ISSN 00310301, Paleontological Journal, 2010, Vol. 44, No. 2, pp. 188–191. © Pleiades Publishing, Ltd., 2010 Original Russian Text © A.V. Khramov, 2010, published in Paleontologicheskii Zhurnal, 2010, No. 2, pp. 64–66.

A New Lacewing (Insecta: : Grammolingiidae) from the Upper of Mongolia A. V. Khramov Moscow State University, Moscow, 119992 Russia Borissiak Paleontological Institute, Russian Academy of Sciences, Profsoyuznaya ul. 123, Moscow, 117997 Russia email: a[email protected] Received April 2, 2009

Abstract—A new lacewing species, Leptolingia shartegica sp. nov., (Grammolingiidae), from the Upper Jurassic of the Mongolian SharTeg locality is described.

Key words: New species, lacewing, Upper Jurassic, Mongolia. DOI: 10.1134/S0031030110020103

INTRODUCTION rowing towards the base. These fossils represent a new species, compared below with other species of the The SharTeg locality in southwestern Mongolia is genus. This is the first finding of Grammolingiidae one of the most important Jurassic Lagerstätten. outside the type locality and one of taxa that allow Diverse fossils of plants and , including mem comparison between Daohugou and other Asian bers of groups rarely recorded in the Jurassic, such as localities. dipnoans, labyrinthodonts, theromorphs, and mam mals, were collected in this locality. Other fossils found there include crustaceans, fishes, turtles, and dino MATERIAL saurs. The group represented by a particularly great The material is stored in the Borissiak Paleontolog number of taxa is : members of 22 orders and ical Institute of the Russian Academy of Sciences about 200 families were collected. The composition of (PIN). Photographs show the bestpreserved frag insects indicates the Upper Jurassic age of the deposits ments of parts and counterparts. (Gubin and Sinitza, 1996). About forty fossils of neu ropterans were found in the SharTeg locality, suppos edly representing the families Mesithonidae, Osmy SYSTEMATIC PALEONTOLOGY lopsychopsidae, and Kalligrammatidae, which are Family Grammolingiidae Ren, 2002 preserved mostly as scattered wing fragments. Genus Leptolingia Ren, 2002 In addition, four wings similar in shape and vena Leptolingia shartegica Khramov, sp. nov. tion, two forewings and two hindwings, were found in Plate 12, figs. 1–5 the locality in close proximity to each other. This sug gests that they belong to a single individual. Sc and R E t y m o l o g y. From the SharTeg locality. are positioned in parallel, not confluent until the wing H o l o t y p e. PIN, no. 4270/5001, almost com apex; the costal field has two rows of cells in the forew plete left forewing; Mongolia, Gobi–Altai Aimak, ing and one in the hindwing; MP and CuA have southeastern boundary of the Aj Bogd Range, Shar dichotomizing branches; between A1 and A2 of the Teg locality; Upper Jurassic, SharTeg sequence. forewing, there are more than one row of cells; nyg D e s c r i p t i o n (Fig. 1). A large lacewing. The mae are absent and the trichosors are present. These wings are abundantly pubescent (large longitudinal characters strongly suggest that the specimen in ques veins show traces of setal bases arranged in several tion belongs to the family Grammolingiidae, which rows; transverse veins have one row). Nygmae are was recently described from a Chinese locality near absent. The trichosors are present. the village of Daohugou (Ren, 2002). The The margin of the forewing is convex basally, should be assigned to the genus Leptolingia, since A1 slightly concave medially, and gradually becoming ends proximal of the beginning of the first branch of convex again towards the apex. The apex is somewhat Rs, whereas, in other genera of this family, Litholingia pointed; the tornus is in the middle of the posterior and Grammolingia, A1 continues further. Members of margin of the wing. The lengthtowidth ratio of the this family have wings of the kopepterygial type, nar wing is 2.6. The costal field has two rows of cells. The

188 A NEW LACEWING 189

h Sc

R1

MA

MP Rs CuP CuA с A1 (a)

Sc (с)

(b)

Fig. 1. Wing venation of Leptolingia shartegica, sp. nov., holotype PIN, no. 4270/5001: (a) left forewing; (b) right hindwing; (c) setation (bases of setae shown as dots) in subcostal area of the left forewing. Scale bars: (a, b) 1 cm; (c) 1 mm. humeral vein is discernible close to the base. Sc and R1 costal field has one row of cells. As in the forewing, Sc are parallel along the entire wing length, converging and R1 are parallel and converge only at the very apex. only at the apex, dividing into several short branches. CuP lacks branches; A1 and A2 are shortened, with one Sc and R1 are connected by abundant transverse veins. row of cells between them; A3 is not developed. The abundant transverse venation is also wellpro Measurements, mm. Forewing length, 54; nounced in other parts of the wing. Rs has seven forewing width, 21; hindwing length, 56; hindwing branches. MA is fused with Rs at the base of the wing width, 18. and almost immediately branches from it. MP bifur Va r i a t i o n. Several minor differences between cates once distal to the point of deviation of MA from the left and right forewings and hindwings are found in . bifurcates distal to the bifurcation of , Rs CuA CuP the posterior branches of veins. For instance, in the opposite the first bifurcation of . bifurcates dis Rs CuP right hindwing, R5 bifurcates not far from the point of tal to the branching of MA from Rs and proximal to the deviation Rs, whereas, in the left hidnwing, R5 does first bifurcation of Rs. The anterior branch of CuP dis not bifurcate; in the fragment of the right forewing, the tal to the bifurcation of CuA is also bifurcating. A1 is penultimate posterior branch of Rs (probably R6) long, terminates just proximal to the first branching of bifurcates close to its beginning, whereas, in the left Rs. Three rows of cells are visible between A1 and A2 forewing, the posterior branches of Rs bifurcate closer closer to the base of the wing. A3 is poorly developed. to the margin of the wing. The distal margin of the wing is indented with a num ber of small incisions (Plate 12, Fig. 5). Comparison. L. shartegica is easy to distin guish from L. jurassica Ren, 2002 by the following The anterior margin of the hindwing is curved more combination of characters: MP in the forewing bifur evenly than that of the forewing; the hind wing is more cates distal to the branching of MA from Rs; A1 ends elongate than the forewing (the lengthtowidth ratio virtually opposite the first bifurcation of Rs (in is 3.1); the tornus is displaced towards the base. The L. jurassica, MP bifurcates opposite the branching of

PALEONTOLOGICAL JOURNAL Vol. 44 No. 2 2010 190 KHRAMOV

Plate 12

1

2

3

5

4

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Explanation of Plate 12 Figs. 1–5. Leptolingia shartegica, sp. nov., holotype PIN, no. 4270/5001: (1) left forewing; (2) right forewing; (3) left hindwing; (4) direct impression of right hindwing; (5) notches in the distal margin of left forewing. Scale bars: (1–4) 1 cm and (5) 2 mm.

MA from Rs, and A1 ends distinctly proximal to the ACKNOWLEDGMENTS beginning of the first branch of Rs). The new species I am grateful to A.G. Ponomarenko (PIN) for valu differs from L. tianensis Ren, 2002 in the presence of able advice and to A.B. Mazin (PIN) for help with the three rows of cells between A1 and A2 in the forewing, photographs. the presence of two rows of cells along the entire costal This study was supported by the scientific program zone, and the more elongate A1 and A2 in the hind “Evolution of the Biosphere.” wing. In addition, L. shartegica differs from the two mentioned species in the sickleshaped expansion of REFERENCES the forewing’s posterior margin. 1. Yu. M. Gubin and S. M. Sinitza, “Shar Teg: A Unique M a t e r i a l. In addition to the holotype, an almost Mezozoic Locality of Asia,” Mus. North. Arizona Bull. 60, 311–318 (1996). completely preserved right hindwing and fragments of 2. D. Ren, “A New Lacewing Family (Neuroptera) from another forewing and another hindwing. They all the of Inner Mongolia, China,” Ento probably belong to the same individual. mol. Sin. 9 (4), 53–67 (2002).

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