, D. deshayesi et Dufrenoyia furcata ont été identifiées. La transition entre la Zone à deshayesi et celle à furcata observée ici est un cas unique dans l'état actuel des connaissances, notamment en raison de la cooccurrence d'ammonites des genres Deshayesites et Du- frenoyia. Ces ammonites sont décrites et les corrélations avec d'autres régions du domaine téthysien sont présentées. Mots-Clefs : Ammonite ; Aptien ; biozonation ; Pays Basque ; Espagne ; Téthys. 1. Introduction nantly siliciclastic sedimentation of the Errenaga and Lareo formations with a major intercalated The Basque-Cantabrian Basin is located in limestone, the Sarastarri Limestone rich in north east Spain (Fig. 1). It is a peri-cratonic rudists, orbitolinids and corals (LERTXUNDI & GAR- rift basin related to the opening of the Bay of CÍA-MONDÉJAR, 1997; MILLÁN et alii, 2005, 2007; Biscay (MONTADERT et alii, 1974). The opening GARCÍA-MONDÉJAR et alii, 2009) (Fig. 2). was dated in the Early Aptian (MONTADERT et alii, The outcrop of these sediments extends 1979; THINON et alii, 2002), and more recently from Iribas in the east to Ataun in the west in the Early Barremian (SIBUET et alii, 2004). (Fig. 2) and they have been described in detail Four main subsidence pulses have been by GARCÍA-MONDÉJAR et alii (2009). Since these detected in the Basque-Cantabrian region du- two formations were reported to contain ammo- ring the Aptian and Albian (GARCÍA-MONDÉJAR et nites in previous studies, a careful search was alii, 2005). The very thick succession of sedi- made by the team led by Professor J. GARCÍA- ments in the Aralar Mountains span in age from MONDÉJAR of the Universidad del Pais Vasco, Bil- the Barremian-Aptian boundary to the top of bao, Spain, which resulted in the gathering of a the Early Aptian and has provided an important large amount of new material. Although most of ammonite record which underpins the geoche- the specimens are crushed, they have permit- mical results and interpretations of OAE-1a ob- ted the biozonation of these formations based tained in this succession (GARCÍA-MONDÉJAR et on the ammonite occurrences. The present alii, 2009). paper, therefore, presents a systematic descrip- The Aptian succession in the Aralar Moun- tion of these Early Aptian ammonite faunas tains (Fig. 1) has been described most recently together with their biozonation. by GARCÍA-MONDÉJAR et alii (2009). In this region, the succession consists of the predomi-

1 Geology Department, Faculty of Sciences, Birjand University, PO Box 79, Birjand (Iran) [email protected] Manuscript online since December 31, 2011 [Editors: Bruno GRANIER & Christian C. EMIG]

163 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

Figure 1: Location of the Aralar Mountain in the western part of the Basque-Cantabrian Basin in the northern margin of Iberia and south of the Bay of Biscay (modified after GARCÍA-MONDÉJAR et alii, 2009).

Apart from some records of the presence of MARTÍNEZ (1982) in Lérida, MORENO-BEDMAR Deshayesites deshayesi LEYMERIE in localized (2007) and MORENO-BEDMAR et alii (2009) in Bar- areas (e.g., Aralar: DURVENOIS et alii, 1972; celona, and AGUADO et alii (1997) in the Betic FLOQUET & RAT, 1975), the only important des- Cordillera describe Early Aptian Iberian ammo- cription of Early Aptian ammonites was made in nite occurrences outside of the present area. the Cuchía (Santander) region in the north- Although ammonite occurrences are relatively western part of the basin (COLLIGNON et alii, few in the Aralar succession and restricted to 1979). These authors identified several species certain horizons, they are sufficient to permit a of Prodeshayesites, Deshayesites, Cheloniceras more precise ammonite biozonation to be made and Vectisites corresponding to the four zones of the Early Aptian in this region than has been of the Early Aptian in England (CASEY et alii, possible hitherto. 1998) (Table 1).

Figure 2: Geological map of the Aralar Mountain showing the Errenaga, Sarastarri and Lareo formations, and sections of Iribas, Igaratza, Urkillaga-Iberondo, Eskisabel and Ataun (modified after GARCÍA-MONDÉJAR et alii, 2009).

164 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

Table 1: Correlation chart of the Early Aptian biozones in the Tethyan and Boreal Realm. 2. Outline stratigraphy an important marker in the determination of the tectonic structures of the Sierra de Aralar. The top of the non-marine Wealden series, Although no ammonites have been found in this which rests upon an Upper Jurassic succession, Formation, it is dated as Early Aptian desha- is overlain by marls with ammonites including yesi-furcata zones transition on account of the Valdedorsella sp. of latest Barremian age. These faunas in the uppermost Errenaga Formation sediments in turn underlie an ammonite and or- below and the Lareo Formation above. The bitolinid-rich marine Early Aptian succession. Sarastarri limestones indicate a stepped The detailed stratigraphical succession is descri- drowning in two different phases in the Desha- bed by GARCÍA-MONDÉJAR et alii (2009) and is yesites deshayesi-Dufrenoyia furcata transition shown in Figs. 3 - 4. Zone (MILLÁN et alii, 2007). In the lower part of a. Errenaga Formation the immediately overlying Lareo Formation at Ataun, there is a specimen of Dufrenoyia sp. The Errenaga Formation comprises shales, already of basal furcata Zone age. marls, siltstones, sandstones and marly lime- At Igaratza, assemblage 9 in the basal Lareo stones. There are significant thickness varia- Formation also contains Deshayesites, and spe- tions in an E-W transect, from Iribas (69 m) to cimen LEIZ-1 is a Dufrenoyia cf. lurensis KILIAN. Igaratza (200 m) and Ataun (482 m) (Fig. 3). Thus, the Sarastarri Limestone marks a pause There is no lateral continuity of exposure; thus in terrigenous sedimentation in the deshayesi- correlation of the three sections has been based furcata zones transitional interval. At Iribas, the on the lithologic similarity of subunits, control- Sarastarri Limestone is overlain directly by the led sedimentological changes, vertical arrange- Late Aptian San Gregorio-Artxueta Units (Fig. ment of facies associations, TOC curve trends of 3). The Formation terminates in an areally selected intervals and ammonite occurrences extensive 20 m thick micritic limestone unit (GARCÍA-MONDÉJAR et alii, 2009). containing corals and rudists (Gorrin-txabola). Various fossiliferous horizons containing It is capped by an erosional surface overlain by ammonites of the Deshayesites oglanlensis, the Lareo Formation. Deshayesites weissi, Deshayesites deshayesi zones and Deshayesites deshayesi - Dufrenoyia c. Lareo Formation furcata transition zones of the Early Aptian have The Lareo Formation is dated with ammoni- been identified and correlated (Fig. 4). tes initially as still within the Deshayesites b. Sarastarri Formation deshayesi-Dufrenoyia furcata transitional Zone and subsequently in the D. furcata Zone. This The Sarastarri Formation consists of interca- unit shows a variety of facies and it wedges out lation of limestones, marls, and marly limesto- from west to east (GARCÍA-MONDÉJAR et alii, nes that extend at least 21.5 km from east 2009). The Lareo Formation is overlain by the (Iribas section) to west (Ataun section) and Upper Aptian San Gregorio Unit which cuts thicken progressively westward, from Iribas (53 down to the Sarastarri Limestone at Iribas (Fig. m), to Igaratza (150 m) and Ataun (178 m) 3). (Fig. 3). The limestones stand as a prominent mappable feature in the field and thus provides

165 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

Figure 3: Lithostratigraphic correlation chart of the Lower Aptian sediments of the Aralar, sections of Iribas, Igaratza and Ataun, with indication of the units and ammonite assemblages described in the text. d. Upper Aptian formations width (U), ratio of umbilical width to diameter (U/D), primary ribs in the umbilical area (PR) The units overlying the Early Aptian succes- and all ribs (primary and secondary) on the sion comprise the Upper Aptian San Gregorio ventral side (SR). Because some specimens are Formation in the west, made up of marly lime- incomplete and not all dimensions can be mea- stones and marls, and the Artxueta Unit in the sured, the ratio of whorl thickness to whorl east, made up of shallow-water Urgonian lime- height (WT/WH) is also given. stones (FLOQUET & RAT, 1975; LERTXUNDI, 1997) The following abbreviations are given for (Fig. 3). They make up a broad carbonate inner localities. ATAN= Ataneta locality, IRIBAS ramp (Artxueta) grading downdip to the corres- section (Errenaga Formation), AIZMUSU= Aiz- pondent outer ramp (San Gregorio). musu locality, 2km east of the URKILLAGA-IBE- 3. Systematic description RONDO section, Ataun area (Lareo Formation), The present study stems from the syste- ER= IGARatZA section (Errenaga Formation). matic sampling of the Lower Aptian succession (material collected loose from surface debris], in the Aralar region. Every genus and species ERG= IGARatZA section (Errenaga Formation). which have been found are described briefly, (fauna collected in situ), IRI= IRIBAS section and their occurrence and distribution in the (Errenaga Formation), IMA= ATAUN section stratigraphical column of the study area is (Errenaga Formation), NAZ= Nazca Maizagi discussed where appropriate. There are some locality, 2km east of the URKILLAGA-IBERONDO genera or species new to the basin. The classi- section and 0.5km north of the Aizmusu loca- fication follows that given in the Treatise on lity, Ataun area (Lareo Formation), SANGRE= Invertebrate Palaeontology, part L (WRIGHT et ESKISABEL section, Ataun area (Lareo Forma- alii, 1996). tion), URKI= URKILLAGA-IBERONDO section, Shell measurements (in millimeters) are Ataun area (Lareo Formation), URG = Urgoxo given in the following order: Diameter (D), locality, IRIBAS section (Errenaga Formation). whorl height (WH), ratio of whorl height to dia- The palaeontological material is housed in meter (WH/D), whorl thickness (WT), ratio of the Departamento de Estratigrafía y Paleontolo- whorl thickness to diameter (WT/D), umbilical gía, Universidad del País Vasco, Bilbao

166 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

(UPV/EHU) and the assemblage and specimen series to the umbilicus. numbers apply to that institution. Discussion: The separation of some genera of the Aconeceratinae is based on the presence Suborder Ammonitina HYATT, 1889 of ribs and denticulation of the keel. However, it Superfamily Haplocerataceae seems that this division is not useful in practice ZITTEL, 1884 (WRIGHT et alii, 1996). Thus Aconeceras, San- Family Oppeliidae DOUVILLE, 1890 martinoceras, Sinzovia, Theganoceras and Subfamily Aconeceratinae SPATH, 1923 Gyaloceras are grouped as subgenera of Acone- Genus Aconeceras HYATT, 1903 ceras. The subgenus Aconeceras s.s. is known Type species: Ammonites nisus d'ORBIGNY, principally by immature (incomplete) shells or 1841 nuclei that lack the mouth-borders. Therefore, Generic characters: Involute oxycone with it is not clear whether lappets and rostrum flattened or gently convex sides that narrow comparable to those of the subgenus San- above to a hollow microscopically serrated cari- martinoceras occur, of if the edge of its peri- na or keel. Ventro-lateral shoulders are distinct stome follows the growth-lines as in Falciferella. or broadly rounded; umbilicus with angular rim Occurrence: The genus is recorded from Eu- and low steep wall; flanks almost smooth or rope, north east Greenland, Algeria, South Afri- bearing sickle-shaped, forwardly inclined striae ca, Madagascar, Australia, Argentina, Nepal and or faint flattened ribs. Suture line with narrow Antarctica (subgenus Theganoceras) (WRIGHT et trifid lateral lobes and tall lateral saddle and alii, 1996) and Iran (RAISOSSADAT, 2002). many secondary elements declining in regular

Figure 4: Stratigraphic correlation of the Errenaga Formation from Iribas to Igaratza and Ataun sections. Note the east to west thickening of the unit.

167 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

pls. 4-6, fig. 11a-c from the Aptian of St. Dizier, Subgenus Aconeceras (Aconeceras) France (collection École des Mines de Paris). HYATT, 1903 Material: AIMUSU-2, 3; URKI-11 Aconeceras cf. nisoides Description: High whorled, involute, oxyco- (SARASIN, 1893) nes, whorl section sub-fastigiate, ventro-lateral shoulder rounded but distinct, sides parallel, Fig. 5 A flattened, keel distinct. cf. 1893 Oppelia nisoides (SARASIN), p. 155, pls. iv- Measurements: v, fig. 10a-c; text-figs. 3, 5. cf. 1924 Oppelia nisoides (SARASIN): SPATH, p. 311, pl. 26, figs. 4a-b; text-fig. B9. cf. 1958 Sinzova nisoides (SARASIN): SAZONOVA, p. 129-130. cf. 1961a Aconeceras nisoides (SARASIN): CASEY, p.

125-128, pl. 26, fig. 3-5; text-fig. 41a-c. cf. 1995 Aconeceras nisoides (SARASIN): KEMPER, Discussion: A. haugi differs from A. nisus by taf. 8, fig. 4. possessing ventro-lateral shoulders. According Lectotype: SARASIN (1893, pls. iv-v, fig. 10a- to RENZ (1982), Sanmartinoceras groenlandi- c) from the Lower Aptian of the Haute-Marne cum (see CASEY, 1961b, p. 131, text-fig. 42) is (collection École des Mines de Paris). distinguished from Aconeceras haugi by more Material: NAZ-6, 7; URKI-1, 2, 3, 4, 5, 6, 8, pronounced costae, a high-serrated keel and a 9, 10, URKI-C-1 spiral depression. Description: Involute oxycone, whorls high Occurrence: France, Venezuela (RENZ, 1982) and compressed, sub-rectangular, nearly con- and England (CASEY, 1965). vex sides, ventral margin with a low keel in Distribution: In Aizmusu locality, 2km east middle part, test nearly smooth with trace of of the Urkillaga-Iberondo section, Ataun area, fine ribs. Lareo Formation and Urkillaga-Iberondo sec- Measurements: tion, Ataun area, Lareo Formation (Fig. 6). Subgenus Aconeceras (Theganoceras) WHITEHOUSE, 1926 A. (Theganoceras) sp. Fig. 5 D Material: IMA-7, 9 Description: Two crushed specimens are to hand. Ribs fine, falcate, smooth around the um- Discussion: Suture line and rib pattern are bilicus and more distinct near the venter, diagnostic for the species. A. nisoides differs Venter with a low keel. from A. nisus by possessing a wider umbilicus Discussion: WHITEHOUSE (1926) recognised and more feeble ornamentation on whorl flanks. the genus Theganoceras as a distinct genus, Occurrence: England (SPATH, 1924; CASEY, but CASEY (1961b) regarded it as subgenus of 1961a), France (SARASIN, 1893; ROMAN, 1938), Sanmartinoceras BONARELLI as does WRIGHT et Germany (STOLLEY, 1907) and Russia (SAZONOVA, alii (1996). Theganoceras is separated from 1958). CASEY (1961a, p. 126) also mentioned Sanmartinoceras by its denser and finer ribs. that the species is reported from Sweden and Occurrence: England, Germany, South Africa Zululand. (Zululand), and Antarctica (WRIGHT et alii, Distribution: Lower part Nazca Maizagi loca- 1996). lity, 2km east of the Urkillaga-Iberondo section Distribution: In Ataun section, Errenaga For- and lower part of Urkillaga-Iberondo section, mation (ammonite assemblage 1, Figs. 3 - 4). Lareo Formation (Fig. 6). Superfamily Desmocerataceae Aconeceras haugi SARASIN, 1893 ZITTEL, 1895 Family Desmoceratidae ZITTEL, 1895 Fig. 5 B-C Subfamily Pseudosaynellinae 1893 Oppelia haugi SARASIN, p. 156, pl. 4-6, fig. CASEY, 1961a 11a-c. Genus Pseudosaynella SPATH, 1923 1961a Aconeceras cf. haugi (SARASIN): CASEY, p. 128, text-fig. 40g-h. Pseudosaynella sp. 1973 Aconeceras haugi (SARASIN): KEMPER, p. 42, Fig. 5 E pl. 2, fig. 4. 1982 Aconeceras haugi (SARASIN): RENZ, p. 21-22, Material: ERG-5-4, ER-22 pl. 1, fig. 14a-b. Description: Discoidal, involute, with flatte- 1995 Aconeceras haugi (SARASIN): KEMPER, pl. 2, ned sides and sharpened venter. Umbilicus nar- fig. 4. row with distinct rim, shell smooth or with near- Type by monotypy, SARASIN, 1893, p. 145, ly simple sigmoidal rib pattern.

168 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

Measurements: (2002) believes the latter species is characterised by a different range of morphological variation in spite of overlap with that of H. feraudianus. This reasoning does not make any sense. DELANOY (1990) suggested that probably H. feraudianus and H. soulueri (COQUAND in MA- Discussion: Rib pattern is the character that THERON, 1878) represent morphotypes variation could be used for diagnosis. The costation is of a single species. very like Cleoniceras PARONA & BONNARELLI, Distribution: Eskisabel section, Ataun area, 1895. However the later genus belongs to an Lareo Formation (Fig. 6). Albian assemblage fauna, while Pseudosaynella is an Aptian form. Because of compression the Superfamily Douvilleiceratacea specimens look like Aconeceras, but their rib PARONA & BONARELLI, 1897 pattern is distinct. Specimens can be compared Family Douvilleiceratidae with P. cf. raresulcata (d'ORBIGNY, 1841) (CASEY, PARONA & BONARELLI, 1897 1961a, p. 171). Distribution: In Igaratza section, Errenaga Subfamily Roloboceratinae Formation (Fig. 7). CASEY, 1961b Genus Roloboceras CASEY, 1954 Suborder Ancyloceratina Type species: Ammonites hambrovi FORBES, WIEDMANN, 1966 1845, Early Aptian (forbesi Zone) SE England. Superfamily Ancylocerataceae Generic characters: whorl shape semi-roun- GILL, 1871 ded, blunt ribs with tubercles around the umbi- Family Ancyloceratidae GILL, 1871 licus, umbilicus deep, suture line simplified with deep ventral lobe, body chamber half whorl in Subfamily Helicancylinae HYATT, 1894 length, mouth border plain with a shallow umbi- Toxoceratoides ? sp. lical sinus. Fig. 11 I-J Discussion: CASEY (1961b) recognised the genus based on its whorl shape and tubercula- Material: SANGRE-22, 31 tion on the flanks. Description: Specimens are fragments of Occurrence: Southern England, France, crushed shafts, more or less curved; shaft with Spain (CASEY, 1961a; MORENO-BEDMAR et alii, spherical to sub-rectangular whorl section; ribs 2009). straight and single, one row of tubercles on each side of venter. Roloboceras cf. hambrovi Discussion: Curved shaft and two rows of tu- (FORBES, 1845) bercles are characteristic for identification. Tuberculation is not clear in the specimen. The Fig. 5 G specimens are not complete, and show simi- cf. 1845 Ammonites Hambrovi FORBES, p. 354, pl. larity to Tonohamites in its curved shaft. xiii, fig. 4. Distribution: San Georgio section, Lareo For- cf. 1906 Ammonites Hambrovi FROBES: SINZOW, p. mation (Fig. 6). SZIVES & MONKS (2002) descri- 162-163. bed the genus from Late Aptian-Early Albian as cf. 1930 Cheloniceras hambrovi (FORBES): SPATH, p. well. 444-445. cf. 1954 Roloboceras hambrovi (FORBES): CASEY, p. Family Hemihoplitidae SPATH, 1924 114. cf. 1961b Roloboceras hambrovi (FORBES): CASEY, p. PATH Genus Hemihoplites S , 1924 179-182, pl. 29, figs. 5-6; pl. 30, figs. 7-8; pl. Hemihoplites sp. 30, fig. 3a-b; pl. 32, fig. 5a-b; text-fig. 54, 55a-b, 57a-b. Fig. 5 F cf. 2006 Roloboceras hambrovi (FORBES): ROPOLO et Material: SANGRE-27, SANGRE-34 alii, pl. 11, fig. 2. cf. 2008 Roloboceras hambrovi (FORBES): ROPOLO et Description: Shell evolute, sides compres- alii, pl. 6, fig. 2. sed, umbilicus deep and umbilical wall vertical. cf. 2009 Roloboceras hambrovi (FORBES): MORENO- Ribs straight or slightly curved, simple or bran- BEDMAR et alii, fig. 10E. ching; primary ribs mostly single, starting at cf. 2009 Roloboceras hambrovi (FORBES): MORENO- the umbilical margin, secondary ribs intercala- BEDMAR et alii, fig. I, B-D, fig. II, A ted between primaries starting in the lower part Lectotype: British Geological Survey of the flank. Museum, collection 2295, the larger of the two Discussion: The rib pattern and whorl specimens figured by FORBES, from Atherfield section are the characteristics for genus and Clay Series, Lower Lobster Bed. species identification. The San Georgio speci- Material: ERG-3-8, 3-12 mens show similarity to H. feraudianus (d'ORBI- Description: Two crushed specimens are to GNY, 1841). This species is similar to H. vari- hand. Whorl section sub-rounded to sub- costatus RICCARDI & AGUIRRE-URRETA, 1989, in its rectangular, venter rounded, primary ribs growth and rib pattern, but AGUIRRE-URRETA straight, single, thick, between each primary there are thinner ribs. Tubercles around the

169 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02) umbilicus are prominent. sites KAZANSKY, 1914, Neodeshayesites CASEY, Discussion: The specimens show similarity to 1964, Dufrenoyia BURCKHARDT ex KILIAN & R. horridum (RIEDEL, 1938), but this species has REBOUL, 1915, Burckhardites HUMPHREY, 1949, more depressed adolescent whorls and tuber- and Kuntziella COLLIGNON, 1962. cles that are more prominent still. BOGDANOVA & MIKHAILOVA (1999) added a new Occurrence: England (CASEY, 1954, 1961c; genus Obsoleticeras BOGDANOVA & MIKHAILOVA, SPATH, 1930), France (KILIAN, 1913; ROPOLO et 1999, and used Paradeshayesites KEMPER (1967) alii, 2008), Caucasus (ROUCHADZE, 1933), Russia instead of Prodeshayesites CASEY, 1961b, and (SINZOW, 1906), USA (HUMPHREY, 1949). indicated that Neodeshayesites, Burckhardtites Distribution: Igaratza section, Errenaga For- and Kuntziella are possibly assigned to the mation (Fig. 7). family. ROBERT & BULOT (2005) have assigned Neodeshayesites, a genus restricted to south Roloboceras cf. saxbyi CASEY, 1961b and central America, to the subfamily Acantho- Fig. 5 H hoplitinae, but there are problems with this approach which will need addressing. Here the cf. 1961b Roloboceras saxbyi CASEY, p. 188-189, pl. taxonomic classification in WRIGHT et alii (1996) 30, figs. 1a-b, 2a-b; text-fig. 57c-d. is followed, but, the three genera Turkmenice- Holotype: Natural History Museum, London, ras, Deshayesites and Dufrenoyia are common 46590, Atherfield Clay Series, Lower Lobster in all classifications. However the suprageneric Bed, Atherfield, Isle of Wight. classification has been under discussion for Material: SANGRE-11, 13, 28, 30. some long time and is still unsatisfactory. Description: Whorl section sub-rectangular to oval, ribs uniform, single or bifurcate from Subfamily Deshayesitinae umbilical edge, lateral tubercles not very STOYANOW, 1949 distinct. Genus Deshayesites KAZANSKY, 1914 Discussion: SPATH (1921, p. 313, quoted in Type species: Ammonites deshayesi LEYMERIE CASEY, 1961b, p. 189) believed that R. saxbyi in d'ORBIGNY, 1841. was comparable to Cheloniceras gottschei (KI- Generic characters: Discoidal, sides and LIAN, 1902). Detailed examination shows they venter flattened or gently convex, ventro-lateral differ in ornamentation and suture line and and umbilical borders poorly defined. Involution Cheloniceras gottschei shows sharper ribs and a varies from one-fifth to a little more than one- venter more flattened venter. R. perli also third the diameter. Sculpture of sigmoidal ribs, shows similarity to R. saxbyi but differs from which tend to lose elevation at mid-flank, the latter species in its more close and uniform divisible into primaries, commencing at umbi- ribbing and less complete tuberculation (CASEY, licus, and secondaries, which are intercalated or 1961b). branch from primaries on whorl side. On venter Occurrence: England (CASEY, 1961b) ribs are of uniform relief and are bent forwards Distribution: Eskisabel section, Ataun area, in an arc: some species have a smooth siphonal Lareo Formation. band for a short at period young stage. Mouth- Superfamily Deshayesitaceae border plain, inclined forwards, sinuous, with shallow embayment at umbilicus, well-marked STOYANOW, 1949 on test especially near aperture, where it Family Deshayesitidae STOYANOW, 1949 stands out as a sheaf of hairlines. Suture line The family is restricted to the latest Barre- with trifid, fairly deep first lateral lobe and mian to the latest Early Aptian substages. Du- simplify auxiliary elements with only a gentle ring this interval, an increase in the involution retraction towards umbilicus. Body chamber of the whorls, flatting of the venter, appearance about half a whorl in length. of tubercles and a reducing curvature and wea- Discussion: The genus is derived directly kening of the ribs at the venter can be followed from the latest Barremian genus Turkmeniceras in this family. Some genera with their short TOVBINA, 1962. KAZANSKY (1914) introduced time ranges and distinct characters are used in Deshayesites with A. deshayesi d'ORBIGNY as Lower Cretaceous biozonation on a global scale. type species. Originally Deshayesites comprised Deshayesitidae is accepted as a family species which are now distributed among (WRIGHT, 1955; CASEY, 1957; MIKHAILOVA, 1958). Deshayesites, Prodeshayesites and Dufrenoyia RKELL A et alii (1957) considered that the family (CASEY, 1964). Later KEMPER (1967) proposed A comprised three genera, Deshayesites K- Paradeshayesites KEMPER, which is similar to ZANSKY URCKHARDT ILIAN , 1914, Dufrenoyia B ex K Prodeshayesites. DELANOY (1995) considered & REBOUL, 1915, and Burckhardites HUMPHREY, 'Prodeshayesites' as a subjective synonym of 1949. CASEY (1964) split the family into two Deshayesites. 'Prodeshayesites' is limited to subfamilies Deshayesitinae and Matherocera- east Greenland, England and North Germany. tinae. Subsequently new genera were proposed In both East Greenland and eastern England it RIGHT for the subfamily. W et alii (1996) followed apparently precedes true Deshayesites (CASEY, ASEY C 's subfamily suggestion at systematic 1961a, 1964; KELLY & WHITHAM, 1999). Howe- level, including genera Turkmeniceras TOVBINA, ver, there is evidence in both Greenland and 1962, Prodeshayesites CASEY, 1961b, Deshaye- England that the earliest Aptian with Deshayesi-

170 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02) tes oglanlensis BOGDANOVA is missing in terms of marine sediments (KELLY et alii, in prep.). The taxonomic status of Prodeshayesites is questio- nable; KEMPER (1995) considered it to be a synonym of Deshayesites and stressed the affinity of 'Prodeshayesites' with some Turkmenian species of Deshayesites. BOGDANOVA & MIKHAILO- Discussion: Fine ribbing and the coiling VA (1999, 2004) proposed a new genus Obsole- pattern are the characters used for recognition. ticeras which is similar to Paradeshayesites or Identification is not easy because specimens Prodeshayesites. It is suggested that Deshaye- show similarity to Turkmeniceras and D. sites be retained and all other related genera oglanlensis. In fact the species could be a grouped in its synonymy, or possibly regarded transition form between Turkmeniceras and as sub genera. The use of Paradeshayesites or Deshayesites. D. tuarkyricus differs from D. Prodeshayesites is one of publication priority. oglanlensis by its more dense rib pattern. It is Obsoleticeras cannot be separated from Desha- supposed that D. tuarkyricus has been reported yesites. Parahoplitoides SPATH (1922, p. 111) is form Turkmenistan only and is an endemic and an objective synonym of Deshayesites having local species, but based on GONNET et alii the same type species and was proposed in (2000), GARCÍA-MONDEJAR et alii (2009) and this ignorance of KAZANSKY's genus. Reference in study, it is possible that D. tuarkyricus is Canadian literature to species of Deshayesites present outside of the Kopet Dagh basin. Even (MCLEARN, 1932; WARREN, 1937) is based on of if the latter specimens are not D. tuarkyricus, the Albian genus Subarcthoplites CASEY, 1954. they accompany D. oglanlensis and are of the CASEY (1965) believed that the Colombian spe- D. tuarkyricus Zone age. cies described by RIEDEL (1938) as Deshayesites Occurrence: Turkmenistan, Iran and Spain. stutzeri, D. nodosus, D. rotundus and D. colom- Distribution: Iribas section, Errenaga Forma- bianus are most closely related to Dufrenoyia tion (Fig. 8). than to Deshayesites. Occurrence: The genus is characteristic of Deshayesites cf. oglanlensis BOGDA- the earlier Early Aptian and reported from many NOVA, 1983 places Europe, Russia, Africa, Arctic, America and Iran. Fig. 5 J Deshayesites cf. tuarkyricus num. nud. cf. 1979 Deshayesites oglanlensis BOG- DANOVA: BOGDANOVA, pl. 1, fig. 5. BOGDANOVA, 1983 cf. 1983 Deshayesites oglanlensis sp. nov.: BOGDA- NOVA, p. 136, pl. 1, figs. 5-9; text-figs. 5-6. Fig. 5 I cf. 1995 Deshayesites oglanlensis BOGDANOVA: DE- nom. nud. cf. 1979 Deshayesites tuarkyricus BOG- LANOY, p. 74, pl. 9, fig. 1. DANOVA, BOGDANOVA, tab. 2, fig. 2. cf. 1997 Deshayesites oglanlensis BOGDANOVA: cf. 1983 Deshayesites tuarkyricus sp. nov., BOGDA- AGUADO et alii, fig. 7e. OGDANOVA NOVA, p. 132, tab. 1, figs. 1-4, tab. 2, fig. 4. cf. 1999 Deshayesites oglanlensis B : cf. 1999a Deshayesites tuarkyricus BOGDANOVA: AVRAM, p. 441, fig. 4a-b. CECCA et alii, fig. 6c-d. cf. 1999b Deshayesites oglanlensis BOGDANOVA: ECCA cf. 1999 Deshayesites tuarkyricus BOGDANOVA: BOG- C et alii, pl. 1, figs. 2-4. DANOVA & PROZOROVSKY, pl. 3, fig. a-c. cf. 2000 Deshayesites oglanlensis BOGDANOVA: cf. 1999 Paradeshayesites tuarkyricus (BOGDA- GONNET et alii, pl. 1, figs. 1-2; pl. 2, fig. 4. NOVA): BOGDANOVA & MIKHAILOVA, p. 527. cf. 2004 Deshayesites oglanlensis BOGDANOVA: RAI- SOSSADAT cf. 2000 Deshayesites sp. gr. tuarkyricus BOGDA- , p. 123-124, fig. 4G-H. NOVA: GONNET et alii, pl. 1, fig. 3. cf. 2004 Paradeshayesites oglanlensis (BOGDANOVA); OGDANOVA IKHAILOVA cf. 2004 Deshayesites cf. tuarkyricus BOGDANOVA: B & M , p. 214, pl. 7, fig. 4; RAISOSSADAT, p. 123, fig. 4F. text-figs. 22-23. cf. 2004 Paradeshayesites tuarkyricus (BOGDA- Holotype: St. Petersburg Museum, N° 12/ NOVA): BOGDANOVA & MIKHAILOVA, p. 212, pl. 7, 9442. from Bolshoi Balkhan, Turkmenistan, fig. 7; text-figs. 19-20. figured by BOGDANOVA. Holotype: St. Petersburg Museum, N° 1/ Material: IRI-13, 19, 54, 55, 56, 57, 65, 76, 9442, from Lausan, Taurkyr, Turkmenistan. 87. Material: IRI-1-35, 36, 44, 83, 85, 86, 102. Description: Some specimens are compres- Description: Whorl section sub-rectangular, sed; whorl section sub-rectangular, venter venter flat or broadly convex, coiling being mo- rounded and narrow, flanks a little flattened, re open in the last whorl. Ornament consists of umbilicus is around one quarter of the diame- ribs, dense and fine, primary ribs originate from ter. Ribs fine and dense, palmate, fasciculate, the umbilical edge and are more coarser and ribs stem from well-marked peri-umbilical bulla, stronger in the upper third of the whorl flanks, their relief is weaker in the middle part of the secondary ribs start from mid flank; every pri- sides. Regularly every two primary ribs com- mary rib is accompanied by two secondary ribs. mence from one bullae, bifurcate or trifurcate, Thirty three primary ribs and seventy secondary secondaries start from mid flank. Twenty seven ribs are present at thirty five mm diameter. primary ribs and sixty one secondary ribs are Measurements: present at thirty mm diameter.

171 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

Measurements: Holotype: St. Petersburg Museum, N° 18/9442. from Bolshoi Balkhan, Turkmenistan, Lower Aptian, Deshayesites tuarkyricus Zone. Material: IRI-61, 62, 90, 95, 116, 117. Description: Whorl section sub-rectangular to oval, flanks parallel, Primary ribs nearly sharp, originating at the upper part of the umbilical wall and cross the flank in a light sigmoidal pattern, secondary ribs branch from primaries in mid flanks. Discussion: The species appears in the oglanlensis Zone and can be distinguished by its Discussion: Fine, dense ribs and peri-umbi- density of ribs which is less than in other spe- lical bullae are characteristic features of this cies which appear in this zone such as D. species. DELANOY (1995) reported the density of tuarkyricus, D. oglanlensis and D. weissiformis. ribs in Deshayesites oglanlensis as being more Occurrence: Turkmenistan, France and important in the separation of this species from Spain. Deshayesites tuarkyricus. AVRAM (1999) belie- Distribution: Iribas section, Errenaga Forma- ved that the specimens described as Deshayesi- tion. tes oglanlensis by DELANOY (1995) are more closely related to Deshayesites planicostatus Deshayesites cf. gracilis CASEY, 1964 (BOGDANOVA, 1991). The Spanish material stu- Fig. 5 L died here looks like Deshayesites callidiscus CASEY, 1961c. Review of the description of D. cf. 1964 Deshayesites cf. gracilis sp. nov.: CASEY, p. oglanlensis and D. callidiscus from the pu- 324-325, pl. 47, fig. 10. cf. 1999 Deshayesites gracilis CASEY: AVRAM, p. blished papers (for example CASEY, 1964; BOG- 444, fig. 3C. DANOVA & MIKHAILOVA, 2004) show they are mor- Holotype: Natural History Museum, London, phologically very similar. In my view they might C3034, Atherfield Clay Series, Crackers, be conspecific, despite the difference in age, or Atherfield, Isle of Wight. D. callidiscus is a descent of D. oglanlensis and Material: ER-76, 79, 108, 173, 183; ERG-3- they are phylogenetically related to each other. 9, 3-10. However, this view needs further study. Description: Specimens are crushed and in- Occurrence: Earliest Aptian in France (DELA- complete, umbilical wall vertical; Ornament NOY, 1995) and recorded in an assemblage consists of ribs; narrow primary ribs start from ammonite fauna of forbesi or weissi Zone age. middle part of the umbilical wall, sigmoidal, Deshayesites oglanlensis is a characteristic form secondaries intercalated between primaries, of the D. tuarkyricus Zone in south Spain free or attached, starting from lower third of (AGUADO et alii, 1997), Romania (AVRAM, 1999), the whorl flank, some fusing with adjacent pri- Turkmenistan (BOGDANOVA, 1979, 1983) and maries. Iran (RAISOSSADAT, 2004). Measurements: Distribution: Iribas section, Errenaga Forma- tion (Fig. 8). Deshayesites cf. antiquus BOGDANOVA, 1983 Fig. 5 K

num. nud. cf. 1979 Deshayesites antiquus BOGDA- NOVA: BOGDANOVA, tab. 1, fig. 4. cf. 1983 Deshayesites antiqnuus sp. nov. BOGDA- Discussion: CASEY (1964) based the species NOVA, p. 138-139, tab. 2, figs. 5-6; tab. 3, fig. on an incomplete specimen. The species differs 8. from D. forbesi var. koeneni CASEY, 1964 by its cf. 1999 Deshayesites antiquus BOGDANOVA: BOGDA- deep umbilical wall and steep-sided ribs. D. NOVA & PROZOROVSKY, pl. 2, fig. e. multicostatus TOVBINA, 1963, shows similarity to cf. 1999 Deshayesites antiquus BOGDANOVA: ROPOLO D. gracilis, but the former species has stronger et alii, p. 178, pl. 16, figs. 1-3; pl. 18, fig. 1. sigmoidal ribs and a flattened ventral area. cf. 1999a Deshayesites antiquus BOGDANOVA: CECCA et alii, pl. 1, fig. 1. Occurrence: England (CASEY, 1964), Roma- cf. 2000 Deshayesites antiquus BOGDANOVA: GONNET nia (AVRAM, 1999). et alii, p. 129, pl. 3, figs. 2-4. Distribution: Igaratza section, Errenaga For- cf. 2010 Deshayesites antiquus BOGDANOVA: mation (Fig. 7). MORENO-BEDMAR et alii, p. 292, fig. 11A.

172 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

Figure 5: Early Aptian ammonites of northern Spain. A.- Aconeceras cf. nisoides (SARASIN, 1893), Lareo Formation, NAZ-7; B-C.- Aconeceras haugi SARASIN, 1893, Lareo Formation, AIZMUSU 2, Lareo Formation; D.- Aconeceras (Theganoceras) sp., Errenaga Formation, IMA-7; E.- Pseudosaynella sp., Errenaga Formation, ER-22; F.- Hemihoplites sp., Errenaga Formation, Sangre-34; G.- Roloboceras cf. hambrovi (FORBES, 1845), Errenaga Forma- tion, ERG-3-8; H.- Roloboceras cf. saxbyi CASEY, 1961b, Errenaga Formation, Sangre-11; I.- Deshayesites cf. tuarkyricus BOGDANOVA, 1983, Errenaga Formation, IRI-83; J.- Deshayesites cf. oglanlensis BOGDANOVA, 1983, Errenaga Formation, IRI-57; K.- Deshayesites cf. antiquus BOGDANOVA, 1983, Errenaga Formation, IRI-90; L.- Desha- yesites cf. gracilis CASEY, 1964, Errenaga Formation, ER-76.

173 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

NOY, p. 74, pl. 5, fig. 2. Deshayesites cf. bodei 1999 Deshayesites weissiformis BOGDANOVA: AVRAM, (von KOENEN, 1902) p. 440-441, fig. 3A-E. 1999a Deshayesites weissiformis BOGDANOVA: CECCA Fig. 9 A-B et alii, p. 278, pl. 6, figs. 2-3. 2004 Deshayesites cf. weissiformis BOGDANOVA: cf. 1902 Hoplitidae bodei (von KOENEN), p. 221, pl. RAISOSSADAT, p. 124, fig. 4. 9, fig. 1a-c. 2004 Paradeshayesites weissiformis (BOGDANOVA): cf. 1924 Parahoplitoides bodei (von KOENEN): SPATH, BOGDANOVA & MIKHAILOVA, p. 212-214, pl. 7, fig. p. 433. 3; text-fig. 21. cf. 1960 Deshayesites bodei (von KOENEN): DRUS- Holotype: St. Petersburg Museum, N° CHITZ & KUDRYAVTZEV, tab. 1, fig. 4. 7/9442, from Taurkyr, Turkmenistan. cf. 1964 Prodeshayesites bodei (von KOENEN): CASEY, p. 358-359, pl. 58, figs. 4a-b, 5a-b; pl. Material: ER-2, 13, 14, 28, 55, 129?, ERG- 59, fig. 3; text-fig. 127a-b. 10-1, 20-3, IRI-3, 4, 11, 15, 20, 21. 24, 32, 42, cf. 1967 Deshayesites bodei (von KOENEN): KEMPER, 45, 46, 59, 77, 81, 101, 102, 103, 104, 105, tab. 12-13. 109, 110, 114, 115, 107, 108, ATAN-11. cf. 1971 Deshayesites bodei (von KOENEN): KEMPER, Description: Some specimens crushed and pl. 30, fig. 3. incomplete. Whorl section sub-rectangular, cf. 1973 Deshayesites bodei (von KOENEN): flanks parallel, flat and broad venter, umbilical GLASUNOVA, pl. 91. area is between one third and one fourth of the cf. 1979 Prodeshayesites bodei (von KOENEN): COLLIGNON et alii, p. 147, pl. 2, figs. 1a-b, 2a-b. diameter. Ornament consists of ribs; primary cf. 1995 Deshayesites bodei (von KOENEN): KEMPER, ribs start from the upper part of the umbilical taf. 3, figs. 3-4; taf. 5, fig. 4. wall with tubercles around the umbilical margin, cf. 1999 Prodeshayesites cf. bodei (von KOENEN): bifurcating in the lower quarter of the flanks. KELLY & WHITHAM, p. 87-88, fig. 3a-d. Secondary ribs are of same thickness as the cf. 2004 Deshayesites bodei (von KOENEN): BOGDA- primaries; some intercalating between primary NOVA & MIKHAILOVA, pl. 2, fig. 1. rib branches, starting near half way up the Lectotype: The example figured by von flank, free or attached to primaries. All ribs are KOENEN (1902), pl. 9, fig. 1a-c; text-fig. 127a-b, curved forward on the upper part of the flank, from the Lower Aptian of Timmern, Brunswick but are straight at the ventral area. Twenty (selected by CASEY, 1964). eight primary ribs and fifty eight secondary ribs Material: ER-9. are present at a diameter thirty seven mm. Description: One crushed specimen. Whorl Measurements: section sub-rectangular, flanks parallel, venter nearly curved and broad. Ribs sharp, primary ribs originate from upper part of umbilical wall and cross the flank in a sigmoidal curve; secondary ribs branch from primaries near to the umbilical margin. Discussion: D. bodei is similar to D. fissicostatus, but differs from the latter by a wider umbilicus and coarser ribs. CASEY (1964, p. 359) indicated that at some stages of growth, the two species are very similar, but believes that since D. bodei is used as subzonal index it is better to keep the two as distinct.

However, the presence of D. bodei confirms the fissicostatus Zone. Discussion: the species is identified by its Occurrence: Spain (COLLIGNON et alii, 1979), fine and dense ribs. Shells are similar to D. England (SPATH, 1924; CASEY, 1964), Germany oglanlensis and sometimes can not be easily (KEMPER, 1967), Russia (DRUSCHITZ & KULRYVTZVA, separated at the same dimensions as D. 1960; GLAZUNOVA, 1973). weissiformis and D. oglanlensis. However the Distribution: Igaratza section, Errenaga For- number of ribs in D. weissiformis is less than D. mation. oglanlensis at the same diameter. Occurrence: Turkmenistan (BOGDANOVA, Deshayesites weissiformis 1983), France (DELANOY, 1995; CECCA et alii, BOGDANOVA, 1983 1999a), Romania (AVRAM, 1999) and Iran (RAI- SOSSADAT, 2004). Fig. 9 C Distribution: Ataneta locality, Iribas section, 1983 Deshayesites weissiformis sp. nov.: BOGDA- Igaratza section, Iribas section, Errenaga For- NOVA, 34, tab. 2, figs. 1-3; tab. 3, fig. 7; text- mation (Figs. 7 - 8). figs. 3-4. 1995 Deshayesites weissiformis BOGDANOVA: DELA-

174 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

Figure 6: Ammonite range chart of the deshayesi-furcata transitional interval Zone in the composite Lareo Formation succession.

175 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

Figure 7: Ammonite range chart of the weissi, deshayesi, deshayesi-furcata transitional interval zones in the Erre- naga Formation of the Igaratza section.

Deshayesites cf. consobrinus Material: ERG-5-8, 28-2, 33-15, APAR-16. (d'ORBIGNY, 1841) Description: Whorls sub-rectangular, flanks parallel to convex, umbilical wall nearly vertical. Fig. 9 D Ribs narrow, sharp, sigmoidal; primary ribs originate from umbilical wall, secondary ribs, cf. 1841 Deshayesites consobrinus (d'ORBIGNY), p. 147, pl. 47, figs. 1-3. attached or free, single, occurring between a cf. 1964 Deshayesites consobrinus (d'ORBIGNY): pair of primaries and appear at one-third up the CASEY, text-figs. 123-124. whorl flank: rarely more than one secondary cf. 1973 Deshayesites consobrinus (d'ORBIGNY): between each pair of primaries in the last GLANZUNOVA, tab. 88, 89, 90. whorl. cf. 1979 Deshayesites consobrinus (d'ORBIGNY): Measurements: BOGDANOVA, p. 159, pl. 2, figs. 3-4. cf. 1999 Deshayesites consobrinus (d'ORBIGNY): BOGDANOVA & PROZOROVSKY, pl. 3, figs. g-i. cf. 1999 Deshayesites consobrinus (d'ORBIGNY): ROPOLO et alii, pl. 19, figs. 1, 4. cf. 2000 Deshayesites consobrinus (d'ORBIGNY): GONNET et alii, pl. 5, fig. 1. Discussion: D. consobrinus differs from D. cf. 2004 Deshayesites consobrinus (d'ORBIGNY): BOGDANOVA & MIKHAILOVA, p. 202-203, pl. 4, deshayesi in having less sigmoidal ribs and less figs. 1-3. overlapping whorls. ROPOLO et alii (2000) Lectotype: Laboratoire de Paléontologie, revised collected specimens from its stratotype Muséum National d'Histoire naturelle de Paris, area that agreed with CASEY'S lectotype. AVRAM d'ORBIGNY collection 5597a, Lower Aptian, La (1999, p. 452) assigned a few specimens to D. Bédoule (Bouches-du-Rhône, France) (selected Bogdanovae sp. nov. BOGDANOVA & MIKHAILOVA CASEY, 1964). (2004, p. 209) described a new species D.

176 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02) kemperi (BOGDANOVA & MIKHAILOVA, 2004) deri- (GONNET et alii, 2000), Russia (GLAZUNOVA, 1973) ved from D. consobrinus. The figured specimens and Turkmenistan (BOGDANOVA & PROZOROVSKY, indicate that both new species are very similar 1999). to D. consobrinus and might be just intra- Distribution: Igaratza section, Errenaga For- specific variations. mation (Fig. 6). Occurrence: England (CASEY, 1964), France

Figure 8: Ammonite range chart and biozonation for the lower part of the Errenaga Formation of the Iribas section.

177 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

Distribution: Ataneta locality, Iribas section, Deshayesites forbesi CASEY, 1961c Igaratza section, Errenaga Formation (Fig. 7). Fig. 9 E Deshayesites cf. euglyphus 1845 Ammonites Deshayesii LEYMERIE: FORBES, p. CASEY, 1964 353, pl. 13, fig. 2. 1947 Deshayesites deshayesi (d'ORBIGNY): ARKELL, Fig. 9 F-G p. 172-173, fig. 14a-b. cf. 1964 Deshayesites euglyphus sp. nov.; CASEY, 1961c Deshayesites forbesi sp. nov.: CASEY, p. 593, p. 336-337, pl. 52, figs. 1-4; pl. LVI, fig. 1a-b. pl. 81, fig. 2a-b. cf. 1979 Deshayesites euglyphus CASEY: COLLIGNON 1964 Deshayesites forbesi CASEY: CASEY, p. 314- et alii, p. 148, pl. 2, fig. 4a-c. 317, pl. 47, figs. 1-7; text-fig. 104g, 106i, cf. 1999 Deshayesites euglyphus CASEY: BOGDA- 110a. NOVA, pl. 2, fig. 7. 1999 Deshayesites forbesi CASEY: AVRAM, p. 449, cf. 2004 Deshayesites cf. euglyphus CASEY: RAISOS- fig. 6F-H. SADAT, p. 125. fig. 4I. 2009 Deshayesites forbesi CASEY: MORENO-BEDMAR cf. 2010 Deshayesites cf. euglyphus CASEY: et alii, fig. 9E. MORENO-BEDMAR et alii, fig. 11D. 2010 Deshayesites forbesi CASEY: MORENO-BEDMAR et alii, fig. 11C. Holotype: Reading University, Geology Holotype: British Geological Survey Museum, Dept., N° 6958, from Atherfield Clay Series, 30918, Atherfield Clay Series, Crackers, Atherfield, Isle of Wight, UK. Atherfield, Isle of Wight. Material: ER-1, 3, 4, 5, 6, 16, 17, 40, 62, Material: ER-50, 51, 65, 70, 125, 127, 135?, 73, 85, 103, ERG-6-8, 6-10, 6-13, 7-1. 137?, 138, 139, 140, 142, 143, 144, 145, 146, Description: Early whorls cannot be seen 147, 149, 153, 154, 156, 158, 161, 168, 169, because specimens are incomplete. Ornament 170, 171, 175, 177, 178, 180, 181, 185, 186, of ribs; primary ribs start from upper part of the ERG-1-1, 2-1, 2-3, 2-4, 2-5, 2-7, 2-8, 3-1, 3-2, umbilical wall, secondary ribs attach to prima- 3-5, 3-14, 3-15, 3-16, ERG-4-1, 5-1, 5-2, 5-3, ries or are free, bifurcating and originate in the 5-6, 5-9, 12-1, 17-1, 18-1, 20-2, ATAN-25, IMA lower third of the flank to mid flank, ribs more 5, 6, 8, 10, 12. flattened and projected forward on the venter. Description: Whorls sub-rectangular, flanks Measurements: parallel to convex, umbilical wall nearly oblique; umbilical width is between one fifth and one quarter of the diameter. Ornamentation consist of ribs; primary ribs sharp, nearly sigmoidal; secondary ribs bifurcating, appear at mid flank, rarely some primary ribs are single in later growth stages. Discussion: The pattern of rounded and Measurements: flattened ribs in the ventral area is a characteristic feature for species identification. According to CASEY (1964) there are some similarities between D. euglyphus and D. kiliani SPATH, 1930. The Spanish assemblage is of the D. euglyphus morphotype. Occurrence: England (CASEY, 1964), Spain (COLLIGNON et alii, 1979) and Iran (RAISOSSADAT, 2004). Distribution: Igaratza section, Errenaga For- mation (Fig. 7). Deshayesites cf. punfieldensis SPATH, 1930 Fig. 9 H

cf. 1930 Deshayesites punfieldensis SPATH, p. 431- 432, pl. 16, fig. 3a-b. cf. 1964 Deshayesites punfieldensis SPATH: CASEY, p. 338-341, pl. 46, figs. 1-3. Holotype: British Geological Survey Museum, 30915, from the Lower Lobster Bed, Atherfield, Isle of Wight (F.W. Simms University of the Discussion: The diagnostic feature for the Basque Country). species is density of its ribbing. D. forbesi Material: ER-77, 81, 89, 101, ERG-3-9, 3- shows similarity to D. deshayesi in rib pattern, 10. but the latter species has strong and more Description: Whorls sub-rectangular, sides uniform ribs and also a vertical umbilical wall. flattened. Ornament of sharp ribs; primary ribs Occurrence: England (CASEY, 1961c), Roma- curved from mid-flanks, commencing from the nia (AVRAM, 1999).

178 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02) upper part of umbilical wall; secondary ribs becomes more regular. Twenty two primary ribs start from the upper third of the flank, in 37 mm diameter. intercalated between primaries, mostly free or Measurements: attached to the primaries. Discussion: The rib pattern is a character for identification. There is similarity between D. vectensis SPATH, 1930, and D. punfieldensis, but the latter species has a wider umbilicus, is more coarsely ribbed and is less flattened on the venter. Hitherto, it seems, the species has not been recorded outside of England. Occurrence: England (CASEY, 1964) Distribution: Igaratza section, Errenaga For- mation (Fig. 6). Discussion: The Spanish specimens are comparable with the holotype figures in the rib Deshayesites luppovi BOGDANOVA, 1983 pattern. S shape rib pattern is a good character for identification. Some specimens resemble Fig. 9 I Deshayesites forbesi and there is not much 1952 Deshayesites aff. dechyi PAPP: LUPPOV, 203, difference between Deshayesites forbesi and tab. 7, fig. 1. Deshayesites luppovi expect in their suture lines nom. nud. 1971 Deshayesites luppovi sp. nov.; and this feature cannot be seen in the studied BOGDANOVA, 22. specimens. It seems that the rib pattern in 1983 Deshayesites luppovi sp. nov.: BOGDANOVA, p. Deshayesites forbesi is more regular than 139. tab. 3, figs. 1-6. Deshayesites luppovi. Deshayesites forbesi 1997 Deshayesites cf. luppovi BOGDANOVA: AGUADO et alii, fig. 7f-g. differs from Deshayesites pappi BOGDANOVA, 1999 Deshayesites luppovi BOGDANOVA: AVRAM, p. 1991, by the secondary ribs, which start in the 447, fig. 5f-g. upper part of the whorl flanks in the latter spe- 1999 Deshayesites luppovi BOGDANOVA: BOGDANOVA, cies. pl. 1, fig. 10. Occurrence: Lower Aptian (weissi Zone) in 1999 Deshayesites luppovi BOGDANOVA: BOGDANOVA north Caucasus and in Turkmenistan (BOGDA- & PROZOROVSKY, pl. 3, fig. f. NOVA, 1971, 1983), Romania (AVRAM, 1999) and 1999 Deshayesites luppovi BOGDANOVA: ROPOLO et Iran (RAISOSSADAT, 2004). alii, p. 178-179, pl. 16, figs. 4-5. Distribution: Igaratza section, Iribas section, 1999a Deshayesites luppovi BOGDANOVA: CECCA et alii, pl. 1, fig. 8. Errenaga Formation (Fig. 7). 2000 Deshayesites luppovi BOGDANOVA: GONNET et alii, pl. 2, figs. 1-2; pl. 4, fig. 2. Deshayesites cf. weissi 2004 Deshayesites luppovi BOGDANOVA: RAISOSSA- NEUMAYR & UHLIG, 1881 DAT, p. 125, figs. 5A-C, L. 2004 Deshayesites luppovi BOGDANOVA: BOGDANOVA Fig. 9 J & MIKHAILOVA, p. 207-208, pl. 2, figs. 8-9; text- cf. 1881 Hoplites weissi NEUMAYR & UHLIG, p. 179, fig. 14. pl. 46, fig. 1; pl. 47, fig. 1. 2010 Deshayesites luppovi BOGDANOVA: MORENO- cf. 1902 Hoplites (Deshayesites) weissi NEUMAYR & BEDMAR et alii, fig. 11B. UHLIG: von KOENEN, p. 207, pl. 45, fig. 1. Holotype: St. Petersburg Museum, N° 23/ cf. 1960 Deshayesites weissi (NEUMAYR & UHLIG): 9442, from Bolshoi Balkhan, Turkmenistan. DRUSCHITZ & KUDRYAVTZEV, p. 310, pl. 1, fig. 1. Material: ER-20, 24, 43, 128, ERG-8-5, IRI- cf. 1971 Deshayesites weissi (NEUMAYR & UHLIG): 5, 7, 8, 9, 10, 16, 17,22, 23, 25, 26, 30, 43, BOGDANOVA, p. 22. 47, 49, 51, 52, 60, 63, 66, 67, 73, 74, 79, 80, cf. 1977 Deshayesites weissi (NEUMAYR & UHLIG): 88, 89, 94, 96, 97, 100, 106, 112, 118. BOGDANOVA, p. 47, pl. 1, figs. 1-4; pl. 4, fig. 6. Description: Most specimens are crushed cf. 1999 Deshayesites weissi (NEUMAYR & UHLIG): AVRAM, p. 439, fig. 2a-c. and incomplete. Whorl section sub-rectangular, cf. 1999 Deshayesites weissi (NEUMAYR & UHLIG): whorl height is greater than whorl width, BOGDANOVA & PROZOROVSKY, pl. 4, figs. b-c. thickest in the lower part of the whorl flank, cf. 2000 Deshayesites sp. gr. weissi (NEUMAYR & compressed. Umbilical wall vertical, umbilical UHLIG): GONNET et alii, pl. 6, fig. 1. width is about one quarter of the diameter. cf. 2004 Deshayesites weissi (NEUMAYR & UHLIG): Ornament consists of sigmoidal ribs, strongly RAISOSSADAT, p. 127, fig. 4L. curved backwards on the lower third of the Syntypes: Both the specimens figured by flanks; primary ribs start from the upper part of NEUMAYR & UHLIG (1881) are lost; one was in the the umbilical wall, thickest in the lower and SCHLOENBACH collection in the Königlichen Geolo- upper part of flanks; secondaries at first are gischen Landesanstalt, Berlin, and the other irregular and in mode of intercalation between was in the Geologischen Reichsanstalt, Vienna. primaries, either in ones or twos, starting from A neotype should be designated from the lower third of the flank, some fusing with appropriate German material. adjacent primaries. With further growth, ribbing Material: ATAN-2, 10, 20.

179 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

Figure 9: Early Aptian ammonites of northern Spain. A-B.- Deshayesites cf. bodei (von KOENEN) 1902, Errenaga For- mation, ER-9; C.- Deshayesites weissiformis BOGDANOVA, 1983, Errenaga Formation, IRI-4; D.- Deshayesites cf. consobrinus (d'ORBIGNY, 1841), Errenaga Formation, ERG-5-8; E.- Deshayesites forbesi CASEY, 1961c, Errenaga For- mation, ER-137; F-G.- Deshayesites cf. euglyphus CASEY, 1964, Errenaga Formation, ER-17; H.- Deshayesites cf. punfieldensis SPATH, 1930, Errenaga Formation, ERG-3-9; I.- Deshayesites luppovi BOGDANOVA, 1983, Errenaga For- mation, IRI-94; J.- Deshayesites cf. weissi NEUMAYR & UHLIG, 1881, Errenaga Formation, ATAN-20; K.- Deshayesites spp., Errenaga Formation, IRI-53.

180 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

Description: Whorl height greater than whorl ribs are more regular. Twenty two primary ribs thickness, subparallel flanks. Ribs start from and forty four secondary ribs are present 21 umbilical wall, S shaped, bifurcating in the mm diameter. lower third to mid flank; some secondary ribs Measurements: are free and unattached to the primary ribs, projected forward in ventral area; secondary ribs are about twice in number to the primaries. In small specimens, ribs are stronger and more dense. Measurements:

Discussion: Rib pattern and suture line are characteristic for species identification. Desha- yesites dechyi is similar to D. forbesi in rib Discussion: Curved compact (dense) ribs are pattern, but the latter is more evolute and has characteristic for specific separation. However less strong ribs. D. dechyi differs from D. their density is less than that of D. tuarkyricus consobrinoides SINZOW, 1909, by a less convex and D. weissiformis. The rib pattern of D. weissi venter and from Deshayesites consobrinus by is similar in some cases to D. tuarkyricus at having irregularly branching ribs (BOGDANOVA et small diameter, but at larger diameter the ribs alii, 1979). In whorl section D. dechyi is similar are coarser in D. weissi and differ from D. to Prodeshayesites bodei. The main difference tuarkyricus. BOGDANOVA & MIKHAILOVA (2004) between these two species lies in the suture believe D. weissi is a transition form between line. The suture of D. dechyi is characterised by D. tuarkyricus and D. weissiformis. narrow and high elements. Moreover the rib Occurrence: Germany (von KOENEN, 1902), pattern of D. dechyi is denser than D. bodei. Russia (DRUSCHITZ & KUDRYTZEVA, 1960), Occurrence: Early Aptian northwest and Turkmenistan (BOGDANOVA, 1977), Romania north Caucasus (dechyi-deshayesi Zone) (DRUS- (AVRAM, 1999). CHITZ & KUDRYAVTZEV, 1960; BOGDANOVA et alii, Distribution: Ataneta locality, Iribas section, 1979), Caspian region (weissi-deshayesi Zone) Errenaga Formation (ammonite assemblage 11, (BOGDANOVA, 1977) and Kopet Dagh (RAISOSSA- Figs. 3 - 4). DAT, 2004). Distribution: Ataneta locality, Iribas section, Deshayesites dechyi PAPP, 1907 Igaratza section, Errenaga Formation (Fig. 7). Fig. 10 A Deshayesites cf. callidiscus 1907 Parahoplites dechyi; PAPP, p. 171, pl. 9, figs. CASEY, 1961c 1-5. 1952 Deshayesites dechyi PAPP: LUPPOV, p. 204, pl. Fig. 10 B 7, figs. 2-4. cf. 1961c Deshayesites callidiscus sp. nov.: CASEY, 1960 Deshayesites dechyi PAPP: DRUSCHITZ & p. 594, 507, 609, pl. 80, fig. 10. KUDRYAVTZEV, p. 310, pl. 1, fig. 6. cf. 1964 Deshayesites callidiscus CASEY, p. 327- 1977 Deshayesites dechyi PAPP: BOGDANOVA, p. 50, 328, pl. 49, figs. 3a-b, 4a-b; pl. 51, fig. 3; pl. pl. 2, figs. 1-5. 53, fig. 2a-b; text-fig. 114a. 1979 Deshayesites dechyi PAPP: BOGDANOVA et alii, cf. 1971 Deshayesites callidiscus CASEY: KEMPER, pl. p. 5, pl. 1, figs. 1-5; pl. 2, figs. 1-3; text-fig. 2. 22, fig. 4. 1999 Deshayesites dechyi PAPP: BOGDANOVA, pl. 2, cf. 1979 Deshayesites callidiscus CASEY: COLLIGNON figs. 3-4. et alii, p. 149, pl. 3, fig. 4a-b. 1999 Deshayesites dechyi PAPP: BOGDANOVA & cf. 1999 Deshayesites callidiscus CASEY: BOGDANOVA PROZOROVSKY, pl. 4, fig. h. & PROZOROVSKY, pl. 6, figs. e-f. 2004 Deshayesites dechyi PAPP: RAISOSSADAT, p. cf. 2000 Deshayesites callidiscus CASEY: GONNET et 129, fig. 4M-N. alii, pl. 6, fig. 3. 2004 Deshayesites dechyi PAPP: BOGDANOVA & cf. 2002 Deshayesites callidiscus CASEY: MIKHAILOVA, p. 203-204, pl. 3, figs. 6-8. BARABOSHKIN & MIKHAILOVA, pl. 1, fig. 1. Lectotype: K7593 Geological Museum of the cf. 2004 Paradeshayesites callidiscus (CASEY): BOG- Geological Institute of Hungary, Budapest DANOVA & MIKHAILOVA, p. 211-212, pl. 3, fig. 1; (selected by BOGDANOVA from PAPP collection pl. pl. 8, figs. 1-2; pl. 10, fig. 1; text-fig. 18. 9, fig. 3), Lower Aptian, Lavaschi, Daghestan. Holotype: Natural History Museum, London, Material: ER-36?, 58, 59, 63, 64, 83, 88, 91, 48836, Atherfield Clay Series, Crackers, 92, 113, ERG-19-4, 20-1, ATAN-15. Atherfield, Isle of Wight. Description: Whorl section sub-rectangular, Material: ER-11, 21, 25, 27, 29, 32, 34, 37, semi-involute; umbilical area is about one 42, 44, 47, 52, 53, 61, 62, 93, 100, 126, 133, quarter of the diameter. Ribs sigmoidal, ERG-3-13, 6-7, 8-1, 8-2, 18-7, 18-16, 21-3, maximum elevation and strength at lower third 21-4, ATAN-7, 8, 16, 19, 31. of sides, one or two secondary ribs are Description: Whorl section sub-rectangular, intercalated with primaries; in last half whorl flanks parallel to convex, umbilical wall nearly

181 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02) vertical, umbilical width is between one fifth on the mid flanks, starting from umbilical wall, and one quarter of the diameter. Ribs low, flat secondaries bifurcate from mid flank. On the topped in the upper part and around the venter, venter side ribs are nearly straight. primary ribs originate from just above the um- Measurements: bilical wall, secondary ribs, mostly single, originate at the mid flank, rarely more than one secondary rib between each pair of primaries in the last whorl. Twenty six primary ribs at umbilical margin and forty eight primary and secondary ribs in ventral margin at 28 mm diameter. Measurements:

Discussion: D. normani shows similarity in rib pattern to D. spathi CASEY, 1961c, but differs in its more rectangular venter, fewer secondary ribs and absence of smoothness in middle of the sides. Occurrence: England (CASEY, 1964), Roma- nia (AVRAM, 1999). Distribution: Igaratza section, Errenaga For- mation, Eskisabel section, Ataun area, Lareo Formation (Fig. 7).

Deshayesites cf. saxbyi CASEY, 1964 Discussion: Flattened ribs in the upper part of the flank is a character for the species. D. Fig. 10 D callidiscus differs from D. kiliani SPATH, 1930, in cf. 1930 Deshayesites consobrinoides (SINZOW): its stronger, denser and more sigmoidal ribs. SPATH, p. 427. The whorl section is also more rectangular. D. cf. 1961c Deshayesites sp. nov.: CASEY, p. 507, callidiscus shows affinity with D. grandis (SPATH, 609. 1930), but the latter species has denser ribs cf. 1964 Deshayesites saxbyi sp. nov.: CASEY, p. and degeneration of ribbing does not extend to 349-350, pl. 53 fig. 1a-c; pl. 56, fig. 6a-b. umbilical margin. D. callidiscus shows similarity Holotype: Natural History Museum, London, 46587, Atherfield Clay Series, Crackers, to D. topleyi (SPATH, 1930), but the former species has less sharp ribs. Atherfield, Isle of Wight (SAXBY collection). Material: ER-46, 54, 74, 118, 172, ERG-18- Occurrence: Spain (COLLIGNON et alii, 1979), 2, 25-3, 26-3, 28-2, SANGRE-13, 28, 30. England (CASEY, 1964), France (GONNET et alii, Description: Umbilical wall nearly vertical, 2000), Russia (BARABOSHKIN & MIKHAILOVA, 2002) sides parallel. Ribs strong, primary ribs and Turkmenistan (BOGDANOVA & PROZOROVSKY, 1999). sigmoidal, originating from just above the um- Distribution: Ataneta locality, Iribas section, bilical wall, secondary ribs in one or twos, Igaratza section, Errenaga Formation (Fig. 7). mostly attached but some free originating at the lower or mid flank. Deshayesites cf. normani CASEY, 1964 Discussion: D. saxbyi differs from D. latilobatus (SINZOW, 1909), in its less flattened Fig. 10 C sides and ribbing which is sharper than D. cf. 1930 Deshayesites sp. nov.: SPATH, p. 431, pl. latilobatus. D. saxbyi might show similarity with 17, fig. 5. D. consobrinoides SINZOW, 1909, but the latter cf. 1961c Deshayesites sp. nov.: CASEY, p. 507, species has more regular ribbing which rarely 609. bifurcates. cf. 1964 Deshayesites normani sp. nov.: CASEY, p. Occurrence: England (CASEY, 1964). 344-347, pl. 50, fig. 7; pl. 54, fig. 1; pl. 55, Distribution: Igaratza section, Errenaga For- figs. 2-4; pl. 56, fig. 3; pl. 57, figs. 7-8; text- mation (Fig. 7). fig. 121. cf. 1999 Deshayesites normani CASEY: AVRAM, p. Deshayesites cf. involutus SPATH, 1930 450-451, fig. 7C-D. cf. 2000 Deshayesites sp. gr. spathi / normani Fig. 10 E CASEY: GONNET et alii, pl. 5, fig. 2. Holotype: SEDGWICK Museum, Cambridge, SM cf. 1930 Deshayesites involutus SPATH, p. 432. B27036, Atherfield Clay series, Crackers, cf. 1961c Deshayesites involutus SPATH: CASEY, p. Atherfield, Isle of Wight. 609. cf. 1964 Deshayesites involutus SPATH: CASEY, p. Material: ERG-12-4, SANGRE-23. 310, pl. 45, figs. 1a-c, 4a-b; text-fig. 107. Description: Whorl section sub-rectangular, cf. 1999 Deshayesites cf. involutus SPATH: AVRAM, whorl height is nearly equal to whorl width, um- p. 454, fig. 8d. bilical wall vertical; sides flattened, venter cf. 2004 Deshayesites cf. involutus SPATH: RAISOS- nearly convex. Ornament consists of ribs, pri- SADAT, p. 129-130, fig. 6A-C. mary ribs thick, sigmoidal, projected backwards Holotype: British Geological Survey, N°

182 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

30919, from Hythe beds, Hythe, Kent, UK. Measurements: Material: ERG-26-2, 26-8, 34-13, 34-8, SANGRE-2. Description: Umbilical wall vertical with rounded shoulders, umbilical area is around one third of diameter. Ribs weakly sigmoidal in shape, primaries smooth at mid-flank and strong near the umbilical margin and at the ventral area, costate near the umbilical wall. Discussion: Ribs are strong at the umbilical Bifurcate or trifurcate, secondary ribs irregular, margin with low elevation at mid-flank and with start in the mid flank. evolute coiling are features characteristic of this Measurements: species. Measured dimensions similar to the holotype. Occurrence: England (CASEY, 1964), Roma- nia (AVRAM, 1999), Turkmenistan (BOGDANOVA, 1977) and Iran (RAISOSSADAT, 2004). Distribution: Ataneta locality, Iribas section Igaratza section, Errenaga Formation (Fig. 7). Discussion: Rib pattern is comparable to figured specimens of Deshayesites involutus Deshayesites cf. multicostatus and differs from Deshayesites grandis in finer SWINNERTON, 1935 ribbing. The incomplete specimens are insufficient to permit precise identification. Fig. 10 G Occurrence: England (SPATH, 1930; CASEY, cf. 1935 Deshayesites multicostatus SWINNERTON, p. 1964), Romania (AVRAM, 1999) and Iran (RAI- 31, pl. 1, fig. 1a-c. SOSSADAT, 2004). cf. 1961c Deshayesites multicostatus SWINNERTON: Distribution: Igaratza section, Errenaga For- CASEY, p. 508, 523, 569, 570, 609. mation (Figs. 6 - 7). cf. 1964 Deshayesites multicostatus SWINNERTON: CASEY, p. 304-305, pl. 43, figs. 5a-b, 6. Deshayesites cf. planus CASEY, 1961c cf. 1973 Deshayesites multicostatus SWINNERTON: GLAZUNOVA, p. 130, pl. 84, fig. 1. Fig. 10 F cf. 1999 Deshayesites multicostatus SWINNERTON: AVRAM, p. 441, fig. 3b-d. cf. 1961c Deshayesites planus CASEY, p. 609. cf. 2004 Deshayesites cf. multicostatus cf. 1964 Deshayesites planus CASEY: CASEY, p. 323, SWINNERTON: RAISOSSADAT, p. 132, fig. 5M. pl. 57, fig. 5; text-figs. 112a-b, e. cf. 1971 Deshayesites planus CASEY: BOGDANOVA, p. Holotype: Natural History Museum, London, 22. C36366 (SWINNERTON collection), from the cf. 1977 Deshayesites planus CASEY: BOGDANOVA, p. Sutterby, Marl of Sutterby Lincolnshire, UK. 52, pl. 3, figs. 1-5; pl. 4, figs. 7-8; text-fig. 4a- Material: ER-45, ERG-23-1, 23-2, 28-5, b. ATAN-3, 4, SANGRE-1, 2, 18. cf. 1999 Deshayesites planus CASEY: AVRAM, p. 445, Description: Whorls sub-rectangular, sides fig. 5c-e. flattened, venter convex. Ornament consisting cf. 1999 Deshayesites planus CASEY: BOGDANOVA & of slightly rounded ribs; primary ribs commence PROZOROVSKY, pl. 4, figs. g, i. cf. 2004 Deshayesites cf. planus CASEY: RAISOSSA- from umbilical wall, are sigmoidal, at first and DAT, p. 130, fig. 5J-K. straighter in the last whorl of adult specimens. Holotype: British Geological Survey, ZM Secondary ribs single or in pairs, interposed 1667 (CASEY collection), from Atherfield Clay between primaries mostly free in early whorls Series, Atherfield, Isle of Wight, UK. and attach in last half whorl. Material: ER-8, 12, 31, 41, 48, 60, 84, 94, Measurements: ERG-12-2, 16-1, 18-5, 19-1, 22-3, 22-4, 22-5, 28-7, 34-1, 34-3, 34-4, 34-6, ATAN-24, 26, 28. Description: Whorl section sub-rectangular, thickest in the lower one third of flanks; umbilical area is one third of diameter. Ribs narrow; primary ribs originate from upper part of umbilical wall and cross the flank in a sigmoidal curve, maximum elevation at lower third of flank, minimum elevation in middle of flank. Tu- Discussion: Dense ribs in ventral area and bercles are apparent around the umbilical small umbilicus are characteristic of D. margin as a result of thickening of the ribs. One multicostatus. It is distinguished from D. or two secondary ribs are intercalated between deshayesi var. strigosus CASEY by its sharper each pair of primaries. and denser ribbing and more stronger curved ribs. The Spanish specimens are comparable to CASEY's figures. Occurrence: England (CASEY, 1961c, 1964),

183 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

Russia (GLAZUNOVA, 1973), Romania (AVRAM, Measurements: 1999) and Iran (RAISOSSADAT, 2004). Distribution: Ataneta locality, Iribas section, Igaratza section, Errenaga Formation, Eskisabel section, Ataun area, Lareo Formation (Figs. 6 - 7). Deshayesites cf. deshayesi LEYMERIE in d'ORBIGNY, 1841 Fig. 10 H

cf. 1841 Ammonites deshayesi LEYMERIE in d'ORBI- GNY, p. 288, pl. 85, figs. 3-4. Discussion: Because of the similarity in cf. 1899 Hoplites deshayesi LEYMERIE: ANTHULA, p. ontogeny in the early stage of growth in Desha- 108. cf. 1914 Hoplites (Deshayesites) deshayesi yesites and Dufrenoyia, small specimens might KAZANSKY, p. 100-103, pl. 6, figs. 81-83; pl. 7, show similarity in their flattened venter and figs. 100-101. effacing ribs in the ventral area. However the cf. 1960 Deshayesites deshayesi (d'ORBIGNY): DRUS- presence of Deshayesites in these beds sug- CHITZ & KUDRYAVTZEV, p. 309, pl. 1, figs. 2, 5. gests a correct assignation to Deshayesites for cf. 1961c Deshayesites deshayesi (d'ORBIGNY): these samples. Some specimens are weathered CASEY, p. 508, 523, 538, 593, 609. and rib numbers cannot be counted. The Aralar cf. 1964 Deshayesites deshayesi (d'ORBIGNY): material is comparable to specimens of D. CASEY, p. 295, pl. 43, fig. 3; pl. 47, fig. 9; pl. deshayesi in the Natural History Museum, Lon- 51, fig. 6 (see for extensive synonymy). cf. 1971 Deshayesites deshayesi (d'ORBIGNY): BOG- don, numbers C4041, C71943-5, C71447-52. DANOVA, pl. 3, fig. 6; pl. 4, figs. 1-2. Occurrence: England (CASEY, 1964); Russia cf. 1971 Deshayesites deshayesi (d'ORBIGNY): (DRUSCHITZ & KUDRYAVTZEV, 1960; GLAZUNOVA, KEMPER, pl. 29, fig. 7. 1973); Turkmenistan (BOGDANOVA, 1977, 1991) cf. 1973 Deshayesites deshayesi (d'ORBIGNY): and Iran (RAISOSSADAT, 2004). GLAZUNOVA, p. 120, pl. 76, fig. 1. Distribution: Igaratza section, Errenaga For- cf. 1977 Deshayesites deshayesi (d'ORBIGNY): BOG- mation, Eskisabel section, Ataun area, Lareo DANOVA, p. 55, pl. 3, fig. 6; pl. 4, figs. 1-2. Formation (Figs. 6 - 7). cf. 1979 Deshayesites deshayesi (d'ORBIGNY): BOG- DANOVA, pl. 2, fig. 6. Deshayesites cf. consobrinoides cf. 2004 Deshayesites deshayesi (d'ORBIGNY): RAI- SOSSADAT, p. 130-131, fig. 5D-E & 5H-I. SINZOW, 1909 cf. 2004 Deshayesites deshayesi (d'ORBIGNY): Fig. 10 I AMEDRO & MARTIN, p. 80, pl. 1, figs. 11, 13. cf. 2010 Deshayesites deshayesi (d'ORBIGNY): cf. 1909 Parahoplites consobrinoides SINZOW, p. 3- MORENO-BEDMAR, et alii, fig. 11, E-G, I; fig. IV, 4. I-L. cf. 1947 Deshayesites consobrinoides d'ORBIGNY: Lectotype: Muséum National d'Histoire ARKELL, p. 170, figs. 18, 14b. Naturelle de Paris, N° 5579c. Selected by CASEY cf. 1961c Deshayesites consobrinoides d'ORBIGNY: (1961c) from d'ORBIGNY's surviving syntypes. CASEY, p. 508, 523, 609. Argiles à Plicatules of Bailly-aux-Forges, Paris cf. 1964 Deshayesites consobrinoides SINZOW: Basin, France. CASEY, p. 302, pl. 44, figs. 5-6; pl. 52, fig. 2; text-fig. 106j-n (see for extensive synonyms). Material: ERG-18-19, 25-5, 25-7, 27-1, 27- cf. 1971 Deshayesites consobrinoides SINZOW: BOG- 2, 28-4, 28-6, 30-1, 30-9, 30-11, 31-1, 31-4, DANOVA, pl. 3, fig. 3. 31-5, 33-1, 31-7, 31-9, 31-12, 32-1, 32-2, 33- cf. 1973 Deshayesites consobrinoides SINZOW: 5, 34-10, SANGRE-2, 3, 14, ORL-B-1, 2, 21. GLAZUNOVA, p. 123, pl. 77, figs. 1-5. Description: Umbilical wall nearly vertical, cf. 1979 Deshayesites consobrinoides SINZOW: BOG- umbilical width is between one third and one DANOVA et alii, pl. 3, fig. 3. quarter of the diameter. Whorls sub-rectan- cf. 1999 Deshayesites consobrinoides SINZOW: BOG- gular, flanks parallel to convex, venter sub- DANOVA, pl. 2, fig. 1. cf. 1999 Deshayesites consobrinoides SINZOW: BOG- truncate. Ribs narrow, sigmoidal; primary ribs DANOVA & PROZOROVSKY, pl. 6, figs. c-d. originate from just above the umbilical wall, cf. 2000 Deshayesites consobrinoides SINZOW: secondary ribs commence forward of the prima- GONNET et alii, pl. 3, fig. 1. ries at the mid flank, rarely more than one cf. 2004 Deshayesites consobrinoides SINZOW: BOG- secondary between each pair of primaries in the DANOVA & MIKHAILOVA, pl. 1, figs. 4-6. last whorl. Twenty four primary ribs at umbilical cf. 2004 Deshayesites cf. consobrinoides SINZOW: margin and fifty primary and secondary ribs in RAISOSSADAT, p. 131-132, figs. 4O, P, 5F, G. ventral margin at 33 mm diameter. Lectotype: KARPINSKY Museum, St. Petersburg (one of the specimens collected by SINZOW, 1898), N° 17727, from the Lower Aptian of Saratov, Russia. Material: ERG-18-17, 25-3, 26-1, ORL-B-7, 12, 15.

184 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

Figure 10: A.- Deshayesites dechyi PAPP, 1907, Errenaga Formation, ER-58; B.- Deshayesites cf. callidiscus CASEY, 1961c, Errenaga Formation, ATAN-16; C.- Deshayesites cf. normani CASEY, 1964, Errenaga Formation, ERG, 12-4; D.- Deshayesites cf. saxbyi CASEY, 1964, Errenaga Formation, ER-118; E.- Deshayesites cf. involutus SPATH, 1930, Errenaga Formation, ERG-34-8; F.- Deshayesites cf. planus CASEY, 1961c, Errenaga Formation, ER-94; G.- Deshaye- sites cf. multicostatus SWINNERTON, 1935, Lareo Formation, SANGRE-2; H.- Deshayesites cf. deshayesi LEYMERIE in d'ORBIGNY, 1841, Errenaga Formation, ERG-31-12; I.- Deshayesites cf. consobrinoides SINZOW, 1909, Errenaga For- mation, ERG-18-7.

185 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

Description: three crushed specimens are to Occurrence: England (CASEY, 1964). hand. Sides nearly parallel; ribs sharp and Distribution: Ataneta locality, Iribas section, strong, sigmoidal near ventral area, primary Errenaga Formation (Fig. 6). ribs start from upper part of umbilical wall, secondary ribs are intercalated between prima- Deshayesites cf. grandis SPATH, 1930 ries. Fig. 11 B Measurements: cf. 1930 Deshayesites grandis sp. nov.: SPATH, p. 427-429, pl. xvii, fig. 2a-b. cf. 1961c Deshayesites grandis SPATH: CASEY, p. 508-510. cf. 1964 Deshayesites grandis SPATH: CASEY, p. 308-309, pl. 43, figs, 1a-b; pl. 44, figs. 1-3; pl. 51, fig. 7a-b; text-fig. 110c. Discussion: Deshayesites consobrinoides is cf. 2010 Deshayesites grandis SPATH: MORENO- characterised by the presence of smooth BEDMAR et alii, fig. 11J, fig. V, B. external sides in the early stages of ontogeny Holotype: British Geological Survey Museum, and is very similar to Deshayesites deshayesi, Geological Society collection 2300, from but possesses coarser ribbing. Another feature Atherfield (probably from Bed IV or V), CASEY is that Deshayesites consobrinoides is more said evidently from the Scaphites Beds (group strongly evolute than average examples of V) of Atherfield (F. W. SIMMS collection). Deshayesites deshayesi (CASEY, 1964). Material: ER-136, ERG-8-3. Occurrence: Europe (CASEY, 1964), Russia Description: Two crushed specimens. (GLAZUNOVA, 1973), Caucasus (BOGDANOVA et alii, Ornament consist of ribs; primary ribs start 1979), Turkmenistan (BOGDANOVA, 1999) and from upper part of the umbilical wall, feebly Iran (RAISOSSADAT, 2004). curved; secondaries are irregular and Distribution: Igaratza section, Errenaga For- intercalated between primaries, start from mid mation (Figs. 6 - 7). to upper one third part of the flank, most of them attached to primaries; all primary and Deshayesites cf. geniculatus secondary ribs are flattened and rounded in CASEY, 1964 upper part of the flank. Discussion: The only character for Fig. 11 A identification is rib pattern in the studied speci- cf. 1964 Deshayesites cf. geniculatus sp. nov. mens. SPATH (1930) stated that the species is CASEY; p. 307, pl. 46, fig. 5; pl. 51, fig. 9. closer to D weissi than to D. consobrinus. The Holotype: British Geological Survey Museum, species is common in the type locality and a ZM 1937, Ferruginous Sands, Scaphites Beds, sub-zone is named on the occurrence of this west of Whale Chine, Atherfield, Isle of Wight. species. SPATH believed that the species appears Material: ATAN-29, APAR-18. in the upper part of the deshayesi Zone and Description: Whorl section sub-rectangular, even occurs in the early Late Aptian. On the whorl height is more than whorl width, thickest other hand, CASEY (1964) mentioned that D. in the upper part of the flank; umbilical wall grandis is limited to the deshayesi Zone and its vertical. Ornament consists of ribs, thick, record in the early Late Aptian is due to the recurved backwards on the mid flank. Primary confusing of large examples of Dufrenoyia with ribs start from upper part of the umbilical wall; D. grandis by SPATH. secondaries are irregular and intercalated Occurrence: England (SPATH, 1930; CASEY, between primaries, start from mid to upper one 1964). third part of the flank, some fusing with Distribution: Igaratza section, Errenaga For- adjacent primaries. mation (Fig. 7). Measurements: Deshayesites cf. wiltshirei CASEY, 1964 Fig. 11 C

cf. 1964 Deshayesites wiltshirei sp. nov.: CASEY, p. 305-306, pl. 53, fig. 4; pl 46, figs. 8-9; pl. 51, fig. 10. cf. 1999 Deshayesites cf. wiltshirei CASEY: BOGDA- NOVA, p. 352, fig. 6. Holotype: SEDGWICK Museum, Cambridge, B 27096, Ferruginous Sands, Atherfield, Isle of Discussion: Occasional bifurcation might be Wight (T. WILTSHIRE collection). a character for the species. D. geniculatus Material: ER-77, 81, 89, 101. differs from D. wiltshirei CASEY, 1964, in its Description: Whorls sub-rectangular, sides thick and curved ribbing. SPATH (1930) flattened. Ornament consisting of primary and described the species as a transition form secondary ribs; primary ribs curved in mid part between D. punfieldensis and D. vectensis of the flanks, commence from upper part of SPATH, 1930.

186 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02) umbilical wall; secondary ribs start from upper fine and curved on flanks and nearly straight on one third part of the flank, intercalated between venter; primary ribs originate from the umbi- primaries mostly free at early whorls and some lical margin, some single. Secondary ribs start attach at last half whorl. from one-third of the lower flank or at the mid Discussion: The rib pattern is a character for flank, intercalated and mostly attached to pri- identification. There is similarity between D. mary ribs. Twenty four primary ribs are present wiltshirei and D. punfieldensis, but the later at thirty three mm diameter. species has a wider umbilicus, coarser ribs Measurements: which are less flattened on the venter. Hitherto, the definitive species has not been recorded outside of England. Occurrence: England (CASEY, 1964); Spain. Distribution: Igaratza section, Errenaga For- mation.

Deshayesites spp. Fig. 9 K Discussion: The specimens occur in a deshayesi Zone assemblage. The rib patterns A large number of crushed and juvenile spe- and number are interesting and similar to early cimens have been collected. Some of these are deshayesitids. The specimens can be compared perhaps determinable at species level. The only to HUMPHREY's figures of Burckhardtites importance of this material is that they are gregoriensis (HUMPHREY, 1949, pl. 11, figs. 3-4) present in the deshayesi-furcata transition Zone and it is suggested that this is the specific discussed below. identity of the Aralar specimens. This species is Genus Burckhardtites HUMPHREY, 1949 recorded also from Turkmenistan (BOGDANOVA & MIKHAILOVA, 2004). Type species: Neocomites nazasensis BURCK- For a long time it has been assumed that HARDT, 1925 Generic characters: Discoidal, compressed, Burckhardtites is limited to America, but the whorl height and width increasing during recent record from Turkmenistan shows that its ontogeny. Sides and venter flattened with distribution is more widespread within the ventro-lateral shoulders. Ornament consists of Aptian Tethyan belt. The present record might primary sigmoidal ribs on the whorl flanks; allow us to determine the migration pattern of secondary ribs are intercalated or branch from this genus from west to east of the Tethys. primaries. Ribbing is retrorsive on the lower Distribution: Igaratza section, Errenaga For- part of the flank, convex forward near mid mation (Fig. 7). flanks and again convex backward on upper Genus Dufrenoyia third of the flanks. On venter ribs are of BURCKHARDT ex KILIAN & REBOUL, 1915 uniform relief and nearly straight. Suture line Type species: Ammonites furcatus SOWERBY seems to be of parahoplitid type. in FITTON, 1836. Discussion: Fine and irregular ribs are Generic characters: Semi involute, whorl characteristic for the genus. On the body section sub-rectangular to oval, with flattened chamber ribbing becomes very sharp. HUMPHREY and angular venter, umbilicus relatively wide; (1949) introduced the genus for all forms of costation consists of ribs; primary ribs rise in Neocomites nazasensis from the Aptian Rio the umbilicus margin, with light tuberculation, Nazas, Mexico. However Neocomites disap- secondary ribs are intercalated, starting at the peared in Hauterivian and it is believed Burck- lower third or middle flank, most are free; ribs hardtites is more related to Dufrenoyia, than it cross the venter isometrically, weak or disap- is to Neocomites (WRIGHT et alii, 1996; BARRAGÁN pearing on venter and forming denticulation on & MAURASSE, 2008). Burckhardtites is probably ventro-lateral shoulders. Suture line consists of derived from Paradeshayesites (BOGDANOVA & four lobes, a bifid ventral lobe, an entire umbi- MIKHAILOVA, 2004). Burckhardtites differs from lical, inner lateral and dorsal lobe. Dufrenoyia by lack of clear tubercles on the Discussion: The genus formerly considered ventro-lateral bend, dense ribbing and sharp to be a parahoplitid is now assigned to Desha- double S shaped ribs on the flanks. yesitidae proposed by STOYANOW (1949). Inter- Occurrence: Mexico (HUMPHREY, 1949; ruption of ribs in the ventral area is good cha- BARRAGÁN & MAURASSE, 2008), Turkmenistan racter for genus level identification. Ribs are (BOGDANOVA & MIKHAILOVA, 2004). thicker than Deshayesites in same diameter and Burckhardtites sp. are more regular. The genus differs from Deshayesites in its Fig. 11 D flat venter, thicker more regular ribs and tubercles on ventro-lateral shoulder. The genus Material: ERG-34-7, 34-22 Kuntziella COLLIGNON, 1962, previously known as Description: Whorl section sub-rectangular, a subgenus of Deshayesites, shows similarity in venter flat. Ornament consists of ribs, dense, some stage of growth with Dufrenoyia, but with

187 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02) higher whorls, flatter sides and without Holotype: British Geological Survey Museum, marginal tubercles differs from later genus in Geological society collection, Hythe Beds, the juvenile stage. The Early Albian homoeo- Hythe, Kent (W.H. FITTON collection). morph genus Neodeshayesites (recently Material: ERG-34-14. assigned to the subfamily Acanthohoplitinae by Description: Shell discoidal, whorl section is ROBERT & BULOT (2005) in its early stage of sub-rectangular to oval, truncated, forming growth resembles Dufrenoyia but in its later angular ventro-lateral margins. Ornament stage of growth the venter is flattened with rib- consists of ribs; primary ribs rise in the umbi- bing continuous across the venter with slight to licus margin with light tuberculation; secondary moderate tuberculation at the ventro-lateral ribs intercalate starting at mid- to upper part of margins' Stenhoplites SPATH, 1923, is an the flank, most are free. Ribs cross the venter objective synonym. isometrically, feebly flattened in the upper part Occurrence: Europe, Russia, Turkmenistan, of the flank and ventral area, with feeble tuber- Iran, Japan, USA, Mexico, Venezuela and culation on the ventral shoulders. Colombia (HUMPHREY, 1949; STOYANOW, 1949; Measurements: DRUSCHITZ & KUDRYAVTZEV, 1960; CASEY, 1964; BOGDANOVA, 1979; WRIGHT et alii, 1996; RAISOS- SADAT, 2004; MORENO-BEDMAR et alii, 2010). Dufrenoyia cf. furcata (SOWERBY, 1836) Fig. 11 E-F Discussion: The flat venter and flattened ribs in the upper part of the flanks and ventral side cf. 1836 Ammonites furcatus SOWERBY in FITTON, p. are characteristic feature. Dufrenoyia furcata 339, pl. xiv, fig. 17. SOWERBY was confused with Dufrenoyia dufre- cf. 1902 Hoplites furcatus SOWERBY: von KOENEN, p. noyi d'ORBIGNY by some authors; full discussion 202. cf. 1915 Parahoplites (Duferonyia) furcatus is found in CASEY (1964). D. furcata differs from SOWERBY: KILIAN & REBOUL, p. 34. the latter species by possessing less cf. 1923 Stenhoplites furcatus (SOWERBY): SPATH, p. overlapping whorls and coarser, sharp and 147. broader ribs. D. furcata differs from D. lurensis cf. 1925 Dufrenoya furcata (SOWERBY): BURCKHARDT, KILIAN by its more involute whorls and less p. 17, pl. 10, figs. 12-13. strong ribs (BOGDANOVA & MIKHAILOVA, 2004). cf. 1940 Dufrenoya furcata (SOWERBY): SCOTT, p. The species is well known in Aptian successions 1021. in Tethys and a characteristic faunal element cf. 1949 Dufrenoya furcata (SOWERBY): HUMPHREY, p. 120-121. for the latest Early Aptian. It has been proposed cf. 1960 Dufrenoya furcata (SOWERBY): DRUSCHITZ & as a zonal index on account of its KUDRYAVTZEV, tab. 1, fig. 7. geographically widespread occurrence (BOGDA- cf. 1964 Dufrenoyia furcata (SOWERBY): CASEY, p. NOVA, 1979, 1983; BOGDANOVA & TOVBINA, 1994; 378-382, pl. 62, figs. 2-3; pl. 63, fig. 1; pl. 65, KEMPER, 1995; HOEDEMAKER et alii, 1993, 1995, fig. 1a-b; text-figs. 134d, 135-136 (see this 2003). reference for full synonyms before 1964). Occurrence: England (CASEY, 1964), cf. 1971 Dufrenoyia furcata (SOWERBY): BOGDANOVA, Germany (KEMPER, 1995), Russia (DRUSCHITZ & tab. 51, fig. 10. cf. 1979 Dufrenoyia furcata (SOWERBY): BOGDANOVA, KULRYVTZVA, 1960), Turkmenistan (BOGDANOVA, tab. 51, fig. 10. 1979, 1999), USA (SCOTT, 1940; HUMPHREY, cf. 1982 Dufrenoyia furcata (SOWERBY): RENZ, pl. 1, 1949). fig. 7. Distribution: Igaratza section, Errenaga For- cf. 1995 Dufrenoyia furcata (SOWERBY): KEMPER, taf. mation (Fig. 7). 2, fig. 2. cf. 1999 Dufrenoyia furcata (SOWERBY): BOGDANOVA Dufrenoyia cf. mackesoni CASEY, 1964 & PROZOROVSKY, pl. 8, figs. d-e. cf. 2004 Dufrenoyia furcata (SOWERBY): BOGDANOVA Fig. 11 G & MIKHAILOVA, p. 219, pl. 12, figs. 1-3; text-fig. cf. 1964 Dufrenoyia mackesoni sp. nov.: CASEY, p. 28. 397-398, pl. 52, fig. 4a-b; pl. 62, fig. 5a-b; pl. cf. 2010 Dufrenoyia furcata (SOWERBY): GARCÍA & 65, fig. 4a-b; text-figs. 140d, 144. MORENO-BEDMAR, p. 129, fig. 1, A-P2, fig. 2, A1- X, AA-AM. Holotype: British Geological Survey Museum, cf. 2010 Dufrenoyia furcata (SOWERBY): MORENO- 108189, Hythe Beds, Shepway Cross Quarry, BEDMAR et alii, fig. 11, K, M; fig. VI, D-E; fig. Lympne, Kent (CASEY collection). VIII, D-E; fig. IX, A.

X Figure 11: A.- Deshayesites cf. geniculatus CASEY, 1964, Errenaga Formation, ATAN-29; B.- Deshayesites cf. grandis SPATH, 1930, Errenaga Formation, ER-136; C.- Deshayesites cf. wiltshirei CASEY, 1964, Errenaga Formation, ER-77; D.- Burckhardtites sp., Errenaga Formation, ERG-34-7; E-F.- Dufrenoyia cf. furcata (SOWERBY, 1836), Errenaga Formation, ERG-34-14; G.- Dufrenoyia cf. mackesoni CASEY, 1964, Errenaga Formation, ERG-34-11; H.- Dufrenoyia cf. lurensis (KILIAN, 1888), Errenaga Formation, SANGRE-6; I.-J. Toxoceratoides ? sp., Lareo formation, SANGRE-22.

188 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

189 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

Material: ERG-33-4, 33-5, 33-7, 34-11, Measurements: SANGRE-17? Description: Shell discoid, transverse section is sub-rectangular. Sculpture consists of ribs of narrow and moderate relief, curved; primary ribs start at the umbilical margin; intercalated secondary ribs start from mid-flank, most are free, the ribs form feeble denticulation on the ventral-lateral margin. Measurements: Discussion: The form is similar to D. furcata, but differs from the latter in its more evolute whorls and coarser, less flexuous ribbing and fewer secondary ribs. Occurrence: England (SPATH, 1930; CASEY, 1964), Germany (JACOB, 1907), France (KILIAN, 1889), Mexico (BURCKHARDT, 1925) and USA (SCOTT, 1940; HUMPHREY, 1949). Discussion: The species differs from D. predu- Distribution: Eskisable section, Ataun area, frenoy CASEY, 1964, by its weaker ribbing. Lareo Formation (Fig. 6). There are similarities also between the D. 4. Ammonite biostratigraphy of the mackesoni CASEY, 1964, and D. stenzeli HUMPHREY, 1949, the latter species possessing Aralar Early Aptian straighter ribs and a higher umbilical wall. Ammonites provide one of the most precise Occurrence: England (CASEY, 1964). biostratigraphical tools for correlating marine Distribution: Igaratza section, Errenaga For- Lower Cretaceous sediments. For much of Early mation (Fig. 6). Cretaceous time there was a separation into Tethyan and Boreal Realms, with distinct Dufrenoyia cf. lurensis (KILIAN, 1888) endemic ammonite faunas, which sometimes Fig. 11 H makes long-distance correlation difficult. However, some ammonite genera are of wide cf. 1888 Hoplites lurensis sp. nov.: KILIAN, p. 681, geographical distribution and this helps to pl. 20, fig. 2a-b. resolve some of the correlation problems. cf. 1889 Hoplites furcatus Sowerby var. lurensis: Because the Early Cretaceous ammonite KILIAN, p. 209. cf. 1907 Hoplites (Parahoplites) lurensis KILIAN; successions of Western Europe have been JACOB, p. 354, 357. studied in more detail than those in other areas cf. 1913 Parahoplites lurensis KILIAN: KILIAN, pl. 7, of the world, the stage stratotypes are situated fig. 8a-b; pl. 10, fig. 5. in this region and the ammonite zones proposed cf. 1925 Dufrenoyia lurensis (KILIAN): BURCKHARDT, there are normally accepted as the "standard" p. 17. biozonation. The Aptian Stage boundaries and cf. 1930 Dufrenoyia lurensis (KILIAN): SPATH, p. index faunas in Europe have been discussed 436, pl. 15, fig. 4. most recently by ERBA (1996) and by OWEN & cf. 1940 Dufrenoyia lurensis (KILIAN): SCOTT, p. 1022. RAISOSSADAT in GARCÍA-MONDÉJAR et alii (2009). cf. 1949 Dufrenoyia lurensis (KILIAN): HUMPHREY, p. Work by the Lower Cretaceous Cephalopod 121, 124-125, p. 8, figs. 7-8 & 12-13. Team of the SCS has improved the standard cf. 1964 Dufrenoyia lurensis (KILIAN): CASEY, p. biozonation for the Mediterranean area of the 382-383, pl. 63, fig. 2; pl. 64, figs. 3-4; text- Tethyan Realm (HOEDEMAEKER et alii, 1990, figs. 137-140. 1993, 1995, 2000, 2003; RAWSON et alii, 1999, Holotype: From the Upper Aptian (Garga- REBOULET et alii, 2006, 2009) (Table 1). In this sian) of the Carniol (Basses Alpes), France section of the paper, the current division and (Sorbonne collection). biozonation of the Early Aptian sub-Stage will Material: SANGRE-6, 7 be discussed and an attempt is made to Description: Whorl section is sub-rectangu- correlate successions throughout the whole of lar, becoming narrower towards the venter. the northern Mediterranean Tethyan belt, Ornament consists of nearly straight ribs. Pri- stretching eastward to Iran. mary ribs rise in the upper part of umbilical wall, with light tuberculation. Secondary rib 5. Current definition and biozonation intercalation is rare. Ribbing commences at the of the Early Aptian Stage lower part of the flank and cross the venter The Early Aptian ammonite zonation used isometrically with ventro-lateral clavi, Shell here is that refined for the northern region of flattened in the upper part of the flank and the Tethyan province (HOEDEMAEKER et alii, across the ventral area. 2003; REBOULET et alii, 2006, 2009) (Table 1) and is relevant to the Spanish succession in Spain. This scheme embraces some of the Early Aptian deshayesitid zones first established in Mangyschlak (BOGDANOVA, 1971; BOGDANOVA &

190 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

TOVBINA, 1994; TOVBINA, 1968, 1980). A diffe- Mediterranean Tethyan belt. The weissi Zone rent ammonite zonation has been proposed by has been identified in north Germany, Romania, CASEY (1961c) and CASEY et alii (1998) for the the Balkans, Tuarkyr and Kopet Dagh (BOGDA- succession in the Lower Greensand Group of NOVA & TOVBINA, 1994). AVRAM (1999) believed southern and eastern England. This region the forbesi and weissi zones are only partially belongs to a more Boreal province, but his coeval and that the weissi Zone commences scheme is applicable more widely in northern with the callidiscus Subzone of CASEY continuing Europe and Greenland (e.g., KELLY & WHITHAM, to the base of the grandis Subzone of the 1999) with some local modifications (e.g., deshayesi Zone of CASEY. KEMPER, 1976). Both schemes for the Early The author (in REBOULET et alii, 2006) Aptian reflect the occurrence of ammonite considered that D. weissi is a nomen dubium genera and species, particularly deshayesitids and suggested, either to find a type from speci- that are inter-provincial in their geographical men of the SCHLOENBACH collection in Berlin range (e.g., RAWSON, 1981; KOTETISHVILI, 1988; (Germany) or to replace D. weissi with Desha- MEMMI, 1995, 1999; SEYED-EMAMI et alii, 1971; yesites planus in the Standard zonation. These IMMEL et alii, 1997; RAISOSSADAT, 2004). The two species are found together in England as correlation of CASEY's scheme (CASEY, 1961c; well as the rest of Europe in sediments of this CASEY et alii, 1998) with that of the age. This suggestion is under review. Mediterranean Tethys has been discussed by Deshayesites deshayesi Zone OWEN & RAISOSSADAT in GARCÍA-MONDÉJAR et alii (2009) and is shown in Table 2. The D. deshayesi Zone of CASEY (1961c, 1964) and CASEY et alii (1998) is accepted by Deshayesites oglanlensis Zone most authors (e.g., BOGDANOVA, 1971, 1979; CASEY (1961c) proposed a biozonation for the BOGDANOVA & TOVBINA, 1994; BOGDANOVA & Early Aptian of England. In ascending order, the PROZOROVSKY, 1999; DRUSCHITZ & KUDRYAVTZEV, earliest Zone is that of Prodeshayesites 1960; DRUSCHITZ & GORBATSCHIK, 1979; HANCOCK, fissicostatus divided into an earlier Subzone of 1991; MEMMI, 1995, 1999; RAWSON, 1983; P. bodei and a later P. obsoletus Subzone. SHULGINA, 1996). CASEY (1961c), CASEY et alii These two subzones have been reported also (1998) suggested two subzones; an earlier from East Greenland (KELLY & WHITHAM, 1999). Cheloniceras (Cheloniceras) parinodum CASEY, KEMPER (1995) considered that the species of 1961b, and a later D. grandis. The co- Prodeshayesites are similar to species of Desha- occurrence of the first appearance of D. yesites reported from the Deshayesites deshayesi and Cheloniceras (Cheloniceras) tuarkyricus Zone of Turkmenistan by BOGDANOVA could be useful in the recognition of the weissi (1971, 1983). He considered Prodeshayesites to and deshayesi zones boundary. BOGDANOVA & be a synonym of Deshayesites and this view is PROZOROVSKY (1999) considered the deshayesi accepted here. Moreover it seems that the Zone in Transcaspia could be correlated with genera Paradeshayesites and Obsoleticeras of upper part of the English deshayesi Zone. BOGDANOVA & MIKHAILOVA (1999) are subjective Dufrenoyia furcata Zone synonyms of Deshayesites. BOGDANOVA & TOVBINA (1994) correlated the Deshayesites Dufrenoyia is a widely distributed genus and tuarkyricus Zone (now Deshayesites oglanlensis D. furcata is the index fossil for the latest part Zone) in Turkmenistan with the fissicostatus of the Lower Aptian in the Mediterranean Zone in England and the tenuicostatus Zone in region. CASEY (1961a) suggested Tropaeum Germany. RAISOSSADAT (2002) considered the bowerbanki CASEY as the index species for the geographical occurrence of D. tuarkyricus BOG- latest zone of the Lower Aptian in southern DANOVA to be restricted to Mangyschlak England, but Dufrenoyia furcata also occurs in (Turkmenistan) and Transcaspia. Therefore he that zone and Tropaeum is facies restricted. suggested that the more geographically 6. Biozonation in study area widespread species D. oglanlensis BOGDANOVA to be a more suitable index fossil for this Zone and The vertical distribution of ammonites in the this has been accepted (HOEDEMAEKER et alii, Errenaga and Lareo formations is shown in the 2003; REBOULET et alii, 2006). Recently, D. range charts (Figs. 6 - 7 - 8). A representative tuarkyricus has been recorded in the Errenaga sample of the ammonite fauna from the Aralar Formation in the Aralar Mountains (GARCÍA- sections is illustrated in Figs. 5 - 9 - 10 - 11. MONDÉJAR et alii, 2009). a. Deshayesites oglanlensis Zone Deshayesites weissi Zone In the Errenaga Formation of Iribas section CASEY (1961c) introduced a Zone of D. typical fauna of the oglanlensis Zone has been forbesi and divided it into four subzones; D. collected in-situ (assemblage 10, Fig. 3). The fittoni, D. kiliani, D. callidiscus and D. annelidus assemblage ammonite fauna (IRI-2 to 134) in ascending order. The Deshayesites weissi includes: D. cf. oglanlensis BOGDANOVA, 1983, D. Zone was proposed for the roughly equivalent cf. tuarkyricus BOGDANOVA, 1983, D. weissi- time period in the rest of Europe and the formis BOGDANOVA, 1983, D. luppovi BOGDANOVA, 1983, D. dechyi PAPP, 1907, and D. cf. antiquus

191 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

BOGDANOVA, 1983. 1930, D. cf. normani CASEY, 1964, Pseu- The lowest occurrence of ammonites in the dosaynella sp. and Roloboceras cf. hambrovi Errenaga Formation is in the Igaratza section at (FORBES, 1845); a typical weissi Zone fauna. approximately one m above the base of the unit At a height of 256.0 m above the base at (assemblage 6, Fig. 3) and consists of an Ataun, IMA-1 to 17 (assemblage 2, Figs. 3 - 4) indeterminate Deshayesites. A zonal age cannot is of definite weissi Zone age and includes D. be determined, but the basal Errenaga Forma- forbesi, Aconeceras (Aconeceras) cf. nisoides, tion at Igaratza is Early Aptian and there is no Aconeceras (Theganoceras) sp. evidence of the presence of Barremian In the Igaratza section, within a thickness of sediments. The interval corresponds probably to 20 m, the ammonite occurrences indicate the prolific fauna of D. oglanlensis Zone age successively the first appearance of the D. found at Iribas. weissi Zone fauna, a typical D. weissi Zone No ammonites have been found in the lower assemblage, the D. weissi-D. deshayesi zones part of the Errenaga Formation in the Ataun boundary and a relatively thin development of sections but an oglanlensis Zone age is the D. deshayesi Zone. The distribution of the probable on sedimentary subunit correlation ammonites is shown in Fig. 7. evidence (Fig. 3). Higher in the lower part of The view that the weissi and forbesi zones the Formation, assemblage 1 (Figs. 3 - 4) at are coeval (BOGDANOVA & TOVBINA, 1994) is 185.0 m above the base of the section, contains supported by the occurrences within the weissi a poorly preserved fauna (IMA-A-1 to 14) with Zone fauna in the Aralar area. This fauna D. cf. weissiformis and is probably of oglan- includes: D. cf. forbesi CASEY, D. cf. planus lensis Zone age. CASEY, D. cf. callidiscus CASEY, D. luppovi BOGDA- In the Aralar district, an oglanlensis Zone NOVA, D. dechyi PAPP together with species of fauna is present at the lower part of the Roloboceras including R. cf. hambrovi (FORBES), Errenaga Formation. The genus Procheloniceras and R. cf. regale CASEY. The occurrences of SPATH, 1923, occurs in the oglanlensis Zone ammonite horizons in the Errenaga Formation throughout southern Europe. Although absent are few and widely separated and thus, there in the Aralar district, it has been recorded in are gaps in our knowledge of the ammonite sediments of this age in the Betic Cordillera of succession in the Aralar area. However, in southern Spain (AGUADO et alii, 1997). This is of England, the genus Roloboceras is replaced by significance when considering the fauna of the Cheloniceras (Cheloniceras) at the base of the succeeding D. weissi Zone at Aralar, discussed deshayesi Zone (CASEY et alii, 1998). Based on next. above intrepretation the Aralar weissi Zone could be equivalent of the D. callidiscus Subzo- b. Deshayesites weissi Zone ne only. The best assemblage of ammonites is to be c. Deshayesites deshayesi Zone found in the Iribas and Igaratza sections (Fig. 7). At Iribas, a precise boundary between A precise sediment boundary between the oglanlensis and weissi Zone faunas occurs at weissi and succeeding deshayesi zones is only 23.85 m above the base of the unit recognisable at Igaratza (Fig. 7) where assem- (assemblage 11, Figs. 3 - 4). The weissi Zone blage ERG-17 is of weissi Zone age and ERG- material in this interval (ATAN-2 to 32, URG-1 18, 0.29 m above at 100.13 m above the base & 2) contains D. cf. weissi NEUMAYR & UHLIG, of the section is of deshayesi Zone age. At 1881, D. weissiformis BOGDANOVA, 1983, D. cf. Ataun, the boundary between sediments of the planus CASEY, 1961c, D. cf. callidiscus CASEY, weissi and deshayesi zones cannot be given 1961c, D. dechyi PAPP, 1907, D. cf. geniculatus precision in the lack of ammonite evidence. CASEY, 1964, D. cf. multicostatus SWINNERTON, At Iribas, it is possible that the deshayesi 1935, and D. forbesi CASEY, 1961c. A laterally Zone is represented in 2 metres of sediment equivalent outcrop situated 1 km West has upon equally condensed weissi Zone sediments yielded Roloboceras cf. regale CASEY, 1961c, if differences in subsidence and correlations of and Aconeceras (Aconeceras) cf. nisoides lithological subunits between the Iribas and Iga- (SARASIN, 1893), indicating a weissi Zone age ratza sections are taken into account (Fig. 4). (assemblage 12, Figs. 3 - 4). In the well-documented section at Igaratza Igaratza has provided the most important (Fig. 7), D. cf. deshayesi appears as a rarity at ammonite occurrence series within the weissi 100.13 m (ERG-18-19) above the base of the Zone (Fig. 7). The fauna includes Deshayesites section and then more commonly up to 104.53 cf. bodei (von KOENEN, 1902), D. weissiformis m above the base. This interval of 4.4 m is BOGDANOVA, 1983, D. cf. euglyphus CASEY, 1964, classified only with the deshayesi Zone. The D. cf. planus CASEY, 1961c, D. cf. callidiscus ammonites from this Zone at Igaratza (ERG-18- CASEY, 1961c, D. luppovi BOGDANOVA, 1983, D. 19 to ERG-28-10) include: D. cf. deshayesi dechyi PAPP, 1907, D. forbesi CASEY, 1961c, D. LEYMERIE in d'ORBIGNY, 1841, D. cf. callidiscus cf. multicostatus SWINNERTON, 1935, D. cf. CASEY, 1961c, D. cf. consobrinus (d'ORBIGNY, saxbyi CASEY, 1964, D. cf. gracilis SPATH, 1930, 1841), D. cf. consobrinoides SINZOW, 1909, D. D. cf. punfieldensis SPATH, 1930, D. cf. conso- cf. planus CASEY, 1961c, D. dechyi PAPP, 1907, brinus (d'ORBIGNY, 1841), D. cf. grandis SPATH, D. cf. multicostatus SWINNERTON, 1935, D.

192 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02) weissiformis BOGDANOVA, 1983, D. cf. involutus Deshayesites only and the transitions to Dufre- SPATH, 1930, and in the lower part of the noyia are absent. Sediments of the succeeding succession include elements that have survived Dufrenoyia furcata Zone yield Dufrenoyia only, from the underlying weissi Zone. Deshayesites being absent (e.g., CASEY, 1960; The sediments of the D. deshayesi Zone CASEY et alii, 1998). This suggests that the typically are thinly represented in the upper earliest part of the furcata Zone with its part of the Errenaga Formation (Fig. 4). I take transitional fauna is not represented by the base of the Zone at the first appearance of sediments over much of the European region D. deshayesi, but it is difficult determination of although detailed collecting in the Vocontian specimens in the upper part of the Errenaga Basin (France) and the Cau Section (SE Spain) Formation and in the Lareo Formation above (AGUADO et alii, 1997) may show a similar the Sarastarri Limestone due to the co-occur- transitional interval. Thus, the Aralar succession rence of Deshayesites and Dufrenoyia (Figs. 6 - in Europe is currently unique and reflects the 7 - 8). Elsewhere in Europe, sediments of the extremely rapid accumulation of sediments deshayesi Zone are characterised by species of during this relatively brief period of time.

Table 2: Comparison of the Early Aptian ammonite biozonation in SE England, Mediterranean Region and Aralar (N Spain). d. Deshayesites deshayesi - Dufrenoyia clearly related to Deshayesites, but hitherto, furcata Zone transition the transitional forms have not been found. The Aralar succession provides this transitio- The ammonite fauna in the Igaratza section nal interval in a very thick sediment succession. includes Dufrenoyia mackesoni CASEY, 1964, Within the Errenaga succession at Igaratza, Du- which first appears in the succession at 104.53 frenoyia furcata (SOWERBY, 1836), the zonal m above the base, where it is associated with index of the furcata Zone, appears in ERG-34 at species of Deshayesites. A similar association of 104.82 m above the base of the section asso- ammonites occurs a little higher in the section ciated with Deshayesites cf. deshayesi LEYMERIE (ERG-34) (Fig. 4) and in the Lareo Formation in d'ORBIGNY, 1841, D. cf. involutus SPATH, 1930, throughout the Aralar region. Such an associa- and Burkhardites sp. among others listed in Fig. tion is unique in the known sections in Europe 7. The Errenaga Formation was deposited in an spanning the deshayesi-furcata zones boun- open shelf environment with a local basinward dary. Elsewhere in Europe, there is a distinct polarity from east to west. Towards the top of changeover in the ammonite fauna at the base the Errenaga Formation in the Deshayesites of furcata Zone sediments with Dufrenoyia deshayesi-Dufrenoyia furcata zones transition, replacing Deshayesites. The rapid sedimen- a new regressive episode is deduced from the tation in the Aralar region reflecting the Biscay emplacement of shallow-water carbonate plat- marginal subsidence has preserved the transi- form facies of the Sarastarri Formation (Fig. 4). tion between the two zones. Dufrenoyia is

193 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

Table 3: Comparison of Deshayesites species of the north Spain and other areas in the Mediterranean Province. The co-occurrence of Deshayesites and Du- discussed here is widely applicable throughout frenoyia in this transitional interval might the Mediterranean Tethyan belt from northern eventually be recognised as a distinct Subzone, Spain eastward to the Kopet Dagh region of to be indexed when more extensive ammonite Iran and Turkmenistan. The assemblage fauna material is available. At the top of the Errenaga in the D. oglanlensis Zone of the Aralar Moun- Formation at Igaratza Dufrenoyia is rare and tains resembles that of France and the Kopet does not become more common until the Lareo Dagh. Moreover this Tethyan Zone can be Formation above. correlated with the Prodeshayesites fissicosta- The lower boundary of the furcata Zone is tus Zone of northern Europe proposed by CASEY taken at the base of the Lareo Formation. This (1961c). The next ascending ammonite biozone would probably be the boundary in other areas is based on the stratigraphic distribution of of more incomplete ammonite record. Strictly, Deshayesites weissi and Deshayesites planus. its base should be taken at its first appearance The weissi Zone has long been established, in the Igaratza section in sample ERG-33 at however D. planus shows a greater geogra- 104.53 m. Elsewhere in Europe and Iran, phical dispersion than D. weissi. The next Deshayesites does not occur in the furcata Zone ascending Zone is based on the widespread ammonite fauna (e.g., CASEY, 1961c; CASEY et occurrence of D. deshayesi and is a long esta- alii, 1998; RAISOSSADAT, 2004). The succession blished zonal index species for the later Early in the Vocontian Basin in France (e.g., MAGNIEZ- Aptian. It is capable of widespread recognition JANNIN et alii, 1997), Switzerland (STRASSER et in the European shelf seas as well as in the alii, 2001) and Spain (AGUADO et alii, 1997; Tethyan belt. The successor of Deshayesites, CASTRO et alii, 2001) for this interval is not Dufrenoyia also has a cosmopolitan distribution. controlled by ammonites as precisely as in The genus is more widely distributed than Aralar (GARCÍA-MONDÉJAR et alii, 2009).The co- Deshayesites, especially to the west, where it occurrence of Deshayesites and Dufrenoyia in occurs also in North America and the northern this transitional interval might eventually be part of South America where Deshayesites is recognised as a distinct Subzone, to be indexed absent or recorded only questionably. Deshaye- when more extensive ammonite material is sites species in north Spain show the closest available. similarity to those species recorded from Turk- The first Deshayesitidae (Turkmeniceras) ap- menistan and Iran (Kopet Dagh), France, Ro- peared in the Late Barremian of the restricted mania and Caucasus. Among twenty identified region of the Kopet Dagh and Turkmenistan, species from north Spain, twelve species are while its descendant, Deshayesites, spread to common with Iran and Turkmenistan (Kopet the Caucasus, Europe and even Greenland. Dagh), and nearly the same number of species The Lower Aptian ammonite biozonation with England, whereas seven species are shared based on species of Deshayesites and its des- with France and Romania (Table 3). This indi- cendent genus Dufrenoyia in north Spain cates good marine connections permitting the

194 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02) widespread distribution of these species and dure orientale du bassin de Paris.- Asso- enabling the palebiogeographic interpretation of ciation Géologique Auboise, Bulletin annuel, the Mediterranean Province. Sainte-Savine, n° 24-25, p. 76-80. 7. Conclusions ANTHULA D.J. (1899).- Über die Kreidefossilien des Kaukasus.- Beiträge zur Paläontologie The Lower Aptian in Aralar is subdivided into und Geologie von Österreich-Ungarns und the Errenaga, Sarastarri and Lareo formations. des Orients, Wien, vol. XII, p. 53-159. Representative species of the following genera: ARKELL W.J. (1947).- The geology of the country Aconeceras, Pseudosaynella, Toxoceratoides? around Weymouth, Swanage, Corfe and sp., Hemihoplites, Roloboceras, Deshayesites, Lulworth (explanation of sheets 341, 342, Dufrenoyia are identified and described for first 343, with small portions of Sheets 327, 328, time from study area. Based on ammonite 329).- Memoir of Geological Survey of the assemblage fauna D. oglanlensis, D. weissi, D. Great Britain - England and Wales, London, deshayesi biozones and an important D. 386 p. deshayesi-D. furcata transitional interval in ARKELL W.J., KUMMEL B. & WRIGHT W.C. (1957).- which Deshayesites and Dufrenoyia co-exist is Mesozoic Ammonoidea. In: MOORE R.C. (Ed.), suggested. The latter is currently unique to the Mollusca.- Treatise on Invertebrate Paleonto- Aralar succession and has important implica- logy, Geological Society of America, New tions for the current global biostratigraphical York; University of Kansas, Lawrence, Part L, correlations which need thorough revision. Vol. 4, p. L80-L437. The Deshayesites species of the Aralar AVRAM E. (1999).- The Deshayesites KAZANSKY, Mountains resembles that of France, Romania, 1914 (Ammonoidea) representatives in Ro- Caucasus, Kopet Dagh and England. It is mania, a link between the West-European capable of widespread recognition in the and Caspian assemblages of this genus. In: Tethyan belt as well as in the European shelf OLÓRIZ F. & RODRIGUEZ-TOVAR F. (eds.), Ad- seas. This indicates good marine connections in vancing Research on Living and Fossil above areas and could use in the paleobiogeo- Cephalopods.- Kluwer Academic/Plenum graphic interpretation of the Mediterranean Press, New York, p. 437-462. Province. BARABOSHKIN E.J. & MIKHALIOVA I.A. (2002).- Acknowledgements Новая стратиграфическая схема нижнего апта Среднего Поволжья [New stratigra- I thank Dr Hugh OWEN (Natural History Museum London) for his valuable comments phical scheme of the Lower Aptian of Middle during my research on these ammonites and Povolzhie].- Stratigrafiya, Geologicheskaya Korrelyatsiya (Stratigraphy and Geological the manuscript. I thank Prof. GARCÍA-MONDÉJAR Correlations), Moscow, vol. 10, n° 6, p. 100- and Dr MILLÁN (both from Pais Vasco University, Bilbao, Spain), for help in sampling, stratigra- 123 [in Russian]. phy and drawing of first drafts of some figures. BARRAGÁN R. & MAURRASSE F.J.-M.R. (2008).- Lower Aptian (Lower Cretaceous) ammonites I would wish to thank Phil CRABB of the Natural History Museum, London, for the excellent from the basal strata of the La Peña Forma- photographs of difficult material. I am indebted tion of Nuevo León State, Northeast Mexico: biochronostratigraphic implications.- Revista to Dr H. OWEN (Natural History Museum, Lon- mexicana de Ciencias geológicas, Mexico, don), Dr O. SZIVES (Natural History Museum, vol. 25, n° 1, p. 145-157. Budapest) and P. ROPOLO (Université de Proven- ce, Aix-Marseille) who acted as referees, for BOGDANOVA T.N. (1971).- Lower Aptian and their useful comments. I also thank Dr. boundary beds of west and south Turkmenia (Stratigraphy, ammonites).- Avtorefera Dis- Christian C. EMIG for his help to edit the manuscript. This research has been supported sertatii na soiskanie uchenoi stepenni by the University of the Basque Country Project kandidata geologo-mineralogicheskih nauk, BTE2003-04823, and the Spanish Science- 30 p. [in Russian]. Education Ministry Project CGL2006-12518 and BOGDANOVA T.N. (1977).- On some Deshayesites is greatly appreciated. of western Turkmenia.- Ezhegodnik Vseso- juznogo Paleontologicheskogo Obshchestva Bibliographic references (Akademiya Nauk SSSR), Leningrad, vol. 19, AGUADO R., COMPANY M., SANDOVAL J. & TAVERA p. 46-70 [in Russian]. J.M. (1997).- Biostratigraphic events at the BOGDANOVA T.N. (1979).- Аммониты семейства Barremian / Aptian boundary in the Betic Deshayesitidae Туркмении [Ammonites of Cordillera, southern Spain.- Cretaceous the Family Deshayesitidae of Turkmenia]. Research, London, vol. 18, n° 3, p. 309-329. In: BOGDANOVA T.N. (ed.), Plankton I orga- AGUIRRE-URRETA M.B. (2002).- Hemihoplitid nicheskii mir pelagiali v istorii Zemli [Plank- ammonoids from the Austral Basin of tonic and organic world of the pelagic zone Argentina and Chile.- Abhandlungen Geolo- in the history of the earth].- Trudy XIX sessii gisthen Bundesantalt, Wien, Band 57, p. Vsesoyuznogo Paleontologicheskogo Obsh- 491-500. chestva [Proceeding of the 19th session of AMÉDRO F. & MARTIN B. (2004).- Les ammonites the All-Union Paleontological Society], Nau- aptiennes des argiles à Plicatules de la bor- ka, Leningrad, p. 152-170 [in Russian].

195 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

BOGDANOVA T.N. (1983).- Зона Deshayesites London, vol. 65, n° 3, p. 262-277. tuarkyricus - нижняя зона апта Туркмении CASEY R. (1957).- The Cretaceous ammonite [Deshayesites tuarkyricus Zone - The lower genus Leymeriella, with a systematic ac- zone of the Aptian in Turkmenia].- Ezhe- count of its British occurrences.- Palaeonto- godnik Vsesojuznogo Paleontologicheskogo logy, Oxford, vol. 1, p. 29-59. Obshchestva (Akademiya Nauk SSSR), CASEY R. (1960).- The Ammonoidea of the Leningrad, vol. 26, p. 128-147 [in Russian]. Lower Greensand. Part 1.- Monograph of the BOGDANOVA T.N. (1991).- Новые виды Palaeontographical Society, London, vol. аммонитов из нижнего апта Туркмении 113, n° 490, xxxvi + 44 p. [New Lower Aptian ammonite species of CASEY R. (1961a).- The Ammonoidea of the Turkmenia].- Ezhegodnik Vsesojuznogo Lower Greensand. Part 2.- Monograph of the Paleontologicheskogo Obshchestva (Akade- Palaeontographical Society, London, vol. miya Nauk SSSR), Leningrad, vol. 34, p. 77- 114, n° 493, p. 45-118. 98 [in Russian]. CASEY R. (1961b).- The Ammonoidea of the BOGDANOVA T.N. (1999).- The Lower Aptian of Lower Greensand. Part 3.- Monograph of the the Mangyshlak Mountains. Stratigraphy and Palaeontographical Society, London, vol. Geological Correlation.- Stratigrafiya, Geolo- 115, n° 495, p. 119-216. gicheskaya Korrelyatsiya (Stratigraphy and CASEY R. (1961c).- The stratigraphical palaeon- Geological Correlations), Leningrad, vol. 7, tology of the Lower Greensand.- Palaeonto- p. 343-355 [in Russian]. logy, London, vol. 3, part 4, p. 487-621. BOGDANOVA T.N., KVANTALIANI I.V. & SCHARIKADZE CASEY R. (1964).- The Ammonoidea of the M.Z. (1979).- Некоторые раннеаптские Lower Greensand. Part 5.- Monograph of the дегезиты Центрального Дагестан [Some Palaeontographical Society, London, vol. 117 Early Aptian Deshayesitidae from central (1963), n° 502, p. 289-398. Daghestan].- Geologica Balcanica, Sofia, vol. CASEY R. (1965).- The Ammonoidea of the 9, n° 3, p. 3-12 [in Russian, with English Lower Greensand. Part 6.- Monograph of the abstract]. Palaeontographical Society, London, vol. BOGDANOVA T.N. & MIKHAILOVA I.A. (1999).- 118, n° 506, p. 399-546. Происхождение и развитие семейства CASEY R., BAYLISS H.M. & SIMPSON M.I. (1998).- Deshayesitidae (Ammonoidea) [Origin and Observations on the lithostratigraphy and evolution of the Family Deshayesitidae ammonite succession of the Aptian (Lower (Ammonoidea)].- Paleontological Journal, Cretaceous) Lower Greensand of Chale Bay, Moscow, vol. 33, n° 5, p. 522-529. Isle of Wight, UK.- Cretaceous Research, BOGDANOVA T.N. & MIKHAILOVA I.A. (2004).- London, vol. 19, n° 4, p. 511-535. Origin, evolution and stratigraphic signi- CASTRO J.M., COMPANY M., GEA G.A. de & AGUADO ficance of the Superfamily Deshayesitaceae R. (2001).- Biostratigraphy of the Aptian - STOYANOW.- Bulletin de l'Institut Royal des Middle Cenomanian platform to basement Sciences Naturelles de Belgique, Bruxelles, domain in the Prebetic Zone of Alicante, SE (Science de la Terre), vol. 74, p. 189-243. Spain: Biostratigraphy of the Aptian–Middle BOGDANOVA T.N. & PROZOROVSKY V. (1999).- Cenomanian platform to basin domain in the Substantiation of the Barremian/Aptian Prebetic Zone of Alicante, SE Spain: boundary. In: RAWSON P. & HOEDEMAEKER P.J. calibration between shallow water benthonic (eds.), Proceedings 4th International Work- and pelagic scales.- Cretaceous Research, shop of the Lower Cretaceous Cephalopod London, vol. 22, n° 2, p. 145-156. Team (IGCP-Project 362).- Scripta Geologi- CECCA F., DHONDT A.V. & BOGDANOVA T.N. ca, Leiden, Special Issue n° 3, p. 45-81. (1999a).- The Aptian stratigraphy of the BOGDANOVA T. & TOVBINA S.Z. (1994).- On southern Tuarkyr (NW. Turkmenistan, Cen- development of the Aptian Ammonite zonal tral Asia).- Rivista Italiana di Paleontologia e standard for the Mediterranean Region.- Stratigrafia, Milano, vol. 105, n° 3, p. 377- Géologie Alpine, Grenoble, Mém. H.S., n° 396. 20, p. 51-59. CECCA F., ROPOLO P. & GONNET R. (1999b).- The BURCHKHARDT C. (1925).- Faunas del Aptiano de appearance of the genus Deshayesites (KA- Nazas (Durango).- Boletín del Instituto ZANSKY, 1914, Ammonoidea) in the lower- Geológico de México, n° 45, 71 p. most Aptian (Lower Cretaceous) of La BUSNARDO R. (1984).- Crétacé inférieur : 1.3.1. Bédoule (SE France).- Rivista Italiana di – Ammonites.- In: DEBRAND-PASSARD S. (éd.), Paleontologia e Stratigrafia, Milano, vol. 105, Synthèse Géologique du Sud-Est de la n° 2, p. 267-276. France. I : Stratigraphie et paléogéogra- COLLIGNON M. (1962).- Atlas des fossiles de phie.- Mémoires du Bureau de Recherches Madagascar (Ammonites).- Service Géolo- Géologiques et Minières, Orléans, n° 125, p. gique - Ministère de l'Industrie et des Mines, 292-294. Tananarive, fasc. IX (Aptien), 64 p., pls. CASEY R. (1954).- Falciferella, a new genus of CCXV-CCXL. Gault ammonites, with a review of the family COLLIGNON M., PASCAL A., PEYBERNÉS B. & REY J. Aconeceratidae in the British Cretaceous.- (1979).- Faunes d'ammonites de l'Aptien de Proceedings of the Geologists' Association, la région de Santander (Espagne).- Annales

196 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

de Paléontologie, Invertébrés, Paris, vol. 65, Paleontología, Volumen de Actas.- Cidaris, fasc. 2, p. 139-156. Elche, n° 30, p. 129-137. DELANOY G. (1990).- Données nouvelles sur GARCÍA-MONDÉJAR J., LÓPEZ-HORGUE M.A., ARAMURU l'espèce-index Hemihoplites feraudianus A. & FERNÁNDEZ-MENDIOLA P.A. (2005).- Pulsa- (d'ORBIGNY, 1841) (Ammonoidea, Ancylo- ting subsidence during a rift episode: strati- ceratina).- Comptes Rendus de l'Académie graphic and tectonic consequences (Aptian- des Sciences, Paris, (Série II, Sciences de la Albian, Northern Spain).- Terra Nova, Paris, Terre et des Planètes), t. 310, p. 661-666. vol. 17, n° 6, p. 517-525. DELANOY G. (1995).- About some significant GARCÍA-MONDÉJAR J., OWEN H.G., RAISOSSADAT N., ammonites from the Lower Aptian (Bedou- MILLÁN M.I. & FERNÁNDEZ-MENDIOLA P.A. lian) of the Angle-Barrême area (Southeast (2009).- The Early Aptian of Aralar (North France).- Memorie Descrittive della Carta Spain): Stratigraphy, sedimentology, ammo- Geologica d'Italia, Roma, vol. LI, p. 65-101. nite biozonation, and OAE1.- Cretaceous DELANOY G. (1997).- Biostratigraphie haute Research, London, vol. 30, n° 2, p. 434-464. résolution du Barrémien supérieur du Sud- GILL T. (1871).- Arrangement of the families of Est de la France.- Comptes Rendus de l'Aca- Mollusks.- Smithsonian Miscellaneous Collec- démie des Sciences, Paris, (Série II, Scien- tions, Washington, vol. X, Smithsonian Insti- ces de la Terre et des Planètes), t. 325, p. tution Publication, n° 227, xvi + 49 p. 689-694. GLAZUNOVA A.E. (1973).- Палеонтологическое DOUVILLÉ H. (1890).- Sur la classification des обоснование стратиграфического расчле- Cératites de la Craie.- Bulletin de la Société нения меловых отложений Поволжья. géologique de France, Paris, (3ème Série), t. Верхний мел [Paleontological evidence for XVIII, p. 275-292. the stratigraphic subdivision of the Creta- DRUSCHITZ V.V. & GORBATSCHIK T.N. (1979).- ceous in Povolzhie, Lower Cretaceous].- Zonengliederung der Unteren Kreide der Nedra, Moscow, 324 p. [in Russian]. südlichen UdSSR nach Ammoniten und Fora- GONNET R., ROPOLO R. & CONTE G. (2000).- Les miniferen.- Aspekte der Kreide Europas, Deshayesites de la Zone à D. tuarkyricus Stuttgart, (IUGS Series A), n° 6, p. 107-116. dans l'Aptien inférieur de Cassis-La Bédoule DRUSCHITZ V.V. & KUDRYAVTZEV M.P. (1960).- (SE France).- Géologie méditerranéenne, Атлас нижнемеловой фауны Северного Marseille, t. XXV, n° 3-4 (1998), p. 125-147. Кавказа и Крыма [Atlas of Lower Creta- HANCOCK J.M. (1991).- Ammonite scales for the ceous fauna of the northern Caucasus and Cretaceous System.- Cretaceous Research, Crimea].- Trudy Vsesoyuznogo Nauchno- London, vol. 12, n° 3, p. 259-291. Issledovatelskogo Geologicheskogo Instituta, HOEDEMAEKER P.J. & BULOT L. (reporters), AVRAM Moscow, 702 p. [in Russian]. E., BUSNARDO R., COMPANY M., DELANOY G., DURVENOIS C.H., FLOQUET M. & HUMBEL B. KAKABADZE M., KOTETISHVILLI E., KRISHNA J., (1972).- La Sierra d'Aralar (Pyrénées bas- KVANTALIANI I., LATIL J.L., MEMMI L., RAWSON ques espagnoles) : stratigraphie, structure.- P.F., SANDOVAL J., TAVERA J.M., THIEULOY J.-P., Mémoire synthétique, Thèses de Doctorat de THOMEL G., VAŠÍČEK Z. & VERMEULEN J. 3ème Cycle, Institut des Sciences de la (1990).- Preliminary ammonite zonation for Terre, Université de Dijon, 240 p. ronéotées, the Lower Cretaceous of Mediterranean 9 pls. region (Lower Cretaceous Cephalopod team, ERBA E. (1996).- The Aptian stage.- Bulletin de Working Group of IGCP, Project 262).- Géo- l'Institut royal des Sciences naturelles de logie Alpine, Grenoble, t. 66, p. 123-127. Belgique, Bruxelles, (Science de la Terre), HOEDEMAEKER P.J. & COMPANY M. (reporters), vol. 66, p. 31-43. AGUIRRE URRETA M.B., AVRAM E., BOGDANOVA FLOQUET M. & RAT P. (1975).- Un exemple T.N., BUTJOR L., BULOT L., CECCA F., DELANOY d'interrelation entre socle, paléo-géographie G., ETTACHFINI M., MEMMI L., OWEN H., RAWSON et structure dans l'arc pyrénéen basque : La P.F., SANDOVAL J., TAVERA J.M., THIEULOY J.-P., sierra de Aralar.- Revue de Géographie TOVBINA S.Z. & VAŠÍČEK Z. (1993).- Ammoni- physique et de Géologie dynamique, Paris, tes zonation for the Lower Cretaceous of the vol. XVII, fasc. 5, p. 497-512. Mediterranean region; basis for the strati- FORBES E. (1845).- Catalogue of the Lower graphic correlation within IGCP-Project 262.- Greensand fossils in the Museum of the Geo- Revista Española de Paleontología, Madrid, logical Society, with notices of species new vol. 8, n° 1, p. 117-120. to Britain.- Quarterly Journal of the Geologi- HOEDEMAEKER P.J. & CECCA F. (reporters), AVRAM cal Society, London, vol. 1, p. 237-250 & E., COMPANY M., DELANOY G., ERBA E., 345-355. ETTACHFINI M., FARAONI P., KAKABADZE M., GARCÍA R. & MORENO-BEDMAR J.A. (2010).- Dufre- LANDRA G., MARINI A., MEMMI L., PALLINI G., noyia furcata (SOWERBY, 1836), ammonite RAWSON P.F., ROPOLO P., SANDOVAL J., TAVERA del Aptiense inferior (Cretácico Inferior) del J.M. & VAŠÍČEK Z. (1995).- Report on the 3rd Perelló (Tarragona). In: MORENO-AZANZA M., International Workshop on the standard DÍAZ-MARTÍNEZ I., GASC J.M., MELERO-RUBIO M., Lower Cretaceous ammonite zonation of the RABAL-GARCÉS R. & SAUQUÉ V. (coords.), VIII Mediterranean region.- Memorie Descrittive Encuentro de Jóvenes Investigadores en della Carta Geologica d'Italia, Roma, vol. LI,

197 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

p. 213-215. Gesellschaft zu Hannover, Bericht 117, p. HOEDEMAEKER P.J. & RAWSON P.F. (reporters), 29-54. AGUIRRE-URRETA M.B., BOGDANOVA T.N., FOZY KEMPER E. (1976).- Geologischer Führer durch I., GARCIA-BARRERA P., GONZALEZ ARREOLA C., die Grafschaft Bentheim und die angren- KAKABADZE M., KLEIN J., LUKENEDER A., RAISOS- zenden Gebiete, mit einem Abriss der SADAT N., SZIVES O. & VAŠÍČEK Z. (2000).- emsländischen Unterkreide.- Das Benthei- Report on the 5th International Workshop of mer Land, Nordhorn, n° 64, 206 p. the Lower Cretaceous Cephalopod team KEMPER E. (1995).- Die Entfaltung der Ammoni- (Vienna, 5 September 2000).- Cretaceous ten und die Meeresverbindungen im borealen Research, London, vol. 21, n° 6, p. 857-860. Unter- und Mittel-Apt.- Geologisches HOEDEMAEKER P.J. & REBOULET S. (reporters), Jahrbuch, Hanover, (Reihe A), Heft 141, p. AGUIRRE-URRETA M.B., ALSEN P., AOUTEM M., 171-199. ATROPS F., BARRAGÁN R., COMPANY M., GONZA- KILIAN W. (1888).- Sur quelques fossiles du LEZ ARREOLA C., KLEIN J., LUKENEDER A., PLOCH Crétacé inférieur de la Provence.- Bulletin de I., RAISOSSADAT N., RAWSON P.F., ROPOLO P., la Société Géologique de France, Paris, VAŠÍČEK Z., VERMEULEN J. & WIPPICH M.G.E. (3ème série), t. XVI, p. 663-691, pls. XVII- (2003).- Report on the 1st International XXXI. Workshop of the IUGS Lower Cretaceous KILIAN W. (1889).- Description géologique de la Ammonite Working Group, the 'KILIAN Group' montagne de Lure.- Thèse, Masson édit., (Lyon, 11 July 2002).- Cretaceous Research, Paris, 458 p. London, vol. 24, n° 1, p. 89-94. KILIAN W. (1907-1913).- Unterkreide (Palaeo- HUMPHREY W.E. (1949).- Geology of the Sierra cretacicum). In: FRECH F. (ed.), Lethaea de los Muertos area, Mexico.- The Geological Geognostica.- Handbuch der Erdgeschichte, Society of America, Bulletin, Boulder, vol. 2, Das Mesozoicum, 3, Kreide Lief 1, 1907, 60, p. 89-176. p. 1-168, Lief 2, 1910, p. 169-287, Lief 3, HYATT A. (1889).- Genesis of the Arietidae.- 1913, p. 289-398. Smithsonian Contributions of Knowledge, KILIAN W. & REBOUL P. (1915).- I. La faune de Washington, vol. XXVI, Smithsonian Institu- l'Aptien inférieur des environs de Montélimar tion Publication, n° 673, p. xi + 239. (Drôme) (Carrière de l'Homme d'Armes). In: HYATT A. (1903).- Pseudoceratites of the Creta- KILIAN W. (ed.), Contribution à l'étude des ceous- Monographs of the United States faunes paléocrétacées du Sud-Est de la Geological Survey, Washington, 351 p. France.- Mémoires pour servir à l'Explication IMMEL H., SEYED-EMAMI K. & AFSHAR-HARB A. de la Carte Géologique détaillée de la Fran- (1997).- Kreide-Ammoniten aus dem ira- ce, Paris, n° 14, 221 p., 9 pls. nischen Teil des Koppeh-Dagh (NE Iran).- KOENEN A. von (1902).- Die Ammonitiden des Zitteliana, München, vol. 21, p. 159-190. Norddeutschen Neocom (Valanginien, Haute- JACOB C. (1907).- Études paléontologiques et rivien, Barrêmien und Aptien).- Abhand- stratigraphiques sur la partie moyenne des lungen der Königlich Preußischen Geolo- terrains crétacés dans les Alpes françaises et gischen Landesanstalt und Bergakademie, les régions voisines.- Annales de l'Université (N.F.), n° 24, 451 p. de Grenoble, vol. 19, n° 2, p. 221-534. KOTETISHVILI E.V. (1988).- Distribution globale KAZANSKY P.A. (1914).- Описание коллекции des ammonites éocrétacées du Caucase. In: головоногих из меловых отложений Даге- WIEDMANN J. & KULLMANN J. (eds.), Cepha- стана со списком форм других классов и lopods - Present and Past, Proceedings of стратиграфическим очерком [Description the Second International Cephalopod d'une collection de céphalopodes des ter- Symposium.- Schweizerburt'sche verlag- rains crétacés du Daghestan].- Izvestiia buchhandlung, Stuttgart, p. 453-468. Tomskogo Teckhnologicheskoga Instituta, t. KOTETISHVILI E.V., KAKABADZE M. & KAKABADZE I. 32, n° 4, 127 p. [in Russian]. (2000).- Steps in evolution of the Early Cre- KELLY S.R.A. & WHITHAM A. G. (1999).- Deshaye- taceous ammonites of Georgia.- Trudy sitid ammonites from the Lower Aptian Akademiya Nauk Gruzinskoy SSR, Tiblisi, (Lower Cretaceous) of North-East Green- (Novaya Serya), Vyp. 115, p. 76–85 [in land.- Scripta Geologica, Leiden, Special Is- Russian]. sue 3, p. 89-95. LERTXUNDI D. (1997, unpublished).- El Aptiense KEMPER E. (1967).- Die älteste Ammoniten del Sur de la Sierra de Aralar.- Trabajo de Fauna im Aptium Nordwest-Deutschlands.- investigación del Programa de Doctorado, Paläontologische Zeitschrift, Stuttgart, vol. Universidad del País Vasco, 19 p. 41, p. 119-131. LERTXUNDI D. & GARCÍA-MONDÉJAR J. (1997).- El KEMPER E. (1971).- Zur Gliderung und Abgren- surco de Aia-Zaldibia (Aptiense inferior, zung des norddeutschen Aptium mit Ammo- Aralar, Gipuzkoa).- Geogaceta, Madrid, vol. niten.- Geologisches Jahrbuch, Hannover, 22, p. 105-108. Band 89, p. 359-390. LUPPOV I.P. (1952).- Нижнемеловые отложения KEMPER E. (1973).- Die Unterkreide im Unter- Северо-Западного Кавказа и их фауна grund der Gehrdener Berge und in der [Lower Cretaceous deposits of the north- Deistermulde.- Bericht der naturhistorischer western Caucasus and their fossils].- Trudy

198 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

Vsesojuznogo Neftjanogo Nauchno-Issle- GES A. (2009).- Biostratigraphic characteri- dovatel'skogo Geologo-Razvedochnogo Insti- zation by means of ammonoids of the Lower tuta, Moscow, vol. 65, 270 p. [in Russian]. Aptian Oceanic Anoxic Event (OAE 1a) in the MARTINEZ R. (1982).- Ammonoideos cretácicos eastern Iberian Chain (Maestrat Basin, del Prepirineo de la provincia de Lleida.- eastern Spain).- Cretaceous Research, Lon- Publicaciones del Instituto Geológico de la don, vol. 30, n° 4, p. 864-872. Universidad de Barcelona, vol. 17, 197 p. MORENO-BEDMAR J.A., COMPANY M., BOVER-ARNAL MAGNIEZ-JANNIN F., BREHERET J.-G. & DELANOY G. T., SALAS R., DELANOY G., MAURRASSE F.J.- (1997).- Un exemple de spéciation lié à M.R., GRAUGES A. & MARTÍNEZ R. (2010).- l'eustatisme : l'apparition précoce de Lower Aptian ammonite biostratigraphy in Schackoina cabri (foraminifère planctonique the Maestrat Basin (Eastern Iberian Chain, mésogéen).- Comptes Rendus de l'Académie Eastern Spain). A Tethyan transgressive des Sciences, Paris, (Série II, Sciences de la record enhanced by synrift subsidence.- Terre et des Planètes), vol. 325, n° 3, p. Geologica Acta, Barcelona, vol. 8, n° 3, p. 225-230. 281-299. MCLEARN F.H. (1932).- Problem of the Lower NEUMAYR M. & UHLIG V. (1881).- Über Ammoni- Cretaceous of the Canadian interior.- tiden aus den Hilsbildungen Nord- Transactions of the Royal Society of Canada, deutschlands.- Palaeontographica, Cassel, Ottawa, vol. XXVI, section IV, p. 157-175. vol. 27, n° 3-6, p. 129-203. MEMMI L. (1995).- Aptian biostratigraphy of ORBIGNY A. d' (1841).- La Paléontologie françai- northeastern Tunisia. A stratigraphical se. Description des mollusques et rayonnés approach of Aptian and Albian of Tunisia.- fossiles - Terrains Crétacés. Tome Premier. Memorie Descrittive della Carta Geologica Céphalopodes.- Masson, Paris, 662 p. d'Italia, Roma, vol. LI, p. 109-120. PAPP K. (1907).- Beschreibung der während der MEMMI L. (1999).- L'Aptien et L'Albien de Forschungsreisen M. v. DÉCHY im Kaukasus Tunisie. Biostratigraphie à partir des ammo- gesammelten Versteinerungen. In: DÉCHY M. nites.- Bulletin de la Société géologique de von (ed.), Kaukasus. Reisen und France, Paris, vol. 170, n° 3, p. 303-309. Forschungen im kaukasischen Hochgebirge.- MIKHAILOVA I.A. (1958).- Дегезитиды из Band III, Dietrich Reimer (Ernst Vohsen), нижнемеловых отложений Дагестана и Berlin, p. 141-174. Центрального Предкавказья [Deshayesitids PARONA C.F. & BONARELLI G. (1895).- Sur la from the Lower Cretaceous of Daghestan faune du Callovien inférieur (Chanasien) de and Central Fore-Caucasus].- Materialy Savoie.- Mémoires de l'Académie royale des kosnovam Paleontologii, vol. 2, p. 21-29 [in Sciences, Belles-lettres et Arts de Savoie, Russian]. Chambéry, (Série IV), t. VI, p. 35-211, 11 MILLÁN M.I., AGIRREZABALA L.M., FERNÁNDEZ- pls. MENDIOLA P.A. & GARCÍA-MONDÉJAR J. (2005).- PARONA C.F. & BONARELLI G. (1897).- Fossili El surco calcarenítico intraplataforma de albiani d'Escragnolles, del Nizzardo e della Desamendi: origen y evolución (Aptiense Liguria occidentale.- Paleontographia Italica, inferior, Aralar-Navarra).- Geo-Temas, Pisa, vol. 2, p. 53-112. Madrid vol. 8, p. 73-76. RAISOSSADAT S.N. (2002, unpublished).- Lower MILLÁN M.I., FERNÁNDEZ-MENDIOLA P.A. & GARCÍA- Cretaceous (Upper Barremian-Lower Albian) MONDÉJAR J. (2007).- Pulsos de inundación Ammonite faunas of the Kopet Dagh Basin, marina en la terminación de una plataforma NE Iran.- PhD thesis, University College carbonatada (Aptiense Inferior de Aralar, London, 337 p. Cuenca Vasco-Cantábrica).- Geogaceta, RAISOSSADAT S.N. (2004).- The ammonite family Madrid, vol. 41, p. 127-130. Deshayestidae in the Kopet Dagh Basin, MONTADERT L., WINNOCK E., DELTIEL J.R. & GRAU north-east Iran.- Cretaceous Research, G. (1974).- Continental margins of Galicia- London, vol. 25, n° 1, p. 115-136. Portugal and Bay of Biscay.- In: BURK C.A. & RAWSON P.F. (1981).- Early Cretaceous ammoni- DRAKE C.L. (eds.), The geology of continental te biostratigraphy and biogeography. In: margins.- Springer-Verlag, New York, p. HOUSE M.R. & SENIOR J.R. (eds.), The Ammo- 323-342. noidea: The evolution, classification, mode MONTADERT L., ROBERTS D.G., DE CHARPAL O. & of life and geological usefulness of a major GUENNOC P. (1979).- Rifting and subsidence group.- Systematics Association Special of the northern continental margin of the Volume Series, London, vol. 18, p. 499-529. Bay of Biscay.- Initial Reports of the Deep RAWSON P.F. (1983).- The Valanginian to Aptian Sea Drilling Project, College Station, vol. 48, stages - current definitions and outstanding p. 1025-1059. problems. In: Symposium Kreide.- Zitte- MORENO-BEDMAR J.A. (2007).- Biostratigrafía del liana, München, Heft 10, p. 493-500. Aptiense del Macizo del Garraf (NE de la RAWSON P.F. & HOEDEMAEKER P.J. (reporters), Península Ibérica).- Geogaceta, Madrid, vol. AGUIRRE-URRETA M.B., AVRAM E., ETTACHFINI M., 41, p. 131-134. KELLY S.R. A, KLEIN J., KOTETISHVILI E., OWEN MORENO-BEDMAR J.A., COMPANY M., BOVER-ARNAL H.G., ROPOLO P., THOMSON M.R.A., WIPPICH M. T., SALAS R., DELANOY G., MARTÍNEZ R. & GRAU- & VAŠÍČEK Z. (1999).- Report on the 4th

199 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

International Workshop of the Lower Creta- graphy.- In: RAWSON P.F. & HOEDEMAEKER P.J. ceous Cephalopod team (IGPC-Project 362).- (eds.), Proceedings 4th International Work- Scripta Geologica, Leiden, Special Issue 3, p. shop Cephalopod Team (IGCP-Project 362)- 3–13. Scripta Geologica, Leiden, Special Issue 3, p. REBOULET S. & HOEDEMAEKER P.J. (reporters), 159-213. AGUIRRE URRETA M.B., ALSEN P., ATROPS F., ROPOLO P., MOULLADE M., GONNET R., CONTE G. & BARABOSKIN E.Y., COMPANY M., DELANOY G., TRONCHETTI G. (2006).- The Deshayesitidae DUTOUR Y., KLEIN J. LATIL J.L., LUKENEDER A., STOYANOV, 1949 (Ammonoidea) of the Aptian MITTA V., MOURGUES F.A., PLOCH I., RAISOSSA- stratotype region at Cassis-la Bédoule (SE DAT N., ROPOLO P., SANDOVAL J., TAVERA J.M., France).- Carnets de Géologie / Notebooks VAŠÍČEK Z., VERMEULEN J., ARNAUD H., GRANIER on Geology, Brest, Memoir 2006/01 B. & PREMOLI-SILVA I. (2006).- Report on the (CG2006_M01), 46 p. second international meeting of the IUGS ROUCHADZE I. (1933).- Les ammonites aptiennes Lower Cretaceous Ammonite Working Group, de la Géorgie occidentale.- Bulletin de l'Insti- the "KILIAN Group" (Neuchâtel, Switzerland, tut géologique de Géorgie (1932), Tbilisi, 8 September 2005).- Cretaceous Research, vol. 1, n° 3, p. 165-273. London, vol. 27, n° 5, p. 712-715. SARASIN C. (1893).- Étude sur les Oppelia du REBOULET S. & KLEIN J. (reporters), BARRAGÁN R., groupe du nisus et les Sonneratia du groupe COMPANY M., GONZALEZ-ARREOLA C., LUKENEDER du bicurvatus et du raresulcatus.- Bulletin de A., RAISOSSADAT N., SANDOVAL J., SZIVES O., la Société géologique de France, Paris, (3e TAVERA J.M., VAŠÍČEK Z. & VERMEULEN J. série), t. XXI, p. 149-164. (2009).- Report on the 3rd International SAZONOVA I.G. (1958).- Lower Cretaceous depo- Meeting of the IUGS Lower Cretaceous sits of the central regions ofthe Russian Ammonite Working Group, the "KILIAN Platform. In: FLEROVA O.V. (ed.), Mezozoiskie Group" (Vienna, Austria, 15th April 2008).- i tretichnye otlozhenija tsentralnykh oblastei Cretaceous Research, London, vol. 30, n° 2, Russkoi Platformy.- Gostoptekhizdat, Lenin- p. 496-502. grad, p. 31-184 [in Russian]. RENZ O. (1982).- The Cretaceous ammonites of SCOTT G. (1940).- Cephalopods from the Creta- Venezuela.- Birkhäuser Verlag, Basel, 132 p. ceous Trinity Group of the South-Central RICCARDI A.C. & AGUIRRE-URRETA M.B. (1989).- United States.- The University of Texas, Hemihoplitid ammonoids from the Lower Publication, Austin, vol. 3945, p. 969-1107. Cretaceous of Southern Patagonia.- Palaeon- SEYED-EMAMI K., BRANTS A. & BOZORGNIA F. tology, Oxford, vol. 32, n° 2, p. 447-462, (1971).- Stratigraphy of the Cretaceous Pls. 51-52 rocks southeast of Esfahan.- Geological RIEDEL L. (1938).- Amonitas del Cretácico infe- Survey of Iran, Report, Tehran, n° 20, p. 5– rior de la Cordillera Oriental.- Estudios 40. Geológicos y Paleontológicos sobre la Cordil- SHULGINA N.I. (1996).- Biostratigraphic chart for lera Oriental de Colombia, Parte 2, Ministerio the Lower Cretaceous of the central and de industria y trabajo. Departmento de eastern European (Russian) Platform.- minas y petróleos, Bogotá, p. 7-78. Mitteilungen aus dem Geologisch-Paläontolo- ROBERT E. & BULOT L.G. (2005).- Albian ammoni- gischen Institut der Universität Hamburg, te faunas from Peru: the Genus Neodesha- vol. 77 (Proceedings of the 4th International yesites CASEY, 1964.- Journal of Paleonto- Cretaceous Symposium, Hamburg 1992), p. logy, Tulsa, vol. 79, n° 3, p. 611-618. 89-93. ROMAN F. (1938).- Les ammonites jurassiques et SIBUET J.C., SRIVASTAVA S.P. & SPAKMAN W. crétacées. Essai de genera.- Masson, Paris, (2004).- Pyrenean orogeny and plate kine- 554 p. matics.- Journal of Geophysical Research, ROPOLO P., CECCA F. & GONNET R. (2000).- The Washington, vol. 109 (B08104), 18 p. stratigraphic position of Ammonites conso- SINZOW I. (1906).- Die Beschreibung einiger brinus d'ORBIGNY (Deshayesitidae, Ammo- Douvilleiceras-Arten aus dem oberen Neo- noidea) in the Lower Aptian of Cassis-La kom Russlands.- Verhandlungen der Rus- Bédoule (SE France).- Géologie méditer- sisch-Kaiserlichen Mineralogischen Gesell- ranéenne, Marseille, t. XXV, n° 3-4 (1998), schaft zu Sankt-Petersburg, (ser. 2), vol. 44, p. 159-165. p. 157-198. ROPOLO P., CONTE G., MOULLADE M., TRONCHETTI G. SINZOW I. (1909).- Beiträge zur Kenntniss des & GONNET R. (2008).- The Douvilleiceratidae sudrussischen Aptien und Albian. Verhand- (Ammonoidea) of the Lower Aptian historical lungen der Russisch-Kaiserlichen Mineralo- stratotype area at Cassis-La Bédoule (SE gischen Gesellschaft zu Sankt-Petersburg, France).- Carnets de Géologie / Notebooks (ser. 2), vol. 47, 48 p. on Geology, Brest, Memoir 2008/03 SOLER Y JOSÉ R. (1971).- El Jurásico marino de la (CG2008_M03), 60 p. Sierra de Aralar (Cuenca Cantábrica Orien- ROPOLO P., GONNET R. & CONTE G. (1999).- The tal): los problemas post-kimméricos.- Cua- 'Pseudocrioceras interval' and adjacent beds dernos de Geología Ibérica, Madrid, vol. 2, at La Bédoule (SE France): implications to p. 509-532. highest Barremian/lowest Aptian biostrati- SOWERBY J. de C. (1836).- Appendix A,

200 Carnets de Géologie / Notebooks on Geology - Memoir CG2011/02 (CG2011_M02)

Descriptive notes respecting the shells figu- North Biscay Margin (Bay of Biscay): from red in plates 11-23. In: FITTON W.H. (ed.), new reflection seismic data.- Bulletin de la Observations on some of the strata between Société géologique de France, Paris, vol. the Chalk and the Oxford Oolite in the 173, n° 6, p. 515-522. South−East of England.- Transactions of the TOVBINA S.Z. (1968).- About the Zone Geological Society of London, (Series 2), vol. Acanthohoplites prodromus in the Aptian- IV, p. 103-390. Albian boundary beds of Turkmenia.- SPATH L.F. (1921).- On Cretaceous Cephalopoda Izvestiya Akademii Nauk Turkmenskoi, Ach- from Zululand.- Annals of the South African gabat, (seriya Physico Technicheskogo, Museum, Cape Town, vol. 12, n° 7, p. 217- Chemicheskogo i Geologicheskogo), N° 2, p. 321. 100-109 [in Russian]. SPATH L.F. (1922).- On Cretaceous Ammonoidea TOVBINA S.Z. (1980).- К вопросы о зоне from Angola collected by Professor J.W. GRE- Acanthohoplites prodromus в верхнем апте GORY, D.Sc., F.R.S.- Transactions of the Туркмении [On the Zone Acanthohoplites Royal Society of Edinburgh, vol. 53, p. 91- prodromus of the Upper Aptian of Turkme- 160. nia].- Izvestiya Akademii Nauk SSSR, SPATH L.F. (1923).- A monograph of the Ammo- Moscow, (seriya Geologiskaya), N° 7, p. noidea of the Gault, pt. 1.- Palaeontologra- 142-144 [in Russian]. phical Society (1921), London, 72 p. WARREN P.S. (1937).- An Aptian horizon in the SPATH L.F. (1924).- On the ammonites of the Cretaceous of the Lower Mackenzie Valley.- Speeton Clay and the subdivisions of the Journal of Paleontology, Tulsa, vol. 11, p. Neocomian.- Geological Magazine, Cambrid- 69-72. ge, vol. 61, n° 716, p. 73-89. WHITEHOUSE F.W. (1926).- The Cretaceous SPATH L.F. (1930).- XLVI. On some Ammonoidea Ammonoidea of Eastern Australia.- Memoirs from the Lower Greensand.- Annals and of the Queensland Museum, vol. 8, n° 3, p. Magazine of Natural History, Oxford, (Series 195-242. 10), vol. 5, n° 29, p. 417-464. WIEDMANN J. (1966).- Stammesgeschichte und STOYANOW A. (1949).- Lower Cretaceous strati- System der posttriadischen Ammonoideen. graphy in southeastern Arizona.- Geological Ein Überblick.- Neues Jahrbuch für Geologie Society of America, Memoir, Boulder, vol. und Paläeontologie, Abhandlungen, Stutt- 38, 170 p. gart, vol. 1, n° 125, p. 49-79; vol. 2, n° STOLLEY E. (1907).- Über ein norddeutsches 127, p. 13-81. Äquivalent der Clansayes-Fauna Südfrank- WRIGHT C.W. (1955).- LXIX. Notes on Creta- reichs und der Schweiz.- Zentralblatt für ceous ammonites.-II. The phylogeny of the Mineralogie, Geologie und Paläontologie, Desmocerataceae and the Hoplitaceae.- Stuttgart, vol. XX, p. 266-270. Annals and Magazine of Natural History, STRASSER A., CARRON M. & GJERMENI M. (2001).- Oxford, (Series 12), vol. 8, n° 92, p. 561- The Aptian, Albian and Cenomanian of Roter 575. Sattel, Romandes Prealps, Switzerland: a WRIGHT C.W., CALLOMON J.H. & HOWARTH M.K. high resolution record of oceanographic (eds., 1996).- Cretaceous Ammonoidea. In: changes.- Cretaceous Research, London, vol. KAESLER R.L. (ed.), Mollusca (revised).- Trea- 22, n° 2, p. 173-199. tise on Invertebrate Paleontology, Geological SZIVES O. & MONKS N. (2002).- Heteromorph Society of America, New York; University of ammonites from the Tata Limestone Forma- Kansas, Lawrence, Part L, Vol. 4, 362 p. tion (Aptian-Lower Albian), Hungary.- Pa- ZITTEL K.A. von (1884).- Cephalopoda.- In: laeontology, Oxford, vol. 45, p. 1137-1149. ZITTEL K.A. von (ed.), Handbuch der Pa- SWINNERTON H.H. (1935).- The rocks below the laeontologie.- R. Oldenbourg, München & Red Chalk of Lincolnshire and their cepha- Leipzig, vol. 1, n° 2, Edition 3, p. 329-522. lopod faunas.- Quarterly Journal of the Geo- ZITTEL K.A. von (1895).- Grundzüge der Pa- logical Society, London, vol. 91, p. 1-46. läeontologie.- R. Oldenbourg, München & THINON I., REHAULT J.P. & FIDALGO-GONZÁLEZ L. Leipzig, viii + 971 p. (2002).- The syn-rift sedimentary cover of

201