Studiesin the genusAcacia (Mimosaceae)-4 A Revisionof the SeriesPulchellae

By B. R. Maslin

Abstract

A taxonomic revision of ,4cdcta scries P lchellae ;s presented Wilh the- exceotion of A. nir.h,llii *hich occurs in eastern Australia, the nembers of it i. \.ri.s are eudernic to south west western Aust|alia. The lwenty-four specics recognized comprise lblty five laxa Twenty-eLght new nanres or conrbrnallonsarc PfoPosec. The morphology of the Seriesis discussedand kets to speclesencl-rnlra- specific taxa are piovided. In most cases tlle taxa arc ilLustretcdand lhcrr distribution nrAPPed.

Introduction

Bentham (1842) proposed thr: first classificationof the genus '{cncrn By in which subgineric catcgorieswere formally named (see-Ross, 1973) ennloving bo"th reretltiie rnd inflorescencccharllcters Benlham divided the'se;us-intosix Seiics.oue nhYllodinousand live bipinnate. The Pulcltellqe, toseilrer \.\itlr 111sg,,rlyot,ph,tltte r.nd Gunmiferae, ale tlie only bipinnatc Seiiesrvhicl.r occirr in Australia -the first two being endemicto tllis region In generirftcrms. the Pulchellae can be distinguished from the Botryocephalae bv"their simple or extremelyreduced racemoseinflorescences, and fron the G'umntiferaeby normally having scariousstipulcs. Althoueh thereare dei]cienciesir Bentham'sclassification it is a convcrlient iv\temlbr g-rotloilrgthe .pccies of thi' r:r\l pentl\. l heretbrc.for tlrcptttpo'c df rhi. rc',iiiorr,Be-nthunr'' lrealnrertl has beett accepled. It sltoul.lbe IloLcd' ho\\cver, tlrat the Pulchell(1eare a diverseancl possibly Llnnaturalgrotrp oi spccies. Unlessotherrvise indicated, thc specimcnscited in this paper are ltoused at tl'reWestern ALlstralianHerbarium (PERTH). In most instancesonly a selectionof specimensis listedfor eachtaxon. The abbreviatiorrsfor herbaria are those siv'enin Index Herbariorum pt l ed 5 (1964),rvilh the addition of UWA for-the Departmcrlt of Botany, University of Weslern Australia (see Taxon l5(8):336,1966) AU illustrationsappearing in this work have beerrdrawn fron.r presscd herbariumspccinrers. Unless otherwiseindicated, the measLuemcnlsgiven in the descriptions below are fion dried herbarium material

Morphology This chapter provides definitions for the more specializedterrns wllich aooearin thc Laxooomicdescriptions below. Many of theseterms arc illus- traied in Figure l. In addition. some of the generalmorphological attributcs of the Pulthellqeare discussedhcre. l. Habil:About half the speciesin the Pulchcllaeare from 0 5 to 2 metres t"tt. fl. remainingtaxa aie either tallcr shrubsfrom 3 to 5 meiresin height (2 species);dwarf, drcct shrubs not exceeding0 6 netrcs in height (17 taxai; or prostrate(3 sPecics). 388 68€ qrrq,r sJr"rlJ,o Surlsrsuoc unluelunpur uc ol pJrlddEsr urJJlsrrll .uon€uelclxo .IeJJqr seJrnb3J,noleq su^orldrrcsapaql ur ,([u5nb:rj ,{L:,1sjna-idt qcrLl,lrrrcy -ntaqnd .1 lulret eq1 orn8rg ur patullsnllr aru :aded irql ur pesn sarifl uinluaui -npur eql selcet?qc crLuouox?t ,esloJleJ 'esJorlu€ Jo. lsotr{ lnJJSnJtE (gulpeuds Jo e r) uorlcerrppu? qt8uelIlErJJdsJ.,{BoloqdroLu rieg :!unluawnpLrJ.Z o1 rradde. osl! sreuunr .sr$rrs urlu€qreq 'JuA '.t?drar./ss'.t) eluos uro{ {i,:!:;fr'::}".r:r:, pue .J.a^ ,DlJLlt .J.t:li Drtn4.y ,.roullu rll-t .y '!.taq)tpaa pup, nttmta,tp JeL DuDlLtoJq 7 aJa papJoceJueeq seq Burra>1cnsqJrrl,{ roJ ExEl a.rnpo.rddnor8 snlt ::]! ,-1"qt Jo $equeur ltg tiru lnq',a.rocleu1 p".,.r"j", llYBl Jlc\1p Jo dnoJb arF o) pJlJlrtseJeq o1 ruedde stelcns ,ell,p ol olqele^" uolllruJoJureq1 rll!A\.slouunJ u"au?rlelqns uorJSruo{ dole^epqcrqt\\ st lans Jo uol1cnpold erll s aD aqfld orll ur eJl.rleal uoruLiocuir fje,rililer y stu.rolJo .aDtptprnd uorrrug.proJ88t d .es eql Jo sernreeJl€.rfcroqdroru J'"tatli':1t"t;,7

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- r!Ed I apuu!d- lrv9nflNn (t) slnvl-l are short, soft, and spreading; the hairs may be either straight or tortuous. Ifthe haiis are directed upwards (towards the apex of the orgar) the i.ndumentum is termed antrorselypubiruloas (ihese hairs are less appressedthan in antrorsely stfigose). 3. Spines: Orly thlee closely related specieswithin tlre Palc/rel/ne-possgss axiflarv spirres.'viz. A. lasiocotpa,A. megacephala.and A pul,htlla Tlrc soines-nremodifled brancheslsee Vassal, 1'970,p.24), occurring at the nodes t^ogetherwith a pail of scarious stipules. The preselce or abselce of spiles, their frequency, and their number pet node li.e. soUtarv or in pairs) are useful taxonomic charactersbut undue reliance shonld not be pliced on them in differentiatingtaxa. In both l. lasiocqrpa and A. ptrlchella spinelessindividuals often occur within varieties which ilpicalll posses5a\ill;ry spines:only one la\on. A lario'arya \ar. villosa, ii con;isiently spineless. Wherr spiles occur on A. lasiocarqq they are always solitary ai tie nodes; likewise, one-spinednodes -are normal lbr A. megaiephala, but a two-spined variant has been recorded for this species. On A. putchetlathe number is variable with both one- and two-spinednodes occurnng.

4. Leaves: Because past authors have frequently applied different terms to the individual parts of compound leaves,the prese[t author's usageof foliar terminologyis here defined. \i\ Jugate-yoked together, in pairs. Used here with reference to the pinnae. fhe word is oiten prefixed by a numeral, for example, 3-jugate ieaves possesstlrree pair of pinnae. See also multijugate a\d unijugate. (ii) Multijugate /eales possessingmore than one pair of pinnae. (rir) Petiote t\e stalk of the leaf, i.e. that portion of the primary leaf axis situatedbelow the proximal pair of pinnae. (ivl Petiolule the stalk of a pinnule. (v) Pinna, pl. Pinnae-pnnary division of a bipinnate leaf. In the des- criptions below. a distinction is made between tbe proximal (lowermost) and distal (uppermost)pair of pinnae. (vi) Pinna rachis-lhe pinnule-beariog axis of the bipinnate leaf. This term was used by Windler (1966) with reference to Neptunla- (vil) Pinnule (: Leaflet)-the ultimate division of the bipinnate leaf' (viil) Primary leaf axis the combined petiole, rachis (when present), and terminal seta. (ix) Raclru, pl. Rachides-lhe pinnae-bearing axis of the bipinnate leaf, situaied above tlie petiole. (Tlis term is conmonly applied to the pinnule- bearing axis of a pinnate leaf.) (x) Terminal seta-the appeldage which telminates the rachis (of a,multi- jugate.leaf) or the petiole (of a unijugate leaf)-see Maslin (1972), for further details. (.xr) (lnijugate /eares-possessing one pair of pinnae.

5. Gl(tnds: Glands are present on the leaves of rtost speciesrn the Pulchellae but sometimes they are quite small and not readily observed. Both gland structure alld position are ofconsiderable taxonomic value in this Series. The only specieswhich Iack glands are A. insolita (doubtfully included in the Pzl- chellae) znd A. mitchellii (but see p. 471). On the basis of gland morpholo gy, the Pulchellae can be divided into two major groups, one with stipitate glands and the other with_ sessile glan-ds. Th; fiit gioup includes A. lasiocarpa, A. megacephala, and A. pulchella; 390 r62 .pu?l3 Jo ocuesold: x :(ele8nftllnu oJe slelcElq tnoqlr,4A srrJros)so^Er[a]pbntun: IOI aDlaqrlndaq] ut spuelSJ€el elrssas Jo uo4nclrrtsrc I alqel

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7. Seeds: Seedorientation in the legume is either longitudinai or transvcrse (to oblique). In the llrst case,the lotrg axis of the seedlies parallelto the lo|rg axis of the legume,while in the secondthe lorg axis of tl'e seedis normal (or inclined up to forty-fi\,edegrees) to the long axis of the legume. The long axis of an orbicular seedis taker as being a line drawn from the middle of the aril and passingthrough the centre point of the areole. A pleurogramoccurs on all seedsexamined. It is either unbrokel (i.e. continuoLrs)or open towardsthe hilum. The area boundedby thc pleurogram ;s termcd the areole(see Maslin, 1972,for further details). The arll is generallydefined as an expansionof the funit:le(Iacksor, 1928); however, as Vassal (1971, p.213). points out, there has been considerable confirsiou in the past regarding the application of theseterms. About half tlre speciesi\ the Pulchelluehave a filiform, or almost filiform funicie which abruptly thickensinto a large fleshyaril. In the rerraining speciesthe funicle tendsto graduallyexpand into the aril without a clear distrinctionbelween the two $truclures.

Relationships within the Pulchellae Tlre seriesPulchellae as circumscribedhcre consists of a diverseassemblage of twenly-four species. With the evidenceavailable to date (which has been derjved almost entirely from comparative rnorphology) it is rtot certain whetherthis Seriesconstitutes a natLlraltaxotomic group. Although a number of natural sub-groupshave been recognised (see Figure 2) the absolutcrelation- shipsbctween thesc have not beendetermined. Data from anatomy,cytology, pollen and seedlingrrorphology, phytochemistry,and other studiesrvould bc iecessaryin an attempt to reve;rlthesc relationships. Such stuciiesare outside tlre scopeof the presentwork.

ConcerningFigure 2 the follorving points shouid be noted:

l. Acqciti ln;olitu. The true position of this specieswithin the genus ,4cnclais somervhatobscure. In this speciesthc uppermostleaves are reduced to pbyllodes,tlrerefbre in the strict senseit shouldbe includcdin thc Phyllodineae rather than the Pulchellqe. However, because,4. insolita has traditionally been jncluded n the Pulchellae,I lTaveacccpted it for the purpose of this rerision, t6t. . oqd rssocr,{zrur"\1, " o nq " ":,:l';11?J#$9,':lii'Jllr";rt:];]:l rluelsrp lq n 1,a.sJ3lJ"J"rlJ t"rl,reruos oq ol J€edd" sercedsaseql acuecselogur pirri e8rrlol palrun .y pluapDuad Iltra^o Jrerll ,{q eJE D|outa)D.tqns puB u)Dy V '(\\olJqg ou) sJtJJdsJodrroJa DuDtu.notq eqt (\1oltq / Jo l.ou) dnoJAttpuoutturup.g-oqt.,:qi,a pue 1rlo1aq ou-) 9 onoJi ollaqrlnd k, Jrlt uJJ,rlaq e.reri,\\oruosSunjncco se 3o lq8noql eq ue) [aq1 aD dLplnd eq:]Jo Japut"ueJ eql rlll,r serltus:E snor^qo-ou',,41snoeu41nuts €,\"q "x"1 eserJlqSnoqtlv J€-elJtou sr ruerll uee,rl3q dqsuoijeler eq1 peleprsuoceJ€ sarceds eseqt selnqu]tu ,Jene,\\oH 'spue13 Jo ltrrSoloqdtoLuJrlt IT?uJLI^\ rrlotled elrssospue so,{"elJlc8nfrun ,(JtuJlstsuof, .j'n ,sr:1re.r"r.1"",*uu -oxBl luelloorur o,{\1elEqs rai,aqt$au.y puB ,ll,tlout .v ,pssatdappDo)v ,t 'puo! 't' atp ue,\a lo'DuDlsstar[.k Jo ltpuoLuLuntp.f.01 pa]"leJ .{1lur1srpec1 plloc }r .ruep :serlruu€ ul?lJeounqlJ,r setcedsJeqlou" sr t:4ott1dotcD:tV- 1ou sr sarre5 .aplpLpfd eql Jo sreqrueru Jaqt_oeql ol seDads oserll Jo drqsuorl"IeJ e\ll eql JoJ Isnsnun elrnb utarlt.repuer qcrrl,r sJalc€Jerlccrurouoxrl ,l/ luirr6our ororu Jo euo ssessodll11a{Jll1./ pua ,lll"taqll8 ,k, ,paoltatp \DDV .Z ',{lru!UE urelJecun selealpur eurl uolorq p :dlqsuorlBlJ.l lsolJ sLqEqoJd selecrpur eurl p:p:rElr.l tle.olJ rrp pelJolq \ep>ds l.los dott,q.l,rd.5re,:rir u,.jrrrugy ; ern3rj llLrvHd r )o uro8 s!ru3s

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lltt,aqtls ol otoqdato6sur V odrDto!sol v Dtl€qrlnd 'v 5. Acacia preissiona and A. lallorian4 appear to be related to each other. The group with which they seem most closely allied is l. drummondii and its allies(no. 7 below). 6. Acacia lasiocarpa,'4 . megacephala,and A. pulchella are the on)y species in the Seriesrvhich have axillary spinesand stipitate glands. These species constitute a natural group and are quite distinct from the remainderof the

'7 . Acacia (lrummondii logether with ,4. luteola a[d A. t'qriq form a natural group whose members share the following characters:obloid to cylildrical flowcr heads, ebracteatepeduncle bases,triangular-shaped calyx lobes, and glands rvhicb are norn.rallysituated on the petiole. As mentioned above, A. prcissiqna and A. tayloriana (perhaps also A. plicqta\ appear to have some alinities witl.rthis grorlp. 8. Atacia brovttiqno together with A. empelioclatla,A. grisea, A. lateriti- cola, A. leiodernn, and A. nigritons form the largest natural group within the Pulchellae. Thesc species are urited by the following attributes: axillary spines abscnt, leaves normally multijr-rgate,gland sessile(never present on petiolc), pedunclesglabrous and with 3 4 fairly conspicuousbasal bracts (except.4. nigricans),flower headsglobular, and calyx lobes oblong, inflexed and slightly thickenedat the apex.

SERTES PULCHELLAE Slrribs (rarelysmall trees)0 3 2 (3 5) m tall, sonretimesprostrate; axillary .?ireJ present or absent. Sliptlcs scarious, rarely spiny. lecre.r bipinnate, rarely ihe uppermost reduced to phytlodes (A. i solittt), articulate on branch, rarely decurlent; pinnae | 3 (4-8) pairs; rachis 1-15 (55) mm lorLg1.tetminal scld scarious,rarely spiny pinnulcson unijugateleaves: (l) 2 10 (12) pairs,on nrultijugateleaves: l-3 (4 5) pairs on proximal pinnaeand 2 10 (12-30) pairs on dlstal piutae. Glant{ sessile or stipitate, situated on upper surface of rachis ancl /or petiole, rarely on pinna rachis. Infl.orescencessimple or some- times shortly racemose; peduncles glabrous or hairy, base ebracteate or with up to 4, + conspicuous bracts: floh,et heaclsglobulat to cylindrical. F/owers 5 (rarely 4) -merous; cal1,-rnormally connate for ca. ] or more its length, rarely t;uncate or divided to base,lobes oblong to triangular; orar1,sessile, rarelt stipitate. Legumes normally somewhat hard and bdttle, rarely firmly cl'rartaceous,flat (rarely plicale); mlrgins normally thickened. Seerls trans- verseto longitudinal in legume;pleurogram continuous or open towards the hilum funiclc filiform or dilated, reflexed below a thickened straight to once- folded aril. Type species:Acacict pulchella R.Bt.

Key to Species based on dried, Jlo|,erlr?g l11aterial. la. Flo\ er headsc)lindrical ... 2 b. flower head\ globular or obloid 5 2a. Pinnulesfla1 oI slightlyrecuwed...... 3 b- Pinnules prominently recurved to revolute 4 3a. Pinrules glabrousor ninutely strigoseon margins. widespread. 20. A. drunmondii (Figs. 28-30) b. Pinnules t conspicuouslyhairy. Widespread 19. A. Yaria(Figs. 2+26) 4a. Flower heads very pale yellow (+ sulphur-coloured),obloid to shortly cylindrical (6-12 mm lonc)l pinnules recurved 10 revolute, 2-6 x 0 5 1 mm, normally glabrous or "parsel! hair]. (Denmark to Cheyne Beach, north to Stirling Range) 18. A, luteola (Fig. 23) b. Flower headsnot + sulphur coioured ,?r if almost so then headsdistinctly cylindrical and DirlnLllecharacters not combined as above. Widespread and variable .... 19. 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(Sl 8IC) BpelJoladura (uerre{ luno^ rsuC ot tselA) uoturuoo toN luosqe selllds J.rElltxe lsno.rqEl? ll3leldNor sol.unp.d p6 6 - ro seurds,{JeJJrxB ,snolt€cs luesq€ luase.rd .q :snouris sel[dr]s:iuoi ru(l €-S.0 E]esIeurluJel:qoue_rq uo a]elnatt.tuse^se.I ()-fl€ 3'C) suo8unddesqns euBr^\o.rp.V ,JZZ (lerxeuu€A\rteN) 'er€l{ seurds,{rellrxE 'lua8und luosqr :tue.seurds+ selndrts:fuol LuuI8t 0I pu€ elplnqns"tos lEurLuret:q3uelqiuole luerrncop ,tl8Lrorlise^€e"I eg -q OI leu selnuurd 8' p"ijl"' so|lLrLr .E/ 6t .... :]]lr*'].." al1u"url:'d (seuo .q elDSnfrllnltl eq1 SuoluE Jnaco ,(tru so^€elele8nfrun .\eJ D) rrsd I u€qt eloul eeuurd L (ele8nftunso^Eal a l) lrud I ,{lluelsrsuoaeeuurd .1r9 ,peJnoloc-ueoJc 9 peaq Jod ojoLu Jo O[ lo \ollel .snoloLlr-gsje,roJl q '€z (zt Brc) Ilucql!ts'v (trlluua( pue ptsn8nvot IroA ruort JrpProos) qrrph ot lrqrurrru uro4 \rr\\oll ..ULU \ crlnuu,cl . s'\rorrLU.t [ 0Z.g :nLroltulu tl I tlottrd:\norqplF.oqruE.rq:p?rqred g Z.r q\\ .reaoj_l Eg b. Leavesarticulaie on branch; petiole presentalthough often very short (:t 1 rnln).... 20 20a. Branchletsglabrous .... 21 b. Branchletshairy ...... 2l 2la. Pjnnules somewhat thickened and paoninendy revolute (these are glabrous variants of nornally hairy species) .... 22 b. Pinnulesflat or slightly recurved 24 22a. Pinnules4-5 pairs; rachis ca. 15 n]nr long; llower headsglobular. (Near lltjl",".,"r" b. Pinnules2 3 pairs;rachis 2-4lnm long;flowerheads obloid to shortlycylindri;d.... 23 23a. Flower heads pale yellow (+ sulphur-coloured); branchlets prominently ribbed. (Aibany to Stirling Range) 18. A. luteola b. Flo\\er'heads deep yellow: branchletsnot prominently ribbed. (Lucky Bay, east of Esperance).-.. 19d. A varia var. paryiflora 24a. Leaveslarge: rachis 10-55 mm long, pinnae 2 5 (6 8) pairs, distal pinna rachis 25 80 n]m long; pinnules 12-30 pairs, obliquely iruncate at base, t sessile(i.e. petiolule extremely reduced). (Pembertonto near Albany) 13. A. pentadenia(Fig. 18) b. Leavessnaller than above; pinnules2-8 (13) pairs, :L rounded at base,petiolule quite distinct ...... 25 25a. Flowers 28 35 per head; gland (small and sessile)normally present on upper surface of distal pinna rachis below LlppermostI 2 pairs of pinnules; braochesprominently libbed; pinnules5 1l mm long. (Albanyto Mount Barker,eastward to West Mount Barren) 12. A. leioderma(Fig. 17) b. Flowers 14 21 per head; pinna rachis eglaDdulose;branch ribs often less prorninent than above (theseare glabrousto glabrescentvariants of normaily hairy species) . 26 26a. Pedunclestwinned on an extremely short racemeaxis,2 6 per leaf axil. (Esperance district) ll. A. nigricaDs b. Peduncless;mple, I (2) per leaf axil. (Albany to Busselton) ... 7. A. browniana 27a. Sepalsfree to base, linear-spathulate;leaves eglandulose. (Eastern Australia.) 21. A. nitchellii b. Sepalsunited; gland presenton upper surfaceof petioleoi rachis often small. (Western Australia) ...... 28 28a. Peduncleshairy to some degree;calyx lobes nofmally triangular ...... 29 b. Pedunclescompletely glabrous; calyx lobes oblong ...... 36 29a. Pinnulesflat to slightly recurved .... 30 b. Pinnulesslightly thickenedand prominently recuwed to revolute ...... 33 30a. Petiole 7-17 mm long; pinnules9 I I x 3 5 mm, + prominentlyrcticulate and antrorselv lrispid below; shrub prostrate. Flowen in January. Rare. (Augrsta district) 16. A. tayloriana(Fig. 21) b. Petiole1- 3mm long; pinnulesnot as above;shrub erect ...... 31 31a. Flowers 40 55 per head; pinnules 2 8 pairs, denselyhairy, mostly above 6 n]m loDg' (Hill River district) .... 17. A. plicata(Fig. 22) b. Flowers14-20 head; pinnllle characters not combinedas above ...... 32 J2x. Pinnules7 1l pairs. sessile(i.e. petiolule+ absent);pinnae 3 6 pairs; calyx lobes trianglrlar:peduncles + conspicuouslyhairy. (Near Augustato Witchcli{Te) 14. A. suhracernosa(Fig. 19) b. Pinnlrlesnormally 3 7 pairs, petiolulepresent; pinnae normally 2 pairs; calyx Iobes oblong; peduncles rpafsel) hairy. (Nannup .o VoLnl chudalup) 7c. A. brownianavar. obscum(Fig. 12) J3o. Flouer headsdrstinctll globular;leaves relatively large: rachis 4-13 mln long,-... distal pi na rachis7 20 mrn ]ong.pinnules 2-8 pairsand 4 15 mm long ...... 31 b. Flower heads obloid (i.e. slightly longer than bioad); leaves normally smaller than above (neglectloliage on juvenile plants): rachis l4 nrm long, distal pinna rachis 1.5 8 lnm long,pinnules 2-4 pairsand 2 6 mm long ...... 35 -14a. Flo\\ers 40-55 per head; hairs conspicuousat apex of bracteoles(observe unexpanded infrorescence);peduncles puberulous_ (hairs < 0 5 mnr long.);pinnulesdensely puberu- lous, normally 6-15 mm long; bbruberect. Flowersfrom ArLgustto October. (Hill River disirict) ... 17. A, plicata (Fig. 22) b. flo$ers 20-28 per head: hairs_notconspicuou\ on bracreole.:peduncle, pllose (hairs ca. I mm long): prnnule,iless harry rnd normallyshorler lhan above:shfub pro(lrate. Flowers from Decemberto January. {Sporadicfronr Bindoon to Albany) 15. A. preissiana(Fig. 20) 396 L6t .,4Juq sutS.tou!:u$o)q ot lq8rL ro sno:qel8 ,speosJo^o pesr€l,ot"lnpun 'ruLulJ€p Allqbrlsro tuu (6 y 'Suolqo ,(l,rorr€u srrlr8a1 pa^reu-l 'luacserqelS /) 9'8uo[ f 0g 9I 01 snorq"lS ulLLr(.g.d Z S.I sl7tad :perreu-S ^lturur -ruoJdo:l ssele^r?upue snopreqnd ,{lesrudso1 snorqrlSeqnl ,soqolet"lorpc Jo 'Dlloror snorqelS 3uo1qo olur rlt8uel s]! +-i roJ pepr^rp .Io qfuel +-1 \'t!D) :snororu-S s.rr.rl.,1J elqEu"A 'lcete (po8uBl.I" Flprrds) seues ;oddn lxede 'pexeuer tB pexouurpur 8uo1qo+ ,{llEtuJou (rlJutdacoJJo ?sEqle 8uu ol€urs € ur pe8u€J.I") ssrJosJa,\\ol :snorldrorxrp sapapD,tg sJe,uog(0g) rpr 'telnqol? 0t,0I spodLl.la[ora:furBri ro snorq?l8,3uolruur (02) red sapun -pad SI Z,lrx" E-l :pzJq lsou.Ire,roleql Jo ltxe uror; Sursrreelcunped puoces ? seuDeuos {Euo[ ulLuf ol)-sl\E JuriJe] uoqs ,{.le^eql uo lc€lq lsoluladdn aqlJo ruo{ '(eTdLuls .Jed lrxs ;ulsrJ? elJUnpJd st tt ateLl/r\sap!()luo8D/ uo ldecxe)eulec?t pesuep -uoc,{re,\ e u sanlil pet"lrp ,,{lFurloLrxodu ,snolnraqndsoulneruos lo '3uo1 sno:qe18 ruor (9.0 >) 'e"uurd uorlcunl 8utsu" ,el€]rdr]sprrl, '(peuo{crqt t-l Jo lE ro eug e^reu) ,roleq pe^Jeu-l [IIE\uJou '(Dxallat Jea uo ,roloq pu" e,roqe snolnreqnd) e1elol1rcro snorqelS 'snorn?18o1 uee:8 '1eg 'iuru (E.e) 'o1e,roqo 'srred 9.1-9.0 x (9) 9 y (,{l,rolr"u) o1 3uolqo (,{l,r\ou€u) (11) g-7 s'alnuud :atelo\ra flesr?ds ro snorqelS ,{lleuuou ,8uol ruru (91) stLpp.t ,.rBln8u"ul0l I G. 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raupreD uou'(tEgt)gtr:€ ?rouueu'D.l.seu xe .rroH^.,,r^o)\"ro31)'t 16e6t) v J 'qlew 10I8I) 0€9:Slddns ( log) lr,{.uE'.llod ( rg ){) D aqr\tt Dsaatw ',t\olaq .ZZ€t .oN uorssncsrpees (uees oloqd-Ng) lleuueB l 'rsurlE4snV ',. .,ua; .1rog...S Z0g r;ll'u^\orB U .. {Z) ( u.rorg rl,! .reN) ,, (1) 'Z .J.lrv IIoH .1 :s.ldttuAs (fI8L) t9t:S pe ,ra) UoH ur.:g.g e11aqc1ndelaecy (tI 8rC)BlorlrrJelat .V .6 (uotrrqurdor E.qJnt\l) so|1uurdsnolnprdsrq ,{lesreds slr€d _ Jo 9 Z .q nurrtiq puei{uol LuLu Sl S \rrlJpreuurd le}srp :selnuurd Jo eseqLLToJJ tlrasqe spu€lD (SI Brl) qBlroledure'y .0I (uJJrPg .salnuulct ^ lunorl lsp:[ o] rqrrd edeJ) snolq?lS ,{llsLruouJo strEd OT+ !u|]]req pupiuol LUru srq)E.rEuurd :solnuurd s_rred 0€ 0T Flsrp Jo t I lsoLLueddn.eot Jo eseq te srqr€r €uurd letsrp Jo 6.eJrns reddn uo lueseld (alrssospu€ ll€urs) puDID 0t pEeqr.d 't-tz sre^\og:srsorllu€le Suol ruru 0[ SI sel.unped q (Zl 3U) euBlur.\orq.y .t elqEue^puE pEardsepry'A peeqred lZ Zt sF^\oU:srsegluet€Euol LuLuSl S selrunped E6t (Zt 8Jg)auelu^{orq'y .L 6lqDu€^pue peorclsoprlAsnoroloc -uo. 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I Suol slolllJuarquo slreH Lt (8uol sselro ruLu1.0 8.) oqs.{luuoJrul slelqcu€taluo s]reH .?9[ (tZ 3U) etoolnt'V (e8oexSururts ol quou,q.Eog eu,{aqJ 8I ,reuurcl ol lr€ruuec) osolnpuelSe,tlletlllou alo4ed l€lsrp Jo uotlresur ^\oJaqsrqcel Jo aJe.]rnsreddn uo luesa.rdpuBlS :(pernoloc-Jnqdlns+) ^\ollo,{ elBd spEeqreA\olC .q (/Z 3Il) Erour,|JEdrE^ €rr€^'V 'p6I 'perrdsopr^ _ rpuurd erll Jo uorl.rosur.rolaq (srqclrloslE uauo pue) .lortso Jo lJEJrnsrrctdn uo lueserd(rlrssos pue llelus) puelS :^\ollo,t deop speaq ro^\oll pgg straight (rarely prominently cortracted betweer tbe seeds),yellow to light brown. Seeds (not seen for tar, refexa or var. saDsessllis),longitudinal in legunre(sometimes the proximal and distal seedson var../irgonioidestendrng to oblique),oblongto olbicular, (2)2 5 4 5(6)xI 5 3(4 5)mm, dark greyish to brown, + dult to shiny|,plen'ogrant often prominent,with a narrow opening towardsthe hilum;./,iric1esometimes minute, filiform but often slightlydilated, reflcxed below a thickened straight or ol]ce (sometimestwice) -folded aril. At British Museum, Natural History, there are two sheetswhich could possiblyrepresent type material of A. pulchella. The first is labeiled" Hort. " Kew. (New Holl. Mr. Brown.)"; it is also annotated Mimosa miffoph1,lla, Acocia puldtella"-none of this handwriting is Robert Brown's; this shect consistsofa singleflowering specimen. The secondsheet r,vhich consists offour " specinens(two flowering, one sterile,one in fruit) bearsa label R. Brown, Iter Australiense,1802 5."-Bennett No. 4322, and a secondlabel (in Robert " Browlr's hand) Minosa crrncrta,KinE George II['s Sound. . . " which has apparently beeir attached here in error. .ludging from photographs seel, all thc specinenscontained on tl.]esesheets represent the sametaxon. I have refrained from selectilrg n lectotype liom this material becausethere ls some uncertainty regardirrgthc origin of the spccimerrs. The whole question of Robert Brorvn's typesfot Acaciaspecies described in Aiton's Hortris Kewensis necdsto be thoroughly jovestigated. The original descriptionof A. laxuginosaHort. ex Regei was very brief; no type was cited. Judging from the protologue,I have tentativelyreferred tlris specicslAcacia pulchella is related to both A. lasiocarpa Benth. and l. megacephala B. R. Maslin. These three speciesare the onJy meinbersof the Pulchellae which possess stipitate glalds and axillary spines. ,4.acia pulche/la is readily distinguishedfrom '4. lasiotarpoby its flat pinl-lules,and from A. nega- cephalaby its smaller flower heads see under thesc two speciesfor f'urther details. Keyto varielles' la. Bfanchlelsand llowering peduncles completely glabrous ...... 2 b. Branchletsand/or fioweringpeduDcles hairy to somedegree (hairs sometines sparse or minute ... .. 1 2a. Peduncles2 3 l11lnlong; braclcolelaninae acuminate,dark brown. + promtnenttn thebud; pinnules 2 4 pairs. Rare f. var.subsessilis b. Peduncles7 l5 mmlong: brdcteole laminae o\ratc, neither dark brown nor very prominent in lhe bud .. . .. 3 3a. Pinnulesnormally 3-5 pairs and obovate;spines mostly numerous, (l) 2 per noJe; branchletsoften slightlypruinose, ribs normallynot prontinent. (WidespreadMur- . hrrorrR.rer to Ra\en.lhorpe) .. c. var.ghberrima (Figs.4D-I) b. Pinnulesnormally 5-B paiN and - naffowly oblong; spinesolten ferv, 0-l per node (never 2); brarchlets not pr-uinose,ribs nonnally prominent. (Boyup Blook to Albany; Esperance) .... al.var'. goadbyi (Figs.4J-L) 4a. PedLulclesarising from near apex ol the solitary dxillaq/ spine; branchlets strigose; pinnules2 4 pairs, normally obovate-glabrous 1o glabrescent. (Toodyay and Eneabba districts .... e. vaf. fagonioides(Figs. 4M-O) b. 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Ftgure3-Acacia pkl,:hellaya(- p lchclla. A portion ol branch. B to E_pinnules(B uppersurTace; L .lo$,ersurtlce showing thickened midrib and lateralvejns; D upper surllce;E lo\^ers_ufface_showing obscure midrjb). F Seed(dark brown). G Nocl'eof ( ape,\aluratr\re-Atb.in) tornt {pin ite renrored) rho$ing axillarl .pines r.pr. sripilrle gland rgr.shorl lefnlnal \etc l/t and.lrpLte\ tr). ba.e of pedUncler/,d,. and vefi sl.ofl petioF rpr. H-Node of Moora-CapeNaturaliste fornr.lpinnai reno,edl "ho*,r,g uiirtu.y #lni. (iij, stipitategland (g),.long ,temrinal sera (t).and itipules 1r.1, and elongot.fip"iioijlpj. 4 f|onl 4 3 Maslin1-s3., B-C fron.F. W.-Wenit54: Lj-Errom p."C. witsii lCiiS; r tron B. R. Maslhts.n.; c fronl B. R. Ma.ylin2965; H fron F. G. Snith 1j22. and brrcicolc-lrrminacnormally pronrilenl r.ee Figr.3l. ( and H). lr i\ rhi\ lorm $nrcnlncludes the lyp?ol A. pul,ltcllo tar.lti:pi,lulo Mci5,t. Therei) jojlre evidenceto suggesttltat morphologicalintermediales occlrr betweenthis form and A. pulchella'tar. reflexo B. R. Masiin seepage 401. The above-mentioneddistinguishing characters are often absent frotn individuals occurring. ir-r the Cape .Naturaliste-Albanyarea. These plarts ofte,nlack the pilose hair layer on their branches,the pinnule midrib is not thickenedand tlte lateral veinsare absent,the petioleis often shorter,and the bracteole-larninaeare nornally.ovate.and lesspromineot (se ( Fi-es.:n u,]a C). It is this -formwhich correspondsto tl)e rlpe oi 4. pulchcllaand irhich is some_ times dificult to distinguishfrom var. globerrina Meisn.-see page 403. 400 I0t 'oeuurd pexeger eql ol sroJal leqlrde leleu€^ eql 'pnq eq1uI snonJtdsuoc ',roleq lerJ,\\etuoseJ€ qcrq,r a?u[uEl-e]oelcsJq etrBuruflro?pu" pouleL(llue -ururo:d elrnb pu"'sec€JJnsqloq uo snolnleqnd'sno.rolocsrpar€ qtlq,\ salnu 'eeuurd -urd Jo srred (g) g g Surp"erds ,{q pezuatc"J€qc ore ,{eql ,{?,rtqBIH uJeqlJoNl€a.rD erll uo uoopurg Jo q:uou ssJlruolr{ 0t 01 seJlsruoll{9z lnoq" ruo.IJ pal€lloc ueeq e,\€q sel€rparulelur e^rlelnd eseqJ '\Lgzl alopulJ pLtE glLI qtluts eas) o11at1apd rert pue nxa{at J€A uea,\\loq JnJco sal"rpsurrelul I€crSotorldJoul€ql lseSSnsol ecu3pr^eeruos sr eroql'3^oq? peuoDueulsV 'o aLplnd JB,\ e.roJeqsqluou o,\\l ^F€3u uosees Sur.ro,rou slr surgeq Dxavla.t tB^ ollaqtptl Dttoty (sur8rsu eqt ol palculseJ ere ,{aq},11ar1r1xd ]"^Jo selnuurdoql uo lueserd er" slr"q uoq,u),{\oleq pu" o,\oqEsnolnJeqnd flaug eJBqcrq,r selnuurdsnocnrlS lrq,treruos'snoroloJ -uoc 'eruurd 'seurds 'ulnluaulnpur pu€ pexeger re,reJ rlcu"rq s:lr Jo ernl€u eql ,{q ,{leue^ 1mrdfl eql ruo{ perlsrn8urlsrp sr 11 (,ro1eq 3os) exel eseq:} uee,tloq lncco selerpeujolur:lEqtlseSSns ol ecuepr^alucrSoloqdrour euos st eJorll putf Dlla9lnd JE^ DlldqJryd y ol pe]"131flesola sr ,{leu"^ ,\\eu srqJ 'laqrueldes-prurur pel€fiot ueeq e^"q sorun8elo.rnl"ruur :lsn8ny 01 eunf uroJJ sJeMolJ :pouad Suum.tlpuo 8ut.tato17

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X ,{[nf 0Z uie]se^ loorqsllng Jo N rul / :a/,q 'snso^reu-t .e8uol alseJrueLLrode€s snqnl rr,i/rJ tlrraqrd !lD)unpa.{ urur € 9.1 'elslrclrls.f/rrl, (o^l^ nlels ul) eernelSunluenbrle 'o€lruaqndralltuqns enbuutn ,Luru . 'a?3nl(Ol)g aqtnlad : oexouerenbuuold ,e"le8nf:runaDuald .onbdnbopou Z-S.IxS'sereltrx€ S Z l .E ur (Z)I ,rr./rds rsolrdlalr,{arq pe rlnloqnd/.rr./ :snll€ ul I pE xat .tJ 'U .qI J-V' sd!!-'/'ou'J€A urTS€Itr g BxOUo|JeA lc, var. glaberrimaMeisn. in L€hm., Plart. Preiss. l:22 (1844). Lectotype: In Nova Hollaldia, ad flumen Sual River, (1843),prcus 884 (iso: K ph-o'to- graph seen.\1El . PtRfH lraemcrrr.l Fies.4D-t ,.in A(r.:iaht\pidi\\iIn, {. DC.. Prod 2:455(1825). Z}"r.. Nova-Hollandiaorient'. (holoi L' DL-lhotograph seen;probxble iso: K, this specimenlabelled '. Nouvelle_Hollande C..occdent..Bdie Ju Geograpl-eHerl,. \4us. par. pl-orogr.rplr.een]. Acaciapukh(|la R.Br. var.hispidissinld (A.DC.J Me \n. in Lehn , pl;nt. preissi:22 (1844) es 1()name only, Acacia laktta Hort. ex K. Koch, Allgeln. GatIenz.26:l9j (t858) pro syn. .lcacia denudataLehm. ex Meisn. irl Lehm., Plant. preiss. l:21 (184.1);Lehnr., Dei. Semin. Hort. Hamb. ^(1842)-nonlen-S.rr?orerr^*_ln depressisurnbrosii prdpc Wooctman's-poini, d.l5, ALls. 1839. Herb. ?'?,.'!. No. 893. (Drumnond n.312),i (iio: MEL, pERiH : fragment). Acaciapulchella R.Br. \ar. dptuldata(Lehm. ex Meisn.) E. pritzel. Bot_.tb. 35:3t 0 (1904). Ancia p chcllaR.Br.yar. denuclata(Lehm. ex tr4eisn.)E. pritz_eisubver. trpica F. prirzet, Bot. Jb. 35:310(190,+). non. i1leg. Aca.ia erioc[adaHotI_ ex K. Koch, Allgen. Carlenz.26:197 (1858) pro syn. Acacia dcnudataLehm. ex Meisn. var. r,r-a.lrl in Lehm., plant. preiss. (1g44). pfeston -MeisLr. l:21 ]"l?e-r."In umbrosisad riprm fluvii (Wellington)n1. Dec. 1839.Herb. prei:.s.N6. 90,1."(n.v.) Auttia denudataLehm. plant pfeiss. "Colirur ex Meisn. \Lrt. spinosi.\sih1.tMeisn_ ijr Lelrm.. l:22 (1841.)..-Ttp.: in Horto RaumarnianoMulhusiae Alsatoruln, ubi ilor.entenvttli vele I8213."(n.v.) .4caciapulchella ya(. pritzel R.Rr. denudato.(Lehm. ex Meisn.) E. subvar. spino.\i.\\ina (Meisn.)E. Pritzel,Bot. Jb. 35:31009021). grutkli\ Aca.ia Hort. ex Heufrey.Gdnfs'. Mag. Bot. ,t:177curn. icon. (lg5l). 4""..n.v. . Divaricately branched s/rruib (0 5) I 2 (3) m tall; brqnchlets often spines_ cent, slightly_ to prominently flexuose, nornrally finely ribbed, smooth and normally glabrous,. sometimes. (rarely sparsely_ denselyj pilose,.often slightly pruinose. Slrxer (l) 2 per node, normally unequal in length, 5 l5 (20_25imm loog. Stipules 0 5-l (2) mm long. Pcriule less than 0.5 mm lone: terminal seta | 2 5 mm long, narrowly trialgular, dark brown; 2inna rachis-l 5 16-131 mm long; pinnules 3 5 (6 9) pairs, obovate to narrowly obovate, sometirnes narrowly oblong, 1-5 x I 2 mnr,. norrnall;, subglaLrcous (sometimes very glaucous.),_glabrous, finely 1-nerved. OlanJ 10.5) I J rnm long, glabroui. Peduncles 7 15 mm long (20 rnrr.rrvhen in fruit), glabror.rs; flon,er hiatti 5 g mn diam. at anthesis,.with (20) 25 40 (50) flow€rs. Bracieoler (upper serlesJ 1 1.5 laminae -mm^lo_ng.: small, ovate. Call_r lii6e 1 obscureiy- nerved; pe_tals| 5 2 (2.5). mm long, l-nerved (nerve sornetimes thickened). Leguntei l5-50x4 5mm,glabrous. SeedsI orbiclrlartooblong,2.54 5r2.5_jmm.

Distributiotl a cl h{tbitat: (Map l) South-west Wcstern Australia: a rvrde_ spJead variety most common in the region from Moora southward to Margaret River and Ravensthorpe; it has bee[ collected as far north as the Hutt Fiver (about 80 km north of Geraldton).. This variety grows in a nurr.rberofecological situations: it is very common in the poor sandy scil ofthe Swan Coastal Flain frory ry3r Yanchepto Busselton.it.also_grous in loamy clay nearswamps in the FitzgeraldRiver area, and again jn ltteriticsoil in tfieDarling Range.' WESTERN AUSTRALIA: I4 nrilesNE of Fir:ge!?i-{Biv!r-qrq!!h K. M. (AD, rffestern , Auatl349 K, PERTH); Arstralia, "4. Dntmmonll t56 (MEL, PERTH fragmenr). ana jtO (isol synlype.of A.-pukhello \Ar..glabe]r.ima,MELr: Clcremont, 6 . I . Fitzlg,r.rlds.n., Iune 1899 (NSW 13-2985);K ings_Pa^rk,. Perlh, C. A. Gatdnd.\.n., Jol) 1920;Victorin piains,C. .1. Gartlner^59'l ; 7 -miles E _ofW_anna-mal, A. S._George5934; Vanchep National park,' l. M Jantas209 and 209a;4 5 ni W of Wongrn Hi " towertlsEtphjn, i. f. ^Za.i,;,t::; Ouns- !9l9r'.g_!!o_1ntp:4 _R.Maslin 467 (pilose \cr;anr); pelrh Merropotitan arri. B. R. tr.tortin 1!2q (Fr Mq, PERTH); About 4 km N of Bridserown rowards creenbushes.B. n. Martin 2E47;Near BeaufortRiver,_Albany High$ay,.B R. Maslin 2984(CANB, PERTH); South Subiaco,.,1_. I908;piDarla, ,4. A[ot.t.iro]ts.11.,:O Sepi tS0'1; Y9,-rj!"lr_(MEL);t!_..1-5,Aug. Tone Rt\et, Okllield456b MurchisonR iter, Adfield s.n.(M EL 4a5001;Ab;ut 20 nii W of Ravensthorpe,S Pa st 706:.Below Mt. Madden riservoir, R. ,.r. Sar?." 292 (b, L, MAAS, !!!ff]);_O S t

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-7nd .rr,r-ruo-tj r1ull.tlaqDl3J',\ Surqsin8urlsrpur JJlc.JBrlc "rOnU".,,ro* 3(i11 elcnnpad snoJqelSoql-relu re{r"B tunol\ pu" u,lrole8pug o1 :e,ug 'tqirls 15:ed -JEt\l eql ruo{ sl"upr,^lpur uo fllercedsa ,{Juo seurrlouroser€ ulerll riee^\l ,tTlptpl|1d €q saoue.regrpeqJ pelelar ,41eso1ce.]r "x"l oserlttt?rlt lueJ"dde y 'J"A ..re^ | Jo l"ql rllr,\ peJcdruoc sr oLuL!.laqDl7 Jo uorlEr.rr?,\Iulo:l eql ueqy'r\ 'e1"^oqo,{l^\oue u ot Suolqo 'le6uol '(sJr€d ^I,\\oJJ"upue f11q8rls 6 sno.rournueJoui el€ selnuutdslr pur 'reguot t) 'J?}JS .salLrdils ,{llrl8rlsal",s€pnJJer o"uurd,pulr s1r lerp ur l1?urLuJil'srllJuElq Dtut,t,laqDl8le^ lucrd,{}ruo.rJ s.roJlp {lleuuoLr LuJoJsrql tsolrd sir oj uor'lrppe .1. uI _'s!pu E Z Jo tgql o1 osle sdeq;ed puz outlsstpldsr1 Jo ad,{l eql ol_spuodseJ.rocDLuutaqDlts l"^ Jo ur.roJosolrd ,{lasuJp s[ll esuap ueuo .esJBCis sr lr qSnoloqsuno punolE uro4 suaurroedsur lnq ,{.re,rsr utnluerunpur eql ,ltBru.roN (,{earrg r nos sue_rrur ,{pu1nct1:ud).rncJo op slBnpr^rpul 'snotq€l8 .1o esolrdoluos ,{11mtd,{1o.re,{teue,\ srrl} uo saqcu€Jqaql q8noq}tv 'DtaruaqolS '.rpi. q]r,$ snoru,{uou,{sosll? olE seruEu esoql ,.usretr^JDLulss$ourds 'J"A 'J1e^ px? usreJAIsllpo,tts njDpnuap k, puE,{e{ueH xa 1to11 srpuo,t'.y ,3q.y .y 1o suorlduosep l"ur8uo eqt ruo{ Sur8pnt otu1ss4t1ds1t1 puz otDp uap 'J€,^ 'usra14tr ptopnuap xe urla-I k,3o sed,{1eql uotl"rJu^ Jo e8urtl s:lr urqlt,'lr sepnlcurpue uoxlrl crqdtoru,{lodr atrnb sr DLaltratlolS.mt',ollaqtpd olcoty ern r"ru : r a q orco or eunr (,{"J{ ) "'.i:q#""i:[ :.!3:.,tT'; ;,,\: tr; ; :: W t':;] , Smm

, 10 mm sp

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Figne 4-Acacia pulchellu. AloC \ar. rcflexa. A-Porlion ofbranch. BandC Densely puberulous pinnules (B-upper surface; C-lower surfaceshowing slightly thickenedveins). D to I. var. glabenima: D-Portion of branch. E to G Pinnule variation. H-Legume. I lnt'lorescence. J to L-var. goadbli: J Portion of branch. K Pinnule (upper slrrface). L Pinnule (lowef surface). M to O var. Jagonioicles:M-Portion of branch. N- Flower- O Bracteoles. P to S var. srlrsesrl/^: P-Pinnule (upper surface). Q-Pin- nule (lower surface). R Inflorescenceshowing short peduncle(td). S Portion of branch. g gland; pd-peduncle; .?-spine; r-stipule; r terminal seta. A fron I4/. E. Blackall s.n.; B-C from "/. So ster 445|'D from C. A Garuhlers.n; F fton1 R. A. Sallrey 292: F and I from,.4. Morrisan s.n.,15 Aug. i908; G fro|mB. R. Maslin 2847: H fronr 8. l?. Maslin 1626; J frot]i. Goadb' s.n.; K-L trom B. R. Maslin 2966; M fron C. Chapmans.n.,24 June 1973; N-O fronr "r. Dnnlmond2: l5,l; P-R from E. Pa*in s.n.; S from R. ,. Rojce 3685.

Distribution end habitat: (Map 1) South-westWestern Australia: common around Albany; it has also beencollected from near Esperance. A small 404 90t 're^rU :IJ lloc puoululnrc u",rs,, prlrJ lnq uortducsepJell.rDe s,url?ultlog pat?ader (902d '8t81) rousrol^l 'ra,uU u".{S,, :uollcslloc 'sapnuro8p/..'puoruurnro euo ,{JUope:}r3 (Zi8l) ruBrlluo8 V Jo uortducsop IpuISuo erlt uI 'J3qO13OuI peltelloauaeq J^?q saurn8elarnleru-reeu :,(lnl pu" auuI ur sler\\ou ;poLtadSmun4 puD 8u!.ta,\\olf 'tt6l ounl 'u s llDtutiD\) '€qqEeu3re3N :€16l osJe'(HldJd '/\N ',\\SN 'IEI '1 ') '5{JiZ 'ANVJ 'S) J tro'p€or 9Z I 'l.61ilnf I'trs uDtudDq)J ouurv-eqqPouEuo ro^rd qtru.rsr\orrvaql qtnos :t!08 ,,tplg.r?g 'pEorqued-I?r(pool ',{eiipool lsrrtA 'uaos Jo tsnf -/ 'ErierlsrlvJ N Jo :('Ialr'{ oloqd )) VS|Z puoutwn.to urolsol\ : VI'tVUJSnV NdAlSf,\\ '("qq€0{,la) qnrcs ur"ldpu€s,\\ol ur pri?s,{11r.r8pur (,{eIpool) lseJoJrJJ"t\-qErr"f uI 'sartll?colIIos cqlJel"l qloq ur s,ror8 11 eqqeeug J€eu pu€ ,{3,{pooJ rEau e I pal"JEdes,{lepr,r o,r1 uro:j ,{1uou$ouq st saptotuoBoll"A 'ue^r8 sr "}Ep ,{lrJ€col eslceJdou qcrq,r JoJ (ed,{leq} osl€ prle,roleq ees)suerurceds puoruuln]O eql SulpnlJxe:?rl€rlsnV uJetsed!lse,\\-rllnos (I dBtN) :iDltqoqpLtD uoluqtl$tg 'ultu g.t x 9'crldrTle'enbrlqospJ",rol Surpuelserurleruos speos I€lsrp pue l"rurxo.rd lnq etlll-l8el ur Ieurpnlrgr]ol (elnlzur fFueu) ,rpras speesueer!\leq pe:]cBJ]uoc (,(llueurruord ^ler"r) ,{llq8rls sutS.tDw:snorq?IS 'ruru x (ernt€ul '3uo1 6 I 08-Ot [Vreu) salun8al (pauo)tc]ql l: e^Jeu) pe,lreu-1 rutu slEled :ssel '€lloroc Z-g.I -e,ueu + pu" snorqelS sqnt qfuel + "J r4rJ sre,\\ou €I qll^\ 'slseqlue Jo '3uo1SZ 1? u.r€rpruru sppaq ,/a,rol:eso8uts o1 snorqelS ruur g-y 'eurds I I uo lctrq lsour-raddn ruo{ SursrJEsapunpad'.ald:::$ s Lt salolfal '€los Jo lrxe'olnlr[u '/\ I"uruJel Surpeecxe lou puolD oIJq pe^]ou-I Jo ssel3,\.r3u 'e^oqessala^Jeu 'tu3cserq"l3 ol snorq€13'snoJn€l3qns o1snocn"l8 'I'lllu g. g.l 'sJ]ed '.8uol € x g.g g.Z'el€^oqo Xll"rulou y 7 salnaud utuJ.9-Z sltpD.tDuuld :8uol rurrl g.O otas t.atntr\\t a1o4a4 'Btto1 ruur ,raldris '(eulds I loatuuat I S.0 uo lcerq lsorureddn ruo{ SursueecuaJs3Jogur) 3uo1 urur 'apou Jo lrx" 'poqqrJ 'luecseurds71-E Jed I ,ra&rds qsr,uollo,{01 rlsrppeJ ^l3Jncsqo^ra^ seullauos '(asJorlal .ro asroJlue srr?q) eso8uls flel€rapouj spltlJuo,lq : ^eJE ol 't.rl"rp qsl{\ofle,{sr{rat]g : ru I ecol'll"1uII S.0 g/?,/4speqcuEJq ,(letBcu]ul ( ^\olequorssnrsrp aos 'ueas qderSoloqd :oloq elqEqord) 'ro^!d I ) 4 puawwn.Q uE,1.S,, :;dq QtSl) /8€:I to{ I puo1 qlueg sapi.)1uo8n/DDDV g-1tyy 's8g-$161) g:69 selres ,\\eN ^r"H qJeH ,{erD rluoJ

'ltqp,o8 JELJo sturoJ pel"al eseql SuIUraJuoJpapeou sr uort"ruloJur r3qunC 'o11aqc1nd 'rs^ 'oor33p I1erus 'pue Jessels ol DLut.tJaqolSre^ ot pelelaj eq o1 :eaddr $u€ld eseql :JellElusa.r? ss^Eel eqt'{ooJg dn,(og 'dnlurluEw 'e8ueu SurTJrlS3e's"er? reqlo ur're,\e,roH selnuurd3o sJlud (11) 3 g 3urreaqse,r.rel e8rel ellnb s1r put 'seurds,4re1lxe ,{re1r1os '^\aJ,{F^rl?lal slr 'slolqauelqpaqqrr ,{lluauturord 't"^ 'slcutsrp slr ,(q pezruSocer sr {qpoo8 ocueJedsg pue ,{uEqlv eql uI 'J!qulJJJC ur pJtJalloJuJJq J^Eq \JLunBJl eJnl"{u :roqo}co ol lsn8nv (,{lnl) ruo{ sJet oll :pouad Sutun.4puo Suttatol! Bu,r']s'ss€d rorsaqJ r,,r',rv v s pueLot atDptlu " #sI iliig^'u1;'i"ly";{";fi"t"'"} 5uf,eqJrEoN:965t ^oV O y'ecueredsEJoN sotrllrS:9962 ultlDN V g',{uEqlv're^ru ue8lu) Jo sluEg:S88ZuusolAtI'g'f,e.1'q8rHInt uo dnllrriuBl Jo gS rul0I :6€9pue 8e9 a!FDN'V g',{llnD,{l.oll spre^\ol dnuofo) S [Lr tt :929 ulsDlt lJ g'dnu€,{Eld spre{o] dn,{og rur :€t6l ^oN 'u s,(jPral?rt tllrH relr"g :(pereqlunuun loora S 8 '668tSZ 'ldes 'punos rry }snoIAI -HIUEd',\\SN) 0t€ IqpDoD s,a8roeC8ut) :Vt.IVUISnV NUAlSa,^[ 'sdruE,rs pu€ s{eeJc JDeu see.r" 8ur,{p,ro1 ur s,roJ8 ,{luoujujoc ,{leu?^ srqJ (,no1eq uorssncsrp ees) slclrlsrp dn,(o8 'dnrurfueyq 'eiueu 'rctt {ooJg puz SurJJrlS oq] ur stnoco tklpDoE Jo urJoJ ps^"el No. 154." At Kew (K) there are tltree specimens of A. Jagonioitles all of which are labelled,Swan River, Drummond (photographseen by the aulhor). Two of these specin.rensare mounted together (sheetstamped " Herbarium Hookerianum, 1867") while the third is mounted separately(sheet presented to Kew by the Linnean Society,1915)-tlre iast sheetalso bearsthe inscription "2nd (in a different hand), ser. (collection) no. 154." This last specirnen, togetherwith the right hand specimenon the Herbarium HookerianLrmsheet, are probably part of the samecolleclion; at Melbourne(MEL) thereis a speci- " " fidn labelled 154 which could also belol'rsto this collsctior. As Drum- mond's Secord Collection was not dispatchedfrom tlre Srr'an River until January 1844(two yearsafter the publicatiot of A. fagonioicles),the collection labelled,Srvan River, Drummond II: 154 cannot bc the type. The specimen here regarded as the probable holotype of l. /irgonioides is the left hand speci- men on the Herbarium Hookerianum sheetreferred to above. This soecimen accordsvery well with Bentham'soriginal description. Although Bentham (1864) relegatedl. fagonioidesto synonymy under A. pulchella,Macbride regardedit as a variety ofthat species. It is Macbride's treatmert which is followcd herc. This varietyis distinguishedlrom the other infraspecihctaxa of A. pulchella by its infloresccnccsbeing bornc near thc apex of the solitary axiLhry spines. It is interestingto note that this sameunusual spine-inflorescencerelationship occursalso nr A. losiocorpaBenth. 'tar. epacanthaB. R. Maslin. Other charac- ters uscful in recognizingtar../agonioides include its strigosebranchlcts and ils few, normally obovate, glaucousto subglaucoospinnLrles.

1f. var. subscssilisB. R. Maslin var. nov.-Frgs. 4P-S. ltute.y25 60(100) cm allus;r-anrl/ put)eruli ad strigosi,raro glabri- Srtrdeplurimae, 1 2inquoctuenodo.Pinnae r]||,ijjgatae: pin ltlaeZ1juge,1 5 2 5 x0.5-llrlm,plerutnque glaucac,glabrae ad minute et spaNim ciliolatae. Gllrr stipitata,\:.1rit\bilis. Pedunatli 2 3 mnrlongi. Capl1rldsirinae noD excedentes, l0 19 floribus;cal),x et "orn1,ot-n"troto. ry/?r Kul(efin,Wesiern Alrslralia, 2 Scpt.1934, C. A. Gatdtets.n. (holo: PERTH; iso: CANB,K). Diffuse, divaricately branched shrub 25 60 (100) cm tall; branchlets sometimesspinescent, shaight 1tl flexuose,normally obscurelyribbcd, sparscly to denselyrninutely puberulous to strigose,rarely glabrous. Spiire,rnun.rerous, I 2 per nocle,4 10 (18) mm long. Stipules0 5 I mrn long. Pellolelcss than 0.5 mm long; lerfiinql seta narrorviytriangular, l2 mm long, glabrous; pinna rachis0 5 2 mn long; pinnules2 4 pairs, obovateto [arrowiy obovate, sometimesoblong, I 5 2 5 x 0 5-1 mm, slightlythickened, normally glaucous, glabrousto minutcly and sparselyciiiolate, nervcless ir.bove, midrib sometimes thickenedbelow. 61ard variable,less than or L equallirrgterninal seta,apex lrardly dilated. R,,.e/?Je4-!l! extremely redtced; pecltutcles2 3 mm long, solitary, sparselyto denselypuberr.rlous or slrigose,rareiy glabrous;fou,er heads not exceeding spirres, with l0 l9 florvers. Bracteoles(upper series) variablc, 12(2.5) mm long; clawsshort; laminaetrianguiar to narrowlyovate, nor- and 1-nerved,sonetimes promirent mally clark bror,vn 'l in the bud. Flol.ers attenuatedin the bud; calyx tube 5-:nerved',petdls 1 2 mm long, prominently 1-ner-ved,narrowed at the apcx, Lef:umes seebelorv. Sccrlsrr.v. DistribLttionen.l ltqbilat: (Map 2) South-rvestWestern Australia: although most rccords of this variety are frorn between Narrogin and Rar,eLtsthorfe, it lras been collected as far nofih as Bindoon arrd Wyalkatchem. Acqcia pulche/lavar. srrbstssl/tsis trot particularlycon.rmon throughout its range. WESTERNAUSTRALIA: I n1i N of Wyalkatche,ntowalds Koordd, B. R. Maslin l51i 2 mi S of Narrogintowards Wagin, B. R. Maslin660 (B, CANB, MEL, NSW, N Y PERTH)i About5 km dueNE o[ Bindoon,R. R.Ifa\lin 3264;100 mi N of StirlingRange, Mrli s.n., 1879(MEL 49546);3 nli NW of Onserup,K. Nerber394; Nyabing, E. l,arki s.n.,20JLtly 1970i36 mi w of Ravensthorpe,R. D. 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'el€urqJnl 'EIIo.roc S/ I loj pepr^rp fl,roll€rr Jo qlSuel t/ Z €J 1.,f/r2:snoleu-g sl.rnto/r1 pJdEqs-wedoeurrull'JBeurl s,uEIJ'8uol ruru g I €c'lcaja (pe8Llu;:e 'Suolqo'pexogol Ill"Jrds) seuesJeddn :8uol urur I Er (eJctldacalJo es"q 18SuIr e13urse ur pe8ueur) sorJ3s.ro,rol :snorldJourrpsapae2.tg sre,'\\ou06 08 tlll\\ 'srsoqluElEur€rp Luu 'r€lnqolS'e8ral',roll3,(doep spca! .tar1ollf:s\'\oarpBa 0t-8 'snorq"l8 '3uol 'ser\Nol soLurleurospue ^J€lllos slcr?rqlrs€q urut EZ-gI eql Surpaocxo lllzat3 sapunpad :srxu etu3J"I uorls ,{ra^ arll uo lcEJq lsourJ:t\ol 'euecBl osl€ rlelJo pue lsourreddn u-ro4 Sursueselcunped Pssueprioc Jo 1x! 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I ,{1,uollurt{le,t salfidlls eseq eq] sprE,rot asolrd ,(lilo{s lo 'snoJqts[3 'sepou euos uJo.IJ 'epou ',{.r€llrxe luesq" uelJo red (Z fIaJeJ) s?&?.IS saqcuElq uo u,to.rq-pel 'eseq I 'esoJld 1e u,{orq-feJ8'qtoorus | lot3 tq8rl trluoqs ^lesuepo} flal€ 'peqqu 'snolnpurdrynt1 -Jeporu ,{leuu LrelJostalLpuD,tq :llBf lu Z-l gt,?lt esnJrO I ;!J (,^dSN'1Ehl ') :osr : : olort) Lt!lsDl,1['Ag '0/61 3rV 'crle]tsrlv u.retsolA '",relln1r'{HIUAd ,, 9t9 'Sulpls 8 spJ€,rot uolplureg ts€o selru luertfo) iy ,, :adIJ '(ZZOI) '3U'992 '. Jo 'urlsel^J6I 'Z t t9Z t(I) "rsl,{nN U g slBqdarE8auBITBTV 'u,torq ':lq8turls 'sno.lqul3 tq8rl lsotult sLlrBjeur:u,$o.rq {Izp 'spoes 'urul 'oLr Je^o pesreJ e x 0t 9Z'.099 LrusDN U 8r qteeuaqpunoJb 3tl1 ruo{ potcelloc oJe,r qcrq,r se,r1z,rBurlrnr3 uollducsep sI 8ur,$olloJ eqJ 'pelrelloc 'r?A Jo -€ ueeq suq s4rs,rasgr,r Jo lPuelEur iurlnuJ ,{Jo]cqsllEs oN 'soloJlJ?Jq luouruold sselpue selcunpad3uo1 s1t ,{q '.rp^ ',{urds 'pe^tel pazruSoJers qarq/d Dwt.uaq'lt Jo urroJ puElur IJo^ lleurs eqt qlr,\\ pasnJuoc3q lou plnoqs :seqJ pelcelloc u33q o^"q slPnpl^Ipul snoJ -qe13eruos 'slalqJu".rq snolnreqnd seq ,{1lururouslltssasq s r"^ q8noqtlv 's14edpeareu,{ltueurruord pu" rsp11q.r3,\\oU pel€nuall€'se^BoI 'seurds Il?uls snoreurnuslr opnlcur srlrssasqs l€,\ SurzruSoJeJur lnJosnsroloElrrqc reqlo sre,\\ou 6l 01 0I uio{ J€eq qcrq,{\ sp"erl Jer!\ou slrsssslsoLult2 sll ^q o11aqc1nd'y1o rxel cgrcedsurjurreqto oq] ruo4 paqsrngullsrpsr ,{}oIJE^sIqI 'requralda5 ol ^In[ :potnd 8a!ta,$ol] .lcm

. 2mm. 1 rQ' q #T{ {!'l-M FlIl rB , 6mm . 2mm . lmm

6 '4 ,4at/-, trr+

Figore 5 Acaciamegacephala. A-Porrion of branch, B Seed. C Legume. D_ Node(pinnae renloved) showing solitar) spine (r2), stipitate gland (t,), and nirrow stipule (.!)and lerminal seta (r). E Flower. F Flowerhead. G Recepi"alie(r) with dinror;hic bracteoles(b). A, D-C {rom B. R. Maslin676 (the type); B from G. phillipss.n ; C from B. R. Maslin684.

Flovering ond fitititg period: Flowers from July to September; mature legumcs have been collectcdin early December.

The presenceof stipitateglands. lriLlary spines.unijugate lerves, nrodified racemoseinflorescelces, and_lolgitrrdi nal seedarrangernent, relal,es A. ntega_ cephala to both A. pulchella R.Br. arrd A. Iasiocarpa Berrth. From both tGse speciesit.is distinguisl.redby its long peduncles,prornincntly capirate recept_ acles,and large florver heads each belring 80-90 nlrrouly iurbinate flowers. (\ote. Irr A. l.r:itt&rl,qtar. rillosatlre pedrrnclesare quire Jons and rhe floucr lrcrd,berr 65 75 narroulyrrrrbirrare tlo*err ,ecrf,i,i.ri"iii"ifrrLfr"i details.) In addition to the charactem mentioned above, l. itegoceplula jrs is distinguislredfrom A. pulchellaby sorne$hrt larger pinnules,oftin longer and narrowcr stipulesand terminal setae,ald often by thi nature of its bran"ch indumen-tum. The flat pinnules and larger glaitcisriadily separateA. mega- cephala from A. lasiocorpa. 408 t 60t 'sselarllJe^oN 'ruoql Surl€J€dessJalcB.rBqc reqlo erll 01 ss aslleteuea ^luo uEc euo sercedso,r1 eseql eJnl"u elqpu€AJLlt JSIEJJg saltruutde1nlo,tal o1 Jo Jo 'V 'Je^e pe^Jncer,{lluaurrrrordslr ,{q paqsrn8uqsrp,41pur:ou st pdinotsol '/ -,roq :peleleJ ,(lesolc el" sercadso,rl oseql ^lpelqnopun JA)l Dlpqpd ;o ru,{uou,{s B sTtDdtD)otsDl f pepl"Sor dllensn e^"q sJoqln€ snol^eJd 'lsud aq1ur sroqtnr,{q pasn,{lluenbe.IJ sloru ueaq seq lell"l eql'atuspecaJd sEq euEu Jeujoj orll qgnoqllv sercedseurrs oql o1 3u€uo1eqs? suaulJeds 'jqlueg 't'Jo rseql SurpJEBeJur uorl"trsaq ou 3^Eq Lunnu)tr) sdfl eql Jo 'tl I qderSoloqd s osle pue DdrDrotsDl Jo ed,{toloq3q} peulrusxe8uI^"H '(lsol ueoq3ur^€q ,{lq"runso.rdseurn8el eql) ed,{loloq eqt s" papreSeJoJeq sr uourcads suuar^ eql '(y) ne> l€ punoJ ueeq sutyDtltocotsol f. 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ZZ:I ssrerd]ueld'uqe-t ur roosrer{ (Zt8I)88€:I tog t puo.I qluegwnron.l, DlrDry '(,, 'ellu€rueu 'tflueg Ddrno$ol ?BAH '.re^ru DDnV palleq€l ts3qs iN :oloq) ., ( le8nH) \te/{S,, :adtJ '(f€8I) OtL, '1ue14'lunrE 'le '€ €t Seng }e lpug u|qlueg adrBrolss[BIoBTY A. lasiuLatpacan often be recognizedby the following attributes: branches rnoderrtety10 der)selypuberulous to shortly pilose (hairs often denser and more-conspicuourrltal in .4.pulclttl/o.1, ne'ci glabrou.:arillarS spirrcs solirary. somct nc5abs3nl {,never t$o per nodels oflen occrrrstn A. pu/c/tc/lo1:glaid '.]:.ll1lly reduced:pcduncle. ne\er glabrou5. ,r.ry indumenLurrlornrrll5 dcnserr|d mole consftcuo: Il)tn in ,J.ntlclrclla. Acacio..losiocarp.ris,u r,ery.variabie , specieswith five varietiesrecognized nere. U[ thcse,var. rillo,rtl]11d yar. epqcqnth.tare qllite distinct, but the re_ maining three_varieties _ (var. bracteolitq,. var.. lasioiarpa and iar. sedifolia) prore,d"iffculr may to distinguishin the key be1ow. iomplete nateriaj, i.el no\\'ers,ano.rrurts. rs generaLlyrequired to confidentJyidentify spectmensto vartetalrank- Ker to varietles: l1 h5.75 Flo$€f5 pe- head: dr,llarv spine, Jt,\."nt:pedu|lcles l0_20 rr,rr ,ung; prnrrrr,es , , n,.,. modcrdrer)to den.el) puberulor,:,,".,i'..i"i]r,1;.":;,ni*u:rlllfil i,.,,1,] b. Fl-owels15 50 per head; axillarJ spinesnornrally prcsent_butsonetimes few (if then ptnnutes absent, narrower, normally shorter, and lesshairy than above) 2 10. P(duncle.arj.. g from neaf ape! of .r\i lJfJ .piie: p, nules2 p: ir\ .hurt o l0 nr,.l lolq: legurner ) pilo.c fhar-. .lrghli) int.rle,lr. , Danirr|ag.,rro Bb,lg.nlir al b. peduncresnorsiruatedonspine(i.e.arisingftotnrearaxir),r1.,11ilii1?lili,'.ili]."? 3a. B.acteole laminaeclark brown, 1-Derved,acuminate, I 2.5 nlm long, prorrlnentln bud; pedunclesshofi, the :t obscuiedb\ ,Rou er h*tr ar on tfri,ir; prnni,]e'i2!l parrs; U,.anch hairs normally densearld retro6e; lcgurresglabrousl seetls pearll $hite with motttinss. (Widespread) brown .. .. a. ,o,l l"ait"oraG lFis;.;N_i; b. Charactersnot combined as above ... 4 4a. Flowersnormaily.less per r 2(3) -rhanm," 24 long; pcduncre,head;_pinnules rrorntajl) 2I pairsanct i 2.5 ntm 8lffJJlitiffili srcnde'iregl'nl:s*l|':'*Un*,!y1ffi b. Flo\'ers per 30 50 head;pinnules ncrirally 'normdlly3 6 pairsand 2.5 3 ntrnlong; prnna rachis 3 10 mnr long; pedunclesstour; iegunres glabrous. (B;;i-u".y'io Murchison River) .. ... a. var. fasiocarpa(plgs. Oa-"p) 3a. var. lasiocarpa- Figs.6A-F

Shrub.0.3 l..5 m.tall, either.dense,compact and sonewhat rounded, _"j :l::i$i"g..i:L.,?L:nl,{ -qT*h.dI . brq ctlletsoften spinescent, moderately ":1.:1I. {hrrrs5ornerimes rerror\e) ro shorrlypilose. Spirci l:..rew !1t-o:lrtou\ prllnle ro numerous.somellmes absent.^ to ca. 0.5 mm long..tcrminal sc.ta1.2.5 mm long:pinna rucltis t2),1 l0t20t mm-long.aper noi pinnules (rarely irron,in.ni; reyol:ure flat to slighrly recLLrved;,(2) .3:o (i2) pui.r', (Z) Z S_j glabrous ' t+)^x 9.L-f mm, to moderatelypuberulous 1or hispirlLrfousj. C/ard (l) petluncles 1O 5 mm long. . 4 l3 mm iorrg,normotty -od.rai"ty 'nnif,..i,, to O.ns"iy . shonty pitose:forer trca.ctr5 z mm diarn.,i "i,1, JU!lb.Trlil: )U flowers. Rt.tctcol(snot very prom,inentin the bud, I . 5 2 (3) rrm long, €lil::::.t?^lll.l,,lo:,,t. tlt' ilj 3 /4 lengthof corotia,;b"'r: obs";;l'i .5 2-(2 rnmlong, r_nerved (ner.re somewhai.ihickeledi :;T:l:1i{ifl^/i,l Lif,-,'15,40,s) fl:1:.1:".:^,-:.cJ.i9*:.:11;, r J 4 nrm.uar, glabrous 1rerl rarer; a gtaucousbtoot\11 maryiur Sellow. Sr.,dr orbiculai, z1"Iy: z ) lntn:T*,119""_urlh oram.. gtev-blown. b:::1" 2::, :: !.!.(,i :, .\*^r. rl sourh-w-est westerr Austratia : extendins rrom near bunbury norlh\\ards to the Murchison River. This pggt variety ii ::.11,".i-"" l]r: X"dJ soil of .thecoastal dunes from ".ai'eunUu.! to Ucr|lOton. -bet\.\ec Bulrbury and Frernantletar. ltttio,.arpasomerime, occurseast of the duneswhere it can be rebtrictedto clay_soilsuiamps. ln the "j,'lr, :1"1"^j:"-r:1T:|J^pr]jr, l?,lc.r var.tasiocorpa extendi inland to ine vichity or Inree Jpflngs and bradll; hereit is quite common in sandy gravel. 410 ltv '.petreLl-g 'snolnreqnd '8uol (7) g. slotad ,{Ierncsqo+ ag l xllo) ruu 'pnq 1 S. I I Jqt ur luourruord tou ,{llEuuou sapap,lg sre,rou (92) (fl) qtl/d '(qse4 'ru"rp €Z 9l ueq,r ruru srseqtuBlB ruBrp rurlr spoaq/r orf : snolnreqnd 'Jepuels8) '8uol 9-V ,{lesuep o} ,{lasr€ds luur S t (g.Z) saL)ultpal etrut||r puplC 'luecsarqelS '3uo1 's.uud o1 snolnrequd ,{lesrorlue ruur (g) S.Z t (9-7) g-7 '.luawwold '8uol salnuud ]ou xad? uIuI (e) Z I sllpD,! Dautd :u^\oJq iJep '3rrol lnrn I g. O ops loutw.tal:8uol ulru S. 0 u"ql ssel21.rlrg 3uo1uru (9. 1) saldts 3uo1 urur 'snoJ3unu seu.rrletuos'(slelqJu€rq xed€ I E.0 Zl t 'esolrdJo ol petculsar +) esl?ds ro luesq? .sarlds' u,rorq lq8rl ueuo fllJoqs seuirleujos'(lueled ro 'esJorleJ'es.roJlue sJreq) snolnJ3qnd Ilelerepou ,{1leuron 'asonxeu 'tuecseurds '.(afi ,{ltq8rls ol lrl8rurls ueuo spl?uDtq tlnq : ruerp u 'llDl 'gn.tLls 'Surp"erds'esuoo E. I ot ru (S. l) I e .O peqruErq ^lal"cue^rp

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'sJJlJEJerlJSuttertu:laJrp JoJ uoxBl slrll ees 'I?^ 'll€lo^O -D\ollpas'rr?^ ol peleleJ,{lesolclsour sJeeddnDd.t')olsol 'rnls?htr E g plDloa|rolq r"^ rllr,r set]rug" ouros e^?rl ol sr_eoddeuroJ slqJ- ieloefcrr! lueiruro:d e1rnb sjrurleruos pue sepruurd srrrd seq s,{e,t1e',furrqung puE ellurureJC uee,\\lequr€ld 3o 6 'uloJ lets€oJ uE^\Soql uo sdur€,uslros-,{ulc Jo .requrnu? ulo4 peproJar Jeqtouv 'lurr cgrcedserlursetrnber,{lrtrro srql lErll plnoJ serpnls Iua^sJ',rolaq pelelap erot{ (uu lnoqe) pu"l8 Suol ,{1e,u1e1ers1r pu" pe^reu-I I 'peuola[l] ^lluaurluord T aJe rlcrq,uselnuurd sno.rq€I8 Jo sr?d 9 ot t t"rl^\ -eruosstr '(esloJtars;req) selcunpad puB seqcuelqsnolnJeqnd o] eso8ulsslr ,{q pezuetc"rcllc osl? sI u.IoJ slrlJ '(olnTo^JJ ,{llueururord jo peelsur) pe,lrncer 'JeArU ol leu + eJe selnuurd eql rJcrq,\\ur sJrlcco Dd,t,)olsDl J€,\ Jo ruloJ ? uosrqcrnw eql ot pr8,{\rluou uo:}pl"J3D Jo rllJou ru{ 0z lnoqe ltlorl 'sJrnds ,{lellrxe snoJerunupue stelqcuerq lueJssurdsq1,{\ qnrqs peqcurrq ,41uebo elou 'leqs.r"q 'ral[E:] r,, olur sr"rot8 ndtD)otsD1 J"A 'npa.rE ol sguuds eerqJ 'o 'e8uer r slr Jo lJed puelur uJeq ou eql uI slreloproqtrnJ roJ {1eue,rsrql eas-urlsBIN U g (Jezlud Dso111,t'.r"aqlr^\ seDruule atrlos seq uroJ sltll 'sleplcu"iq E) luecsaurds,{leJr"rs pup (sseleurds,{1er1ua ro) seurds,{.re11rxr me1 'qnJqs 'lcedruoc 'esuep ',\\ol qll^\ pepunoJ l"q,\\euos r: sr s,ro:3 1r seunp l"lsuoc eql uO uoxel elqerJ",le elrnb sr zrllalor spI Je^ Dd,lo)otsDlDDD)V 'raq -ueceO pu" JeqlueAoNur,{lunlEur qceeJpue Joqureldospu? lsnBny ur:leddr 1s;g seurn8el:leqolJo o] aunf luo{ sre,\\oJC.poi,iad 3wl1n.( puD Suuatlol! 'rr?d 'Lue(r leuorlcNrrrcqre) etsunfJo N Lu)9l :(t€96t -r] t{ l , , .,,,,^ltl2 iii!!l""n S :(Cv) (atq!t/,U '1 ',tEAueunf pur ErrESuriipegueer$1o{ :€6ZT appnlJ 'siundsaft€ I ir 'us 'epunuele) e t\l aaiql Jo N tLrr2.9 :ta6l rdes 9 ut)wpDtsH :9€61,rr S' ', jr 'urlrru€'I:(HldEd',l{SN "ISN ''J) la.4ios sprD^\otolEpuellE ')) ttgTruolpleieD V'U'xe,\\rIFI lo IA rrrl 9.9 :(Hldgd LZSI{).llos t' y }o N ru) 9.92 :(6tII Vldn) u s uDSoIoy I ? uolSurrrueJ:(!ed ur /ts6t'l-{I^l) 868 pur (l_8t6t.tEt\J) 988 sslar(t.J 'prolplrnDr€eN :(HIlIEd'IN'E) alsan 'F€AueuntJo rLLr: y'g 'nAeu.SulIAI /g A A ! a LZLutlsDI^t pue ars8uo(tuae.,!\l.B:(Hlugd't3tJ) Z0l u!l:^nt\t'l/g'p^\ellnl,^l sp]e,{ol uolplpre9 pu12 r?&/r1'^t'/').trtd deqrue^:t96J Jo E'uoopurA lru l.Z:862 'o8uEd802 leuorlEN ldos''us )auprD)V ) rEau 8ur[.rrc:(//62€L4\SN) v's ss!lppH J V puesatuDA '/ 'eli(BluerC:(80!6t'fgt{) rrpuoutun.q uretsalA:0t0€ lfranoJ y ruol f ']{) ttt 'uotrelupldf'€rl€rtsnv -plEreD:(HfUEd I Lt uDllv N X eurd ortr^reLuos : Vt.MlSnV NXgISa,\\ T.T l9ng, neryelessto prominently l-nerted. Legumes(5) 10-25 x 3-5 mm, slightly.to promilently udulate, moderatelyto denselypubenrlous to shortly pilose (occ€sionall; glabrous). brown ro black. Seedr lfeu seen,trransverse in legumebur thepro\imal arrd disral seeds of(en longitudirral. 2 2.5 r 1.5 2 mm. uniformllbro\\n. or blacki,huirh brourrmottlings. Dislribtiion and hqbitat: (Map 3) South-$est Weste-rnArLstralia: from near Watheroo southward to near Cranbrook and Ravensthorpe. In the northern part ofits range, var. sedifolia extendseastwards to lear Wyalkatchem ano I ammln. . Tlis variety is conmon in the rocky clay of the Wandoo woodland (Eucalyptus . u'aruloo Plakely) around Bindoon. It also occurs on loamy flats around Moora, but does not extend onto the higher sandplaincountry west of this town where both var. lasiocarpa and yar. bracteolqta occur. To- wards the easternlimit of its distribution (arourrd Tammin), var. sedilblia grows rn sandysoil. WESTERNAUSTRALIA: 2 mi N of, W_y:rikatchen\?. E ti. ,4plinnA; Near Bc\lgarr, "/.,s.,4.,1,,r/I576;N^earMorntBlandM I. H Broot:er2t t8 (AD,PERTH);W;stern Anstr;lia, J. l)t.ummondno.108(MEL);Ioundegjn,YorkEast,MissA.Eatons.n..1890(MEL4968i and,1968.2):Bet$een. Perrh_ano King Ceorge. SoUnd. U. H. Ha t.v..n..Ap.l..tLrl\, 1854 (e\ K).:Nal)ering \ryell.. A. R. Ma,ti 5r3r\4LL. \S\\. pLRIH': 4 nri \ of Kaianrrng towardsDutultey_u,ng, ts.,R- Maslin 648 (MEL, NSW, PERTH);14 ;i N of Wagintowardi ]]gIIgSt!, 1r.4. Maslin656.0.5 nri S of Arthur Rivertowardi Wagin,,. R. Mastin1139 (NSW,PERTH); Alrout 5 km dueNE of Bindoon,B. R.Mastin 32J0;kilgan RiYer.,Oidfield 458(M^E!); MouDt Bakewell (York), Pr"'sr 892 and 894 (MEL); 15 iri S of Tannin, i. D. Rolce9131; Watheroo National Par.k, R. D. Ro),rc9j291 Moir:rt Caroline, Miss J. Sett,alt s.n.,1890 (MEL zt968zl);13 km E of Wagin,p. C. Wikotr6209. Flovering and .fruiting periocl: Flowers from June to September; marure legumesare presenlin \orernber.

. A.t'atia lasiocarpa var. sediJblia is based on A. cycnorum Benth. var. sedifolia Meisn. (1844, p.22); on this page Meisner also described A. cl,cnorum yar. ninutilblict. A1'ter examining a large range of material (including an isotypeof_the above two varieties)it seernsas though both thesenanres should to lpply the sametirxon. - The typeof var. minuti.foliadifers most significantly lrom the typeof \ ar. sedifolioin that it is more spiny,but ascan be seenfron the description above, this is a variable cl.taracter. In the above-citedwork, Meisneraltered-Bentham's original spelling " ,; " ". of the specificepither cycnorum to cygnorlrn The original spellingis retrined by the presctrtauthor. Complex variation palterns occur \,!ithin var. selilblia al'd more work is certainly required in order to elucidatethese. This variety is most closely related to yar. ltsi,t.arpa fronr which jt is distinguishedby its puberulous (rarely gl brous) legurnes,stnaller flowers and flower heads,more slenderand often shorter peduncles.often fewer and shorter pinnu1e.,and .horrer pinnaerlclride.. ilre geograplricalrangc of nese t$o \trleties does not appear to overlap; however,in the region frorn Moora to about Bullsbrook they do occur within a few miles of each other (var.. sedifolia being restricted to the heavier, Joanry or clay soils to the east, while var. Iasiocarpagrows in the sand soilsto the west).

Figure 6 ,4cdrl!' lasiocarpa. A to F-yar. lasio&rya: A portion of branch system. B_ Pinna. C-Bracteole. D Flower. E LegLrne. F Inllorescrnce showing new shoot (ri) arising from axis of lowermost bract on iondensed racenlea}'rs. C to Hlvar.. u//ora: G-Portion of branch. H-Pinna. I to J-var. cpacantha:I-porlion of branch showing inflorescenc_e.po\itioned !n spine. J Pinna. K to M-var. radfolra: plnna. K-inflorescencel L ano M.''. N to | . yar. btac!?otara:N_Seed (pear.lywhite with brownish mottlings). O Legume. P Portion of branch. e,einna. R-Flower. S Bracte;les snowlng prominent, dark brown, l-nerved laminae. T Inflorescence. A frcm K. M. Allan 371; B and D from_R.-,1.Sajlle! 1513:C and F from A. M. James2O8; E_ftomB. R. Maslifl 1457;c-H from B. R. Marlij32l ; I-J from B. _R. vaitii ii,n tne typel; Y-L flo\ B: -R..Mqtlt..656; M fron 8..R. M.astin3230; N-O from n. n. Unrtin ZSZO;V_Q lrom B. R. Maslitl3246 (the type); R-S flonl K. Newbey297:'f ftotn B. R. Maslin 3256. 412

L O^.m d ," & X@w; " #ffi@ s s6 ri-t $-: ,t ----_E-=--wq$ \ f 'q2 $-

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o) /.fi il li*t\ f$ #ti$ w ______--dE..._ W_____E-z_ 3c. var. villosa(E. Pritzel) B. R. Maslin comb. nov Figs.6G-H Acctciapulchella R.Bf. var. I i1i7raE. Pfitzel,Bo1. .Ib. 35:310 (t904). Syll))pes:Di?l:3211 and2066 (n.v.). .5-2 __ Single-ste Imned. openLy branched, somewhat spindly s.4raD, 1 (2 .5) m tall: batk grcy, srnooth but finely fissured towards base.,l)runchlets oftelL pendulous, densely shortly pilose, ligirt brownish. Spll??rabsert. Stirle.r very.narrowly triangular, 2 3.5 nm long. petiole I 1.5 nm long; terhinol Jc/./lincrr to very nrrro$ ly tri! lgular, 1 .5 3 mm long; pinta rachis1.5_7 nnt long: ltinnulcs.24 (5) pairs,3-7 (8) x t-1 .5 mm, moderatelyto denselypr_rber- ulou-.. Gl.ind normally nrinute, apex slightly dilated with a shallow ientral orifice; an additiotal smallerglald oftcn occurson the pinna rachisbelow the lowermost pair o[ pirrnules. Pedunclasspreading to ascending,l0-20 ntn long (excecding lea-\c-s).rnoderxtely to denselypuberulous; flott,er heattslight t6 deepycllorv,7 0 mnr dia r. rr rnrhe.ic,rrirh 65 r5 den.elvprcked floucrs. Brqtleo.les2-2._5 mm Iong; clarvslineal larrinle nano*ly orite, moderately puberulous, yellow to.light brown, + nerveless. Calyx 311 ,l/5 lcngth of corolla, narrowly turbinate, tube 5-nerved:petals 2 2.5 mm lorg, l-nirved. Legumes75-45 x ca. 4 mnt, flat, glabrous,dark brown. Secdsn.vl

Distrihution and habitat: (.Map 3) South-west Western Australia: kr]own only from the Nanson-Howatharraregion (about 30 km nortll of Geraldton) wherc it grows in rocky loam.

WESTERNAUSTRAI-IA: Howatl'rarra, A. M. Ashb!21,18; Moresb) R.nge, A. M. Ashby 4586.4587.401/rYglgsbl .Lqnge:_{:!.B,ri".S,g.p1p-t",anJ8:8[.rrr\t;r NJrr.or,B.R. M,l\tltt)164 and ll49 (MEL, PERTFI);28 km from Ceraldtonrowards Northanipton, North WestCoastal High\,,a\,. B. R. Maslin 3321. Flowering and .fruiting periotl. Flowers fron June to Ausust: lrTlmature lcgume\are nre\enti Scprenrber. Acacia lasiocarpa yar. rillosc! is readily distinguished fiom the otltcr varieties_by its large flower headswilh 65 75, denselypacked, narrowly tur_ binate llowers. Other cltaractersuseful for recogniiion include its dinsely and shorily pilose branchlets,long rarrow stipulei and terminal setae,quiti large pinnules,rclatively long peduncles,and lack of axillarv spines. Furiher uork corrldindicate tlrat thi. ld\on warrrrrrsspecific rank.' The habit of yar. illosa is very distinctive. It is an openly branched, sornewhatspindiy shrub, nornally grorving to 1.5-2 m tali; iti branchleti are often pendulous,and the foliage is dark green exceptfor the light green new shoots; the trunk can reach 5 cm in dianeter at ground level. the-very spreadingpinnae are caducoustowards the base of the branches but thc stipulesalrd terminal setaebases are persistent. yariety , .This is closely related to var. lasiocarpo. The axillar.y spiues in the latter are often sparseand sometjmescompletaly absent 1N.8. var yi1losa rs spineless). Spinelessindividr.rals of rar. lasiotarpa oflen cccur on the coastalsand dunes between Bunbury and Geraldton;thiy aredense, f rounded, intricately brarrchedshrubs, not ercecding0.5 m in hejght. Tiiis habit i; quite diflerert from that of var. yil/usr (secabo\,e). In addition to its habit and_Iarger ffower yar. yillosa, heads, is dislinguished from var. lasiocarpa- by its broader, often longer, moderatelyto denselypuberulous pinnules. * . TIre variely lil/osa shares a number of characters wtth A. megacephala. Both la\a hare qLritelong peduncles,relatively large pinnules and flower hea-ds.very .narrow.stlpules at)d terminal setae,often pendulous branchlets, and a restricted distribution in the Geraldtol disrrict. They are readily distinguishedby the absenceof arillary spirresand the reroluti, puberulor.rs pinnules of var. rll/osa. 414 --t

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3e. yar. epacanthaB. R. Maslio var. nov.-Frgs.61-"I I-,',rre.\_30 50(70) cm al tus; ,?rrl antrorsepu beru li ad strigosi. ,.tplrd?10 20mm longae. Pr?r.7eunijugatae; pitln lac 2-ju9ae,(4.5) 6 l0 mm Iongae,revoluiae, glabrae ad spar:sinr antrorsepuberulae. 6/rnr stipitata,minvla. Pcdnc li spinarumaxillis bractearumsum- marum(raro infinrarunr)exor'ientes. antro^e sljigaJ,. Brccteolaeln alabastroprominentes, Callr et corollas-nervosa. I.egnni a pletumquecur\a ad circinata,brevirer pilosa, ,Scmrr?d (propenrarufrrJs) In legume IongiLudinalir, orbtcularia, contplJrdla. Tlle:: 1! kq _Sof Badgingarratowards Dandaragan, Western Australia, 3 A|g. 1973,,. ll. Maslin3247 (holo: PERTHT iso: CANB, K, MEL, NSW, NY, PERTH).

Rather dense, f rou nded s/rrrD,30 50 (70) cm tall, to 150cm diam.; branches slightly flexuose, moderately antrorsely puberulous to strigose, brown. Spine.r l0-20 mm long, bracts2 (inflorescencearising from axil of upper,rarely lower, bract). Stipules I 2 mm long. Petiole \ess than 0. 5 mm long; terninal seta I 2 mm long, dark brown; pitl q ra(his l-2 mm long, apex 0 5 1 .5 mm long acuteand normally dark brown1.pinnules 2 pairs,(4. 5) 6 l0 mm long, glabrous to sparsely antrorsely puberulous. Gland fiinfte. Inflorescencessimple: pedunclesarising from axil to uppermost (rarely also lowermosl) bract on spine,T 8 mm long, moderatelyantrorsely stri1ose: fower headsbright yellow, ca.6 mm diam. at anthesis(10 mm in freshstate), with 23-26flowers. Brqcteoles prominent in bud, ca. 2 .5 mm Jong,prominently inflexed, glabrous to sparsely .5 puberulous; .clawsca. I nlm long; laminae narrowly ovate,ca. I nrm long, brown. Cal1x trDc 5-nerved(nerves f thickened):petafu2 mn long, l-nerved (nerve thickened). Leguntes normally curved to circinnate, 15 40 x (4) 5 ? mm, moderately to denselyshortly pilose (hairs slightly inflated, I trans- parent), dark brown. Seeds(near matlrrity) Iongjtudinal in legume, + orbi- cular and llattened,3-3 5 mm diam., Iight brown. Disttibutiona .1 hahitqt: (Map 4) South-westWestern Australia: restricted to an area fron near Dandaraganto Badgingarra(about 200 km due north- nofil'rwestof Perth). This variety appearsto favour lateritic loam or clay soils.

WESTERNAUSTRAT,IA I North of Dandaragan.C. A. Goftlner9285 l5 km N of Badgin- garra,P. G. l.I7(",?3838 (AD, BRI, PERTH).

Flowering and tuiting periocl: Flowering specimens have been collected in August, while a fruiting specimen(with near-matureseeds) Jras been collected in November. _The_pedunclearising from the axil of the uppermostbract on the spine readily distinguishesyar. epqcanthqfrom the other infraspeciflctaxa oi l. lasiocarpa. It is interesting to note that this same ul]uiual inflorescence arrangement occurs also in A. pulclrclla R.Br. var. fagonioides(Bentb.) Mac' Dnoe. This variety appearsto be most closelyrelated to yar, brqcteoletqB. R, Maslin. Both thesetaxa have similar pinnulesas weli as dark brown terminal setaeand pinna rachis apices. In addition to its inflorescencearrangemenr. 416 LII (8 8rJ) EdrEr,{sEpdsqns 'q (dnllrrcepief ot 01 unotedoH r€eN) 'tq;rerls slqcer euurd xode pLIe eles :snolir^,tlssllep Jo',\\olrq Ieulturel seqcu?rq:8uolruru 9 [ trlltruuoll'srladt € pue 6^oqe,{ileq,{lesuepso|1ould €Z ,{\ol5qqrrprlu z ',{urds peuaFrql 1?qtl^\ ro/pw .(rlell ,{lesuepselnuurd :iurpuease ,{liBturou lou salndlls q (6 3rl) elrdrlsl.lrnrer dsqns J (l.utsrp erie8ur6p€{ :uapptt\l lunotrN pue uopiH ol uuo)n)) srred ! t 'snornElSpuE ssola^rell+ luerserq€]8ot snorclElFselnuuld :pexeuarpuE,{ulds seJndlls €l , \JtJJd\qns ot iJ) 'relrrurs ^.rel sr "x"1 cgrcedse:1ureseql1o tr3o1oqd-roru1ttl.U puB ecLIeJsoroUuI3rl1 'suB8ro 't' Jo ozrs o^rl€lor erll ur secuerellrp IqBIIS lo.l ldacxg ttttow ulqll\\ pazruSoter e.rEs.relcE.rErJJ e,\u"lo8e^ ur,{lueruud Sutragrp setcedsqnseerql 'serceds l"ql Luo.U uJtow 'y qsr1n?u\sp ,{Iprer 'speesesJe,\Su"Jt pu"'spu"l8 Jelo]l3dslrss3s 'seulds 'Jo^o,\\orl ,{r"llrxe eq] :'tB lI D alpll1d V rlll\\ fliuluE u? 1se33l1splno,\\ Jo IcB[ 'selnuuld 'se^€el spur13 rr:1or1odpue :ltU el"Snliun Jo uolssessoderlJ 'sJlnuuld -IeqrunuJ31?Jl3 ueuo puB 'l11o.tocpue x,{l"c pe^.reu Jo 'spEerl ,{ltuaururo.rderout 'serunSolpul, selJunped,{rreq,{lasuap Je,\\oUle8lEl 'seprqceJ 'lrqeq l€r{,reulos e"uurd pue selndtls lo8uol s1t,{q setcodsasaql qloq ruo.U peqsrn8urlsrpsr lt,!rcLuDDIJV spue18relotlad olrssaspu€ sel"el etr8nftun lrrql ,{q pelrurl eJB"x"1 eatrll esaql :utlsut|)j'g ltlaclitaL!'Vpun 'B 't uflsEtr{ U ossatdap rl]l,\\ seIlIuUB ]sesolc sll enerl o1. sJ"eddE }l lnq J€elc lou sr aD a!)ud serJ;s eql urqJr$ ll,tlout F Jo uolllsoo p?xe erlf '1tle tqBtnrts P:ttl -1crql ,,tpdn.rqru? ^\oFq 'JJ'JoJrIgalJtunJ:annllq eq1 sp-tt,no1 Sutuedo Pexeuel 'Ic"lq 'urru 'clldrTle ,,(o.u?u ? tllt^\ LuoJSofiald :Iu[ls Z x t-E.Z ot Suolqo 'euln8al ur enbrlqo ,,{11q8r1sol osJe,\suBr:}spras paue{tlql 'sp33s ur3^\]aq 'snolll^ 'sp33s pelc"rluoc 1ou sutS.tour:u,\\orq-,{o.r8 ,{[esuop rono pesIEJ '1cg '!uur 'SuolqolJEp 'JI}10q ,{pq8rls 9 g x 0t gt fl,uorr"u pu" pr"q l€rl,\\euos sawnBaT 'ewllrdad,llesJrdsro snorqtl8 r(.rz,ro:tueJsalq"l8 ol snotq"l6'p€^rau 'qfuet '8uol -T troqt .IoJ st"uuoc ulru 9.7-9.1 slotad :pa^reu-S 's1.roJq€13tl z-Zl I 'xedE fllueuruo:ci eqot aql le esolrd,{luorls pu? pexeuulpeue{clql ,{llq8rtsa.r" qcrq,r seqol Jt€lorllr Suolqo olur q13ua1sll ti I t/ T roJ papr^rp 'EIIoror qt8uel r,qrr :snorotu-g sratrolf esolld ,{llroqs ()1 Jo €i Z-Zll 'snorq"l8 snolnJoqnd'a^?Juoc 'pexeuul aBuru?l :pel€llp .{llq8tls s,\\€lc:8uol 'sJa,rou u.fujg. sapaDn.lg pJlJ€d ^lJsu;p; sz-sL qll\\'slsJqlue te ul€lp l-l 'J"l[qot8 'snolnJeqnd'8uol urlu g spDaqra,rolf :e]"lorlc slJ?rq I?s"q tuur 8'(slalqJu€rq pue spr",tol red g 7 {1a:er) f.nylos sapunpad 0Z-g Jo lrxe ;eel'JBlnJrrJ 'eJISSes :aldunss uap.olfq urErpuLu g . 0 3c :eeuuldJo uoruosul ,roleq elorledjo ec?,JJnsreddn uo petEnlrsprrlt xedz 1e pe,tor:tu ,{lenbqqo '(culuacxa a^reu) ,{\olaq pe^Jeu-i ^lluaulruo.Id Jo sseloiJau 'e,\oqe ssele,\.Ieu 'snor.r€,f. 'snocn3l3 'teg'utur 'cudtlls lunlueurnpur ol uear8 g-1 x (0I 8) g g I 's]Ied ,{lenbqqo ot Suolqo ,([,rorr"u + , t saTLtuld lpex}AeJ ol :]q8r"r]s xad" 'e^oq" '8uol peqqrJ,{larncsqo = Jo elerel rutu gl g slLp Duutd'.JtedI aDawd '.IElnBu"ul :pexeueJ01 tq8reJls'3uol ruru (9) t-l ,{l,\\orrDuol JeeuII'snou€csJo furds olas puttutal lauol wut alotlad '.altuurdrq satoaT asolrd ,{1goqs o1 'pJxageJ 7'3uoI 1 'lBInBu€r.Il snolnroqnd ol lq8rEJls urul 8 Z (9. I) ,{I,\\oJt"u,{JeA 01 ?uolqo ,{l,roJ.r"u'snolr?cs Jo ,{utdssa7rr,75 'luasq" seuldsdJBlltx" :snouE^ '(,rolte.{ unlu3runpur squ) poqqu ,{leugsapuuq: "1 ulc 09 SI gr?JrlsII?tuS '('^o).tiory '/ t' :(tuaLu8uJ HJUEd :osr) 89t€ 'ZIt:SE 'qf 'lezllrd 't slal1 :sadtrtuls (t061) tel tog E llrJour ElJBcy

'eurds,{rr11xu eql uo peuortrsod sr ecuecserogurerll teq:l lJ€J aql ol sJeJeJleqltde 1e1eur:,1eq1 'sJelc".r"qcolun?al slI pu3'selcunpodeso8l.rls 'te8uol sll 'Lunlueunpur rlcu"tqs]l tq.otnloaJtn,tq'r"^ ruo.UpaqsInBullslp st o4luttcoda'lr-t' b. Pi,rnLle..glabn,.r.abo,e. "par.el)hairy be , .1ofnlall).Jntrorcebos dnd u" on ntdrgrn\.I 7 pai15.rrorntall) t ) J i llnt Iong:btan.her pJberLloL.:te.rrrnal .etr andapci ol pinnaracbis ollen reflered. (Near Manypeaksio -rer

4a. sLrbsp.moirii-Flq. 7 Shrub l5 40 cm tall, dividing at ground levelinto many slender,spreading to erect branches;brnlcles obscureiyflexuose, nloderately altrorsely' pubeni Ious (occasionallywith an additional la1er of short, pil-osehairs).- ,Stpa1c,r larrow]y oblong to very narrowiy trirLflguler.(l .5) 2 415; tnrn long, srratgnr to slightly reflexcd, slightiy thickencd especixllyto\\ards tlle hrse. Zea|es with indumentum as on branchcs(except for pinnules);tenninql setq naffow1y trlanglllar, 1 5 3_mm iong, normally reflexed torvards the apex, thickened (especiallytowards.the base),sornerirncs :l spiry; pinua ruchii (5) 7 l0 mn long, . apex_triangular slightly thickerredrnd rellexed lsometirnei straight); pinnules pairs, .5 .5 .5 .3 7 ,a obiong to obliquely elliptic, 1 3 x I I mm, green to subglaLrcoLrs,glabrous and nervelessabove, prominentlv i_nervec{"and sparselyto moderatelyantrorsely puberulous below (hairssom'etimes restricted to margitr). Peduncles8-13 mm 1ong, detrsely I retrorsel] puberulous; fower heods6,8 mm diam. at antl)esis,wjth l6 25 flo*ers. Cnl1-i 2/3 length of corolla; petels 2 mn long, connate for 2/3 their length. Seels obloing toelliDtic.ca.3x2mm.

. 5mm. , 4mm

.3mm.

yliii9.tit&./ \'? AJP CD ,6 '4 ,4azl" nr+

F,igur,eT ALa':i.,]noiii s\bsp..noitii. A portion of branch system. B Node showing strpure1i). ter'mrnatseta (/) and pinna.rachis(p/) $,ithrefiexed apices. C pinnule(uppei suflace). u ptnnute(to\ er surlacesho$ing nridrib). f Inflorercence. A from l(. NenDrl, 1275;B E from l{. Newbej,t26gD.

Dis.tribution and lnbitat: (Map 5) South-west Western Auslralia: southern regrons troln tLre \icinity of Manypeaks (about 40 km nofth_east of Albany) to about 30 km east of Ravensthorpe. This subspeciesfavours areas of said over laterite. 418 -.1

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JO Alrursr^aql ol unolodoH .reeuuro+ seJE pelculsol l"q.{\eulosr ur 8nu -Jncco :?rl"JJsnVulelsa/r\ lse,r-rltnos (S l'Jcr\l) :lonqoq puD uottnE.ljsre 'urur 'Suolqo Z x g. "c s/aas snollr^ Z ,8uol ,'([esuep"rau43a7 q]3ue1:reql 71 1 ec rc1,al€uuoc tunt g.7 splad .srserltue 7 :€lloroc Jo q13uel f/ Z-Zl I r,{pJ sro.{\ou qt!^\ .ruerp luru '8uol 0Z L I le 8 9 spDaLl.ra,rrog1 :snolnreqnd .{[esuep urur (OI) sapunpad .n\opq 'e^oqe 'snocn"lSqns 0Z EI pe^.rou-I ss![o^.reu o1 ueelS ,snollr^ ,{lesloJ}uE,{JlJorls 'tuur 'cr1dr1le .'(lesrop (S. x (0t ,{lenbr1qoo1 Fuolqo ,{1,to.rre'tl 'sJrEd € Z I) 8) 9 E ri sd\uuld :peuettl.1 pu€ Suolqo ,{l^\oJ.rEu xed? '3uo[ t-€ i lq8r"Jts = ruru slLpD.touutd I aseq]E peue{lrql {Dq8rls snou"cs ,lrl8r€Jls '8uol 91 9 'rDln8u"ul 1nq ruur (9) t E ,(l,uolr?u ot JEeutl,la.r Tozzza;a7:sno11r,t ,{lioqs ,4les.ro.tlu€flasuep sa,raaT oseq Iltrl8rTs snorrecs,lq8rerjs '3uo1 'r"ln8u"r.u _ lE_pJlJIJrLLt 1nq urur fl,tro.l-rru f,j:,r ot Suolqo ,{1,notreus.?rd45 .sno1l,r 8 t 'eso,nxog Allroqs Alasuop tou saq)uDtq'.saLlau?tgSurprejds,{llq8rls.ro tcala ,reJ .TIe] " olur Je^elpunor? 1e Sulpr^rpJo peuruels-olSurs tuc 09 Of qnlqs '..:l?3 gLrti)q^aN )1 '0r.6r,(?4 0€ ,p,lp4snv u,,.r..^ ."runo lir$f?. i.:j^{*"'.rti 'snqrou-.z -eB rI'€solnqoln-,l,rldrJ rrnroqndasuep tp"',;:;z:t:r;:r;::t"3rrli:Jilr':'": Prons l v t aoltluuttl:Dnuol rrlru gl,g un)Duuldstr/rr,/:eso]ll^ Jo]liojq esJolluEesuep Dlloll ae8volmlu g-t ,?r1rd4s' tsollr^lotr^erq asuop 7(/r_r :sntle Lur 09 0€ ).alnjJ g .-8u--.^ou .dsqnsurlsel,\l .A g udruc,tsup.dsqns .q,

.urts"l .U uo^r8eJ" exel oseqtJo srelcrruqcSurqsn8urlsrp orlJ .g -,roleq .dsqns l Dd.rp)t[sppdsqns pu" urls€J^l U g optdtlst,untat uee,\\leqotBtp3rrr]elur ()1 '"ualuJ sselro eJoul ell s.rr7odd"rr.//or{ dsqls I"c6oloqdJoL[ uo pos"g .JoqrueJao pu" Jeqrle^oN ur luasord serunfel ernlurrl :(lsn8nv) ,{1n1o1 ,(r:1,,quo4 s]o,roTJ :poLlad 3L!tt1n4pao qut.t,ttro1i : - "0" ",,",,'.ort jit'; ro'dnroau6rN rru0z rnoqv ELz r,b.t.1D N x,0 ,t) l i"*,i ;t f lill";i"J ;8iii.9ij JoES rlu I l :(HIdEd.,,16SN..13N ggtt ttrDhrV.i,acuujoasg spje^\cil adioql 'sue^eu.Jo rul :(HtuEd .aN!.)) 'dnBLrnLuerrl.Jo E 0€'odroqtsue^eutnoqv IosTt lsr,r\ry g s rul sz : LglL ) D9 Sl ,, Jo r1'\S)fued dnrr3ooqy' tVl]VdISnV NdfJSA,^[ Jerdacuttup (about 40 km east of Ravertsthorpe). A specimen of subsp. dasycarpo,'seenby Bentham and housed at Melbouue (MEL.49407), is re- corded as being collectedfrom Cape Arid (east of Esperance);there is no collector given on this sheet. As this locality is 250 km eastof the known range of aubsp. fillosa, and as recent intensivecollecting. in the Cape Add regfun has failed to reveal this subspecies,it is probable that the Melbourne sGet is incorrectlyannotated. This subspeciesoccurs in a vadety of habitats: around Hopetoun it grows in deep white sand in tall open shrubland with Banksia sDiciosaR.Br.; at East Mount Barren it grows i[ stony quartzite in 1ow heath vegetation;while around the Eyre Range it grows in sand over laterite in tall open shrubland with Eucalyptus tetragora (R.Br.) Benth I E. H ,47'/in2120;40 to 70 km E of WESTERNAUSTRAI,IA: 'f. East Mount Barren, Ravensthorpe,-of Danielts.n.,5 ALrg.1972; Jerdacuttup, R. Edtisto sn- 1972;About I 5 mi NE No TreeHill. EastMoLrnl Barrcn arca, B. R. Maslin977 (CANB, MEL, NSW, NY, PERTH);2 rniN of Hopetoun,K. Neiber,3304,Cape Arid, no colleciorgiven {MEL 49407). Flovetiftg and.fruitixgperiotl. Florversfrom May to August; mature legumes are presert in late October. Acociq ntoirii subsp, dasyc:arpais more closely related to the typical sub- speciesthan to subsp. re(utt)istipLtl.tB. R. Maslin. Fronl the former taxon it is distinguishedby its denselyshortly villous branches and pinnules, its somewhat ionger stipules, terminal setae, pinnae rachides, pinnules, and pedurrcles,and its \traight (rot reflexed)terminal selaeand pinna raclrisapices. Aca(ia nrcitii s-rl:sp.dasycarpa grows to a slightly taller shrub than subsp. noirii; also it is not as prolilically branched at the baseas the typical subspecies. The subspecific epithet refers to the densely hairy legumes which are particularly prominent on specimensof this taxon but not diagnosticfor it rvithin the speciesl.,7oili1-

4c. snbsp.recunistipula B. R. Maslin, Nuytsia 1(3):258-260, f.3 (1912). Tttpe: ' ' " 9 milei north of The Humps (whichis ca. 10miles due north-eastof Hyden), WesternAustralia, I5 July 1970,B. R. Maslin 570 (holo: PERTH)"-ft.q. 9. Compact shrub l5-45 cm tall, dividing at ground level into a number of erect branches; brant'has L flexuose, moderately antrorsely puberlllous. S/i,iilL,Jspiny, 3 o n n long, reflexed. lealcs spa$ely antrorselypuberulous (exiept for pinnules);letmiuol sL14 f spiny, I 3 mm long, straiglltor somewhat rellexed;plnla /dLl/l.rca. 5 rnm long, apex oblong + flattenedand straighl or reflexed;pinntles 3 5 pairs, narrowly oblong to obliquely elliptic,2 4r I l 5 mm, slightly thickened,1lore or lcssglaucous, glabrous to sparselyantrorsely puberulous(hairs normally restricledto the margin), neryelessto inconspicu- ously l-nerved. Pecluncles5 l5 mn long, llormally moderately retrorsely nuberulous; flover lrcadsca. 5 mm diam. at anthesis,with 15 17 flowers. Calyx ca. 1/2 length of corolla; petals 1 5-2 mm long, connatefor 112 their lelgrh- Legumes 15 40 x 4 5 mm. Scel.r not seen in mature state. Distribution and hubitqt: (Map 5) South-west Western Australia: about 300 km due south-eastof Perth from Kukerin to Hyden then south-eastto near Mount Madden (40 km north of Ravensthorpe). A collectionof subsp recurristipLa has also been nade from the Dandaragan-Badgingarraarea, about 200 kn rorih of Perth (seeB. R. Maslin 300'l). A similar distribirtion pattern occursit A. Jrct iana W. V. Fitzg. subsp.irllnor B. R. Maslin; however, ihe northernpopuliltion of tltis trtorr is centredaround Wongan Hills. Future samplingof suitablehabitats in the it]telveningregion may determinewllether these occurrencesrepresent real or apparent disjunct distdbutions. The Badgingarraspecimer of s\bsp. recurlistipLtladiffers from the southernpopu- latio-nsln thai its branchesare less flexuoseand its indumentum is slightly denserand more spreading. 420 -I

IZt .r s'rruoue* ou e qd srrrr eu pr€c oq:u,:,,: J:f;ry) :);l,3iltrii?':iTlu,"i,'"1 ,(t ql"eueq rellrl Suoure e1u:lsordnuesSur,tor8 punoJ uaeq seq -T 'Jei.e^\oH lr,tlleuors€JJo ctllloleI lo dpu€s uo uor]"lege,\ ur"ldpuus Suouru secrld lros 'lJo.ro EI uedo ur qnlqs lc"druoc ue se s,rol8 olnd4stt.tnca.t.dsqns ,{11eur-Iog 'unlueunpur I: l3suep e e,\rLlpuE 'snocn"l8 ssel,{ll€Jeueg ,Moleq pe^.reFl 'Jequrnu 3, ,{pueururord ur.l3t"al8 ueuo'Ja{clrll 3J€,{eq} sarcedsqnsod,,tl 3q] ul '.1 .xede ::lueJeyp elrnb err sercedsqnso,rl aql Jo solnuurdeqJ pexege:,{11ud II -ulluord oJoru€ seq,{lerurou 1rpuu olndpslttntat dsqns;o urql re8uq.ttlqatts ,E 'uorlrpp" ,Dfdlls!,ttn)a.! AJIeurrousr /irlor, dsqus Jo srLlo".t"uurd eql uI 'c 'dsqns ,esBq ur sBpJxJUeJ.{ltuJurLuoJd Jo ,{urdste,reu ere ,{aq1 aql lE p3uolcft{l o .{llq8lls pue tuou[uo.rd e:e sarcadsqnsad,{l eql uo s3lndls eql qanorltly odtot 'dsqns 1r -frrp ol u12t4!llrloLu dsqns ol pe:lBleJ,{lesolr oloul sr ,selJunpadsl1ol 'e€les 1r -n.reqnollesro.rleJ puE pexegal ua13opue,{utds ssal ro 'seqcuejq lBuIluJal ]"q,reulos aJolu snolnJeqnd ,{leslol:}u€ .dsqns ]I seq Dlnh$t,r.tnD,! :}Brll uI 's 'r3que^oN ur eJnl€Lurur speas ^ llrls :^tnt ol ,{€I,{ru()4 sJa,\\olC ;pouadSutunl puo 3u1tatr47 -r o*::\n ,{ e)?T Lu) .clrLlsuMol E Lt:6911aostt/rl'rrn.,D , d Bur) ere.I Jo i\A"rr, ,;t:i:Jrit':;40 stDtlu!tO r\t 0lpt Jo,i^i\ rur Bt :bhieJ,?,a.,r'S '.8Un(rlqlunuJo t.drdrl.LU;rol s :tOOt ltllct1h[t, g crrEdu'bpeu\Fe$ot r,piErppupc.uorJ url S.SZ:rzg u]foh d g l( 'ulln) rLU :bgsullsDn y g '. '/4 Jo'duL) S i lncqv sdunH oLIl. Jo N rru g:66a1/89 VA BuluuoJ 11 3 rlEJ sptp\^ot:drcqtsuo\Ex urol_] rLu /l tnoqv :vl.Ivufsov NuSIsE,\^ sl

puerLr appulr o 7r,rlro{ c-s '.62t Buno,t o ,!Ji {!'{#n !t.'","T;1tc"';1jlfirya-5,$ J( 'o^re^oLun'e-r D (qrrprruBur.roqs e.,Brrns -^q) ;1fl1,:?1""t1""'l(ll$l?,"j:1"tfi""1"".,;,1 -E e.ue.seloguJ (I re^\olC-J (l) eles puc (s) selndrls ,{utds Bur^\oqs .Dl11d1ts1,r.Lrrr,leuruuel .dsqns epoN a lu€ld Jo uorlrod raY\o-j v tlLtotu ntnty 6 inFil7 c '€-- ]I l1 e. R dl /1 '( # ---l-ilU F____i.Z_ V p r& S: 7m II /'#;+.H a ffi t 'uL'r-T-

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e ! U .---muF- o uus 5. Acacia depressaB. R. Maslin nom. nov. based or A. echin.rtaB. R. Maslin-,Frg. l0 (3):256 (1912), AtutiLt?chitlata B. R." Maslin,Nuytsia 1 25'l, fre.2 non G. Don, Gen.Hist. 2:402(1832). 4?.. 6 mileseast of Kukerin,Western Auslralia, 20 Dec.I964, K. N(rrbc! 1620" (holo:PERTH; iso: K, MEL, NSW). Dwarf, n.ruchbranched, compact, cushion-iike,prostrate shrub,2 5 x 50 (or more) cm., vegetativeparts sparselystrigose to glabrous; branchlets+ erect, spinescent,terete; axillary spinesabselt. .tl,/r.//eJnarrowly triangular, 0.5-1 mnr long, somewhat thickened especiallynear the base. Leut'esbi- pinnate, glabrous to glabrescent;petiole 3 4 mm long, lerete, te llinal seta 0 . 5 I nm long, sonrcwhat thlckened; pinnae I pair; pinna rachis 2 4 mm long, terete,apex somewhat flattenedl' pinnules 3-4 pairs,narrowly oblong to asymet- rically narrowly obovate,2-3 x ca. 1 mm, flat, smooth, concolorous,obscurely 1-ierved, Glend situatedon upper surfaceof petiole normally near insertion of pinnae; sessile, small, circular. InflorescerceJ simple; pedtucles solrtary, 10 20 mm long, sparselystrigose to glabrous, basal bracts 2; flover lteacls yellow, globular, 3 4 mm diam. at anthesis, with 12 l5 flowers. Bructeoles lessthan 1 mm long; clawsshort, glabrous: laminae concave, ciliolate. F1or,r,r,r 5-merons; cqux ca. 1 /2 length of corolla, dividedfor ca. 1 /4 its length into ob- long to triangularciliolate lobes which areslightly inflexed at apex,tube glabrous and obscurely 5-nerved;petals l'5 mm long, obscurely l-lerved, glabror-rs, Lagunes f- Iirmly chartaceous,narrowly oblong, ca. 10 x 3 mm, prominently raised over seeds,moderately strigose,dark browf; margins not contracted betweenseeds, somewhat thickened, light brown. Seedsnot seenin mature staIe.

Fteore 10 Autitt depre.ssa.A Portion of branch slstem showing prostrate habit. B No'de showing sripules(s), terrninal seta (l), prominent petiole (p) and minute gland (g). C Flower. b Flo*er head. E-Receplacle (r) with bracteoies(r) F-Legume valves G Leaf axisshowing position of pinnerachides (p,'); ninule, sessileSland (g) pronlinenl, tereteDetiole (r). A-f rbrr K. Ae.h,r 1o.20 (l he rl pc) ; (l from V. D. li,ht.lh l-0rn\l B R \'la'lt

DistribLttion artd lruhitat: (Map 11) Sorith-west Western Australia: known only from the type locality which is a lateritic hill betleen Kukerin and Tarin Rock. -\

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20mp .. @ F 6 a ,lu,/n /?7+

Fiqwe11 Acacianavbeli. A Portionol branchs)slenr' B Slrb-spine\cenlbfanchlet *;in t*o l"uu.t,note few-pinnules, minute petiole (,aJ and 5onre\that Ihickened pinna rachis (p,:i. c pto*it. D-Iirflorescenceshowing pe,luncle arrsi,rg- from a\il. of xppermosl 6.,t"t (b) ot tt,eu.tv .ondensedraceme axis, an,j i newbhoot (r,) developingin axil of lower_ mostbract. E-Seed (darkbfown). F LeguN€ ;-ii;; k. t;'"r;r liii (ttretvpej; c-o froin K Nrl'be12821 ; E'F lron, K Arel'ber2821D'

Distributionand habitat: (Map ll) South-westWestern Australia: southern regiorr.frorn lhe vicinil)'AlthoLrsh of Ongerrtp eJ\l to nedr Rarensthorpe,uttdnorth to Niabinc rrrd Hrden. di5lribuiedover I reasottrbl)side area, Al tteuir.rii, ttoi comtnonrhroughout it. range This specie'.appearsto be confined to areas of sand over laterite, often in associationwtth Eucalyptus retlunca Schau. ot E. fAkata Txrcz. WESTERNAUSTRALIA: Hyden district,8 km N of Dragon Rccks'.-N 6 Mar"dtt lztiit: 4 n i N of Chillinupiool, K. Newbev2821 and 2821D; 20 n]i NNw of Ongerup' f<.'fv"n,bnI lzz8: ,auout35 km dtre-Woll Ravensthorpe, P. G. Wilson1l l0 (CANB, PERTII) Fl(Nefing a d fruiting perlorl. Florvers froru JLrly to August; mature seed has been collectedin Novembcr. The exact taxonomic position of A. newbeli withir the seriesPulchellqe is not clear. Although it superliciallyresembles A. brovnianaH- Wendl., the unijugateleaves, petiolar gland, and inflorescencestructure readily distinguish it fio; that species. ,4c4ri?ner'6e1,1 appears to have its closestamnities with A. dLr)rcssoB. R. Mxslin frou which it is separatedby its habit, subspirescent branihlets, mirute petioles,and normally fewer pinnules. Acacia newbeyi also has some afllnitieswith A. noirii E Pritzel, but it is readily distinguished from that speciesby its minute spitules and petiole, shorter and broader pinnae rachides,normally fewer pinnules,flower structLlre,somewhat smaller llouer head..urrd glabrou. nedtlnclc..,^, -!

9Zl ',{:o1sr11 'runosntlil leerls srrlJ oso8ttls D\JDV pelgtoesr I?Jnt"N ,,'1u1'J 'f/ ,, qsrlug le 3uo sl'rg U Dioltn Jo IcuelpuI ed,{l s" pepr"SeJeq ,{TqISSodplnoc J3 qorq,{\ JoLl:tr1Elucsord et11()1 u,rou{ leerls ,{luo eqJ puelSug ur pel"^ullc te 'srl"J}snv sluuld uo pesEq,{lluepr^3 eJa,\\ ]rq ur palf,alloJs:lu![d o] Lurqfq p. perldde ere,m srrll Llr rrr\\org fq pesn seruuuDDo)v ^\aLreqJ (€l8I) !t(' "Z )lo,r 'J 'rg 99t:9 p3 ,re){ lJoH llv ur peqrJJsepsts,r d DtDllD oDo)y lu 'DLrDrL!.tro,lq qr p. Jo ru,{uou,(sIEJn}tlc -ueuou lalEI B Srrrcq'31?Lurlr8elJ] sr eur€u s.{ur-l 'Je,\a,roq'.llIr1 DSoEt.tls se ',rorr k, orl uox"t s[l] o] e^€Llsrol[]n? lsour (609I) pllri& xe 'qronH PJJToJJJ'( Inun Jduog ar la uou 8l) JB d |?tDllt) t uo p3s8qsr DlrDlu,ro.tqr elusu ?rll

I!I3 ST pcploJ-aJuopauel3rlll ,{1}dn:qu- €,r\oleq pexeuerapunf :LUV|Z I x g.Z I a1oar, :Lrinlrqerll spJE,rolSuruedo ,roJl"u " qlrM ro suouurluoJ 'tuaururoJd a 'Lr,'\\orq 'ulul ruurEo.ltl)ld:,{Lrrris lrep ()1lq8rl S.Z g.l x g.g S.Z'Jrldrllo()1 Suolqo '3un8oT ur anbllqo ol esre^su€rl (.tyuo oryatutalu.t Je,' pLtz DlDlLrttiotq 'JE^ 'peu3{crrtl 'spoos ot sE) .r/7.)as u,{orq lqirl uo3^\toq palt?rluoc lou 'Lr^\orq 'snorq€13'speas ,{1leuuorr,saiS.rzia :qsr,(el8 ,{[oilr rl1 ]q8IT le,\o ,1, 'tru 'Lum 'Suolqo U€p pcsrBr L-g x St 0l ,{l,roJ.reu sa.rrrSa1 snoJqelS tr,lo :ecu€rl']addcelircunJt + E pnq-Je,uou3nr,ll8 xedr 1epexeuur,{li"rulou'pe^Jeu-l 'snoJq"l8'3uo1 r-uurg.Z g.l slotad:u,\\oiq ro,\\olle,{pue luecserqrlS ol 'pc^.rJLr-E'sJqol )rep Jqnl o1€loITlc,{lleuuou paue{ctrlt dllq8lls pexouur Suolqo lL 'elloror otur rltSuelslr t/ I ur roJ pepr^rp Jo qt8uol ti Z Zl I xllm :sno.rau-S .r.r.),11o1!slor\\ou lZ Zl rtlr^r'srs3qtu":l? Lu€Ipurtu g-t'.r?lnqoF spDaq.ta,rogl 'c :alulorliJ pLr"t-e slcErq lesgq'(.o,tDsqoJe^) snoluaqnd ,{lesJedsJo snorqelS '3uo[ rurrl (02) 'lrxe red sapunpad leldurrs saru sa.el/iq 'ur"rp u]iu _Jl 'olrsses9l S Z 0) I 2.0 :(o.nnsqo nt) ru3qt ^\olaq urlu I ot ro oeuurdjo Js"q l" roqtro srriJBr oo",Jrns.reddn uo pefir]'].s puolg 'elelorTrf,Jo snolnreqnd o1 snor 'snocnelSJo ',roloq 1O -qal8 pu" snoJolocuor,{leJeJ pe^Jeu-l dlauu puB ueeJ8trl8rJ 'e^oqB 'elnlo^er 'orlLr ssale^Jou= prrB uoor8 ,(Jl"rurou ot lcg g.€-l x 'oll?^oqo lJEp 'aEuurd 6-Z ,{nrlSrlsro rndrlla 01 Suolqo (,,{1,$o.ueu) lElsrpuo sJred (ZI 'eeurrrd uo srzd g salnuutdlaeuurd y4srp uo 3uo1ruur il L Z 'oeuuidl"urxold l (OESZ) gt 9.1 lErurxordLro 3uo1 uru (9-S.t) S.e 9.1(1\ sttlctu DLtutdlsJtEd ( Z i) a'uutd :8uol lulr[ (Ol 9 I stqrD.t:(solnuurd roJ tdecxe) seqcuuJquo su,{lFrurou runlueunpur 'el"uurdrqra,lra7 alElorJrc'u,roJq 'snou?cs 'Surpuecs? 'snoanp"J JJEp sorurt3luosJo pelcapal o:l lueled saurlaruos 'ruur (6.0) x g.f-l (9 . 'r"Tn8uerrtfl,\\orr€u.ro Srrolqo,{T,rorr"u ol 9.0-Z.O 0) 'snous^ J"eurl saladlls luesqr seurds,{r"llrxB :snolq"l8 ,{[eJ"l rurllueunpur 'peqqrr t][el ,{lerncsqo+ s}q)Lo.!qlSur.rslclrs ^llsuors€oco w Z-t.O qtuqs 'rB rt,/?, uo pos€g (928I) Fuards( sred) .ao '3ellr-Luou '(ZZB) U F 'y]rur'l (8t8I) Ipnfl uou tuttz loreg uoH tuBld Lunu3 Dso8uttDt)oxv '(uaes otoqd L{{) t.aqs uo ueur.edspueqljel ra,rol (IZ€t euuog) 'osuellulsnv.ro1['u.\^olg >J:,],QoaN (608I)plll,^A xe lo qlunH S Z08l 'eu"d !Z lduo{ uou'ed,{toelr o} sE ord (fl 81) S9}:t pe.^\e) lroH'J }lvut rg'>J oJD!/t)D|)DV 'rg 'lpur,AA u otDttD v rro peseq-(6191) 61:Z "rolJ H Eusluaorq BITBTV',

'€eJe qnJaduo eql ruo{ ,4Fe1ncr1red'stueld uerleJlsllv uJelsel\\ suorlJelloJelqenle^ ep"u seq sree,{ 'oq,r 'leq,ne51 Jo ,{rilur .ro1 ua) lt\ Jo Jnouoll ll peLu"u sr sercedssrql '( uuroc sred ,{eq,tep uey) seJu?qrnlsrpI€luoLuuorrnua .raq10 ro eJu Sur,rolloJ ,{Trp€eJel€urureS sposs :sjee,( 8 S lnoqe Jo u€ds eJrl € s€rJ sercedssrr{J JelauErp ur ruc 0gl-00l ruo{,{JBA seqsnqeq:} pus'(ll"t urJ 09 0S)losuap puE Ja.\\o[sr ?tr.1.11au y'spuEl -orr",l pJr?Jlo-rluos3 e 's?ar" posodxJololu uI ralourt|rpur urc puE 'Surp"ards OZt 09 llel ruc 00T 0g qnJrls peqcuerq ,{luodo e olur s,roJ8 lr erarl,r pu€l -q[Jcs eell"trAlJo uorlzte8e,rJo,rol eril Suorue srncco ,41lerurouserceds srql " " has a Robert Brown, lter Australiensc label bearing Bennett distribution number 4321 and it consistsofa mixture of two taxa, vrz. A. brownianaa\d A.luteola B. R. Maslin. Thesetwo speciesoccur sympatrically at Albany (King George Sound), a locality visited by Brown in 1801-02 There is no direct evidenceto suggestthat Brown actually consideredthis sheetlvhen describing A. (iliatq, nor is thereanything to dellnitelyindicate the origin ofthe specinretrs contained tllereon. As this sheetconsists oi a mixture of two species,it is necessaryto selectone specimenas the nomelrclaturaltype, thus fixing the name A. browiiana. Becauseol the uncertaintysurrounding the specimenson tbis sheeta lectotypecannot be designated;instead, a neotypeis selected. Accord- ingly, the lorverleft hand specimenon this sheetis here chosenas the neotype of ). tiliata. This specimeragrees with Brown's (very brief) description. At'at:ia browttiena is related to A. grise.t S. Moore, A . Iqtcriti@la B. R. Maslin, and A. nigticcrnr(Labill.) R.Br. (seethese speciesfor di{l'erentiating clraracters). The above species,together wtth A. empeliocladaB. R. Maslin and A.leiodern.tB. R. Maslin, form an inter-relatedspecies gror"tp (see p. 394). Altlrough A. btov' iafia is very polymorphic, it can usually be recognizcd bv a combination of thc folLouingcharacters: brarrches normally hairy; Jeaves relativelv srnalt (but see belo\\); flowering pedunclesnormally glabrous and slender'(thickeningconsiderably when in fruit); flower headssmall, globular, norrnally with lesi than 20 flowersi petals quite prominently l-nerved, often inflexedat the apex giving the flower-btldsa + huncate appearance:legumes relatively small. glabrous. As stated above, tile leavesin ,4. brottnianaare relatively small. How- ever,the foliageonjLrvenile plal.lts as well asthat at the baseoftbe main brancl]- lets on mature shrubs is often considerably enlarged. These larger leaf dimensionsarc given in parenthesisin the descriptionabove and below Following extensivefield work it became possible to recognize lour varieties under A. brofnione, Horvever,due to the variable natrrre of this speciesand the relatively minor characters distinguishingthe intraspecific tixa, the key presentedbelow rray not work in all cases. AIso, there is some indication tlrat morphological intermediatesoccur between var. brov iItts ard var. obscura. This adds to the diliiculty of keying out thesevarieties.

Keyto varieties:(Negiecl leaves on juvenileplants.) la. Pinnuleshairy belorv(hair densityvafiable), prominently rec rvedto revolute,2-4 pairs:distal pinna rachis 2 4 mm long. (Stirlingand PofongurupRllgu"utl. int".*.Oiu b. Pinnulesglabrbus of ciliolate(if sparselyhairy below then pinn'rles flat to slightlyre- curvedanl ofienexceeding 4 pairs,also distal pinna rachis above 4 mm long) 2 2a. Mostdisial pinna rachides not exceeding5 mm; llinnulesnormllly 2-3 pairsand 2 4 nlm lolrg. (Albany to Busselton) .... a var. brornianl b. Many distalpinna rachides ce. 1 cm or nlorclong; pinnulesollen abcve3 pairs and/or above,lmm long ...... 3 3a. Branchhai$* antrorse;pinnules normally flat and glabrolrs...... 4 b. Branchhairs* patent; pinnules normally slightly recurved and tor ciliolate 5 ,la. Pinnulesglitucous and concolorous. (Mogumber 1() Bindoon) .. d. vaf. endlicheri b. PinrLrlesdark green above, light green below. (Albany to Bu.t"l,on]. uo.. b.ornniono 5a. Flo\\'eringpeduncles glabrous; dwarf shrub 0 3 0-4 m iall, spreadingvegetatively by s,rbterfaneanrunners. lsooradic in Jarrah lorest from Bindoorl ," \:.i,lr. ?l;li!l*, b. Flowering pedunclesnormally sparselyhairy at base;tall shrub to l-2 m, runnersabsent (Karri oi Karri-Jarrah forest from Nannup to near Northcliffe) c. var. obscura

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"Typical "Weslern Chamcter form" fornl"

Raclr;slength rlrnr) l-2 (4) 2 4 (5) Di'1lr.pinna racl'i. length (mm) 1 5,3 (5 15) 5 12 (to 30) (6 Pinnulipr.r. tperdi.tal n rrnarach.) 2 3 (4-8) 2-s 12) Pinnulelength (mm) 21 28 Clanddi.lrneler (nrrr'J 0.3 0.4 05I Pedunclclen{lh (1rnr) 5-10 10 15 Flower head diaDleterat anlhesis(n1n1,\ 56 Nllmber of Ilowersper flower head ... 12 t5 14 t',7

" " " Table2 Difi'erencesbelween the typicallonn and the westernform" ol A browniana ;;;. L'rt",;t;;;. Nb. Pu.uto.te.sloi ieaveson juvenileplant and thosetowards the baseof fiain branchletson maturesllrubs given in parenlhesis. " " Glabrous variarts of var. brottnicrno westernform do occLlr. These mav strnerficiallvresemble ,tr. leiodermabut are distirguished by their less orornineltlv ribbed branches,snaller leavesand flor'verheads, and eglandulose distal pinnaerachides.

7b. var. intermedia(E. Pritzel)B. R. Maslin comb. nov. Fig.12P-tl Acaciastrisosa Link \ar. intetmediuE Pritzel,Bot Jb 35:312(1904) . Ze.too?erSulkev's Feak,(Sukiy Hill nearCranbrook-B- R. Maslin) Stirling Rang€, Z. ,re^ 3002(iso: PERTH) z",iia ciiiataR.B(. \ar. internedia(E. Pritzel)Macbride, Contr. Grav Herb.Harv N.S. 59:9(1919). Shrub0 5 1 m tall, d ividillg a1ground levelirto a few spreadingbranches ; branchesmoderately to denselypuberulous (hairs sonewhat antrorse)to his- oidnloLrs. Stiptrles12 x ca.0 3mm. R4cris1 5 3(4) mm longl'pinna rathis I : : r+ Srnrtn lortqon proximalpinnae, 2 4 (15)mrl long on distalpinnae; (12) pairs pinnae,2 3 (4) -rnm"oinnult,.y 2 oairs on piorirnal pinrrae,2-4 on distal x I prorninerttlvrectrrr ed to rer olute, somewhatthickeled, dark greenabove, lisht sreen below. pttbcrtrlous(\ometimes glabrous above) G/anrlsituated on uiolr-surface of rachis irnnrediatelybelow insertion of pinnae; circular to slichtlv oblong.0 3-0 5 rnm diam.,lip pronrinent,orifice well defined. Perfurr- ci.J 7110 mm-long. glabiorts:llt)ver lleadswith l4 l8 flowers Pelalsinflexed at apex gir'ing flouei-bud !r+trunaate appearance. Legumes20 35 x 7 mrn, litlrt bro-rrn ro grcyi.h. S,, .1s2 5-J r l 5 mm. light brown. DisttibLllionand hqbilttt: (Map 6) Soutl.r-westWestern Australla: commol] below thc sunmits in the Stiriirrg Range (also occurring in the Porongurup Range but lesscommon) rvhereit grows in shallorvsandy or rocky soil.

FtsLtre12 Acaciebrcttnr./rd. A lo F \ar. etdliLheritA.Io D Pinnulevarialion. E Base ol_Dlant\bo$inc subteffaneanrunnef. F Flower bud G 1() O var' brctniana: C- Por'tronol brane:hsJstenr. H Inflorescencel to J Pinnulevariation ("typicai lbrm"). "typical (K jrLvenileleaf; I- mature leaf) M- it and L Leaf siliouettes of form" "Wesiern Pinnule ("Western forn'). N and O Lea[ silhouettesof ]brm" (N matlrre leafr O ju\eni c lerf). P to l)-\at. intermedid:P Flower showinginfiexed petals Q rnd lt Pinnules. S Portior! ol branch systen. T and U--Leaf si]houettes(T-mature ieaf: U rurenilelecf). V to ZZ |ar. obscurttV and W Pinnule\rariation (V-sliShtly i".u'r"al-W-nutl. X to Z-Lerf silhoueltes(X and Y matureleaves; Z juvenileleaf). ZZ Porlionof branchsystem. AandFlrom B. R. Maiin2956: BfromB.n Mttslitr32o3;CfromB R Mdytr 4; D from tl/.E. Blackalls.n.:Efron Mar! Waftu onsn|, GfromB. R. Maslitt2930;HfronB.R Marlrlr2864; I ffom S. Paust101; Jffom,|-.D/eh3378;KfromB 1?Marll// 29254; L fronl a. n. uotlii 29631M fronl E. M. Be nett 2852; N-O from B. -R. Ma:lrr 2863; P-Q fron R T. La pc lllr R fronl B. R. Ma:|it 2609;S lrom Maxwalls.n.;T'U fron B. R X.laslin zsr,o,va;dXlrcIB.R.Maslut2893;WandZZfromF M C. Schoch501Y-Z fron B R Ma\Lin 2864. 428

I zx

a"ad- rer&s%

fi'_-E6t- ffi^

d U

+K- W

"m"ffi*- --m#- Z-

I ,I l- ii'rr 1 ]=ffi'*'",*-flfi-F f il t{ i ii /! ------i // /i --"'-i.il,a:'rO V f,8V 13il ,fi] U'ijM - uut- WESTERNAUSTRALIA: 37 mi peg,Albany-Borden road, A. S. Geotge6395; Porongurup Range,R. Z Lange13lt SukeyHill. eastof Cranbrook,B. R. Maslin2609 (K, PERTH); Turn---olTto NancysPeak, Porongurlrp-Mount Barker road, B R Maslin2960 (CANB, PERTH);Soulh West Australia, Maxwell s;tt. (MEL '19669);District North WestPlan- tagerrei,-E. Pritz?l 350 (AD, K (photographseen) svntvpe);Chester Pass, N. H Spetk s.n.(UWA I l3?). Flovering ancl.fruiting period: Florvers from July to August; mature legumes presentin late December. The combinationof smallleaves, promilertly recurvedto revoluteplnnules which are sliglrtly thickered and normally moderately puberulor.tsbelorv, ard a restrictcddistribution, serve to distinguishvat. intermetliafrom the other infraspecifictaxa of l. browniqna. This variety appearsmost closely allied to tlre typical lorms of var. l:rovt'nian(laldvar. endlicheri(Meisn.) B. R. Maslin.

7c. var. obscura(A.DC.) B. R. Maslirr comb. et stat. )1ov. Fig. 12V-ZZ Acaciaobsc ra A..DC.,M6nr- Soc. Phys. Gendve 6:605 t.3 (1834);G. Bentham, Flora Austral 2:418(1864)-not as to description,see discussion below. T.tre Jardinde Grenier,prds cendvc,19 Mar. 1833(holo: G). Spreading,openly branchedshrub, 1-2 m ta.ll; btanchesnornally Jinely ribbed, moderately to densely puberulous to hispidulous (hairs straight or q) hooked at apex, often tubercule-based).Stipules 2 3 x 0 4-0 6 (0 mm. Rqthis3'5 5 (10) mn.rlong; pinna rachis 1 5-3 mm long on proximal pinnae, (3) 5 13 (25) mm long on distal pinnae:prrruies 2 (3) pairs on proxinralpirnae, (2) 3-7 (.8 l0) pairs on distal pinnae,oblong, 2-3 5 (6) x ca. I 5 mm, slightly iecurved, dark green above, light green below, ciliolate (sometimeswith a fcw addjtional hairs below). Glancl situaled on upper surface of racllis 0 5-1 mnr belorvinsertion of pinnae; circular to oblong, 0 5 0 8 mm diam., lip and orifice quite prominent. Pedtorclesl0 15 nm long, sparscly puberulous cspeciallynear base, sometimesglabrous; .flower lteads with 14 20 llowers. Pctals inflexed at apex, normally giving flower-bud a truncate appearance. Lectrhes tvtd seeclst't.v. DistribLnio and habitat: (Map 6) South-west Western Australia: Warren districLfrom Manjimup westto Nannup and south to Mourt Chudalup (10 km north ol Poirrt D'Entrecastrealrx). Occurs as an understoreyshrub in Karri (Eucalyptus ditersicolor F. Muell.) or Karri-Jarrah (EucallptLts mctryinatcr Donn cx Sm.) foresl. WESTERNAUSTRALTA: Nannup-Manjimup rcad, D. Churcltil!o17 (UWA); 1 6 km W o[ DonnellyRiver crossing on Manjinrup-Nannuprold, .B.R Matltn2864 (BRI, CANB, K. MEL. PERTH);12 8 km S of Pembertonto$ar'lb Northcliffe, r. ,R.Marl//r 2893; Mount ChrdaluD(which is ca. 10 km N of Poinl D'Entrecastreaux),B R. Maslin2900; Warren Riverdistrict, F. M. C. Schoch50. Flovering periotl; Septcmber to October. Bentham (1864, p.418) misapplied the name A. obsctca. Of the nine collectionscited by Bentham under this specicsfive have beenexamined by the Dresentauthor, viz. DruntmondIV;18 and Pretss885 (4. preissiana (Meist.) b. R. Maslin). DrnnnondY.l7 (A. empelioclctdaB. R. Maslin), Cordon River, OldfieldandDruntmondll:153 (A. lateriticol.lB. R. Maslin). Anotherspecimen (which I have not seen)is probably A. nigricans seepage 441. Bentham gave no irdication that he had seenthe type of A, obsctra The confusion iurrounding the applicationof the name A. obscurQhas persistedto the present day. After inspectinga large range of material, including.the type of l. obicura,I can seeno iustificatiot in maintainingthis name at the specilicrank. Acaciq hrovniana var. obscurais closely related to the typical variety from which it is distinguishedby the following characters:branches puberulous to hispidulous-hairs shofi, at right anglesto the branch surface and either 430 f

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8. Acacia griseaS. Moore, J. Linn. Soc. (Bot.) 45:174 (1920\. Type: " East ". ofKatanning; Stowarcl,166 (holo:BM n.v. but checkedby A. S. George, Australian Botanical Liaison Ollicer, 1968)-F?S..13 Small s/rrrb to ca. 0.5 m tall. branchesinconspicuously ribbed, greyish, denselyand uniformly puberulousto hispidulous(hairs 0 10.3 mrr-rlorg);

. 3mm

. 10mm

,(" gr STir'

Figore 13 A.ada glrJea. A Portion of branch system. B-Flower. C Inllorescence. D and E Antforsely puberulouspinnules (D-1ower surface; E upper surface). F Leaf showing + imbricatepinnules and small,sessile gland (g). All fron ,(. Newbey1283. 432 E"F tsnololoruor ^aq1 slloloelS + urql Jlr '{e\} Jr\o'(oq u'' (luo :\\olrq uerrS rrrLlSrl :rE urt\pl^t ( u\rr|/\r.,,11'apr''--"i-io'iii'o1 "ooiu'g rqt d B 'rlnuurd tt4t DttDlltnDtBb ul lrqr^5iuoi pue 5roqn.u:::F lreo '(cn1c" r'ourln rlc 'elctrurolq g.I w afu,LI:(\rifuuld ro1 1d:cxo)sotlcu"rq uo ,t "1"1l"^Y'lf:l (.),1rr'7 .rrorIIc '\resrEos'snollEJS'(ploiqu[u I 01pu" Suolqo'{llcuolslcco) 'ru,.,r * y '.rcln8Lruur,{1uor;lu '{re't '{lletuJou-s'lrldrls :}uosql; i.g-7.g 6 'snolnp'dsrLi.pltt : srieripeseq-a1n5raq 1,1 (3ttot-'-t--t ot) esoltd seurds"[:"u1irxi piqqIJ a.r,'tvuiu llll\\ psqlolJfl:sllJp (llIluloll q'l{JJ3 ol ll\\olq. ilEp 1o B olul I11uoururo.r,-la1rllb s.rq.rio.ig :sJlJJuurq lcoJo o:] SulPBolds:]o,l3qunu '1131 (9 q/l-rys yJ,r.e1punor8 1u Surpr,tip:o peuturels-e18ursrori:lre ru l) l-9 0 lttl\ol\ 'ZL6l '(AN 'I3hl ) UNVJ-'''"q,t :o\r :Hldfd cloq)-LtSz >I I 'rd.s '€rl€rrsnv LLrrtI :drg 8i ,l,i;i;,1\-:i;^,i's' 'rrhbt-nt lobrqtuuoqJo s 'Luru tLLDr'/ &r',S lsoltd z x E t ! z enbllqo pt n5re^su}'l]r^ultunFrl '.r'u ntott'l :riuolqo lgn1\rr'{/o) :eerellr-s errpolu Ia^ €rqel3 61 9i 6! 6g ,uiuE; i&eF 'rr . qr €\olnqolnopt',to I it nrrq'id or"'' rrq\rls r€uollutL 0f cl 'ull4l,J rdlJ rquog tZ '. :rrlnprd''qur pPrP\rnJrr -r,,p1",7.'piipjq., uridr. eu 3r'dur 1r"r-',r'nfo nEd''rPlnlo^!- 'oB;nl u''d- 'tet-ni-(5) tt'l1l]:till-"]d unrluurd "/'7u le^ e€uatd+ (8) s-Z ou,'1o1.,p* n,nu 7-1 lrad sr{2rvi :eEBnf-(tl \l)-aouutd:.elEuuldrq,n oJ- aellnuserrllrxP :eSio1urur 1E.9jf1 l. (s r'2r'U atiurcts:rlnprosrq l-iuin6 rsotrdasuep in6tunr:1d tlttutt)t srttlPtLr l) I s 0 orse err€dord'(re8r) Brr:z rerrsnv","lJ9lilqi:"t:%d4:'{PJg"'::"it))}'t"#:i:'Y;::fi 'i€I^l'log 'ralooH uou '"-'".il;i?jrl t (zE8t)tsgt l 8l s,srlrnJ : qlu'g tt"" tx-eulluedres 'rcl Furlre(]: -H]XEd :osr) ,. (6SIt'(1)-litool)zlt,se unf Lu iou srsouart sll3l'l'{l!'ne- .rriip"ii:qnH.. 1r)'r-r q1 ioa'1"tt1ra f tuDa'toqrl-^ 1ut1 DsoStusD!)o)v 'tl tr3 ^ou ds ulls?l'{ U g Etorl}lra}El BITETY'6

'(nuotu\utq f u1,'tllPuEltlr lJLLllnJ polnqir lsrP sI .aiuel u\\oLll Jtll oaq.tt-vt detlJ\o lou op \JrJJLl\o.\ l J:Jql lo leJIqdElSoJd '\\otJq puP J\oqn qnolnlJqlru^lJuu Jl[,lJlqn tJllrutlld+cnololoJiloJ'Jtf,)IJq -ur'1Rq\ eLLro<-pltralrrqt (1tqdrlspue sJLunBJlJ'oltd 'l! t9 rrro{ ;1pua16 'H ::lJ-Jd'"rqlsl Dasltn pritstnButtsLp st 11 llLlDru.llolq t'

mrn long; rachis 2 4 (5.5) mrn long, prominently ribbed above; termiflalsetq rarro\\,ly-triangular to linear, I 2.5 mnr long pinnae (.1)2 (3) paus; pttrna rachis.2 6 rnn long on proximal pinnae,.5- I 5_(20)mm iong on'aistal piinae, ribbed above, apex I 1 5 mm long_and + flattened;piniules | 2 (5) pairi on proximal pinnae,2 5 (8) pairs on distalpjnnae, nrrro* ly oblong to nairbwly ovate,2 5-7 (9) x I 2 mm, ,r flat or recurred to rerolute, dark-greeniutcl i shiny_above,light green arrd duil belou'. sparselyhispidulous, fini1y t_nerved. Gland situ.atedon. Lrppersurface of rachis belorv insertion of distai pinnae on 2-jugate leaves,.subsequently absent if pinnae3 pair: sessile,circular io oblong. 0 5-0.7 nm diam. .InJlorescencc.isimple; petiunclesl-2 per axil, (10) l5_3'd nm,long. glabroirs,(rarely puberulous),. basal bracts 3 and ciliolate;flott,er hc.a:ls-h.g\t- y_eJlow (sometimes cream), globular, 7 10 n.rmdiam. at anihesis, witlr 24 36 flowels. BrecteolesI.5 mm iong, shortly pilose. dirnorphous: lower series(arranged in a sirrglering at baseof receptacle)reflexetl, o'blong; upper series(spirally arranged)ercct. cla$s nrrrow, gradurll) erpanded inio thickened inflexed laminae. F/orr.,r's .5_-rnerorLs ; caly.t Z, 3 tengtli of coroila, divided for 1 i4 t i3 its length into oblorrg inflered thickenediiliolate lobes. rcadiJy..separatinginro nrrrro\\ly oblong-ipathulate sepals,tubc glabrescen.i ard )elLo\\ or brorrn; pttals 2 2.5 nrm long, co_[[ate for li2 tl;ir length, glabroLrs,1-nerved, thickened at apex; orarl glabrous. leguincs some hai lra.rdald britrle.narroull oblorrg-30 :0.,, 5 t0 mm. tlar ro iliglrrll unAL,taie, rarscrrover scc(l..gtabrou\ lo ntuJeralcl)ptlo\e. dark broun: it,a3rirsnot contracled between seeds, thickened, light brown. Seerl,rtrarrsverse to obliquein legume,oblong,2 5 4-5 r 2 mm, light ro drrlt broun; pteurogri,i ui,tha narro\ qk.lc .2 :on.lilruglrs,ol- openingtouards rhe lrilum: 2.5 r I rnrn; Jr./r/(/Lslrghtl) drlateLl,relle\ed below a gradually thickened onr:e_folded crean-rcolourcd arii.

Dis,tributiona d hqbitat: (Map 7) South-wesl Weste.l Austra]ia: confined to,lareritic,foil i,t Jrrrah lE^utalyptusnaryinut.r Donn ex Sm.) and Varri (A La-tq,qUa K.Br. e\ Lrndt.)torest lrom near Muchea southwardto t1.ievicirity of Pembertonand Busselton. WFISTERN 4!rlT_ry4L.-l!,D!v9.!li.]sup,.ty. ,E._Btackull s.n., Sepr. 1937; Western Ausrratia. J. Drumnntld(.lt):r.s-3. (ryLEL, and n.167(MEL,; Near Co ie, ,t. r.i, (MEL, \!Y) iii l1't-i'l!4ii: Jtmup,M. Ko(h2460 PERTH);.10-nxE of Ha]\e1,B. R.Mastin 4-13; 13 kn1 W-;f Nannup-Busselton road on SabinaRoid-, B. R^.^M.aslin2921j (BRt, C,a.i{C, pllf Hl; j .Z k.., { ol Bg)"..yp.19y.u.{sBunbury, ,. /?.MdJl1/7 28t6; 2 4 Im \t of Uaniinup rowardii\annup, 4. R. Ma:!i!l2861: Kelmscort, A. Morriton\n, it SeprlsqTand 20 jrtl teOO;CiiaT,i R;.yet,Oldli(lrl-556 (M!!); Bedfor.dale.It M.. C. Scioch:te: Morningiin-S.pr.M'it., R. r. W^illiamss.n., 2 OcI. 1932(UWA 1135);Dweuingup, J. H. Wi is s.n., te+t'iVrl 19719).

Flovering and .fruiting pcriod. Flowers fron late May to Octobcr: marure legumesare present from October to December. P_ritzel(1904, p.3J2) _ describedthis new speciesas A. strigosa"by Link var. Dorealls. However, Prilzel's name was never rvidely adopted subsequent botanists. Hooker, ( 1.852)erroneously referred this taxon to L t.ycnorum Bent1.r., a specieswhich is treated here as a taxonomic synonym of A. tqsioiirii Benth. ,,^..u:1,1,:-._lllt,l,o-lll .':o.opr'"0the.name A. obscurq A.DC. lsee page 4JU); ot the nrne collectionswhich he cited rrnderthis species,at leasttwo are A. lateriticolq, yiz. Drtunntontl II:I53 and Olclt'ield,Gordon Rwer. . Acac.iq-late licola_is cjcsely allied to both A. bror,niqnaH. Wendl. and A. enpeliocladoB. R. Masiin. From the former speciesjt js distinguishedbv longer peduncl.-s,Iarger.liower heads, type of brlnih ildumentum (;.G;;i and to some degree,pinnule morphology (the pinnules are nor.ira y eiihJi larger or more recurvedthan thosein ,4.broruiana\. The geogrlphicairangei 434 :osolld lsni er€ soqJueJq e'41npo1:totlatltua p. uO podole^ap ,{pood sr re{e1 snolnprdsrqerll seLurtoluos'Je^a,\\o{ :sllpq pespq-elncreqnt'snolnprdsrll puE solrdjo ernlxrur u Jo slsrsLrocDp)tl.tatDl t/ uo ulllueurnpur rl3utrq eqf 'qBueAo lou op serJ3dsod\l esaqlJo se8u€.rJectqdEl8oo8 u,\\ou{ erJf serunSolre8uol stI puE 'solnuurd snolnprdsrqflasJEds 'J3:uorl5 'ra/!\c1 ,{11erele3 slr 'seprrlc€r aeuutd esolnpuclSJ'let.roqs ueuo slr '(,\\oloqaes) Lunlueornpurqcuerq str ,(q .opopo1adwa. peqsrn8urlsrpsrrlo)!t!.laql/,ta^e^\oH y ol ;e1ruls,{lplcgroclns 'o8lEl 'sJIaur'rpid plD!itJaJDlt/ repueJselnuurd pa,uncer uelJo ,tle^tl?leJoql pu" '3uo1 'a5r€[ snoJq"lS uo euJoq spueq r.,rog.,no11o,{1q8r1 ,{lltrurou aql 'lsa,r pur?lsue'qlnos eql o-ls.rnrco 'bulIlads Dut2lutt\o.tcly' cllq,\\ lcIJlSlp Sltll -]o q:UoupslnqIJlSIp SI DIo)4IJalDl |/ ,{11uraueg lJulsrp Lrol.requred-Cn urr lur:6 cq1 ur dul.rslo srrJadso,\\l aserllJo '(t68t ta.S "u'suos!.t.toN ruo{ :0061ILnf ''r's 'J lI ,/ O 0Z losl.!.tow y Luor.l:l puB g :'u s llDpDlg 3 /11 ruo.t)rJ pu€ g : l l/ //r.rrr!' t, LLro{y -sorouul '(snolnpiclsrqpue asolrd) sreq q.uDrg Luels,{sqruerq uonrod 'eLunSo'I D jI Jo :l (T (erp.Jrnsra.\\ol IIv) uortBD€^olnuurd J ot v Dlolt!)atol oDDv lI e nfi! J,$int

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-____EE_r_ they lack the additional layer of shorter hairs. Pilosehairs are not ahvays developedon A. browniana,but if presentthey are normally shofier than on A. lateriticolq;also thesehairs are mixed with an antrorseiypuberulous (not hispidulous)layer. The specificepithet alludes to the preferenceof tbis speciesfor lateriticsoil.

10. Acacia empeliocladaB. R. Maslin sp. nov.--FE. l5 " ,4caciaabscun A.DC. var.moiiona E. Pritzel,Bot. Jb. 35:ll2 (1904).4"e.. Hab.in distr.Stirling pr. CapeRiche in clivisrupestribus montis Melville flor. m. Jul-(D.3489)." (iso:PERTH fragment). Acadaobsaua auct. non A.DC.: tsentham,Flora Austral.2:418 (1864), p.o parte a5 to fDrummondY .l'7.l Ftutex | 2 nl altus; /-.?r/imodice ad densepilosi; spinaeaxillares nullae. FrIb bi pinnata:thachis (2) 4-7 (9) nlln longa;pl"nae 1 2 (.3)-jt).-ae.pinnulae pinnaru]nl proximalium 2 4-jugae,pinnarum distaliun (3) 4-1o-jugae (foliarum unijugatarum 3 T jugae), 4 l0 x I 3 mln, planaevel recurvaead revolutae. Glarr sessilis,in paginasupera rhachidis; glans minorin rhachibuspinnarum distalium infra insertionem parlrm l-4 slrmmorumpinnularunl Capr'ralaglobnlosa, 25 ,l3-fioribus.Florue 5 rnefte: ca!)'cislob- oblongi;pcld1d glabra. Lagttmina15 25x 5-6 mm,parce ad densepilosa. ,Serrrlain legunrinetransversa ad obliqua, ryze:About 1 .5 kn] dueNW of MountBland, about 60 km dueSW ol Hopetoun,south coasfof westernAustralia, 30 Aug.1973, B. R. Maslitl3.184 (holo: PERTH; iso: B, BRI, CANB,K, MEL, NSW,NY, PERTH). Shrub | 2 m tall, either single-stenned or dividing at ground level i|rto many spreadingto erect branches branchesribbed, dark grey to blackish, red to bro\,vnon young shrubsand also often on branchletsof mature plants, moderately to denselypilose (hairs tubercule-based); axillary spinesabsent. Stipules varral:le, normaliy narrowly oblong, 3 4 x 0 2-l l.rm, often caducous, glabrous or sparsely pilose, slightly thickcned at base. le.ife,r bipinnate, indumentum as on branches(except for pinnules);petiole 1 2 mm long; raclrrs (2) 4-7 (9) mm long, promirently ribbed above; terninal setanarrowly oblong to narrowly triangular, 1-2 x ca. 0 2 mm, often caducous,somewhat thickened at ba,se;pinnae I 2 (3) pairs; pinna rachis2 12 mm long on proximal pinrrae, 10 30 mm long on distal pilrrae (7 25 rnm long on unijugateleaves), ribbcd above, apex car.I mm Iong: p[rutules2 ,1 pairs on proximal pinnae, (3) 4 10 pairs on distal pirrrrae(3 7 pairs on unijugateleaves), narrowly oblong, 4 10 x l-3 mm, flat to recurved(revoiute upon drying), dark green above,slightly fighter green below, normally glabrous, finely 1-ncrved. Gland situated oI upper surfaceof rachisbelow insertionof pinnae,absent from unijugateleaves; a small gJand(0 3 0 5 mn diam.) normally occursbelow insertion of upper- most I 4 pairs of pinnulcs on distal (rarely proximal) pinrra rachis; sesiile. circular,0.5 0 8 (1.2) mrr diam.,lipyellorv and prominent,orifice orown and shallow. lnfloresceacessimple; petluncles1-2 per axil, (10) 15 20 (25) mm long, glabrous,basal bracts3 and ciliolate; ,4ower headspale yeJlowto cream, globular, 5-7 mm diam. at anthesis,with 25 43 flowers. BracleolesI mn long, moderatelypubelulous, dimorphous; lower series(arranged in a single ring at baseof receptacle)reflexed, oblong, apexinllexed; upper serics(spirally arrangcd)erect, claws narrow, expandedinto narrow inflexedlamirae. F/or,ers S-merous;cely.\ 1 i2Iength of corolla, divided for ca. 1 /5 its lergth into oblong inflexed . thickenedciliolate 1obes, tube slightly anguiaryellor.v or bror n and sparselypuberulot$, petals ca. 2 mm long, connatefor l ,/2 their length,glab- rous, l-nerved; o|ary glabrous or tonlelltose al apex. Legumessomewhat hard and brittle, narrowly oblong, 15-25 x 5 6 mm, flat, raised over seeds, sparselyto denselypilose, dark brown; marginsfrol contractedbetweei] seeds, thickcned. ,S?e,futransverse to almost oblique in legume, oblong to .-iliptjc, 2 5 3 x 2 mm, tan, shiny; pleurogrctmprominent, open towards the hilirm; areolectr. 2 x I mm;.frrlclr: slightlydilated, reflexed below a graduallythickened once-foldedaril. 436 'sesrnoc -rele,\\ SuoI€ Iros d17lcur s,roJ8 1r €er" ueJ]"g lurlolN tsEE eql tlr :elrJe]"I ^pues I Jo pu€s ur s,ro]? 1r ue:reg turlol^JtserlA pu" plr"la lunol^tr 3ssq oql punoJt 'oqcrUJo :uerj€g lLrnot\JlsalN.Jo sedols elrzlJenb aqt uo osl" pus ed"J le olll^lal,ril l-unow Jo sedols o:lrloSuodsd{JoJ eqt uo s,nor8 }r :suortrpuoJcrqdepa ;o ,^larJE^" saleJelol DpoporyadulaoDo)y' 1?1serll uo.rJ m{ 9l ]l]oq€ u€ql sJour SuurnJco loleu :ueJreg luno14trlseE 01 pJe,rls"e eqcrg edr3 ruo:g suot8er + uJerllnos :erTuJlsnv uJelsei& lser!\-qltos \L dBl^l) :tDllqoq pup Ltolnq!.tJslo '916 uusDl\t U I tuo4 N-I :us 11r,rr.rrv ruo{ H:f6rl a oat S'r.,,jrorJD : SZ8,fr./nrN , Luo{ g : t906r.)Jry O yuro+3:rI:A puouu rrrorJ :r€6€ .t)trt!!tt g uro4 :(ed,{l eqt ,.lsnl1,r y Luo{ '(seu.s O T C C t8tt 5r'srrEll I'g-V reddn) soloelr€rg N ol'I (seuesre^\ol) eloelrerg ) -le,rol-{ I qtrulrq esolrd I lpernolo3 uel) pees H orun;o'I 5 eauoasalogul I uals,{s qcusq Jo uorlrod-g (ereJrnsro,rol ll?) uorteue^ elnuurd-C ot A salnuurclJo eseq te (t) puelS elrssesSlr|roqs srq.cr €uurd lptsrp Jo errJrns )addn V DpDtrolladtaaDlrnl SI nStJ

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q)8 w_@_ffi WESTERNAUSTRALIA: SwanRiver Colony, .l DrummondV:17 (MEL, PERTH)' Half-wayup West Mounl Barren,/. .S.Oeorye 1'791; Mourlt Melvillenear CapeRiche B. R. Manin 1062(AD, K, MEL, NSW, PERTH);BetweeD the HamersleyRiver and East Mount Barren,B. R- Maslin916i MettiersBrook on the westside of west Mount Barren, Maxttells.n. (MEL ,19,169):Mount Bland,K. NevbeJ825; FitzgeraldRiver Reserve,R. D Rotte 9063;22lni southof Ravensthorpe,-4. Wittwet,393^. Flotering and fruiting period: Flowers frorr JLrly to October; young legumes appear in early October and mature jo late November. Bentham (1864, p.418)misapplied the name A. obscuruA.DC. (seepage 430): of the nine collectionswhich he cited under this species,at least one is A. ampelioclqda.viz. Drunnond \ :77. Pritzel (1904, p.312) describedthe presentspecies as A. obscuraA.DC. var- moiriqna. However, this name was never widely adopted by subsequent botanists. Pritzel'sepithet was not usedas the specillcephithet for A. enpelio- c/rzdcbecause confusion may develop between this name and A. noirii E. Pritzel, which is a valid species within lh.e Pulchellqe. Judging frorn morphologicalcriteria, ,1. enpaliocladaappears to lic be- tween A. lqteriticolnB. R. Maslin and A. leiodermoB. R. Maslin seebelorv. The speciesalso beals some resemblanceto A. nig cans (Labill.) R.Br. and A. bror'filanaH. Wendl. The leaveson A. entpeliocladaare quite variable. Soms plants arc coo- sistentlyunijLrgate while others have from one to thrce pairs of pinnae on the same bush; the pinnules range from flat to recurved(revolute upon drying). Plants with perfecllyflat pinnulesare known only from Cape Richc: the tyle ol A. obscura\ar. noiticttlq and also DruntmondV:17 (seeabove) are included here. This forn never has consistentlyunijugate leaves, rvhile its pinna rachis is somelimes eglandulose. In mary respecls,e.g. large leaves and flower' hcads,flat pinnules,and glard morphology, this form resemblesA. leiodernta from which it is distinguishcdby its pilose,less prominently ribbed branches. Further east from betweenWest and Easl Mount Baren, some plants have consistentlyr-rni.iugate leaves, and the pinnulesvary from concaveto reclrrved. These plants bear an overall resemblanceto A. laterititola-see this species for further details. The spccilicepithet refersto the blackish colour of the mature branches. However, the brancheson juvenile plarts and also ofteit tlte brancltletsol') mature shrubs are red to brown in colour.

11. Acacia nigricans(Labill.) R.Br. in Ait. f., Hort. Kew. ed. 2, 5:465 (1813) as to name only. Ftg. 16 igtitdns Labill., Nov. Holl. Plant. Specim.2:88, t.238 (March 1807),non Vahl Mimosa " ". ll80-/). T),pe; Habitat jn ter'ra Van-Leuwin (lecto: righr hand specimen on sheet labelled " Nova Hollandiae ora austr-o-occidentalis.Herb. Webbianuln. Ex Herb. Labil- Iardidre" FI, photograph seen). Acacia nttaefolia Link, Enum. Plant. Hort. Berol. 2:4,14(1822), nom. ilieg., based by inpli- cationon,4. /?tgr-icdn.!(Labill.) R.Br. llcada iEnontaK. Koch, Allgem-Gartenz. 26:195(1858). D?er n.v. Acaciapatltadenia Hort. ex K. Koch (1858)non Lindl. (1832),pro syn. fAcaciaobs(uru auctt. non. A.DC.: (?) Benfhanr,Flora Austral. 2:418(18621), pro parre asto R. Brown.Goose lsland Bay, n.v.: Willis 1953,p- 26 and 1959,p. 100.1 Shrub 12 rn ta.lli brancltes normally prominently ribbed, moderately hispidulous to glabrescent,hairs denser along ribs and normally tubercule-based; axillary spines absent. ,!l?r1es narrowly oblong to narrowly triangular, 2 3 x 0 3 0.5 mm. Zeares bipinrate, jndumentum as oo branches (except for pinnules); r.tchis 3 9 (13) mm long, ribbed above; pinnoe (l) 2 (3) pairs; pinna rachis 24 (7) mm long on proximal pinnae, 5 1l (15) mm long on distal ptnnae; pinnules 2 (JJ pairs on proximal pinnae, 3-5 (8) pairs on distnl pinnae, oblong to some\ahat elliptic or obovate, flat to slightly recurved, glabrous or 438 6t '869€arloy y ruo4 g '.goOOIuaq41 tuo{ :90t2 'S o , ;r.11 a PtDag / ruo{ J'.5t8 )nD8u,tt)S N'gruo4 H pu" C'g'.9991 PtPtlX y LLro4( puEV '(qr) srxe alua.er pesurpuor eql Folpuotqns isPrq ,{relrlospue (pd) elrunped Surpuolqns(qq) rre:q leseq,{re}rlos 8ul^\oqs lesur ecuarserouul (erelrns re^\ol :ereJrns raddn-J) uolttlre^ alnuuld ot eLunSo'I 'urolsisH lsel '(1)C C '(putliJ qruprq Jo uo|Uod I (rcl) seprq.€r eeuurdJo ospq pu€ eles lEuruuel Jo ^\or^ dort-teaLr) pu!13 8ur^\oqs (-\\el^ eprs) srxe Jae'l-V sub)].t8tt 2DD)V' gI eJr'ff|:f

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Distrihtrtionand habitat: (Mtrp 6) South-rvestWestern ALrstralia: from Barker Inlet (50 km due west of Esperance)to Cape Arid (120 km due eastof Esper- ance); also recorded lor Middle, Mondrain, and Sandy Hook Islands in the RechercheArchipelago. Acaciu nigricans appeirrsto be restricteclto near coastal regioDsin either sandy or rocky soil. WESTERNAUSTRALIA: Mount Merivale,Esperance, "/. S. Bead 2305:Barkers Inlet, E. M. Cannirg702'7 (CBG); Esperance, D. Churchills.n.,l6Aus. l956 (UWA ll42);About 50km eastof Esperance,R. H. Kuchcl1656 (AD, CANB,PERTH, UC); Near Cape Arid, Maxv,es.n., 1875 (MEL 49470);Duke of OrleansBay, A. E. orLhard1316; Cape Le Grand NationalPark, R. D. Royce8690;5 mi N of Esperance,R. ,. Roj,ce3598; Condingup Road jLnrction,570 miles on Coolgardie-Esperancehighway, -E. M. ScrlmgeoLrr835: Middle lsland, RechercheArchipelago, J. H. Williss.n., 22 Nov. 1950(MEL 49461);Lucky Bay, 35 km ESE of Espefance,P. G. Wilsoil10006 (B, K, MEL, NSW, PERTH). Flot:ering and fruiting period. F]owers from July to October; legumcs appear in early October and reach maturity in November and December. A(.tcia igricqns (Labill.) R.Br. rvas based on Minoss nigricans Labill. At Florence (FI), where much of Labillardidre'scollection is housed, there are two sheetswhich can be regardedas type naterial. Both thesesheets are " " labetled Herb. Webbianun. Ex Herb. Labillardidre ard pinned to each is part of Labillardidre's handrvritten manuscript description of Mimosa nigricans. One of these sheets cardes two specimens of A. nigricans while the otber has only one rather depauperatespecirnen of this taxon; thc right Jrand specimenon the lbrmer sheet is here selectedas the lectotype. This " sheetis annotated(not in Labillardidre'shand) Nova Hollandia ora austro- " occidentalis and althor.rghthis annotatiol is in a form diflerent fronr that " ", i[ tbe protologue, i.e. Habitat in tcrra Van-Leurvin it urdoubtedly rcfers to the samelocality, that being EsperanceBay on tl,e south coltst of Westerl Australra. A Central Arnericar species, Miilosa nigricans, was described by Martin Vahl in Ecolog.Am.3:37 t.29 (1807);however, no exactdirte of pnblication can be found for this work. Labillardidre'sdescription of tho WesternAus- tralian taxon, Minosa nigricqns,appeared in March l807, and it is lrcreassumed tlrat Labillardidre'swork antedatesthat of Vahl. As Acqciqnigricans (Labill.) R.Br. was based on Mintosa nigricans Labill., the correct author citation for " this plant is (Labill.) R.Br."; however,if it is subsequentlyfound that Vahl's work was publishedfirst, the author citation would become "R.8r.". When Robert Brorvn nrade the above new combination, he apparently " applied the name to a plant from the Albany area (perhaps R. Brown, Iter " Australiense,1802 5. King George III Sound Bennettdistribution number 4320, which is housed at BM-photograph seenby the author; this plant is here describedas A. leioclermaB. R. Masiin). Brown's descriplion of ,4. x€ricar,r, although brief, stronglyindicates thut he r\as referringto A.leiodermq even tlrough Mimosq nig cans was given as the basionym. Most authors, including Bentharn (1864), seem to have followed Brown's intcrpretation arrd thus lrave misappliedthe nan.re,A. nigricqns.Ironically, Robert Brown appearsto have collectedthc true,4. nrgricars(see below). Parstauthors have generally consideredA. ruteefoliq Link to be a synotrym otlA. nigricans(Labill.) R.Br.; this approach is also adopted here. However, it should be noted that there is somc uncertaintyconcerning Link's intention " when lre pnblishedA. rutuefoliathus: 5534A. rutaefolia. Ait. Kew. 5. 465. Hab. in Australia. l. T. Elegansplantula floribus flavis. Foliola inaequalia." On page 465 of Ait. f., Hofi. Kew. ed. 2, vol. 5 (1813)Robert Brown made the new conbination, A. nigricans(LabiJl.) R.Br. and proposedthe cornmon " " name, Unequal-\\jog'd Acacia for this species(presumably to reflect the discrepancyin size betweenthe proximal ard distal pair of pinnae). Link's "Foliola "Urrequal-wing'd", inequalia" presumablyrefers to Brown's and it is therefore assumed Link intended that the name A. rutaefolia should replace l. 440 IW .SEDIUCEJ .I 'sJIBq isec oEUurdlElsrp SrruEeq-pu"l8 puB'sp?oq re,lroglo8t"l r{cu"JqroSuoi s -uacse;ogurelduirs s1tfq sun4tEtu ruo4 poqsrnBullstpsr opoltotladuraatto:ty 'srrEq 1; S pasIrq-alncjeqnl8uu€oq seqcuElqpeqqll ,{l}ueululoJds?Ll osle 1I plIE t 'snorfel ur s,ror8 oslB sorJodssrLIl sLto)t,t8lu y o1t1 slrrads srql u l?lsuof,-JEeu 's3^rt!ieJ.}sosolc lserreu ,{llBcrqderSoa8sr oltoltotladtua t' .stltrrrt?& t=j8,u"n, o 'sJJtrJJ.;joUul Jpuurdler\rp dtrue;q-pue16trllnurou puc .ppJq rJ^\oUrrSrnl :i eldurrsslr fq pezruSocersr sJrf,rdsJJllul Jq] tnq'(ittorqElS fTJlJldutocsI tiJlLI,^a) U DLaJapoDly' clqLuos3r,{ll€rJuJedns uBt stnl.rittt suIJoJluocsaJq€lD F'(e€uulo Jo 3qlJo uol asllraql u ,!\olequrul pe}rrurs,(11r:rurou st qclq,$)puE[3 luaulutold aroru ueuo sll puE 'slr?q e I pos?q-elncreqnl8uu€eq soq3u"Jqpeqqlr ,{ltuoulLuoJdeJou ,{11eur:ou u slr dq parJsrn8urlsrpoq uBc sLlD)t.tBtu 'e^oq" patlol]uetu sralaPJl?LJJerJl ol ,' 'DuDru'K).tq u uorlrpp" sJqcu?rq snoll-lpldslqq]r^\ stluoJ esoq:],{lelcedse 't I 't' sr pesnjucc eq ol ,{lelrT lsour sr .trrJl,/trir qclqr\\ rpl't\ salcadseLlI 'Jg 'urTseJi{ ' 'Y D >JD aWUd puir g DlDLld DEaLuy' ql[e8 Dd,,tD)otsDl 'I 'ulls€t{ V 'uIlsEI^ll ' 'V U g DSoLu o.tqns V U gltaqtrdu V IpLu1 DtuapDlLldd uo s Jncco seop urall€d IeJnlcnJtsJo ]Jos oulese\t lnq'aDlla?lnJ eqt ur LtoLuuoa li Jou olE seruec€JpecnpeJ esolll epou ]€d sp?all le,{\ou xls ol o'{\l Jo slelsnJJ Ji 'lrx! ur Sortlnsersnql Jtol qcEeur .rncco,{€Lo soluecBJ 3}€utq eserJlJo eelql ol .{ euo ,{.relllos e ,{q papuolqns osl" sI qcrLl^\sIxE eurec?.ruotls Ala^ lc€Jq Il€rus 'selcullpocl u uo polEn}rs^ltE:]sip':lcErq lts"q fJElllos s ^q popuelqnsrlJuo o^U Jo lslsuoo ,{oql : setuaJ€Jpecnpe.I eJB tunl.tSlu k' uo s3cuecselollul erJI 'sorceds l( eseqtJo ^u" dtllo^o 1ou seopo8ue: lectrlderSoe8u,lrou{ slt osJ€ ( :(,lloleq oas)]LroLrie8u€Jr? ocuecseJoUUt s:tt ut ,{lluecgtu6rs:tsour sJaJIP ,rrrJl,/t/t/ p 't' '"xel 'g 'rilJsEtrN 'B eseql tuou ullsal,{ d DLutaPolal fl puE d DPDI) -f -o4adtua'y ''lpue,^A DaDrurto.lq ()1pel?leJ ^losolc aq ot s]"eddB:]l p€oLI H f' 'czrs 'urnluslunput u Je,\\oUred sre,\\ogJo requnu pu" elnuuld qcueJq sll Jo II ,{lrsuep eq1 ur,{llurcedse sercedsalqcr:e,t l€rl,\\euros E st sLttDl,tSlttrt)otr.ruo,

II -tt8tu 'y ol selJddp,'(lur€uec plrp a,Ltsueqe.rdruoc sr olo.louSl lsourl" 1rnb ,' Jo S. uorlducsepeqt '(stsrxoeuo od^l aql uaas e',.eq q8noqlly 'D.tn)sqo'V JD lou 1 1r ruo{ tuoleJrp lr reprsuoctou plp luEriluogl?ttl Pelels osle rlco):puclSul -( pecnporlur pesoddns se.\\ srrlt : Dludpoluad au.rcLr uro4 uoeq e^"rl o1 tuEld V sl eql repun suepr"S u"LuJaC ur u,ror8 sree,{ l"re,\es loJ s?,rr satcadsslrJl lerll 'g9gl) 'DiDtou8t lr pet?ts (961d qco) V 3o uorlducsep IeuISuo oqt uI ',{lrl"col el srql }t; 3ur.trncco ortoty aleuurdrq,{luo eql sI sltll leql p. runueqreH eql sJequlaul ^q elPplt{ uo FuqcalloJ lee^er v,^[ Jo J"]s Pu€lsl'uollJolloc q e,\rsuelurllroceJ esneceq suott.r8tu / ,{lq€qojd }sotu sI 1I snll UJJS10u J. e,req1 q8noqlF :D.lnsqo'V ropun (8ltd't98I) luEqluag,{qpetrc:i (pu"lsl e. elpprl uo sr qcrq,n),{rg pu?lsJesooD ruoU uorlcelloau,'\\olg ueqou "'oslv .I '(T9t6t 'IEIN) -toq1n" eqt ,{q ueasuooq seq pu"lsl elpprtr ruo{ uoturrods lueserd { 'uru]puow ]rqcno^ " o8eledlrlcJveqcleqcJu eql uI spu"lsl loog ,{puz5 puE 'elpprJ uror-l,.rrrsgo selrc(00I d '69-6I)puE (SZ d 'eEOt)srllIIN'eldruExe roC 'DJt1)SqOl ,/ Y peIIeArdeeq UoIJO SBq ,trrr?.lti, t' el-LI}3rF lJullapotal t' sE paqllf,sep erJq uox€t sun!.t8!u'tl eql jo uoqecrlddesruroql esn€ceg eqt ol euiru Jo .F 'sllo rJ\seJ:Jq1 tJodd S pr ol ecueprle ou sr eJeql'JaAa,\\oH (DLuoporal k' et) saDl,!8ta t' s" u,\\org p' geqol ,'(qpegerdrelursrur serJeds oql loJ eureu,treus Sutsodords€,rl ?q lerl} 'p ',{la^rt"urell? ',. ro ^\a) lly go g9p e8vd o1 elqeroJor,{ll?Jr}Due}s,{s sE,\\ qclq,la 1n 'o.t1olao1n.r't'sarJeds ,, 't ,reu r Surqr.rcsaps€,r )url lEqt pon8r" eq plnoc tl 'euteu ettuutt8e[r u,uo srq Parldd€sItu,4llenlct sBrl 'uoxet srlt ol Dllo/>Dltu eureu eqt 8ur,{ldd€ uI DutapoDl 'y {urI t' '.Ig ]I sE pequcsapareq sr qcrq,^\lurld eql ol oruEu?q] parldd" ,{llueJ"ddeeq JI 'e^oq" d (\rql,1) suDlJStu uoD"urquror ^\eu eql epElu u,rorg uorl,r pauort -J t, 'ro,\e,toq -uoru sV snonuledns Sureq oleulrll8elJl sr eluBu s.lul-I :srtrtlti, J; 12. Acacialeioderma B. R. Maslin sp. nov. Frg. 17 IAcacianigrica s auctt.non (Labill.)R.Br.: R. Brownin Ait. I, Hort.Kew. ed.2- 5:465 (1813);K. Koch,Allgem. Gartenz. 26:195 (1858); G. Bentham.Flora Auslr'al.2:418ll864l.l [A..tciarutacfolia aLrctt. non Link: Link, Enum.Planl. Hoft. Bercl2.441 (1822); W- Aiton, Ann. BlunistereiGarlenbisitz p. 58 + tab (1827)-l hulex | 2 nl altLrs;rdml manifeste costati, glabri; spinae axillares nullae. l'rlr, bi- pilnar^: pilnae\1) 2 (3\-jugae:pinntrlaa pinnarunr ploximalium 2 (3)-jugae,pinnarum distal- ium 4-8 (I3)-jugae,51l x 1.5 3 (4) mn! planaevel interdumparum recurvae, glabrae. G/darsessilis, in pagina supera rhachidis inlra insertionem pinnarum posita: plerumque glans minorin rhachidibuspinnarum jnfra insertionem parum 1 2 sumnorLrmpinnularun. Pr.rrr- rali 10 20 (30)mn longi,glabri. Capintlaglobulosa, 28 35-lloribus.Flotue s-:me-a:e ca[rcis /16l oblongi;petala glabrc. Leeumina30 40 x 6 7 nn, gl.abra.Senine in leglrmetrans- versaad pdrurnobliqua, 2 3 x 1 5 2 n1ln. TJ??r2l km lrom Denmarktowards Mount Barker,Western Austraiia,2l Sept. 1972, B.R. Matlitl 2955(holo: PERTH; iso: CANB, K, MEL, NY).

Sltrtrb 1 2 nt tall;. branchesprorninently ribbed, norrrally red to brown, smooth, glabrous; axillary spincsabsert. St?r/c,! linear to tarrowly triangu- lar,2 5 4 x 0 3-0.5 nm, slightly thickened especiallyat base. Zcalcs bi- pinnate, glabrous btrt rachis sometimespubcrulous aboye: petiole | 2 mm )ong; rachis 3 6 (8 5) mm long, prominently ribbed above (rib f vertically dilated toivards the terminal scla.);teruinol seta narrowly triangular, l 5 2 (3) mm long, slightly thickened at base: pinnqe (1) 2 (3) pairs; pinna racltis 3-4 (6) rrrn long on proximal pinnae, l0 30 (40) mm long on distal pinnae, prominently ribbed above, apex 0.5 1 5 mm lorg; pinnules2 (3) pairs on proximal pinnae,4 8 (13) pairs on distal pinnae, nornally narrorvly oblong, 5 I I x I .5 3 (4) mn, flat or occasionallyslightly recurved,dark greenabove, light green below, n.ridveinline, a second shofi less colrspicuousncrve often diverges from the pctiolule. Glan.l sifuaLedon upper surface of rachis below inscrtion of the pinnae; a smali gland normally occurs on upper surfacc of distal and mcdial pinnae rachides (rrormally absenl from proximal pinnae) below uppernost I 2 pairs of pinnules;sessile, 0.6 I nnr diam.. lip and orificc prominent. Inflor<'stentessinple peclnttles I 3 per axil, l0 2b (10) nrnr long, glabrous, basal bracts 3 4 arrd ciliolate; flo,ter lteads globular, 7 l0 mn diam. at anthcsis,with 28 35 flowcrs. Brat:teoleJI I . 5 mm lons. sparsclyto moderatelypuberulous, dinorphous: lower series(arranged in I single ring at base of receptacle)reflexed, oblong 1() obovate,0 5 0.8 mm broad;_upperseries (spirally arranged) erect, claws linear, Iarrinae pear-shaped and inflexed. Floi|ers S-merous;calyx I i2lenglh of corolla, diviiled for i/5 its length into somewhat imbricate inllexed thickened ciliolate lobes. tube often dark brown glabrousto glabrescentand obscurely5-nerved; petals 2 2. 5 mm long, connate for 1 i2 213 thelr Jengtlr,glabrous, + obscurely5-nerved; of4ry tomentose at apex, occasionallyglabrous. Zegarres somewhat hard and brittle, narrowiy oblong, 30-40 x 6 7 mm, flat, raised over seeds,glab- rous, dark brown, ntargins not contracted between seeds,thickened, light brown. Seedstransverse to slightly oblique in legume, oblong ro cllipiic 2 3 x 1.5 2 mnl tatr to grey-brown; pleurogramprorninent, continuous or with a_narrowopening lowards the hilum ; areole1 . 5-2 x I n.rm;.fizicleslightly dilated, reflexedbelow a :h graduallythickened olce-folded aril. Distributio and hahit.tt: (Map 7) South-west Western Australia: common in the Albany, Mount Barker, and Manypeaksdistrict but has been collected as far east as West Mount Barren. This speciesfavours sandy loarl in Iow Jying areas nea[ watercoL]rses,but it also grows in lateritic s"oilson higher ground. Under L nigricans,Meisner (1844) cited two specinenscollected from Mount Bakewell,which is near York, about 9o km eastof Perth. Duplicates of these plants_(Prer,rs891 and Drumntoncl314) are at Melbourne (MEL); botlr are l. .sr'l&e,l/i Meisn., nol A. leioderma. tvn '0261 ^oN'us png uos!1/1,1 nnaEtu,rS LnqDH V Q 'tAI I/ !r',{uBqlv:IZll -( ',{ 'e8plr8 ue8lpj racldn tuorJ peor sleed {utN Suole salltu :989L Iotr Y ',{ueqlVE 'ciu€d 'rnoqrpH t 'stuBIJIr/A Jo 3 uoerD :(13I,{) /88 .r. .)-rd Itr{ou ssatuLrdreau : ,{rgri,N y '[ou) qlnos selrlu :(Stt6t puE u \ S luno^'purlos T€tZ .gntg]o 'dnrn8uorod0I I tt6t tA ht) lla;rxD!{ s,a8roeC3ur) :6S62 ulllDN A g sprB,\\ol ro)rEg lltnol,\ Jo g rul ! :00it zr?,0/r-l siurtrrls qtnos :(Hldgd 'OV) peny y',{uBqtv Lluou a./l V 6961 H 'ra^IU Jo Lrr) rnoclv'.(VA\n) runot :22662tro2, plero8zlrl SZ 'tghl)Lulvoa) 1's)Eed,{uEy{ { S f re^\ot :(Hld3d 8Iu Puoutun.to I'.{uolo].r.^lu ut,\\s:(HfU3d'9BJ) 6/n/ J SutnuDJ lsta'turod s.rellrl^J:(urcs qdlrSoloLICI lsqlunu t/{ J'rnoqreH trog 'osuall?r1snvto !punos tri{g)oz€t uortnqrrlsrplteuuefl s z0gl retl utto.tfJy III e8roaD 8ur):108t dnrnsuorod:9 LZ ulltlv H E -I'IeA uollrc :VI'IVUJSnV Nlt ISl,^^ I lr./rrg S /'e8ur>l -. L

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SI rfit ,1{r, -t fl /t/ - *r- iij -l 'il I. G./ st Fk)trering and Jruiting periorl: Flowers from August to early November; both young and mature legumesare presentin October and November. Acctcio leioderma has previously becn known as A. nigricans. This is becauservlren Robert Brown made the new combination, A. nigricans (LabiIl.) R.Br. (basedoo Mimosa nigricansLabrll.) he apparentlymisapplied the name " ", to a plant from the Albany area (perhaps, King Georgc III Sound Bennett distributior number 4320, which is housed at BM-photograph seen)which is here describcd as A. leioderntu. Acacia nigricans (sensu lectotypico) is quite a different speciesald is restrjctedto the Esperancedistrict which is about 380 km due east-northcastof Albany. Link proposedtl're name A. rutaefoliapresumably as a nomen novum for A. nigrit'ans (Labill.) R.Br. Link's name is illegitimate, being superliuous. In applying his name to the Albany plant (.A. leioderna), Link has actually misapplied his own illegitimate name. For a full discussior see under y'. nrgrlcans. Acatia laiodermdis a mcmber of the L brovniqno H. Wendl. group of species(see p. 394). Its closestrelative is A. enpeliocladaB. R. Maslin from which_it is most readily distinguishedby havirrgglabrous, prominently ribbed Drancnes. Vegetatively A. leioclerna car sometimcs resemblc ,4. gllDt'rril Meisit. and thc glabrous lorms of :1. broyniqnq H. Wendl. var. bro)t't1iafia see these taxa for distinguishingfeatures. The soeciflceDith€t refers to the snooth br;ruches.

13. Acaciapentadenia Lindl., Edrvards'Bot. Reg. l8:t.1521(1832). Lectotlpe: Nov. Holld. Ba.rtcl-central fragment on sheel (CGE-photo seen)-Fig. l8 Meisn.in Lehm.,Plant. Preiss. 2:205 (1848). Aca.ia" biralandulosa "3rd 4?.r SwanRi\er, Dnmnot . coll.lL No. 97."(iso: MEL, NSW sphalm. colle.- lion", PERTH). 'l A@dancilii Hort. ex Seelnan,Verh. Gartenbau Ges. Wien 1846:72 (1846); fide K. Koch, Allsem.Gartenz. 26:195 (1858), "Neillii " pro syn.sub. L pentadeniaLtn'Jl. Type:n.v. 'lall shrub or snoll lree,2-5 (1) m higlr; Dzirltsnrooth. grey to brorvn; brcrclrc,iprominently ribbecl, glabrous or rarely glabrescenl;axillary spine; absent. Ie4re"r bipinnate, glabrous to glabrescent;petiole 2 3 (5) mm longs rdchis 1O-55 mm loltg, promiirently ribbed abovc; pinnae 2 5 (6 8) pairs (proxin.ralpair ofterrcad uco us) ; plirra racfus3 l0 mn iong on proximal pinnae, 25 80 nrrr long on rernainingprn:nae; pinnules 2 4 pairs on proximal pinnae, l2-30 pairs on remaining pirrrrac,oblong (nay become + triangLrlardue to folding of margins),(l 5) 3-6 x I 2 5 mm, flat or sligiltly recurved,glabrous or sparselyciliolate, finely l-nerved, dark green above, light green below, apex somctimesprominently inllexed, base obiiquely truncatc, petiollrlc + absent. Gland situated on upper surface of rachis a shofi distance below insertiorr of pinnae; present or absent on petiole belorv proxirnal pair of pinrrae; sessile,circLrlar, 0 5 0 9 mm diam., lip prominent. Inflorest'ent:es of (I) 2 4 (5) short axillary racemesbearing (I) 2 4 florverheads, each raceme, as uell as the peduncles,subtended by a brown ciliolate bract; tatentc qxis Iessthan I to 4 mn lorrg, glabrous;petltncles 6 20 mm lcng, glabrous;./oIcr /zcarlsglobular, 5-7 mm diam. at anthesis. -Flolers 5-merous: cou\ | /3 to nearly1 ,t2length ofcorolla, dividedfor lcssthan 1 /4 its lcngthirrto inflcxed f thickcned ciliolate slightly imbricate lobcs, tubc brown glabrous to sparsely puberulousand nervclcss;petals 2 5 mm long, obscurelyl-nerved, yellow, glabrous,tube slightly contractedtowards the base;orarl glabrous. legi.irres somewhathard and brittle, narrowly oblong, 15-50 x 3 4 mm, flat, slightly raised over seeds,glabrous, dark brorvn; tndryifis rlot contracted between seeds,thickened. yeliorv. Seedslorgitudinal ir legume, oblong, 2 5-3 x 2 444 9nv '|fln) 'rllr^(rrcd rdrs 'lrPtrrur(l I'u :r^o8totlll/\) us rt6l s'u.'!1t!,4a'AN ')H '8\kJr 'B\ Jo/l|\ zl z t't'4pto'\eqrcI:tHl Uld ZZ5Zlttl'DA tr g \eMll:lrHlsPoJ qlnos ilreuruog tul :(Hld3d'IUS) ulltDw g'rnoq.reH^pullA II sp.reAorelodlei^ 3 S.tT 9682 'qlnosd spre^\o1egrtoqtroN tul : r(lV ' tt s UDED.!2uo7/1 o ood lunol ,l :('lgt l) ,{ :N'uolreqrred')oorgS 9. I s96l 9 *ft WaX 8ra'.;tII D)19 1 7'alodlBy' Jo A\S selru 9 i're^r{ t: Aeecf:eefrLip971 L,tup.rnS loet :(CV) 58091nltplg /l/')nrLuuec puE IuEqlV '(]1/) t' Jlra^rd 'N '/ ''tl\tN uee^\}eg:(HJlad LtiZ t.tqsv uouueqs rEeN :VI'MI-LSnV NdaJ,Sf,\\ I l ',rolaq ( uIS xe uuoo oi?utS.tDtu g) qBJlLrf uI ul€ol clllle]?l uo 33s lseJoJ Pu? € sdru€^\s JEeu rln]co osp,{errl lnq ( llen1N'f .tolusua P snldtp)ng) $eJo! uJe>l pot"rcoss€ ,{llEru]oN (fueqly rteu) ,{?q.IoJ o] uo aqued uroJJ sooISoJ rllr,r tl ursqlnos :"rl".rlsllv urolse^N lse^\-qlnos l8 (1?w) :JDllqDq puD uollnqulslo 'iI.I" p3^.lnc Si peua{crqt € ,{\oleqpexeueJ 'pauellEu 'uoqs a/r/r/1/ :unlrq eql spJ",\\o:lSuruado 'u,{olq 'rutu ,\\orrsu " rllr,r Jo snolrurluoc'lu3urLuord tuo.tSotnald : ,{uIqs IqAII

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Flou'ering qnd fruiting perio(l: Flowers from September to October; mature Iegumesoccur from Decemberto January. . The type of.A. pentadaniais ilt.Cambridge(CCE) and a photograph of it l]as been seen by the author. This sheet carries three separateiragments: the central one, consistingof a singleleal three flower heads,and a minute portion of stem, l]as been chosen as the lectotype. One of the remarnrng lragmentsis a leafolrll.uhile the orher lcon,i.ting ol rheupper porrion of; branchtprobabl5 represents the stunted form of A.-Tcntu,ltnia$trich is relerred lo belo$. Acatia pentaclenia is typically associated with Karri (hence ' lorest the comrnonntme. Karri Warrle ) u hereir rrormrllyform. trrlj.dense thiclers. Howe!er. lt some areab. particularly il the Walpole-Dennrarkregion, L pentadeniqgrows near swampsand on lateritic loamin Jarrah forest. Under thesecondilions it is often a stuntedshrub (to about 2 rn ttll.) aud its pinnules are oflcn \nralleralld rnoreirrfle-red tharr norrnal. Acaciapentadenia is most closelyrelated to A. sttbracemosaB. R. Maslin. These laxa, besidcsbeing the tallest speciesin the pulchellee,also possess larger leavesthan any menber of this Series. Other characterscolllllol to them include reduced racemoseinllorescences ald obliquely trutrcate, + s_essilepinnules. Both speciesare associatedrvith the Karii forest, but iheir distributionsdo not overlap. Acaciapenrodetia is readily distineuishedfrom A. sttbracemosaby its glabror.rsbranches, leavcs, und peduncles"(seel. sr.i6- rqcenosafor further details).

14. Acacia subracemosaB. R. Maslin sp. nov.-Frg. 19 .. . Ftntex2 3 m alrus:razl densepuberuli ad hispiduli;spinae axillares nullae. Folra brplnnata;rri./.rir 1,5-55 mm longd:prl/ioe 3-6-)ugae; rhachis pi natumprcximalium 2 6 mnr longa.arlrrum l) )U finr tonga;piln laeptnoa(un proximaliul]1 2 3_jugae, aliarum 7 ]3_ lugae.obto gaeact i ifllngutafes,I 6 x 1.5 3 mn1,modice puberulae, basibus oblique truncatis,petiolulis - nullis. Gt rr sessiljs,jn paginasupera rh;chidis hiud procul sufra pinnasproximales posita. .tnftotesrcntia racemosa 6revis aiillaris,2 4 (6)-capilata-f1.raepeltn;li 7-17mnr longi. modice puberulr ad hispiduli.Capl/r/1d globulosa, 15 20-lloribus. 5_ mefae;r.r/,rir ,,bl triangrLlore..Lcg nilta 15-40x4 6mm,spalsiDlpuberula. ,.tcmr)/a in leguminetransver-sa ad obliqua immatora interdum longitudinaiia), + oblonga, 2 2 5 x 1.5

11".,^ll-ry of Karridaletowards HaDletin Bay, western Ausrratia, t9 Sepi.1972, a. R. Maslin2871 (holo: PERTH; iso: BRI,CANB, K, MEL, NSW,Ny, pERtHr. _ Tall, spreading,dense slrurb. 2,3 m high; bark srnooth, grey to light brown,. _lenticels prominent; branchts I'nely ribhcd. denseli puberulJus to hispidulous(hairs normally tubercule-based): axillary spines;bs;nt. ,Sr"- ules promioent on young shrubs,narrowly triangular,2 4.5 mm long, spread- ing, curved, sparselyciliolate, light brown. leales bipinlate. in

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Acacia subracenosa is closely related to A. pe tqLleniqLindl. These are the tallest members of tlr'e Pulchelloe, reaching 3 metres and 5 (7) metres respectively,also their leavesare larger than any other speciesin this Series. In addition, both specieshave obliquely truncate, _ sessilepinnules, and reducedracenrose inflorescences. A(acia subrqcenosais readily distinguished from A, pentadeniaby its Jrairy branches,leavcs, and peduncles,its solitary leaf gland,its solitaryaxillary raceme,and its triangularcalyx 1obcs. Although both speciesoccur in the Karri forest, their distributionsdo not overlap. Some of the larger-leavedindividuals of A. brownionaH. Wendl. var. obscura (A.DC.) B. R. Maslin may superficiallyresemble ,4. subracemosq espcciallysince the indumentumon both thesetaxa is quite similar. However, A. subrat'entosamay be recognizedby its f sessile,obliquely truncate pinnules, its triairgular calyx lobes, and its soiitary basirl peduncularbract. The specificepithet refers to the reduccd racenose inflorescences,a char- acter not particularlycommon n lhe Pulchellae(see p. 392).

15. Acacia preissiana(Meisn.) B. R. Maslin comb. et stat. nov.-Fig. 20 prcissiana Acacia" obscura A.DC. var. Meisn.in Lehm.,Plant. Preiss.1:20 (1844). Type: In region.interior. Ausiraliae rnefid-occid. m. Oct. 1840He$. P/.eAr-No. 885" (iso:K pholographseen, MEL, PERTH-fragment). Acaciaprcissiana Lehnr., Del. Sem.Hort. Hamb. (1842),nom. nud. lAtaciaobstura auct. non A.DC.: Bentham,Flora Austral.2:418 (1864), pro parte as to DrumnandlV:18 and Pr"6.! 885.1 Prostrate, multislemmed shrub; branchesto 40 cm long, linely nerved, brown to greyish, indumentum a mixture of pilose and hispidulous hairs (rarely glabrous); axillary spinesabsent. Stipult.soblong to ovate, 1-1 .5 x 0 6-1 mm, somcwhat obscurely I 3 nerved. Zeales bipinnate. iodumentum as on brarrches(except for pinnules):prtiofu 7 2 (5) tnm long, terete ruchis (2) 4 10 (20) mnr long, obscurelyribbed above; terminalseta 1-l .5 mm long. straight or reflexed: pinnae 2-3 (5) pats; pinna rac,4rs3 5 mm long on proximil pinnae,7 20 mm long on distal pinrae, obscurelyribbed above; pinhtles 2 3 pairs on proximal pinnae,3 7 pairs on distal pinnae, rrarrow]yoblong, 4-6 x ca. I mm, revolute,pilosc to puberulous,nerveless and uneven(in dried state) above,prominently 1-nervcdbelow. G/4nrlsituated on upper suface of rachis normally halfway betweenthe pinnae (may occur immediatelybelow insertion ofpinnae);sessile,0.30 .5 mm diam.,lip and orificenot prominent. lxfoler- cencessimple; peduntles solitary, l5 25 (30) mrn long, moderarely piLose (hairs spreading,straight, ca. 1 mm lorrg),rarely glabrous,base ebracteate at anthesis;/oler /readsycllow, globular, 6 7 mnr diam. at arthesis,with 20 28 flowers, Brqcteolesca. I nm long,ltispidulousto glabrcscent.F/olfer,r 5-mcrous: col1,x | !2 length of corolla, divided for ca. I /4 its lergth into triangular ciliolate or glabrous lobes which are neithcr thickened nor infiexed,5fi5)- nerved,tnbe glabrous;petals 2 mm long, sparsclyhispidulous at apc\. 5(15)- nerved(midrib promioent, two marginal ribs lessprominent);olary glabrous, Lagunes and sccrls-see belolv. Distributiortancl hahitat: (Map 8.) South-wcstWesterl Australia; occurrins sporadicalJyfrom Bindoon southwards to near Albany. Acacid Vreissiana grows in laterilc (or white sand over laterite) in Jarrah (Eucalypttts norginata Donn ex Sm.) forest. WESTERNAUSTRALIA: Near Albany, W E. BlaLkolls.n., Der L9t7;C()llie L. Diels 2175;Waroona, Diels and tuitzel250i--Wesrefll Au.tralia, J. Dnunmonrl(tV): l8 (MEL, NSW,PERTH); Woololoo, M. Koch.1600 (A1). MEL, NSW,PERTH); 62 n]i peg,Creal NorthernHighway, F. Lullfitz 5920:Abour 12 mi N of Bindoon.B. R. Mastn;2618and 2619:19.2km E of Collietowards Darkan, B. R. Ma.\lin3201 (CANB, K, Ny, PERTH); MountTalbot, O. H. Strget s.n.,23Jan. 1921. Flowering periocl: December to JatrLlary. 448 -!

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Although A. preissianais nornally hairy, a glabrous variant has been rccordedlrom Mount Talbot which is near York (seeS(trgent s.t.). Reliable fruiting material of A. preissienahas not becn collected. The following is a descriptionof somelegumes and seedscollected from the grorld beneath-8. R. Moslin no. 261Eard 2619'.Lcgumes somewhat hard and brittle, narrowly oblong, 15-30 x 4 6 mm, Ilat, slightly raised over seeds,glabrous or pilose, dark brown; nqrgins not contracted between seeds,thickened. SeedsIorrgitudinal in legume,oblong, 3 5 4 x 2 mm, brown'.pleurogram qite prominent, continuous: areole ca, 2 x I mm: .funicleless than 1 mm long, abruptly expandedilrto a thickenedstraight aril.

"Near 16. Acacia tayloriana F. Muell., Sth. Sci. Rec. 2:l5l (1882). lrpe; the Blackwood-Rivcr." (holo; }dEL 49742; iso: PERTH) Fig. 21 Prostrateshruh; brancltcsto ca. 30 cn long, finely nerved,sliglttly angular towardsthe apex,brown, indumcntLrma nrixtureof short piloseand hispidulor-rs hairs; axillary spines absent. StiprlleJelliptic, ca.2.5 x 1 nrm, obscurely 3-nerved,ciliate. leares bipinnate, indumcntum as on branches(except for pinnnles);petiole l 17 mm long, ol'tenslightly verticallyflattened ; rachisT l7 mm long, narrower than petiole; terrninqlseta linear, ca. 3 mm long: pinnae1-3 pants,pinna raclis 15-20rrm long on proximal pirrrrae.l5 22 mm long on distal pinnae,ribbed above;pinnulcs 4 5 pairson proximai pirrrrae,3 5 pairs on distal pinnae,elliptic to narrowly elliptic,9 l1 x 3-5 mm, 1lat, I antrorselylrispid on margin and below(hairs tubercule-based), 1- prominently reticulate, midrib pro- minent, 2 3 lessconspicuous veins divergilg liom the petiolulc. Glanclsitttated, on upper surface of petiole eitller immediately beiow insertion of proximal pinnaeor sonredistance belor,v them; sessile,0 8,1 nrm diam., lip not promin- ent. Inflorescence,rsimplc; peduncles solitary, 25 50 mrn long, sparsely to modcrately pilose (hairs spreading,slraight, ca. I mn1 long), base ebracteale at anthesis; /?or1'?rlteads cream, globular (l obloid in thc bud), 7 8 mm diam. at anthesis,\'ith ca. 22 flowers. Brqcteolesca. 1.5 mm long, ciliolate,claws graduallyexpanded into slightlyinflexed concave laminae. Flor,erss-lr-terous; calyx I i3 I /2 length of corolla, divjded for ca. I i3 its length into triangular ciljolate lobes which are neither thickened nor inffexed,obscurely 5-nerved, tube glabrescert;p etqls2111t11long, connate for | /2 2/3 the:l lengtl.r,l-nerved, glabrous to glabrescent;orarl glabrous. Legunes aftd seerlsn.v. Fig B DistribLltionand hllbitet:(Map 8) South-westWestern Australia: Known only reti tiom a restrictedarea near the BlackrvoodRiver in the vicility of Nillul (tbout BrE 25 km due north-eastof Augusta). This speciesgrows in gravelor deep grey A-I sandysoil in Jarrah forest (EucalwtLtstncrginatq Donn ex Sm.). 77. WESTERNAUSTRALIA: Sue's Road, I 5 km N of BrocknrarlHighway (east of Nillup), B. R. Maslin3526; Nilllrp, R. D. RolcelE Ion Flon,eringperiod; Jamary. rha A(atia tatlot'ianais a rare species. With the exceptionof the type, there ins( are only two collectionsof it known to the author. C"t This prostrate, summcr flowering speciesappears most closely allied to lon apu A. preissiula (Meisn.) B. R. Maslin liom which it is readily distinguished 10 by its large, flat pinnules-see A. Feissiata for further details. Also the petiolar glands, cbracteatepeduncles and trizrngularcalyx lobes indicate an Tw lsol amnity witlr the A. dnntntonclii Lindl. group of spccies(see p. 394) fron.r which it js most readily distinguishedby its globular flower heads. However, until further material of /. tavlorionais collectcd(il particular,fruiting specimens) rib the exact position of this specieswithin the Pulchellueremains sornewhat Pilr obscurc. on 450 4r5 (€)-6e9tt ISt 'eluuurdrq .e,'oqa s" runluounlpur salDa7 snoJq€l8 ,,roloq puz ut8reur uo 'snouecs'rurlr .t'luasq? et"rlrJ t.0 "J x g.Z-Z.Z-Z satnd\ssalndus saurdsdl? rx? :esolrd '(s1e1qcue.lq ,peqqu ,{1goqsdlesuep uo elrrl,r u3uo s[u:eprde) qsr^eJBI]"p 'qntUs ,tcerE ,{lsnoncrdsuocur saq)Lrq.tq:IIE\ ur I.I I peqf,uerq ,{lrJdo ')'8\vJ:o'r '6t61 hJ}t :HfXEd rotoq)$t6 .taupDtV J dnV 6l prlert\nVur .r\\ 'JSupUuetrD .)d(J .plnprdsrq .LLru r+u f Z r^t ['Bnbtlqo eurrunSolr.r ,r.rrr.rS r\urp pe rJtpolu I \ 0! 0l 'Ete.tld'ElEcrldtlo DutLunnd1DuanEaT rlrtsolr t\J\ rsDulnqJ.stlnled.\nq.tu)utLuord.trnpd,slaurnqg .srlnted '\nqL1u)urulord 'sor€ln8u€ut srrnail JnbLJnrrld.r1rd,:rrde:n6ilnt:r1d srlrOs::ir,ii pe pe Euolqo, tqol-snIlD, :a"rrur-S .'u.,o7y ealrls:,r ."reulnqr .rtlniid (srCuol u.Lu S. I pE) cnq|lueurLrord \td Jn ulDt nirlo.'t1Dtg cnq!oU-SS .eiolnqo,B uirrrrdaj 'rteJq"ln'rteJqEln ,LunJEJnuuld,LunJElnuuld 91 pepp lltueqnorlnJeqno rnuoltiuol lJlrtrtLttr| 9ZSZ 0Z tln)unpadtln)unpal Llrnjoruutns €, I urauorurauoruesul esur erlur runrEuu.rd snqtuuro tnc ]onbrle snqrprqseqrur rourtuJourru suel8sueJ8 tuerle:r!riodmerle :rlrsod srpiqceqisrplqa?ql 'srlrssos'stlt<

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i on branclres(hairs shorter on pinnr.rles);petiole ca. I mm long; rachis 4 13 (20) mm long, commonly channelledabove betweenllrst two pair of pinnae, J, narrower and ribbed above betweensubsequent pinnae. terninal sela linear j to narrowly triangular, I 5-3 mm long' pinnae(1) 2 3 (4) pavs; pinna rathis 2 5 mm lorg on proximal pinnae,(5) 10-20(25) mm long on rcmainlngpinnae, : mm long, pinnules1 2 pai$ on proximal ribbed above, apex brown and 1-2 ( pinnae,2 8 pairs on remainingpinnae, narrowly oblong to narrowly obovate, (4)6 15 x 1-4 mm, slightly recurvedto revolute,light green 10 yellow-green, ! densely puberulous, nervelessor obscurely l-rerved above, l-nerved below. l Glantl sitttated on upper surface of racl'tis, on 2-jugatc leaves the gland is in- serted belorv the distal pair of pinnae, it is subsequentlypresent or absent I if pinnae 3-4 pairs; a small gland occurs belorv insertion of uppermost I 3 I pairs of pinnules on somc or all pinnae rachides; sessile,circular, 0 5 mm I s diam. or less,lip and orifice prominent. Inflorescencessim ple ; 2er/airclcs soli- Lary,20 25 mm lorg, puberulousto glabrescent,base ebracteateat atrthesis; I S

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FrulexO 5 1 m altlLs;,'dml pilosi, raro ghbri; spinaea\illares nullae i'.d1rdbipinnata; /'rdctir 1:5 4 (8) n1mI o11g\pinnae (1) 2\SlJigte; pinnulaeptnnatu nr pro\rmaliun12 3-jugae, oin"uiu"l a;ttitiu"t 2 g ioijugae,' recu.rodad rerolutae,glabrae rel sparsim (raro dense) i"iior.i o.,u".utu."a nlsii'altie. G/darsessilis, jn paginastrperr rhschidrs rnfra insertionem luteola, 'iii':iio;nnaium posita, raro etliam in pettole C,llilrla obloidea ad bretiler c)lindrica, i-^irat iuttt et co,?1/4manifeste 5-neirosa. Iegamrrrodense breviter pilosa Sent)ra in legumineobliqua, brunnea. Tlpe- Porongurup Range, Western Aust|alia, 3 Mrrch 1922' C. A Ga net 1287 (holo: PERTH; iso: CANB). ShrubO'5 I m tall, eithersinglc-stetnmcd or dir'idirrglrear ground lcvel prorninently)rlerved, nerves into 2 3 erect branches; branchesfinely (rf,rely I norrnally yellow, indumentum normally a mjxture-of spa$e to.dense short a pilose and puberulous to antrorsely puberuloushairs, rarely glabrous,green io brown;'axillary spines absent. S/,p?.i1"rnarrowly oblong to narrowly triangular, l 5 3 x 0'2 0 5 mm. Z"dre.! bipinnate' indulllentLrm as on t (8) branihcs iexcept for pinnules); petiole | 2 mm long; rachis l 5 4 mm to lons. often obscurely channellcd above, te nifial .tcl4 narrowly oblong I elliSticor narrorrlyelliptic, I 2 (ll \ 0 5 0 7 mm. \lrrrighlor \ometime5 (: thickened, l-nervcd stightly (rarely prorninentlyt reflexed,flat but somewhat I below'(nervethickened): pinnae (1.)2 (1.)pairs: pinna tathis,2 3.(5) mm long on pror.imllpinnile, 3 61o1mm longoll di'trl pillnae.aper clliptic lo nilrro\^ly ellipticor occrrsionallyoblong I 2 {J} \ 0 5-0 7 mm \lraightlo \rrme\hal reflixed flat but slighily thickined l-ner\cd below (nerre thickened);phnalr's 2 3 pairs on proximal pinnae,2 3 (6) pairs on distal pinnae,narrowly oblong or sometimes-rarrowlyobovate, 2-6 x 0 5 I mm, Lecurvedto revolutc,slightly thickened, dark greei above, light green below, glabrous to sparsely(rarcly densely)antrorsely puberulous to hispidulotts,obliquely narro\\'ed at apex. Glqfid's:itaatedon'upper surfaceof rachis below insertionof distal ard medial pinnae,rarely an additional gland situatedon petiole bclow proximal pinnae; iessile, circuiar to oblong, 0 2 0 4 mm dianr., lip not prominent (hardly protruding above level ofrachis), orifice normally shallow and-not very dis- iinct. Lrfctrescencessimple; pedunclesnornally solilary, l0 20 nl:n long, moderateiyto derrselyshort piloseto hispidulous,rarely glabrous' base normally ebracteateat anthesis;flotier heads\'ary pale yellorv (f sulphur-coloured), obloid to sholtly cylindricalat anlhesis,somelimes , globular prior 1oanthesis, 6 12 x 5 9 mn, with 16 22 (26 30) flowers; rcceptaclel 5-9 mm long (can vary from l 5 5 rnm on a single specimerr). Bracteoles1-1 5 rnm long, spalsely puberulot-ts:ciaws narrow; laminae narrowly ovate,-somewhat ln- flixed.'concave. FLrncr"s5-merous, rrormally minutely pedicellate;cal,,-x I i2 Ienstli of corolla. divided lbr I /3 I /2 its lengthinto triangular ciliolate lobes' prJnrinently 5-nervcd,tube giabrous to nloderatelyshortly pilose;patals 2-3 mrr long, connale for l/'2 their lengtll, promincntly l-nerved (neryes most conspicu;us at apex), glabrous, sparselyhispidulous, or moderately shortly pilosi: o|o|t' nornrrll\ deuselyvillorLs. Legrirrtessomclvhat brittle' narro$'ly bUtong.UO SO x o 8 nrrn. flat. raisedover iecds, denselyshortly piJose,dark broui to grey-brown; ncrryins :lrot contracted bctween seeds, thickened Sccrlsoblique irr legume,clliptic, 3 x 2-2 . 5 n.rm,dark brown, shiny;21t'lrogranr conlinuous or witd a narrow opening towards the hilurr, often bordered by a band of light coloured tissrte;qreole 2 x 1 mn; /inicle fi1iform,reflexed below a normally once-foldedthickened cream-coloured aril.

Ditrrihttri,'rtan,l lnbi!at; \\'lap 8) South-west Wesiern Australia: southetn rcgionsfrL'nr Dettnrrk to Chclne Beach (100 klr1 due northeastof Albany) l'avourssandy or loamy soiL and north to the Stirling Range. Tl.risspecies F] in low-lying s\!ampy areas. IA (p WESTERN AUSTRALIA: Mount tsarkef,W. E. Bla.kall s.n., Sept. 1899;About 8 km G south of Mount Baiker. -Fll.Eichter 16005',Hay River, west of Albany, C A. Gatttncrs.n; Red Cum Pass,Stirling Range,A. S. Gcorge6l14; Porongurup,Mts Knight sn. IMEL A 454 99t '(ed&aq!) '. .A .g L\ZI.D ptnt v j uro4l L6SZultsow LuorJJ-O:ll9 )tuoat S f utorl ):allsDw V g pve 16Z)lDputJ O /,tr ruorJD puE g :6tzt ar!(otr O U.uro4Hpu?v 'sauoaso.rouul .lr,holl-H 'eurnie'l I Lurl{{s qJuetq Jo uoruod 5 J (u,rorq poas (rJEJ.rnsrr{ol) uorlpDe^alnourd-C o1 g )rrp) a '(t) .ld) 1r[1 saprqrere€uuro Jo uortlesLrrJo uoqrsodl)ur BlJslr,:urrurrl rlorled trorls,(r) srrlc?r uo petentrs (a) puBIS 8ur^\oqs (erp.llns .reddn)stlp J?el V o1oa1t1npnty, 97 atiBrg

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Lj wt) 'p0 z 111 ,{t^ ,{tr. :lso €-i zi -ul s 'nL o) IIB 'sll '(p & ,i -st #ffi ,tfi - lPl EUoa-- .X: ^t: ,{ll ,..413u --_EEE_ :l"l {lt oq1 pue l"cupull,{J ,{luoqs o} prolqo rtlor.} e8urr uec speeq ro,rou orll .lpurl tu8ld elaurs " uO qlu?g Daptoto J"A ltpuowwntp .1. pu? ,.usretrAlsu sauo) Pa 're^ 'v ser olnsqo sE paqucsJp ^lsnor^Jtd erel ar11uorleue,t JC V go e8uzr o1 str urrllr,r sapnlcurpu? serJ3dscrqdroruf tcq \\Jutosr st opatry \JDV lod UI ,sercunped LIC er"er.€rqe1,0,,.u,"^oo liflotiffi:"1\1.,Sli: ;ir;,LiffiiHT#l ,{I, -u,rotLl V_uoJJpoqsrn8urlsrp f,ltp?al sr oloalryn)Dh, .gZV d des-Dpafil ,y pu" u3 {oJro^loJtrrlsuasl nuotu,tlu.tq r. :?x":} o,rl Jo aJnlxrur" Jo s}slsuocpu? (^rolsrH l€int"N) runosnJI qs]ttJgeql le pesnorlsr serJadssrql ed,(1elqrssod ]l( .Z .Jg.U Jo SE oqJ lpuel\,\ H Duotulo.ttl Jo Lu^uors"q oqt sr DtotlD D?D)V la, pu" reqolco uee,\\leqslq"ru pulr.leqluetdeso1 lsn8ny ",.rJ ,""ddi::ffffi$ :.requaldes ot dJenlqed etel ruo{ src,\\oll ;potlad Euutn.( puo BurnuoyJ j.'n!li N srllrrr rr^,uEur) orlul ,,,,p'a:{'ffirY ro b JoN 1 {.r::;;iffJ'!f!fr;f.v .dn)urpN K uo JJo-urnl.LlrprSruirqJ Jo gN sslru t :6€zt DIov Q :(lu€ut^ snorqPlb-A\sNflqu]H 'gv) .l.-nlslp .IaW) .u.s tur 'punos blt laztltd E leuaSeluEldlsea_qlroN :(2996i ll rxDJN e8tooC 8ury:(HfUad.)) ultsDry y b)peglonoi -ueeuo4rxl LS1Z 6r apre,{\ol1r€ru tZ IL69Z utlsotlt y €',dn3unLuBrrrfsprr^\ot s)EedFuBJ lJo J ur) Z€:(ZtL6t :r a

€ oinnules from recurved1o prominently revolute. Although this speciesnor- t ir.rallvhas hairy bralches, leaves,peduncles, etc., a glabrousvarialt has been t recoided frotn'the Albany and Stirling Range areas. This varrant rs more f iobust than the typical foim, its branchesare nore prominently ribbed (when c pinnules' dry), and a sessile;land sonetimesocclrrs at baseof the distal pair of I I luteolqis closelyrelated to A. raria B. R. Maslin; it also llas sorue Acqciq c y'. druntmctniliiLindl. From the latter speciesit is distinguishcd afflnitieswitlr ( by the nature of its branch indumentum,its recurvedto revolute'often puberu- pinnules,its short, \'ery pale yellow 1'lowerheads, its prorrinently 5-ncrved lous I ca1yx,and its dark brorvn seeds. c Thc two charactersrvhich can be employcd to fnirly reliably separate c A. ltrteola',t-i,dA. vaia are gland positiol rtrtl flotter head morphology ln A, luteola a gland is always presenLbelow lhe inserrion of the distal pair of I pinnae (i.e. Jn the rachisj. Only very rarely is ar additional gland present 1l telow the proximal pinnac (i e. orr the peliole). Acacia..v4llaon the other b hand ahvayi has a gland on its petiole; only in var. pattiflota is an additional d Eland often orcsenton the rachis. Acqcia lutecla has vcry pale yellow (ruore ir less sulpirur-coloured),obloid to shortly cylindrical llorver heads The f( tvoicai varietvof l. r'allais the only rrernber of thrt speciesu ilh headsapproach- t( ii-! this cotoirr whilc var. par foia is the only member.rrhichoften has obloid sl flo"werlreacls. Other charactersLlscfttl in distinguishingA. lttteola from A' ralla inclucieits somewhattallqr habit, its normally straight,ascending, sparsely divided branches,its often uarrower and less hairy pinnules,its oiten more K oromincntlv rerved calvx alld corolla, and its slightly broaderlegumes '1'dr-"ia li iuteolabegins its flouciing seasonat leastthree months beforeI t"rlia. The specificepithet refersto the very pale yel1owllorver headsI'uhich are typical for this taxon. I

2t 19. Acacia yada B. R. Maslin sp. nov. t Frutex 25 6O (100) clrl ahus: raml pilosi; spinaeaxillares nullae fi)//a bipinnata; pintae (1) 2l3\-iuglae-pinnulae pinnarunr pro\rrrcliunr-2 lJUgrt, pinnrrum distalium2 5 3e plr"""',i re.blutae.pi'.r.,c. rlfo G/.r" 'e"rli'. ir pats-nu'upcr: leriolr i;i:i"q"". Eabrle 'rale' I intruin."riio.. " pinnaftrr pro\ir' rlrult po':t,.glan' a l(rd in ra' rr| ' infraf nnt' d ,,.r.r.oo,rilural. P"/,/,,'r4; ncrurn\ltrepilo'i. rd ba'in 'Ll'J'rthe'r ebrlcer'i Cdfi'l" zufa. 5-rnerae;rall'r'r.r /o,l lfiangulares; p?tdld enervia ad .tiinaii*. int.tau'" obLoLde.i. (\3t n'r:nifc.rel-rrcrro.a. l,ttt,,ti'to l5 J) \ I : Inr'. r:r'icre'rrrr' \' ni,1,t 'ta, irelvP i*"i,,roi in t.gun',;n" tran.ilcrsa ad cbliqua, 2 3 x l 5-2 rlrm, bruinca ad nigra (var. a//iirlr palliclebrunnea). ll

I1,r?. 5.6 km frolr1Denmark towards Mount Barker,WesLerll Australia 2l Sept. 1972, I;. R. lLIasti2919 (holo: PBRTH; iso: CANB, K, MEL, NY' PERTH). di (n Smali s/rr-afi 25-60 (100) cm tall, branches very rarely prostrate (var' pLu|i:1uru),sometltncb suckering; branthas obscurely nerved, indumentum te r'arJous;arillirry rpitteslbseLtl. Lealcs bipinnate,indumentum as on branclles SC {erccDt tbr pinnr.tles): pLtiol€ 1 4 (8) mm long; rachi.s l-1 (5 l0) nm )ong, pr terete ol broadir ribbed rbove: lernindl seta vartorrs pinnae (l) 2 (3) pairs; OI j rinna r.tthis I b (7-9) tnm loug on proxinal pinnae' l 5-10 (12 15) mm pi iong on drstrl pilnuc, rpex r arioits ; 2rnrtrlcs 2-3 pairs on proximai pinnae, ol (5 2-i(7) trirs on clistaI pinnae. normally narrowly obJong,2 4 5 t3) x th 0.5 i 5i3 5) mm, flat 10 revolute, often thickened, normally dark green and cc nerycless above, subglaucous and l-rerved below (nerve often prominently th tlrickened), hairy, rarely glabrous, often obliquely narrowed at apex. Gland sit'r' Gt ated on upper suface of petiole belorv inssrlion of proximal pinnae, sometimes of an atlditional gland occuri or rachis belorv distal pinntLe(var. panifl.)/a); s€ssile, OI circular to oblorrg, prominent to obscure lnfloresceuces simpl1; peduncle[ oc norma)ly solitary, 7 25 rTrmlong, sparsely to densely hairy, rarely glabrous, base 456 L9n eled ,{re,rspoaq Jatrolf:srr€Ll asolrd 3o re,{e1leuorlrpp" u€ qlr,r ,{lLpuorsr?JJo SA, 'snolnprdslq o1 snolnreqnd flesuap '3uo1 utur g sapunpad ',\\oll€qs ecgrJo 'luauluord ' tl 'e"uurd tou fllBuuou dil ruerptuor S.0-E. 0 :s[lc"l ruo4 luesqe Jo SO .rrud lsoru-re,no1jo uorlJesur ,{\olaq elotled go ec"JJns.roddn uo pel"nlrs prr1, -n 'el€lncrde,{l3lnurur'xed" '(peue{cql le pexeuurueUo pu€ ps,rolJeu ,,(lonbrlqo {l: dltueululoJd 01 ,{lel?repoul e,lreu) ,no1eq pe^reu-I 'e^oq€ ssJle^leu '(es.rBoJ '(,&p 'peuelcrql p( l"q,{\euos srBq) snoF.reqnd,{lesro.r}u"+ ueq,r) ueelS e^rTo X 'pe^rncel 'ruur 'al?^oqo ,{lpJ€q ot tBU x ,{I,$otlsu dltq8qs o1 8uo1qo 'e- 'e?uurd Z I 9-€ 'e?uurd ,{l,roJJeu 1r1stpuo srrzd (g) t Z leutxord uo sted g 7 salnuud UI :pexeueJlBq,reluos Jo lr{8rH}s pue 3uo1 tuur Z I l€ln8u?ul seuuouroslo :S. Jrldrllo 01 3uo1qoxede 'e€uurd uo 3uo1 ruru (6), '3 T?lsrp '1eg t'e"uuld'poxeuat l"rurxold uo Suol ruLlr (g) stLpD.tDuultl :peuel.]]I.lt ,{I1q8r1s }"q,raruos '3uo1 l-Z 'Jelngu?u:l s3 :o 1q8rr.rtrs urru 7 I ^l,\\ot.leu ol Suolqo KI \otJgu oDSlDulwot TU :8uol ruu (9 S)t-t .qr1,rrr:8uolulwZ I ap\ad slrEqosolrd uorls'os.r"ds qlr,u 'll pex[u fll"uuou pue snolnraqnd,{1esro.r1ueot snoln]aqnd'(qsr,ro1a,{se,lreu) pa^rou ,{lJug u"lJo saqJuD,tq:serlcu?rq }ca.le olur punoJ8l€ Surpr^rp 'lu3saJd '[?tt Z Ia^eT '7t Jo perljuels-elSurs .reqlre sJalf,ns ruc 0t 0E qft]LIS JJEMA lZ 3!I Elre|'rE^'E6l

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yellow ( 1 cream-coloured),cylindrical, norrrally l0 20 x 4 5 mm at anthesis, witlr (20) 26 35 loosely arranged flowers. Bructeoles1 2 mm lorg, shortly pilose, normally dark brown; claws linear; litminae narrowly elliptic,. slightly inflexed. Ca1l,,r and cctroll.t spanely (sometimes densely) shortly pilose to puberulor.rs(hairs somewhat coarsc), 5-nerved(nerves thickened, sometimes W B WW

, 5 rnm

. 50mm

J w;:, ? ( ,l '1 Lj:* ,7 e{g-' F r I 2 mm r ,t a M s 4 wtsl S | I 5i ,4a+/n tqr+ c a Figve 21 Acacia raria \ar. ratia. A Mature plant showingerect habit and subterranean T runners. B to E Pinnule variation (C- .rpex infiexed). F Leaf axis (upper surf-ace) c showinggland (g) situatedon petiole(p), r-achis(r), tenninal seta(t), and position of insertion 1l of pinnae rachides (pr). G Inflorescence. H Flower. I-Bracteole. A,C-D.FIfrom-8. R. Maslh 2949(the type); B from R. D. Rotce2719; E lron] B. R. Maslin h 2868. r 458 I 6sy 'snolnraqnd 'Ello.toc ,'(lJJnJSqo {LJSuJpol ,{lal"teporu Jo W?'J?I tl Z Zl I -(tr/rJ snolnJJqLrdflosuep '3uo1 urur g.I sdpano.q sre,rog pelced I 'l?cupurl,{c'tto11a[, ,{lesu:p.raqtu E9 qll.{\'srsaq}u"}s ruru spnatl 9t '8uol 9 SXtl , "rr,torf:snolnJeqnd ftosuep !uLu sapunp)J luoururoJda}lnb drT '3uo1 'Suolqo ll-6 'o"uurd uo ruur 9.0 t . 0 : stqcBJruo{ lrresqe 3o rrtd lturrxo.rd;o uol esul\{oleq elorled Jo ece,3:ns:rddn uo palBrlts puolg el"Fcrde ,{letnurur ue uJ:Uo 'xedB pa,rorreu ^lenbrlqo ',\\olaq pe^Jeu-l .{[tuoutnioJd'a^oqe 'ssel3^Jeu'snolnJeqnd,{lesuop'snojolocsrpl" F Jo snololocuoc'poua)i3rrll'pe^Jnc 'olE^oqo -er,{pq8rlsot }"U'mlu x .{11q8qso1 Suolqo'eEuurd 'Jpuurd9.2 9.1 9.t-€ letsrp uo sred S-€ l"rurxojd uo s. gd t-Z salnuuld:poxeuer terl,retuos ol lq8lPllspuP Suol urLU g.q 8uo1qoxede'eeuurd uo 3uo1ruu g 'outtutd 1 1e1srp'pexogel 6 lururrold uo Suol ruLu slrpDt rra?.1:peualJtit

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v P 3. f B I E I h r r4flil' s d r( pinnuie Ftgrve 2s"-Acacia rar[a \at. crcssinerris. A Por'tion of branch system. B 10 D A variation (B and C lower surfaceshowirlg thick midrib; D upper surface). E-Flowef. F-Bracteole. d A, E-F from B. R. Masli 771 (the type); B from M. D. Titdale 362 and B. R. Masli : C-D n from R. /J. Kuchel 1887. 460 t9t 'ex"l o,\\1esarJl uee,\\]3q JnJJo,{euI sel€IpeuJeluIlEclSoloqdloLu lerll 1s333ns ol ecuepl\e eluos sl elaqJ npaowultltp clsqts llpaowwntp 'f 'Ja^3^\oq 'Y qlr^\ trllu+I" ut lseSSns slelJ€Jeqc ecLloJserouuls1I :Dl.toit elr ur uorinlrui s1i selr:crpur,{?olorldJoru J"ol lereua8 s11 uox"] slql uo peJlnbsl sI JarllJru ,{lpelqnopun :ut?Jlscun l"q,'!\oLLlossl xelduoc !!puoututtl,tp 'b'-ot.tDt{Jo,\\ k, eq] urqlr,r slul[D JE,. Jo uor]rsod cturouox€l laaJloc ?qJ 'reqluecec-plur Aq PeJsIqspe^€rl sor.un8allsotu iroqrueldag ol ,{lnl ruo4 sre,tolg ;por-iad3u4n4'ptto 3uuato17 '08 'tr.roN ^\aNreeN :(cv) g tualA'/t1 j/ leoqcnhl 96t lazt!tJ l.ulslc uB^1S:(ISIIv,$n) -loorqsllng 8S618nV'us uot.tduDd9 1 :ZZZ.ullsow V g 'tdrreillqJsp:enor' ised ruorjru1 / rnoqv:(Hfxfd',\N',{\SN'rus) 8lzt ulltDJIYg'eeqcnj^JJogNAenpLUII]noqv:6823uD'IJU"{e^\q:irHuolplEreC-qued '3ed ' alrrll 19 :9961 ldes u s .!aup.tD),/ J'uoopulg rEeN IVI-IYXJSIIV Nd3ISA,^d 'se8ro,lpeor s€ qcns suel?p3qJnlslp uo lloululoJ 'PueJpoo,{\ ellnb sr 1l (oopuElN salulleulos)My{-qErl€f uI (eluelBl Je^o 't, pu€s Jo) iros llluel€l uo s,ro.r8 JI ,r.rt,r selcadseql Jo .I3qureulpatrnqlJlslP ,{lJ3quou lsolu oqt sr ,&eu"^ srqJ ErJroN ,\\3N ol €3qcn}\ t€3u luorJ rllJed JO qUOu :"rlBllsnv ule:ls3/^^lse^\-qlnos (6 d?t{) :lDltqD\ Pao uotlnql4slo 'IIle peploJ-ecuolo 'pouelrrrll " ,\\oloq pexaueJ'uuoJrlg aplunf '.U]nl\q aqt spl",tol uado lq?rerls 'Lllru LuDrSotDld:pees lser urql ral.rzp ,{llq8lls a1oJr, :u^\oJq lq8rT 9. I x Z "c 'Suolqo Jo tulru (uoes^\oJ) spars snolnlaqndfleltrepou o1 flesrtds 9-? x 9t 9t saurta1 snolnreqnd ,(lasr"ds ,{llBuors?JJo ro snolqelS 'pe^lou ,{leJncsqo '3uol lutu g.l'ea sl7tad:snolnreqnd^lasl?ds ol snoJqel8'sselo^leur r..{/rJ '8uol 'poxeuur :lErl,\3ruos'31"^o s"urueT :.Ieeull s,$BlJ:snolnJoqnd ,{leslsds 'sre,\\o!: 'slserllut rrrur saloaDo.tg pe8u"lJs IIJsool . (0r) 0S-0E qtr.{^ l-l.0 'lEcr.rpurl^J',rolle,( l" urru g "c x S€ gI lq8uq spoaqJa$oL/ :snolnJeqnd ol esolrd,{11:oqs^Jl"uors€cto'eso8uls,{lsslollel o1 snolnreqnd,{TeSlo.llol ^lesuep 'ecBJJns dllBrrorsecco.ro ,{lesJeds'8uol ruur sapunp)d ^\oll€tls sculro 'ururp9Z-OI 'sl elorlede,loqe pesrur,{1preq dr1 tuur t.0 2.0'lueutruoJdlou seull}oluos : eeuurd rred uoruesur,roleq olorled ec"J.rnslsddn uo PelEnlIS ea Jo luturxo:d;o '(peuelcrrnJo puDlC xedB le pe,roJJ"u ,{yenbrlqo+ tolr 3^leu) ,^ol3q Pe^Jeu-I '0,\oqE 'snolq"l3 ,(1e:ncsqo i: pu" snocne[8qns ssele,\.roupu€ ueeJ8 IJ?p ^l -{lar€r 'osolrd ,{11roqssorurlautos'snolnreqnd flesloJlue ,{lasuepo1 ,{losreds 'paua{f,rqt 'e1n1o,r.ar .{1tq?r1s ol pe,\Irosr'ruLu (9.t) E.l-9.0 x (El) 0l € (Z) ul 'e1r,ro 'e€uuld 'etuutd -l Xl,rorreu o1 3uo1qo^I,\\orr"u uo s.ttud(9-7) g 7 'srrlJ".r le1srp 'elqell"^ Ieurxojd uo sIEd (€) Z €uurd 3uo1e peceds [3,\\ .lr1nrrld :peue{crqt ,,(pq8rlspuv]"U pexeu3rtprl,{\eruos o:} tqSruJlsSuol urtu E S.I 'epuurd 'eBuuld cqdrlle o1 Suolqo + xods 1u1stpuo 3uo1uur (11 71) 0t-q (E) uo SuoT ruul (y1 g snpo.t ,rrrd :paualcltlt '(ltt[8lTs '1"U 'poxeueJ el IErurxord '8uol 7 'crtdrlle lErl,ralrjos ,(lleuuou ulru 01 ale,\o ro Suolqo Dps lDutLatat qr € I '8uol '.l :e^oq" pelleuu"rlcpur? p"orq ecp,JJnsJeddn uo qtJ lutu (9-S) E Z slt'pD.t 'qsruaatB :8uol ruLu tI alo!|ad stalqtuotg :es€q eqt spr€^\ol ,{313 :}Eq,{\ -euros u,roJq o:l qsrppa: '(as:o:1erJo 'esJoJlut 'lqgreJlssJIErl) snolnJoqnd lnq 'PeAJau ro/pu? Jsolrd (e1qrr:e,r)ulnluerunpur ',roll3f ,{ll€uuou se^JJu ,{l3ug saLpalitq'. seLlcLreJq repuels ,{ueur olur puno.r8Jzau Sutpr,trp'luasq" sle{Jns '(8uol le^el ruc 001 ol seqou"Jqqtr,\\ esnlip seullauos) IIEI ruc 09 ol qnrqs IJaJZ t: 'v s 'zL6r'Fnv 0z 'e,rerrsnv urersea ',{E^\qslJ;1}tY,il"$";BtdS""ti#2L'f"iT'|:XA u( et 'eeuunrq eplllEd'ESuolqo (€sI^ elned) Dullu)S €lnreqnd e.Ipolu p€ Ur 'ele,\Jau esJgds oulunEaT erqElS onbunrald ejnJsqo DlDpd :srltiJe\ra l: )./f/rJ ellsod JT tunrleLLIxoJd LunJEuurd urouorlJasul ?{ur rloDed "radns €ul8ed ul srrl, aelnleqnd esloJlo? 'eElnlo^er 'ae8nl(g 'oe8n anbtunjeld pr? ep^rncgr -t) tunrlelsrp run]euuld f_(€) Z Lun 'selrqgue^ [ Z ',e?uol -riELurxoJdu.rnJeuurd aDlnuuld :€rrtclllla pe ele^o IoA BEuolqo Jrratu,!al Dps 'u ruru (9-S) € Z r{rrr18. rlnr.qnd le^/te rsolld ?r.rr.r :sntF tu. 09 pE enburnlsld ,.rlrg ,{l 9Z ;U ^ou Je,\ulls"tl{ u g sJusB JB^'J6I a

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FtEurc26 Aruda ratio v$. afflnis. A Portion of br'Anchsystem. B Node showing stipules(st), peduncle (pd), and leaf (glandposition arfowed) witlr Jlatrenedpinna rachii apices(a). C Bracteole.D Flower. E- Seed.F to H Pinnulevariatio;i aF uDDer \uf'aie;Grnd H losercurhccl. A-D, F C fronrB. R. Mdrrir 2793(lhe type); E ftom B. R. Mdslin3222:H fromR. T.Lanse 89.

This variety is quite variable, especiallywith respectto the nature of its branch indumentunrand its pinnule size. It is distinguishedfrom the other tlrree-varietiesof A. wri(.tby the fbllorvingcharacters: its pitrnules,peduncles, and llower heads_are normally somerhat longer, (however,there is an overlap of paraneters whcn all the varietiesare corrsideredsimultaneously), its pin- nules are somewhat iess promirreltly nerved below, its pedunclei normally bear rctrorsc hairs (spreadingor antrorse in the other varieties),the pinla rachis apex is usuallymore prominent,and its petalsare obscurelynerved and glabrousto glabrescent(in thc other varietiestlte petalseither have a f thick- ened central nerve and/or are sigrrificantlyhairy). The varietal epithet refers to postulatedrelationsltip betweel this taxon lnJ 1. rlnunntou,lii\t6sp.,l uDtntdii.

19d.var. parviflora(Benth.) B. R. Maslin comb. nov. ,/:8.27

Acacia chummorlii Lindl- var. parrifarc BenIh.,.F-lora_Au5tritl. 21419 \t8tj4). Lectot),pe: \\c,r \4o,rnrts.irren and on llre hill, lo ea\tsaro. Va.rrell :.n. tMI L 4qlbl speiinren.ien by Bentham;PERTH fuagment)

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Disttib ion a d llabitqt: (Map 9) South-tvest Westcrn Australia: most common in the area from near Manypeaks east to tlte Esperancedistrjct; I yar. pqr fioru aiso occurs north-west of Manypeaks to near Afihur River. 1 This variety norrrally grorvsin sand or gravelly sand. I ! WESTERNAUSTRALIA: Point Ann, 20 mi NE of BremerBay, K. M. AIa 345(CANB, : K, NY, PERTH,villous variant); 8 mile^sN_of Esper-ance, ,. M. Ch rchill,l: Flopetoun, ( G. Douglass.n.. Jul]' 1915 (NSW 65957):SoLrrh Sfi fling sandptc irr, ,.1. S. Crurye6239 i Lucky J tsey,A. S George'7166(glabrous varianr). \\'e\t Mounr Bcrren, I .t. 6.diI,. t0031;-Cibso; So^k.Lullfitz 3596].I 6 nri S of Jerraftunguprowards Albany, B. R.Maslrir 991; Near Cair dner River,B. R. Mdyr?1052 (K, PERTH);0 5 n]i\\'est ol YolngRiver cros:rng, Albany-Esper_ anceroad, r. R. Moslin2553:. S.W. Austrclia, Md\rL.// s.n. (MEL 4q157); t37 ni'peg on I AlbanyHighway, K. N(ribe'"36B2; Ar|hur Rt\et, Ol(tiLlJ555 IMEL, ro-synrypeoT,.{. ttruntnnndii-Lincll. vat. patrillora _Bet1th.); About 20 mi N E of Albanyrowards rer ranungup, S. Par./rl533; Ravensthofpe, Ralphe and StanlAd s.n..26 Jlrne 192,1; Cranbrook, Sroiairl l 38: NearMount Le Crand,,,1. S. I4/eston6114. 1 Flovaring an! ft'uiting periot:l: Flowers from May to Septentber; ntatlrre iegLrmes are presgntfrom October to Dccember. Atacio raria var. parvdlora is a viiriable taxon. Thc leaves on the : young plants are gcnerallyconsiderably larger than those on mature bushes: theselarger leaf dimensiorrsare given in parenthesisin the descriptionabove. Altlrough var. parvifora normally has hairy branches,a glabrousvariant has been recorded fronr Lucky Bay, which is 30 km south-castof Esoerancc. Another variant, knorvn only fron Point Anl (60 km south-westof Hopetounl is prostrateand its branchesarc much divided arrd denselyvillcus. TIie habii of-the typical form is norrnally erect(clthough in regro\\tlt the pLxntsmay be diflusewith nore or lessprostrate, spars-'ly dir ided brtnches)lnd the branihes arc puberulousto shortly pilosc. This varicty appearsto be most closelyrelated to var. crasslr1crlrsB. R. Maslin (seethis taxon for details). Acqcia rarict var. parviflora is the only irrfraspecific taxon in wi.iich a gland often occurs belou' the distal as well as the proximal pair of pinnac. This fact, together with thc prominently recurved to revolute pinniries and often obloid florverheads, could lead to conlusion with A. luteolail. R. Maslin. From tlris species,var. port'lflorois distinguishedby the folto\ ing characters: shrub often smallerand morc diffuse; branchesless obvioLrsly neived; petiole slightly longer; rachis often shorter: pinnules rhicker.,blightl\ broad6r.and norrrally more hairy; flower lrcads distinctly yello\.\( E strlph u r-colou red in A. luteola); calyx (and often corolLa)somewhat less promine|tly nerve

20. Acacia drummondii Lindl., S^ketchVeger. Sran Ri\. Colony (Append. to Edwards'Bot. Reg.)XV (lE39i. L.(t('t)p,'. Sran Rirer. Dninm6ni/ s.n. (holo: CGE photo seeo; ?iso; K photo. secn,MEL 49354) Shrtrb0.3-2 m tall: brq cltcsstrigose ro puberLrlous(hrirs retrorse or antrorse), sometimes glabrous; .rriliary spines !rbsent. "S1?r/csnarrowlv oblongto narrowlytriangular, 1 4^mrn.1ong..Lrarcs bipinrrlie. rndumentunl normally.as-onbranches (except for pinnuler);pttiolc. , ter.-te,0.5 5 mni loog racltis 2 8_( | 0-23) mn long, ribbed lbo r e ; rernt inal st ts narrowiy ttiangLi_ lar to oblong-eliiptic,slightly thickcned;pinnoe | 2 13)prtts: piu a iqchis i g (10-13) rrm long on proximal pirrnac. (3) 4-13 t16 21) nrrn long on distal pinnae,apex triangular or obJong-elliptic;?intul(s (t ) 2 (3 4) pairj on proximal 164

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20a. subsp.drummondii Frg. 28 Aca.iadtanuondi Lindl. var. typita E. Prirzel,Bor. Jb. 35:313 (190,1), nonl. illes. _ Openly branchedshrub 30 60 (100) cm tall, norrnally dividing at ground level into many spreadingbranches; hranches finely ribbea (ribs ofien yiilow), sparselyto.moderately reLforscly strigosc (rarely retrorselypuberulous or glab_ rous). red-bro\\rl (grey Jt brse). Rqchis2 3 (5 9) nm long rvitn a sl[care rib above; temina.l_seta. narrowly triangular to oblong-elliptii, slightly thick_ 9r9.d;.4innge(l) 2 (3) pairs:pinna nthis2 4 5 (6 l3) mm orr pror iiral'pinnae, 4 11(ll l6)_nm on distalpinnae, iLper narrouly triangulurrj oblong_illipticI pairs t)inDu/cr.2.tJ 4t on-lrro\imalpirrrr.re.2 3 i5t prir-.o di\lal finnac, nar_ rorvly oblong,(2) J 6 (8 12) x I 2 12.5) mm. slighttylhickerred, gla6rous (rarely.ciliolate),glaucous, l concoloroLrs.nerveleis tb obscurely l-_nerved, apex obliquely narlou'ed. Glo d.not prontinent, situated on Lrppersuriace of petiolc belorvinsertion of proximal pinnae; 0 .2 0.3 mm diarr. petluncles 10 18 mm.long (25 mm in fruit), sparselyto modcrateiyretrorsely puberulous to retrorselystrigose (rarely glabrous); .florer lteadsl0 25 x 4 6 mm ar anrnesls.

, 3mm ,<.\ -\ -\{rA f't;\ 'r' f21

t(" t.q,r/-pr Iil I 'l "- s l..qtp.._prIf r : o{&-n 2 G e S lt h ( 2 .J ; I s rni$; I s Figllre 28 Acacia dtumno dii srbsp- rlrunn.ndii. A- Node showing slipule (sr), peduncle (pd),and leaf(gland position ar-rowed). B Prnn!le(louer\urfecc shciurng obscuie_midrib). C Pinnule(uppcr surface). I)-Portion of brcnch s_\srcrn.E Florveri F Bracteoles. -n G Leal axislupper -surtace) showing gland (g) situatedor1 petiole (p), rachis(r), tenrinal seta (t)! and base pinnae (pr). I of rachides H Inflorescenceshowing retrorscrl, srugose F Deduncle. A, D-C frorr B. R. Maslin 2985t B-C fron M. Koch 118t; H ftarn R. T. Lange 186- al 466

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Ftewe 29 Acacia druktmondii xtbsp. candolleana. A-Portion of branch system. B Sead (light brown). C Node showing stipules (st), peduncle (pd), and unijugate leaf (alandposition rrrowe,l). A an,tC fronr tl L. phitl;p'u A h8 10ob;B fronr B. R. V,r,/r, 1200.

Flowering and fruiting pel'lod. Flowers from July to October; very few un- dehisced lesumes still present on tbe bushes in late Decenber. The alfinities of this subspecies seem to lia lvitl.t subsp. "1cgors ratller than rvitlr subsp. clrunmonclii; however, subsp. cancblleana is readily distinguished from both of these taxa by its consistcntly ullijugate leaYes.

20c. subsp. elegans B. R. Maslin subsp. nov. ?:lg. 30 Aucia ps?udodrunmorrlllHcrincq, Bull. Soc.Hort. Seine1l :194(1853) 4".i n.v Aca(ia Delbiae C. A. Gafdner. J. Roy. Soc. W.A. 9(l):40 (1923). T:'pe: Gad et Herb. no. 626i (holo: PERTH). hutex 1,2 m altus: rdnrli nranifestenervaii, sparsimantror-se puberuli ad glabri. Rhachi.\J 8 (15 2l) nrm longa; r.r.r tuf,?l/rdlr angustelltang,rlaris: pinnue (1) 2 (3)jugae; DinnulaeDtr,na(um provnralium (l) 2 (3)jugae' pinnarun distalium2 4 (5)jugae' glabrae ,rd glcbrntae. 6/.rr?(\e\\r rs.in paginaslrpela rachidis inlfd insertionempinnarlLnl distaliun, praisensvel rbsensin pdgLrlxsupera petiolum infra insertionempinnarum proximalium- adrl/r/,/ c\ lindricir. P.rald plerufiqlle modicesligos^. Lqunina modicepuberula. S?tnna pallide brLLnnea. TrDe: 35 5 km from Denurarktowards Mounl Barker,western Austlalia, 2l Sept.1972, B.'R. Maslin295i1 (holo: PERTH;iso: CANB, K, NY). Shrub 1 2 m tall, eitller sirgle-stcmned or dividing at grourd level into a number of spreading branches: brunchlets quite prominently ribbed, sparseiy antrorsely puberulous to glabrous, reddish brown to grey. Rac,4rs3 8 (15-23) mnr long, ribbed above; terminul set(t narrowly triangular, slightly thickened towards the base; piruna (l) 2 (3) pain; pinna rctchis 3-ti (10) mn lorg on prorimal pinnae, (l) 5 I3 (16-27) mm long on distal pinnae, apex narrowly iriansul.rr and + thickened to\l'ards the base; pinnttles (l) 2 (3) pair's on proxi- mal pinnae, 2 4 (5; pairs on distal pinnae, Qrarrowly) oblolg to elliptic or obovate. (l) 5 12 (11 17) x (l) 24 (5 6) nlm (size variable), flat or slightly recurved, glabrous to glabrescent (hairs rninute), dark green and netvcless (or vcry obscurely nervcd; above, light green and J(2)-nerved below (nerve - e\centric), aper oblique atrd sometimes rnflexed, Glqnd situated on upper 468

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favours loamy or sandy soil in low-lying, often swampy areas. There is, however,a lorm which grows on the rocky slopesof somepeaks in the Stirling Range (seebelow). WESTERNAUSTRALIA: .12nile peg on Great NorthernHighway, tr. M. Allart 294 (CANB,MO, PERTH, SP) [pfobably introduced]; Tenterden, W. M. Atki 3l; Toolbrunup, 700metres, L. Dicls161'7: Drummond l5 (MEL 49184);Blull Knoll, summit,C. A. Gardner 1450;Near Mondurup,Stirling Range, C. A. Gardner16208; Hali-way up BlulTKnoll, SlirlingRange, B. R. Maslin1102 (MEL, NSW, PERTH); Near top of BlulTKnoll, Stirling Range,B- 11.Maslin ll03 (NSw, PERTH);20 mi eastof Manypeakstowards Jerramungup, B. R. Maslin2598,Perongerup IPorongurup] Ranges, Ma-{}rcl1s.n. (MEL 49373);Mondurup Peak.F. Muellets.n., 15 Oct. 1867(MEL 49377);King George'sSound, s\'ta:nps, Ourteld 554(MEL) r About20 mi NE of Albanytowards Jerramungup, S.Pdrls/ 532; 5 lni N of Narri- knp, -R.,. Rq'ce 3741;Toodydy dislrict, roadside, C. L. ,yilson83'7 [probably introduced]; StirlingRange, I km N of baseof BlLLllKnoll, P. C. Wilso 4193(AD, E, L, MEL, PERTI-I). Flovering ancl fruiting pefiod: Flo\iers from July to October; young legunes appearfronr Septemberto November;mature legumesare presentir December, In the past, most autirorsseem to have errorreouslyregarded subsp. e/egars as the typical form of ,,1.druntntonclii. However, slbsp. drunmonclii as here lectotypilied is quite diflerent from subsp. elegans (see subsp. drunnonclii lbr differeniiatingcharacters). Herincq was aware of thesedifferences when hc describedA. pseudodntmntonr/il, this speciesis here relegatedto synoryny " under subsp. elcgars. Becauseof its awkrvardness,tlre rame pseudodrum- " nrondii rvasnot usedby the preselt alrthor as the subspeciliccpithet for subsp. elesans. Gardner (1923) describedA. pelloiae from material collected rear the sumrrit of Bluff Knoll in the Stirling Rarrge. This name is irere regzrrded as being synonymousr.vith subsp. elegair.r,howevcr, it is slightly different from the typical lbrnr, e.g. stipules sometimesa little nore prominent (not spinescentas in tlre original description),pinnules slightly n-Iorerccurved, and the pcdunclesand flower heads slightJyshorter. This may prove to be a montane lorm of subsp.elegans;rt has been collectedfi'om BLull Knoll and Mondurup Hill, both in the StirlingRarge.

21. Acacia mitchellii Benth., Lond. J. Bot. l:387 (18,12). 7,1,pe.'" Interior of N.S. Wales. A4itchell."(n.v.\ Diffuse slryD 0 5 l 5 (2) rr till; branchesslightly resilorrs, obscurelv to moderately ribbed, normally denselypuberulous; axillary spincs abseni. Stipules + triangular, rninute, ca. 0 5 mm long, prominently thickenedat base. Zcayts bipinnate, indumcntum as on brauches(except for pinnules); petiole 0 5-1 (1.5)mm long; rachis 12 (7 15) mm long, terele or ribbed or clrannelledabove; terninal sela oblong to triangular, I 2 mm long, straight to somewiratreflexed, slightly thickened(cspecially towards the base):pinnae (l) 2 3 (4 5) pairs; pinna raclis 1.5 3 5 (5) mm long on proximrl pinnae, 2 7 mnr Iong on distal pinnae,rjbbed above,apex somewhatrcflexed; pinnules 2 3 (4) pairs on proximal pinnae, 2-4 (5-7) pairs on distal pinnae, normrlly oblong to narrowly oblong or obovate, somelimeselliptic, 2 5 (6) i I 2 mm, flat, glabrous, sometimesslightly thickcned, light green to glaucous, 1 con- colorous, very obscurely 1-nelved, obtuse but often minutely apiculate. Glantl absent (but see discussion below). Inflorascenccssimple but sometimes falsely racenroseat ends of brarrchlctsdue to leaf reduction; pecluncles1 2 per axil, 10-20 nlm long, sparsely(rarely densely)puberulous to glabrous, baseebracteate at anthesis; f/ox,€r headspale yellow, globular,5 8 mm diam. at anthesis,with 30-45 flo\vers. Bra(teoks, ca. 1 mm long; claws linear. glabrous to puberulous,laminae ovate,.ciliolate,often puberulous abrxirlly, slightly concave. F/oriers 5-merous,siightly resinous,sepals 213 length 6I petals,shofily uniled at base, linear-spathulate,ciliolate; petalt ca. 1:5 mm long, fice; orarl glabrous. Legumes firmly chartaceous, narrowly oblong, 470 tLv esolrd,{I1roqs ,,{losuep'8uol u-]ru '[Je'rJIossapunpad :eld]llts s u salob[ul '; 'snoncldsuoJul 0Z t ue13o:es€qslr r"eu €les l"urru.r3lJo eJ"Jlls rJddn uo pelenlls u :se,Le3[ele8nfiun uo:e€uurd uor].rosur(,{oloq ocu€lsrpeuros Jo) :}? srqceJ Jl Jo '^\oleq Jo eJ"Jrns reddn uo pelenlrs :se^uel ale8nftllntrr uo puolC pe^reu-l fllueutruord 'a^oq" ssela^Jau'snolnJeqnd o1 snorqelS 'snocnel8 ol uaor8 'peueltrrll 'ujru 'Suolqo 'I leq,\\eruos'elnlo^ol o] pa^Jnoal I'€c x (8) 9 g.Z illl'oJJE\I safuuld : pexeue.r01 puu peuell?U ,{ltcurtsrp sorurleruosxed" '3uo1 lrlSr"rls uru (01) sr4to.rouutd:s.rr"d (t) g aouuttl:pexauerro tq8rErts '8uol 8-E.1 1 z ruru 'lrrJJs3urdsseurrleulos 'elq"r.r"A ,la,r '.esBqJgl S II-g.l 'pr?u loLtlw)at spJ€,{\o}pet€lrp l€q,\\eruos e nb'3rlol u.ruJOZ g slLpu:(rlcu?.rqpu? slxe uorlcunIlB esu" a"uuid ]rEd e r) aloUad: (sJ|luuId.IoJ -l J"elJo Jo I€rurxord luasq" 'l ldecxa) seqcurrq uo se tunluolrlnpur']uarrncop 'el€u0rdrq saazaT :luecsturds :l: Jo snou€cs selndrls luesqE seurds dlelllxE :snor.r"A Iunluatunpul i 'paqqu 'esonxsB ,'(leuu ,ra{rirr.rg :seqJuBlq lcere o1 Surpee:ds,(uEru olur & '8urJe)Jns 'll1?l le,le1punor8 le Surpr^tp sorurle(uos tl|c 09 0Z grrils ll"rus a '(HfUaa :osr :IASN :oloq) qeJ'plo.!a84u 'uot8tJluun) :adtJ '(Ll6l) 9061 '8ztrl I /u 1l tLZ:19 ,^AS N cos ,{oU f'uepre]Alur A /A euE!^rorpalcErv'ZZ Jl ;1 .^tl!q 1r 'l -ISSodsrql 01 parmbeJsr fpnls uy pue13po)npo: fleurell {Jeqc l"crruolsue I -xa u? sE pelerdralureq plnoc qJrq^\ uorssardoperncsqo,{JeA " srqce.rerJl uo srncco eJerll serurleruos 'Jei.e,uoH (acuerue^.uocJoJ fluo uorsr^er slql ur pepnTcursr \IJtLIn\'Dtllosu t Surpnlcxe) aDlpqJTd eql Jo sJJqlueul leqto uI J( s" slqcul ro elorled eql uo slrlcco pue13peugep ,{l.r"alJou ,{lurulreJ esoJ -npu"l8e aq 01 prEsarE sercadssrtl1 jo sa^"al aqt '€^oqe uorlducsapeql uI 'roqtne (q I uqt p petpn$ )D aqlnd Jo seroadsrerllo eqt Jo leql !!ro{ sJeup r11ar1rtir.a'. ur luetu -dole^apJ€JI '(S961) 'osly E Surlpeesloecuenbes eqt l"ssB^ ol SurproJJe dnora p srql ur eJerl,reslaplrnoJ sr '(ao11aqt1n4€qt ur eJe.r) serun8al snoeJ"u"qc 1ou 1( ,{lulg pu" 'speos 'sesEqalcunped el"elcBlqe uor:l"urqluoc 'uorlrpp" Ieurpnlr8uol 'sJedes Jo ll eql uy sarcedssrql ur ^luo punoJ ol" (pelrun lou) ee{ pu€ p '(^\oteq ees esolnpu?l8J 's.r3lf,u€rlc o,\\f 'prl"rlsrly lnq) sa^€el luuuodulr el urelseiAeprslllo sJncco qcrq,\\ seues Jql ur sJrJJds^lrro aql sJlJ J"alJ lou sr aD aq4nd oq] Jo sleqlueu Jerlto eql oJ lt aq)llLu / Jo drrlsoorl€lalaql '( -uosr:es Suue,nog:lx3uoql Suunp seqsnqoq1 uo luJs3jd ueuo e.ru -I 'ornl?Lu slrn{ s,Jeo,{snor,\ard eql ()1soun8ol roJ eurrl Suol " se{"l 1r esnEcaq [. :rlorer\ pu" Jeqolco (lsn8nv) uea,\\leq sJe,roll :pouad SLt4tntl pttD EurDrtoll u !!, -er€rs e. z/61unoJ ees ueessueu.eos oN eqrJo ls'e-qrnosr.c :vuvdlsnv H;Kot (68ltZ .Igr{) .u.s./1114,.1/ ,"uotrr^ ,ro^rl{alouolC :(tgt\) nag ,ll 'J 'suErdlrr€rD:(1f tso^\_qtnos'ztrlSrels t98rz'sa8ued 4oU)tR ,/ )ZB uolsv / l? Jo gN elrtuI lnoq€ auEqsug 'l :(tSt,'l) tlosrapuv ,{toeHFuolp 'pueprod q ou selrtu :VIUOIJIA .€ 'ntZZl'.{p.'!\q8rH 'seuul Jo /'Ued pup.EtZZ,{ur,1r'J y uolDJo a selrurZt r tEuorleN si eSuedrelle.rqrD :(ll_68E,\\SN) 'us ssa8tng'4'seuul ualD rtu 096t/Zl''us '.seI^\El lo E Zt'.te,rq;rH ( rlp,{^\5 :(01699,tSN) 606I/ZI uDutloag7 / ,tnD :Sg'lV^A HtnOS ,t\AN :I 'spuelpoo,rsadQouE ur ur"ol ^Icol puu,{purs urorj peprocal uJJq oslE sBq tl !sdurulrs rzeu s,roJ8 ^luoururo, 11llaq)Jlu!DDnl dr '€rlEJlsnYqlnos puu '?uolcr^ 'salsiA qlnos ,reN :lDllqDl pao uotlnqulslo 'c 3' .lF€ 'l 'pelroJ elEAEppeJnoloJ-ruEoJc ? olur peuJ{Jrqt^l}dnrq" i tlauo pue uJJoJrlU ':lu3urtrloJd ')tc"lq 'urul ,/r?rrJf:snonuquoc uto.tSotnald:llnp ol u,\\o.rq 1.1"p n x fltq8rlsro ot?^oqoSuolqo 'orunSol ur S.t 9.2 9.9 t'crldrlla 'spees l?urpntl8uol spars p3u3{crql fllq8lls erll u3o,\\laq po:lJ"rtuoc tBq^\euos ro lrlSr€Jls su18.tDw:ui\oJq-AJrB o1 u,rorq'snolqelS'el€lnJrl3r floJnJsqououo 'spJasJo^o 3 't€U 'pe^.rno 'rulu .S p3srer t"rl,reluos T ,{llq8rls ()1trlSr"rts / 9 (t) x 09 0Z (SI) a

to hispidulous,basal bracts minute and normally absent at anthesis;jflon'?r heads globular.6 l0 rum diam. at anthesis,rvith 14 35 flowers. Bracetoles I . 5-2 mm long, shoftly pilose;claws linear; laminaenarrow, straightor slightly inflexed, concave. -Flo11el.i5-merous; calyx 2/3 length of corolla, dividcd for ca. 1 1'2its lelgth into oblong ciliolate lobes which sometimeshave a ferv additionalcollspicuous spreadir.rg white hairs at the apex,prominently 5-ncrved, tube glabrescent;petels 2-3 mm long, prominently 1-nerved,glabrous to puberulous; orary glabrous. legarres somewhat hard and brittle, narrowly oblong, l5 45 x 5 7 mm, flat to slightly undLllate,raised over seeds,moder- ately to denselyshortly pilose,dark brownish; marginsnol coltracted between the seeds,thickened, light to dark brown. Seeds(as to subsp.pungens only) longitudinal in legume, elliptic, 4 x 3 mm, black, dull, minut.'ly rolrghered; pleurogran fine, continuous or open towards the hilum. funicle less than I mm long, expanded into a once-foldedyellowish aril which is sonrewhatdilated at the hilum. In his original descriptionof A. dreviqnq, Fitzgeralddescribed the flowers as 4-merous; however, as can be seen from the descriptionabove, they are 5-melous. Acqcia clreriqna is unique withil tlre serresPttlc]rcllae in that its plimary Ieaf axis is not articulated on the branch. Instead it is decurrent and in tiris respectA. drcviqfiqcarr be likenedto the phyllodinor.rsspecies in the series Continuae(Berthan, 1864). The lowermostpair ofpinnae ariseat thejunction of the plimary leaf axis and tbe branch so thai no petiole is developed. In its hairy branches,recurved to revolutepinnules, and hairy, ebracteate peduncles,A, drewianq supcrficiallyresembles A. praissiona(Meisl.) B. R. Maslin and A. plicoto B. R. Maslin. Ac{tciq dretrianois distinguishedfrom both thesespecies by its habit, flexuose,branches, decurrent leaves, and abserce of petiole. Keyto subspecies la. Pinnaeconsistently I pair; tertuinalseta pungent, *^t.;. .1,!rT:oYAHi?31I, ,,rrai b. Pinnaenrore than I pair (a few unijugateleaves may occur anong ihe nultijugateones); lerminalseta not pungent.... 2 2a. Apexof pinnarachis conspicuously flattened and proninentlyreflexed; pinnules 4 6 mnl long;calyx lobes ciliolate and wiih additionalconspicuous, spreading hairs at theapex. (Near Armadale to Bindoon) a. subsp.dreryiana (Figs. 31A-l) b. Apex of p'nna mchisnot conspicuouslyllattened, - straight;pinnules 2.5+ mm long; calyxlobes ciliolate, without additionallonger hairs. (WonganHills; Kukerin to Lake King) .. b. subsp.minor (Figs. 3lL-O).

22a. subsp. drewiana Flgs. 3ll-1 Shrub 30-60 cm talll. branthes quite stout, puberulous, hispidulous or slrortly pilose. Stipules 152 5 x 0305 mrr, base slightly thickened. Rachis 5-20 mm Iong, rigid, prominently tltickened, ribbed above and below (upper rib chanrrelled above); terminal sela variable, linear to narrowly triangu- lar, 2 ll nrm long, slraight or reflexed, rather rigid but trot pungent; ]rr?r?ae

Flgtre 3l Acatiu drewiana. A. to I subsp. drc*,ianat A Portion of branch systent. B io D Pinnule var'iation(lower surface). E Inllorescence. F-Flower (note conspjcuous hairs on calyx lobes). G to H Bracteoles. I Node showing stipules(s), decurrent prinary leaf axis (la), and flattcnedreflexed pinna rachis apices(a). J to K subsp.prrg?rj: I Portion of branch showing I spinescenlstipules (s), and decurrent spinescentterminal seta (tJ. K Seed. L to O subsp. mr'l?or:L-Flower. M to O Pinnule variation (M and N lower surface; O-upper surface). A from l.t/.M. Cat e ;.n.; B, C, E and Ll frcm H. Steadmens.n.; D and I kovt B. R. Maslin 3217;F and c from C. A. Gartlners.n.; J fron\ Jalia Wellss.n. (the type); K f.om ,,1.S. G"orge 5937r L. N and O from,4. S. Ceotge1447 (the lype); M iiom ,41.,. Tindala226 and B. R.

4',72 ttt/ 1'//a'H / tl ONw ffiili a{h #ig 55c1 ffi #@.-[F ._@6r_

cl

uu 0t W a

(l) 2 3 pairs; pinna rctchis4 8 (10) mm long, ribbed above, apex flattened oblong to elliptic 2 3 mm long reflered rnd ribbed bejou ; piniules 2 4 (5) pairs, 4-6 (8) x ca. I mm, recurvedto revolute,dark greenabove, subglaucous below, glabrous to spa$ely puberulous. Petlunclesl0 20 mm loti: flower heqds8 10 mm diam. at anthesis,with 25-35 flowers. Caly-{ loheiciiioLate and rvilh a fervadditional conspicuous, spreading, white hairs (0. 5 2 mm long) at the apex; petals 2.5 3 mn long, glabrous to glabrescent. Legunes 30 40 x 6-7 mm. Secrlsn.v. Dislt'ibutionand hahitat: (Map 5) South-westWestern Australia: occurring in a somewhatrestricted area fronr Wongong Brook (about 35 km SE of perthJ northward to near Bindoon (about 110km NE of Perth). This subspecies appearsto favour sandy loam, often in low-lying areasnear swamps. ry9SlEBN A-USTRALIA:Wongong Bfook nearArmldale, W. M. Can! s.n., 30 May f924(NSW fragment,Perth); Muchea, C A- Gurclners.n., April 1928;Canningion,,.{.7. C/aJ,sn, 1960About 0.5 km E of MucheatlLrn-off on CreetNorthern Highway, B. n. Maslin321'7 (BRl, CANB, MEL, PERTH);About 3.4 km E of Muchelrurn-off on Creal NorthernHigh*ay, B. R. Maslin32208 (8, K, NY, PERTH);Muche1, A. Slceunus.n., A,pd .1928(MEI_49335, PERTH); Dariing Range behind ccsnells, l,. Steznotnms.n., l0 Feb. 1960(NSW fragnrent,PERTH). Florering period: April-J uly. . At atia ,tlrewituta sobsp. drawiann is rlore closcly related to subsp. ninor than to subsp. pungens see these taxa for differentiating characters.

22b.subsp. minor B. R. Maslinsubsp. nov. Fig.3l L-O t-rll.,r nanus,2045 (60)cm altLrs;rdmi plerumque dense puberuli. Rrd.rir 5 1l (t5) mn1 lo11g ; pinue 2 3 (4)-jrqae:' lhachi! pinrarr,? 3 8 mln longa; pinnulae 2 54 nn tcngrc, nranileste revolutae, plerumque n1odicepuberulae. T),p(: Lake Camn] turn-olT, Newdegate-La ke_K ing,road, We.tern AurtralLa, 25 May 196j, A. S. Gcotgr 414- (hnl,r;PfRTH; i.o. CA\8, \5W. PI.RTH,. Shrub 2045 (60) cm tall, sometimessuckering; brarcles slender,densely puberulous, glabrescent, rarely light _brown. Rachis 5 l1 (15) rnm long, rather rigid, not prominently ribbed above or below; terminql set.t nafiowly triangular, 1.5 2 mm long, slightly thickered towards the base, oftel dari< brown; pinnae2 3 (4) pairs; pinna rach.is3-8 mm Iong, apex narrowly oblong to narrowly triangular ca. I mm long slightly thickenedoften dark brown ani straight to slightly rellexed;linnulr 2.5-1 mm Jong, prominently rcvolure, tlrickened, of'ten glaucous, normally moderately puberulous. Fliwer hetuli 6 7 mm diam. at anthesis,with 22 24 flowers. Caly_rmoderately puberulous (lobes lack the additional lorrg, spreadirrghairs of subsp. draiiemal; perats 2 mrl long, puberulousat apex. Zegrlr?.,sand secrlsnot seelrin mllture state. Dist bLttiotlqnd hobitcrt:(Map 5) South-westWestern Australia: this sub_ specieshas been collcctedfron two widely separatedregions, the first about 300 km due south-easlof Perth from near Kukerin eastward to the Lake Kirrg district, and the secondfrom near Wongan Hills which is about 150km noflh-eastofPertlt. Future samplingofsuitable habitatsmay indicatewhether theseocclrrrences reprcsent a real, or only an apparent,disjunct distribution, A sirrilar distributional pattern existsrn A. moirii E. pritzel sLrbsD.recarri,c_ tlpula B. R. Maslin. Acqcia drel,iana. subsp. ntinor grows in lateriie or slndy laterite among dense sandplain scrub. WESTERNAUSTRALIA: 2.2 mi N of WongsnHr r, E. M. Scrjnseuut.2095(CANB PERTH); I I lni E of Newdegate on the road to Llle King, M. D.-TinJalc 226 and _A.R. llotering antl ftuitingpcriod: Flo*er< from abour Vay to Julv; immalure fruiting specinens have been collecredin August and Septerr6er. 4',71 SL'

- .ISI^J: ueesqd'rbo : osr) oN .1o. pro,rrrl?/4.'rel6r.S,1::ls0l::3tli '(8r81)tOZ:Zt0i6i loqd-) !! /s I II sstardlu€ld'rxqeT uI usret DsouarDiqnsrP^ .r8d ( nrqe'I)srDr!)8!a DI)DV Z€ t?./-(uees qd€rSotoqd -) :osr).,'0S oN'/./aq1rC cac Lu',peor€]sn8nv pue ess"A,'ro,uu Zt8t ' 'tZ u€,$S,, . rd..{J QV}]D nO(,:Z ssrard tueld Lurle'I ul'usreytr 1p.raq113BlcBJv 'uoxel sql .roJcrlsou -8erp sr rlcnl,r elos l€uruJet tur8tnld eql o1 sra;er leriltde cgrcedsqnseq1 'srqJErJtlr pJurJJrJq plnoqs rl ^l)rrrnJJE e.rour sd€qrad q8noqlle 'e1es eql peIIEa st 'saa8und dsqns uo sIXe lEurulle} 'el€lnqns 'a8lBl 'tustsrs Jee[ 3.rrlue erl] sur.roJ qcq,r arnlcnrls lueSund ?q:] -uoc ur?ureJoI (z,L6I'[rls€]r'\tr€s) "les l"ultule:]oqt s" ol pejJeJelueeq seq (1uasa:d sr .red euo ,(luo ueq,r ue,re) eruurd jo rr"d lsoLu.reddneql ol 'uorsr^el l€lsrp srxE J"al ,{J"urJd eqt Jo uoruod I€crde eql s[il lnoqSnorqJ '"xBl ''qluag essql uee,rleq ,{lruu€ leeJ ou sr e.reqt :}nq od.tototsol 't, Jo suuoJ oruos solquesal 1r ,{11errgr:dng s:Lradsqnsr:qto rql uro{ lI qsrnBurlsrp,{lrprereelas leurrurel lue8und'o}rlnqns'agr"l eql puE sa^tal el€8tlI -run ,{JluelsrsuoceqJ uox":} erBJ t st sLla8und dsqns ououarp o1:to)y ur-re6r '^oN s'Lt6saE)aaD s F,'reur'uu.l Jo a serl*' vrrv($iS3$ N*JJlt '(.^I!)lzlg oopuDttl sndlp)ng) puBIpoo,r oopol?iA ur |os d11e,trr8 uI palJelloJ se,\\ uerllrJeds l"rusuu"iA aql l?ql sr srrtrl?d dsqns Jo stuoluerlnber lEllqErl eql SuIpJeBaJ pu"rl ot Lrorl?uroJur,{luo erll (quad Jo q ou ul{ S6 lnoqe) lBtuBuuBlA J€eu luo.rj puB (lnJtqnop tell,reLuos sI riclrl,\\ lo uorl?Jol otl]) JID,\xog tuo{ pspJoc?r :"rlEJlsnv uretselA ls3,r-qtnos \.9 o3\\I) :lpllqDqpuo ao!lt1gllls!Q 'uorldrrcsep sorJadsoes s/ars Suol tuur Z !J olloJoJ :(ouot't\a,{Pds.lr1s rc srrrq Surpeerdsl€uorlrpp€ eql tnoqlr,{\)el"lolllc s?go1r.I1rJ pBoriled gl tl sDt$o :BLtoIrum sapunpad esEqsll r"eu Elssl?ulLulel ecqllls lsddn 0l , 'elnlo^or 'strBdJo uo pelBn:lrs puDlD laecseJqq8 fllueutuold t Z salnuald '8uol '. '. ireeull xade urru g.l g. sry)u DLtutdJrEd 1 aouuttl paxegatro lq8rlrls ']uacseurds'el"lnqns 'pr8u'8uolI mru II-OI Dps IDLI|LL.|aJpoxoge] ro trq8terls '8uol 'lu3osaurds ruru 9 E + safd!8 snolnroqnd [IasJoJlLrEsaq)uo,lg '(€6S6t 'aF^xog -tal l :oloq) u s s/1r1 DllntsstN :)dQ 'e!-ll?rqel8 'eelnlo^er 'aE8n|-€-Z 'r8uol LULLr lln)anpa

Distribution: (Map 1l) South-west Western Australia: occurring sporadically from York southward to near Augusta atrd Denmark.

WESTERN AUSTRALIA:6 5nri W ol Penberton,E. M. Bennatt3203 (B, MO,PERTH); Near Nannup, W. E. Bla.kall s.n.,lateDec. 1930;Dryandta, A. L. ClAo 10; Wester.nAus- tralta, J. Drunmotl(l 31,1(MEL)| Denmark, C A. Gartlner407; Penberton- M. Koch 227| (MEL); Between187 and 188mile pegson BussellHighway, B. R. Maslitt476 (AD. MEL- NSW, PERTH); 5 nri S of Witchclille on BussellHighway, B. R. Maslin 1603; Mount Bake, well, P/'ersr891 (MEL); Forest Crove, R. D. Royce 397/: 1 mi from York, O. H. Saryetrt s.n..22June 1924.

Flovering arul lruititg period. Flowers frotr December to March; nrature legumes appear in August and persist on some bushes ultil Dece]nber. Bcntlram (1864, p.417) included the type of A. nigricqns(Labill.) R.Br. yar. subrqcentosqMeisn. (Drunmond Il:151) in the list of specimenscited under A. gilbertii. Mr. L. Pedley,while Australirn Botanical Lirison Officer at Kew in 1972,compared the types of thesetwo taxa (which are mounted on the sameherbarium sheet)and commentedthat they agreewith each other in almost every detail. The present author has inspecteda photograph (frorn Kew) of thesetypes and it is apparentthat they representthe sametaxon. The white, 4-merousflowers, and the small, truncate or sinuate-toothed calycesrender A. gilbertii unique within the seriesPulchellaa. [t is interesting to note the sinrilarityin flower structurebetween this speciesand ihc phyllo- dinous taxa, A. ntyrtifoltu (Sm.) Willd. and A. calostrifoliaBenth. Other interesting characters found in ,4. gilbertii rr1cl\de its cbracteate peduncles, 2 to 8 flowers per head, sessilecaducous bracteoles,stipitate ovaries, and longitudinal seeds. Acocia gilbertii is not closely related to any specics|t tbe Pulchellqe but as it has smooth, glabrous,red to brown branches,and large pirnules, it bears a superficialresemblance to A, IeioclermaB. R. Maslin: In addition to the

Figfie 32 Acacia gilbertii. A poftion ol branch systent. B Node. C -lnflorescenec. D-Flower showing + truncate calyx. E Stipitale ovary. F Gland (g) obliquejy ter- minating the vertically Ilattenedrib on upper suffaceol petiole (p). G-Leglrme. H-Seed. A from R. ,. Rof& 39'77, B, F-H lionl 8. n. ,{r'asrir1603: C-E from Blackall s.r. 476 LLN

,lb/ u.y'c"e/./yt

rq_ sln l'l{wm )L #r% J\w'----Ei!z- c ! *w L s\ \\\-. _, u-__ ---\\ '+s\ V charactersmentioned above, ,4. gilbertii is distinguishedfrom this species by its caducousstipules, longer petioles,normally unijugate leaves,alld the absenceof glandsbetween the distal pinnuleson tlre pinnaerechides.

24, Acaciainsolita E. Pritzel, Bot. Jb. 35:310,f.36 (1904). Slirrlpes; "Hab. in distr. Darling australi pr. Greenbushesin silvis unbrosis glareosisflor. m. August. (D.3835, E. Pritzel PI. Austr. occid. 1013)." (Iso-Pritzel 1013: AD. NSW, W.)-Frs. 33

S,4raDnormally 30-60 cm tall, probably producing s:uckers' branchlets ribbed (ribs yel1ow), green to brown, glabrous or sometimespuberulous; axillary spinesabsent. Sliprl"s + narrowly oblong, I 2 (6) mm long, glabrous or ciliolate, light brown. leares dimorphic, indunentum as on branchlets. Bipinnate.foliage (normally confined to lower part of plarrt): petiole 7 25 mm long, l_ teretc to somewhatvertically flattened,straight, lateral rib normally conspicuons; terninal set{! | 2 mm long; pinnae I pa]r' pinna rachis 20,30 (45) rnm long, ribbed above; pinnules6 l0 pairs, elliptic to narrowly elliptic or somewhatovate, (3) 4 8 x 1 5-2.5 (4) mrr, flat, subglaucous,concolorous, smooth or obscurelywrinkled, nerveless1o obscurelyl-nerved (sometimeswitl.r an obscure second nervc diverging from the petiolule), minutely apiculatei g/azrl abscnt. Phyllodes(confined to upper part of branchcs);linear, l0 60 x I 3 mm, flat, midrib prominent, shortly apiculate (apiculum straight on uncinate);pulvinus 0.5 1 mm long, obscurelyr:ugose; gland absent. Inflorcs- cencessimplc; peduircles solitary. normally confinedto the axil of the phyllodes, 6-12 mm Iong, minutely puberulous,base ebracieate at anthesis;flower heads yellow, globular, 7-9 mm diam. at anlhesis,with 14 l7 flowers. Bracteoles I l 5 mm lorg. I/ou.ers5-merous; calyx 1 i2length of corolla, divided for ca. I /,1 its length into oblong to triangLrlarciliolate lobes, tube glabrousto sparsely puberulor.rsand normally prominently 5-nerved; petqls ea. 2.5 mnr long, glabrous,1-nerved (nerve thickened and prominentat apexof petals). L,girzes somewhat hard and brittle, narrowly oblong, 45 50 x 7 mm, reliculate.flat, slightly raised over seeds, glabrous, light to dark brown; matgins rot (or only slightly) contracted belween seeds,prominently thickened, yellow. Seedstransverse to oblique in legume, orbicular to broadly elliptic,3 3.5 x 3 mm,flattened,dark brown, shiny; pleurogrttntcontinuous; acrole I 5x I mm; lunitle filiforn, reflexedbelow a straight abruptly thickened aril. Distribution and hqbitut: (Map l1) South-west Western Australia: lateritic soil in the Darling Ralge from Dwellingup to the Bridgctowl district: this specieshas also been collectedfurther to tlle east from betrveenPingelly and Narrogirr. WESTERN AUSTRALjA: Collie. C. Antlreyss.n., July 1905(NSW, PERTH) also Oct. 1913;Lowden, M. Koch2152 (.MEL, NSW, PERTH); Approximarelyl4 ni E of Dwellingup, W. Lotrctaea 332; West of Popanyining, l-. Lullfi. 1786 Darkan Wesr, F. L l\ltz s.n., 20 July 1962;Marradong, N. G. Marcha t 4,7 nri E of Donnybrookon roadto Collie.B. R. lAaltin ql5 6.Dt MEL, NSW, PERTH); 45 mi SE of Boyup Brook on road to Cranbrook, B. R. Maslitt 632: Narfogin, .1. "/. Milesi s.n., July l9zl0; Bridgirown, 1. Oh.r? 641 (NSW). Flow-ering and fruiting period; Flowers from late June to August; mature legumes havc been collected in early Decerrber.

The mature foliage in A. insolita consists of phyllodes; the bipinnate leaves which occLlr lower down the branch appear to b,- persistent throughout the life of the plant. Juvenile plants (where phyllodes have not yet developed)

Figote 33 -Acocia /rrdll/a. A-Mature plant with ph),llodesat apex of branchesand bi- pinnate lbliage towards base. B Ph)llode. C Bipinnare lcaf uitLr long periole (p). D legume. f Seed. f flo$er hcdd. A from C. Andrawss.n , July 1905; B-C from N. G. Marcha t 4; D-E from C. AnLlrcv,ss.t1., Oct- 1931;F fron F. LulL.itz1786. 478 6L'

:,,*io'

.-Ei"__

'dffi fiH tr[f,

-- -.-E-- can be recognizedby their relativelylarge, unijugate leaves, their long, eglandu- lose petioles, and their flat, minutely apiculate, quite numerous pinnules. Inflorescencesare generally not developed until the mature, phyllodinous foliage is produced. Jtrdging from some herbarium sheets, it appears likely that A. insolit.t producessuckers; horl'ever, this is yet to be conlirmedby lleld observatiolls. Because l. insolita h'as traditionally been included in the Pulchellae, I have acceptedit for the purposeof the presentrevision. However,it is highly unlikely that the natural amnity ofthis speciesis with the Pulchellae; the develop' ment of phyllodesalone would excludeit from this Series. Also, the eglandu- lose petiolesare very ulrusllalfor this group. The true position of A. insolita within the gen\s Acqcia is somewhat obscurc.

Dubious names Acacia subtilisHoffmannsegg, Verz. Pflkr.rlt. Nachtr. 2:43 (1826). This species was describedfrom vegetativemate rial; it was apparertly clrltivated under the name A. pulchella. Having examined the original description of ,4. subtilis I am unable to apply this name with certainty to any knor,vn nember of the Pu/chellqe.

Excluded species Acacia shirleyctnaDomin., Bibl. Bot., Stuttgart 89:826(1926) Dichrostqch)is "N.W. spicata(F. Muell.) Domin. I have examinedthe type of .'1.slirleyana, Australia: inter flum. Ashburton et De Gray River. legit E. Clement"-PR) and this specimerris dellnitely Dichroslacll)s spicqta. It is interesting to note that jn the samepublication as A. shirleyanowas described,Domin made the new combination D. spicqt{t (based on Neptunia spicatq F. MLrell.) and cited "zwischen the following specimens: Ashburton r-rndYule, Ashburton-Lrnd De Gray River, E. Clement." It will be noted that tbe second locality is identical to the one given on the type of A. shirleyana,

Acknowlcdgements Assistanca with the revision of the Pulch(llae has been received 1'rommany sourcesboth at the institutjonaland personallevel. Although it is not possible to mention everyoneindividually, the autbor extendshis sincerestthanks to all thosefrom whom help hasbeen received. Specialmention must be made of the generosityof the National Herbarium of Victoria for donating to the W.A. Herbarium fragments of maty Acacia type specimens(including some Prl- r:hellae.) I am particularly indebted to Mr. Arthur Court (CBG) for his valuableadvice on sofle nomenclaturalmatters and in particLrlarfor allowing me accessto much of his manuscript malerial on Australian lca./o species. Mr. Alex George is also gratefully acknowledged for checking my Latin des- crrptlons. References BENTT{AM,G. (1842)-Noteson Minoseae,with a Synopsisof Species.Lond. J. Bot. 1:318392. BENTHAM,G. (1855) Mimoseae. PlantaeMuellerianae. Linnaea 26 : 603 630. BTNTHAM,G. (1864) Flora Australiensis,vol. 2. Reeve,London. BINTHAM,G. (1875) Revisionofthe SuborderMimoseae. Trans.Linn. Soc.Lond.30:336- 664. BRowN,R. (1813) Acacia.In W. & W. T. Ailon, Hortus Kewensis.Ed. 2. S: 459 413. London, CANDoLLE,A. P. DE (1825) ,1.4crd. Prodromus Systenatis Naturalis 2: 448 473. CouRr, A. B. (1972) Mimosaceae- In J. H. Willis, A Handbook to Plants in Victoria, vol.2. Melb- Uni. Press. DoN, G. (i832) Acacia. A, General History of the Dichlamydeous Plants, 2: 401 423. London. CARDNER,C. A. (1923) Contribuiionsto the Flora ol WesternAustralia. No. 2. J. Roy. Soc.W.A. 9 (l) | 37 ,13. 480 I8t

'0zn-6LE t\E) 9l '1o{ 'lsnv '(e€souurnSe'I)DlunrdaN sr.tra8 oqt uolsl^at{ U C['ulnqNll!\ 'lOl-L6f 'oSeledrqJrv Jo V-@96I) 'srTrlld tQ) I eHo en,\I V lA rq'rqJeu oql Jo sluEld-(6s6l) H I '| oN'dru JoS rSoaD 'lsnv eJog ps€'I e€ 'eqcleq.eu eql o8elodlq.rv eqI (€96D 'sn'IllA '9901 lJEd Jo H I d'loreg lloq lueld'wntJ:]' Dto!llt p!Jo2V-(608I)'I'J'/doNqqr[rll\ '9tz-I6I :l0I esnolnoJ leN lslH 'JoS "rvssv llng prJErV.p s.ulprS sop rnbrSoloqdrotu opnlg,l € uoIlnqI4uoJ-(IZ6I) f 'I dd'tI 'UV'8'lo^'I 'tuoJ osnolnoJ 'e€leqdoco{.4o9ld + Z€ I 'aElleqclnd 'lsorod'qe'I ^E{ aEre]lur@nC :sguuedlqsBlcscY so'I II 'tr€r\eH 'asolurog 'ell€llsnv le ecrrnej l sell :enbgls€d 1o uelpul suee-tosep sell?lnsur selrew '€lcecvtp seFtueld seperSoloqdroqr ?l ep opnt?ll€ uoqnqlrluo3 (0/6I) '6-I ']rV'9 'uIoJ f'lYssYA dd'tz 1o,4.'[ esnolnoJ lsejod q"'I ^"rJ 'o!tD?y afieR eI suep serrelloJsopets sraFuard sep epnl?.I e uoqnqlrluoJ-Gg6D I lvssv :(z pue "lluqlog -s"r.€cv uErrrJv eql uolle.glsselc ? sprea,roa1g161i I) rr '€cr4\/ Jo H J;ssod qlnos Jo ,{}eraos so4"^lesuoC pu€ uollaolold eJIIplIlA aql q.u"rq eqt .{q pe 'pl€N solteds D12D2Y.r -(ZL6I) 'ssol{ Jo leleN lqnal Z Jo 'slFluaplsroH I nle-gLz:9E qf pg e€llBrNnv e"qd?folfqd eluourS"rg pue sl0I(I '1 q 'DrrDV (1061)'g'lazllxd lazlud'uu€Luga'I E 'rIaNsIsW 'SJnqrueH'902-66I :Z Buelsslejdeelueld uI ,irrr,,-(8t8l) C J '8mqur€H 'IrIIuI"qe'I J O f tuaN$a,\I tZ-€;I "uErssrorde€tu€ld J C'-t q rrrY-(tt8l) d C 'O9Z-n92.(O "rsl{nN snue8oql q salpnls-(Zt6l) g 'NnsvI^I I l-rjrrrz '86td t6t:92 urFog 'zuelreo '8llv 'aEllaqclndrep Sunleqlqv rop sn€ ueIzDIv eI(-(8S8I) 'X 'Hco) '{roi r\eN 'rouJeH 't po 'srurotr.ru€tog .{.ressolC g'NosxJvt '€99t 'uinroDtaJo V-(826I) C 'rarooH 1g,/. 3"W log s,slunJ olzotY-(298l) I l t l-' *,""':: . ", i si I -

I Acacia mesacephala Acacia p ulche I la e var.lagonioides O var.glaberrima I var goadbyi

o I d6*b."""."".... to

o.9Q,^., o oo

o t 4'::""' o

Map 1 DistfibLrtion of Acacia , ecacephala, ,7, pulcheila vat. faganiaidet, \ar. Elabeninn. ,rnd \ar. E.,adb)'i.

482 'sr/rssasgTrr'.r€^ pue ov{).l )e^'Dlptpltld )e^ D apl d Dlrttry lo uorlnqulsrc-Z dEI l

? Ot oou'tlo'o' o

@

o

!t \ .'"",,,'J,r'^"r j"""1 T o3naoo9rvy o vsooy{aO o+

rIro,o!+ t.-frls] rrr-.- f ll LJ I s,,,qs.sqns,e^o"":ii'Tiiir"i'"" \ 1 | ) - L/YxBo'r' 1 ,

rlvcs I P TNDEX

Acacla lasiocarpa var. lasiocarpa O var sedilolia

o A KALGOOFLIE'

"?^ ooov

Map 3-Distributioo of Acacia lasiocarpa var. lasiocarpa, \^t. sedifolia and var, vlllosa,

484 s8?

'DqtuoeDda're( pse DJqoanDrq'&L Dilrwolsol 4tnY Jo v.ogng\rtslo--t dEL{

o 6S,"""', o^ bo o "o oo o

olnuooclvy

eqtue.eda re^ * eletoaroerq re^ O e.lre.o!set R!.ecV Acacia drewiana O subsp.drewiana $ suosppuneens

O subspmoir.i I subsp.dasycarpa

KALGOORLIEO

**';+'j,t'.^""."

Vap 5 Di.trrbrrrionol At,hio tlt?tviunurnd L n,,itii. L8t

'suoxtrStu 'oa$r8 'y 'otolx,idq v pne Dlxbr/ Jo uounqrlsre 9 dejt

all'ruoo0lvy

sueru6!0 et.eov El ees!r6 er.ecVa erncsqoLea S e!pauraru! re^ + !raqc pu€ 1?^o eueluaorq]e^ o euquforq e!.E3v fl\tDEX

;ffiW/,

XALGOOFLIE'

Map ?-Distribution of Acacia etqelioclada, A. lateiticola and A. leiodema.

488 68t

.DuDltoltbt .V plde osow -aJbtqns 'V 'bublsslald 'v 'olorlld 'y 'o! 'l/ 'bpavl apotuad blxbev Jo uo4nqrllsl(-8 dEj I

a3tltoo0lv)

xlor{l KALGOOBLIE'

Map 9-Distribution of Acqcie r)aria.

490 t6t

'lryouunxp t,poty Jo uorlnqr4sr(-ol dey{ Map ll-Disttibution oI Acacia depressa,A. eilbertii, A. insolita and A. newbeyi

492 :u' 'uslel/{ 8rt rrrlsslrld .rBA l6z]t-rd g DUD!rlou' Je^ 9tt 'uslelr'\iJlrJ,salrt r€A €et ''' ''J(f Dtnrsqo oIt ''- - 'lalrev '8€' '9€t €c? oJntsqo 8?i ,' uslell{ rxowaJDlqhs tel 9Lt 'rg d (lllqE I) su€cu8^ts 8€' '''' '''' '7lne suDclJdlu zrt '' tTt ullsEw { { llaqrdou ttt uuerxe5s xe uoH Lllre, ullsEl^l l{ g Dlndlts!^ln3ol dsqns oTt ''.' ullsEl^l BdrEcfssPdsqns 6It '-U { lezllla[ 3 rulour Ltt ' ' 'qlueg rlFqtllur oLt ' - 'g ulls€I I { eleqdt]?8atu , LOb ' '' g €En ullsel,r{ u elmlnl urls€I^l x g Bu|repolal t zw EIocgFelBI It €tt uils€I I d { ttt urlsel,i{ U g oezllrd'g) €soll!^ r€^ ullsel I l{ g ( ustol I) ElloJlpesre^ n' re^ 9tt ullsEJI d'g eqluBrE0e ulls€I/'{ U g a}eloepBIq IE^ 9tl '''- qlu0g softaolsel ffit ,so4tl{Dl L6t IeBeXxe uoH qco) ) xt uoH ,/Dllrl zon ''' rgzllJJI elllosul 8Lt ' qco) >[ DlDroud! 8€t ' 'JA oals\plasly '''' Y zot eroow s e0sl6 zln ' ,{r{uoH x2 lJoH stPuorS^ zw ulwo(l €0t ' 'uslel !4qPDod Mueql!8 sLt ' ' 'glueg saPloll1dof s0t q.otxe opol.oua zot '''' 'trstaltJoH l latlcllpua t* ' ullsel BpelJolladue 9lt '''' l {'g uIlsEI I d g DlDu!4)a zzt a71t)A'-l D)tdlt )B^ 99t ''4:u5goio{tdLo,t te^ 79t '''' 'tlltr.g oaplo^o te^ tsl 'tlTueg )olDu ft^ L9t g clsqns 89t uIIs€I^JX suegela ullsEj,\l g ( usIoIAI)Bueellopue:) dsqns L9n ',' d ultuo( ( usql ) DuoaopuD).E^ L9n 'Iput'l rrpuoLuunrp nw g sur8unddsqns ull

seuEu ol xapul 468 438 444 451 448 448 468 397 409 402 4A 402 405 442 403 399 402 399 401 var. subsessilisB. R. MasLin 406 var. villosa E. Pritztl 414 tutaefolia anct. 442 rutaefolia Link. 438 shirletatsDomio 480 r/rifosdLink ...... ,raJ.borealis E. PtilzEl 433 'rar, brertolia Meisn. var. €rdr?r"ri (Meisn.) Bentb. 43r var. intetmedia E. Pritzel. " . 428 subracemosaB. R. Maslin 446 .rrbrl/s HoffmanDsegg 480 taylorianaF. Muell. ... 450 varia B. R. Maslin 456 var. amnis B. R. Maslin 461 var. crassin€rvisB. R. Maslin 459 var. Darviflora(Benth.) B. R. Maslio 462 Dichrosta.iysspicala (F. Muell.) Domin 480 Mimosa nigricansLabill. .. . 438 Mimosa pulchella(R.Br.) Poir. . 39'l

Publication date of Volum€ I' Number 4 The publication date of Yolume 1, Number 4 was 23 May 1974'

115a9/9t7+-l$ WILLIAM C. IROWN, Gov€rnm.nt Printe.. W.5r€m Aultr.li.