(Coleoptera, Staphylinidae, Tachyporinae) from China

Zootaxa 4105 (3): 201–242 ISSN 1175-5326 (print edition) http://www.mapress.com/j/zt/ Article ZOOTAXA Copyright © 2016 Magnolia Press ISSN 1175-5334 (online edition) http://doi.org/10.11646/zootaxa.4105.3.1 http://zoobank.org/urn:llsid:zoobank.org:pub:4B6393FF-2CC4-445D-9C8E-1435558E07CF New species and records of Ischnosoma Stephens (Coleoptera, Staphylinidae, Tachyporinae) from China

MATÚŠ KOCIAN1 & MICHAEL SCHÜLKE2 1Department of Ecology, Faculty of Environmental Sciences, Czech University of Life Sciences Prague, Kamýcká 1176, CZ-165 21 Prague 6 - Suchdol, Czech Republic. E-mail: [email protected] 2Museum für Naturkunde Berlin, Leibniz Institute for Research on Evolution and Biodiversity at the Humboldt-University, Invaliden- strasse 43, 10115 Berlin, Germany. E-mail: [email protected]

Table of contents

Abstract ...... 202 Introduction ...... 202 Material and methods ...... 203 Descriptions of new species ...... 205 Ischnosoma zipapertum sp. n...... 205 Ischnosoma vicinum sp. n...... 207 Ischnosoma foedum sp. n...... 209 Ischnosoma involutum sp. n...... 212 Ischnosoma yunnanum sp. n...... 213 Ischnosoma abstrusum sp. n...... 214 Ischnosoma gemellum sp. n...... 216 Unnamed and presumably undescribed species ...... 219 Ischnosoma spec. 1 ...... 219 Ischnosoma spec. 2 ...... 221 New records of described species ...... 221 Ischnosoma absalom Kocian, 2003...... 221 Ischnosoma bohaci Kocian, 2003 ...... 222 Ischnosoma bolitobioides (Bernhauer, 1923) ...... 223 Ischnosoma chinense (Bernhauer, 1939) ...... 224 Ischnosoma convexum (Sharp, 1888) ...... 225 Ischnosoma discoidale discoidale (Sharp 1888)...... 225 Ischnosoma discoidale taiwanum Kocian, 2003...... 226 Ischnosoma duplicatum (Sharp, 1888) ...... 226 Ischnosoma evae Kocian, 2003...... 229 Ischnosoma farkaci Kocian, 2003...... 230 Ischnosoma fasciatocolle (Champion, 1922) ...... 231 Ischnosoma fusciventre (Tikhomirova, 1973) ...... 231 Ischnosoma hieroboam Kocian, 2003 ...... 232 Ischnosoma noemi Kocian, 2003 ...... 232 Ischnosoma quadriguttatum quadriguttatum (Champion, 1923) ...... 233 Ischnosoma rosti (Bernhauer, 1923) ...... 237 Ischnosoma splendidum (Gravenhorst, 1806)...... 237 Ischnosoma taibaiense Zhu, Li & Zhao, 2005 ...... 239 Key to the Ischnosoma species of China and Taiwan ...... 239 Acknowledgments ...... 241 References ...... 241

Accepted by J. Klimaszewski: 4 Mar. 2016; published: 21 Apr. 2016 201 Abstract

More than 1100 specimens of the genus Ischnosoma Stephens, 1829, from China and Taiwan are revised. In all, 29 species and one subspecies are recognized. Seven new species are described and illustrated: Ischnosoma zipapertum sp. n., I. vicinum sp. n., I. foedum sp. n., I. involutum sp. n., I. yunnanum sp. n., I. abstrusum sp. n., and I. gemellum sp. n. Additional records of previously described species are reported. The distributions of all the species are mapped. Four species, Ischno- soma dicoidale discoidale (Sharp, 1888), I. faciatocolle (Champion, 1922), I. hieroboam Kocian, 2003, and I. rosti (Bern- hauer, 1923), are recorded from China for the first time. A revised key to the Ischnosoma species of China and Taiwan is presented.

Key words: Coleoptera, Staphylinidae, Tachyporinae, Mycetoporini, Ischnosoma, Palaearctic, China, new species

Introduction

This paper continues earlier studies of both authors, which were started by the first author (Kocian 1997, 2003) and continued in several shorter articles by the second author (Schülke 1998, 2001, 2003, 2007). An overview of the world fauna of the genus Ischnosoma was provided by Kocian (2003), including a revision of available type material, descriptions, and illustrations of all the species not covered in preceding revisions of New World (Campbell 1991) and West Palaearctic species (Kocian 1997). Since that revision, only few species of the genus have been added from Greece (Schülke 1998), Turkey (Schülke 2001, 2003, 2007), and China (Zhu et al. 2005a). Previously, 99 valid species and two subspecies were known from all major zoogeographical regions, with the greatest diversity in the temperate zones of the Palaearcic and Nearctic regions and in the Oriental region. Only few species are distributed in parts of the Australasian region. Records from the Afrotropical region are only known from mountainous areas in Ethiopia and Central and East Africa, mostly along the East African Rift Valley. Records of Ischnosoma species from South America are unknown; the genus has been recorded only from the Nearctic/Neotropical transitional zone from Mexico southwards to the north of Panama. Species of Ischnosoma are easily distinguished from their closest relatives in the tribe Mycetoporini by many characters, including the structure of the mouthparts, the chaetotaxy of the apical spines of the tibiae, the shape of the basal carina of the scutellum, elytral chaetotaxy, the shapes and chaetotaxy of the male sternites VII and VIII, and the aedeagus. The closest relatives of Ischnosoma are the genera Parabolitobius Li et al., 2000 and Bryoporus Kraatz, 1857, which share a similar chaetotaxy of the apical tibial spines. Both genera are clearly separated from Ischnosoma by different mouthparts and primary and secondary sexual characters. The fauna of the Afrotropical region is still somewhat doubtful, as some of the species described as Bryoporus may belong to different genera. Their systematic position requires revision. Extensive collecting in China has been conducted by the second author in the past two decades (together with V. Assing (Hannover), A. Pütz (Eisenhüttenstadt), A. Smetana (Ottawa), and D.W. Wrase (Berlin)). Therefore, we decided to treat the Chinese fauna as a first part, to be followed by a revision of material from other regions. Our knowledge of the Chinese fauna has improved considerably during the last two decades, but it is still far from complete. This is particularly suggested by additional unnamed species of which only female specimens are available. Including the species decribed in this paper, 29 species and one subspecies are known from China and Taiwan, thus rendering this the region with the greatest diversity, followed by the United States with 13 and India with 11 species. Of the 30 recorded species and subspecies, 18 (60 %) are known exclusively from China, one subspecies seems to be endemic to Taiwan. Figure 1 shows that the genus has not been recorded from large parts of China, which does not necessarily indicate the absence of the genus from these areas. The diversity of the Chinese fauna increases from north to south. Yunnan is the province with the greatest number of recorded species (18), followed by Sichuan (8), Taiwan (8), Shaanxi (7) and Gansu (7). Significantly more species are known from the mountainous western provinces (Yunnan, Sichuan, Gansu) than from the eastern, more lowland provinces. This tendency partly reflects the different intensity of field work, but probably will not drastically change even when more material becomes available. Prior to this study, fewer than 400 specimens of Ischnosoma had reliably been recorded from the study area (less than 200 from China, some 200 from Taiwan). In this paper, we add data of more than 1100 recently collected specimens (nearly 1000 from China, more than 160 from Taiwan).

Collections cAss private collection V. Assing, Hannover, Germany cKoc private collection M. Kocian, Prague, Czech Republic cPüt private collection A. Pütz, Eisenhüttenstadt, Germany cSch collection M. Schülke, Berlin, Germany (now MNB) cSme collection A. Smetana, Ottawa, Canada (now NMNST) CAS China Academy of Sciences, Beijing (via V. Grebennikov) CNC Canadian National collection, Ottawa (V. Grebennikov) KUM Kyoto University Museum, Japan (M. Maruyama) MAKB Museum Alexander König, Bonn, Germany (D. Ahrends) MHNG Museum d'histoire naturelle, Genève, Switzerland (G. Cuccodoro, I. Löbl) MNB Museum für Naturkunde, Berlin, Germany (M. Uhlig, J. Frisch, J. Willers) NHMB Naturhistorisches Museum Basel, Switzerland (M. Geiser, E. Sprecher-Uebersax) NHMW Naturhistorisches Museum Wien, Austria (H. Schillhammer) NME Naturkundemuseum Erfurt, Germany (M. Hartmann) NMP National Museum Praha, Czech Republic (J. Hájek, M. Fikáček) NMNST National Museum of Nature and Science, Tokyo, Japan (S. Nomura) SMNS Staatliches Museum für Naturkunde, Stuttgart, Germany (W. Schawaller)

Measurements

ED distance between interior borders of eyes in dorsal view) EL eye length (dorsal view) ESL elytral length from humeral angle to posterior margin HW head width (including eyes) PL pronotal length PW pronotal width

Permanent preparations of abdominal segments and aedeagi were made in Euparal (M. Kocian) or PHP (M. Schülke) on plastic cards attached to the pins or on the mounting cards next to the specimens. The morphological studies and measurements were conducted using STM 822 5410 and Olympus SZH10 stereo microscopes. Drawings of the abdominal segments and the aedeagi were created using a Meopta microscope and a Zeiss drawing tube. A digital camera (Canon EOS 450D with macro lense) was used for the habitus photographs. Micrografx Picture Publisher 6.0 was used for post processing of the illustrations and the compilation of plates. The maps were created using MapCreator 2.0 (primap) software. Additional sexually dimorphic characters. In many studies in the past, beginning with Rey (1883), Ganglbauer (1895), and Luze (1901), the sexually dimorphic shapes of the abdominal sternites were used to distinguish species at least of the I. pictum group. The characters were first only described from non-dissected specimens. Later, starting with Hansen (1952) and Palm (1966), the characters were illustrated from non-dissected specimens. Campbell (1991) was the first who gave detailed illustrations at least of the male sternites VII and VIII, as well as of the aedeagus, which is still the base for distinguishing many similar species. These characters were used also in earlier studies of both authors and other papers cited in the introduction. In general, male secondary sexual characters are the shape and chaetotaxy of sternites VII and VIII, ranging from simple emarginations of sternite VIII to deep emarginations of both sternites VII and VIII, combined with characteristic patterns of clusters of modified setae. Other sexual dimorphisms had been unknown. A detailed study of species of the I. pictum group revealed that some species of this group possess a complex of sexual dimorphisms, which are partly related to wing dimorphisms (males macropterous, females brachypterous or macropterous) and which are most pronounced in the widespread species I. duplicatum (Sharp, 1888): Head (Figs 28, 29). Eyes of males are larger, about 0.25–0.30 mm long (EL in females 0.15–0.21 mm) and

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 203 much more prominent (ratio HW/ED about 1.70, only 1.35 to 1.40 in females). Eyes in males distinctly protruding from outline of head, temples distinctly narrowing behind eyes, maximum width of head across eyes. In females eye size varies between small-eyed brachypterous specimens and macropterous specimens with larger eyes, but the eyes never protrude from the outline of the head. Temples behind eyes subparallel, not converging behind eyes. Pronotum. Same ratio (PW/PL around 1.20–1.25) in both sexes, but the sides more evenly rounded with almost completely rounded posterior angles in females; in males with almost parallel lateral margins in posterior half and pronounced posterior angles. Elytra (Figs 28, 29). Longer (ratio ESL/PL 2.95–2.15 in males; 1.50–1.80 in females) and wider (ratio EW/PW 1.35–1.45 in males; 1.15–1.35 in females) in males. Elytra of macropterous females are distinctly larger than in the brachypterous morph, but not reaching the size of the male elytra. Elytral chaetotaxy varies with elytral size, the number of setae in the doubled dorsal row is about 20 in brachypterous females, up to more than 30 in males. Antenna. Male antennomere II, especially male antennomere III, distinctly shorter than female antennomeres II and III (ratio length III/IV about 0.75 in males, 1.15 in females), while the penultimate antennomeres are more oblong in males (ratio length/width X about 1.3 in males, 1.0 in females). The length of the antennomeres in females is not related to flight capability. Aside from I. duplicatum, species with similar, more or less pronounced dimorphisms are I. rosti (Bernhauer, 1923), I. noemi Kocian, 2003, I. maderi (Bernhauer, 1943), and I. yunnanum sp. n. All these species are restricted to the East Palaearctic.

FIGURE 1. Pooled distribution of the genus Ischnosoma in China (material examined and literature records).

Ischnosoma zipapertum sp. n. Figs 4–8, 56.

Description. Body (Fig. 4) reddish-brown; head, elytra and anterior part of tergites often darker, reddish-brown to brownish; legs, palpi and antenna brownish-yellow. Head 0.40–0.45 mm broad; 0.55–0.59 times as broad as pronotum, as broad as long. Eyes slightly shorter than, or as long as temples. Ocular seta well developed, as long as setae of pronotum. Surface without microsculpture. Antenna 0.96–1.11 mm long. Antennomere X distinctly wider than long. Pronotum 0.68–0.82 mm broad; 0.53–0.65 mm long; 1.22–1.29 times as broad as long, broadest in posterior half. Surface with distinct transverse microsculpture. Posterior angles relatively well marked. Anterior and posterior punctures separated from pronotal margin by 2–3 times the diameter of the punctures. Anterior margin of pronotum without carina. Elytra quadrangular, parallel-sided, along suture 0.4–0.5 mm long; 1.00–1.04 times as broad as pronotum; 0.76–0.83 times as long as pronotum. Surface with distinct transverse microsculpture similar to that of pronotum. Sutural row with 6–8 punctures, discal row with 5–8 punctures and lateral row with 6–9 punctures. Wings reduced. Abdomen parallel-sided, only slightly tapering towards apex, densely puntate. Tergite VII without whitish apical seam of palisade fringe. Surface smooth with very fine transverse microsculpture. Length of forebody 1.39–1.69 mm; total body length 3.1–3.6 mm. Male. Sternite VII with conspicuous modification, in the middle of posterior margin broadly and deeply concave, with medial dent carrying some short setae (Fig. 5); sternite VIII in posterior margin in the middle of posterior margin very deeply concave, with 6–7 rows of modified setae similar to an opened zip (Fig. 6); paramere and aedeagus: Fig. 7.

FIGURES 2–4. Habitus of Ischnosoma vicinum sp. n. (2); Ischnosoma yunnanum sp. n. (3); Ischnosoma zipapertum sp. n. (4).

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 205 FIGURES 5–7. Ischnosoma zipapertum sp. n. Male sternite VII (5); male sternite VIII (6); aedeagus in ventral (left) and lateral view (7). Scale bars: 0.1 mm: a (Figs 5–6), b (Fig. 7).

FIGURE 8. Distribution of Ischnosoma zipapertum sp. n. (circles) and I. taibaiense Zhu, Li & Zhao (triangles) in China (material examined and literature records).

206 · Zootaxa 4105 (3) © 2016 Magnolia Press KOCIAN & SCHÜLKE Etymology. The specific name is composed of the English noun zip and the latin adjective apertus: opened. It alludes to the modified chaetotaxy of male sternite VIII, which somewhat resembles an opened zip. Type material. Holotype, male: CHINA: "Yunnan [CH07-28A], Nujiang Lisu Aut. Pref., Gaoligong Shan, side valley 19 km NW Liuku, 25°59'02''N, 98°42'23''E, 2730 m, devast. prim. for., litter sifted, 10.VI.2007, M. Schülke / Sammlung M. Schülke, Berlin" (cSch). Paratypes (30): same data as HT (12 exs.) (cSch, cKoc); same data as HT, but "9.VI.2007 [CH07-28]", 1 ex. (cSch); "CHINA (Yunnan), Nujiang Lisu Aut. Pref., Gaologong Shan, creek valley, 20 km NW Liuku, 3000 m, 25°58'49''N, 98°41'48''E (bamboo, shrubs, litter sifted), 9.VI.2007, D.W.Wrase [27]", 2 exs. (cSch); "CHINA (Yunnan), Nujiang Lisu Aut. Pref., creek valley 3 km SE Gongshan, 27°43'02''N, 98°41'27''E, 1450–1500 m, (litter, moss, sifted), 5.VI.2007, D.W.Wrase [20] / Sammlung M. Schülke, Berlin", 1 ex. (cSch); "CHINA: Yunnan [CH07-28], Nujiang Lisu Aut. Pref., Gaoligong Shan, side valley 19 km NW Liuku, 25°59'02''N, 98°42'23''E, 2730 m, devast. Prim. Forest, litter sifted, 9.VI.2007, leg. A. Pütz", 4 exs. (cPüt); "CHINA Yunnan [CH07-30], Nujiang Lisu Aut. Pref., Nu Shan, 7 km NNW Coajian, 25°43'29''N, 99°07'57''E, 2420 m, second pine forest with shrubs, litter, bark sifted, 11.VI.2007, leg. A. Pütz", 1 ex. (cPüt); "CHINA (Yunnan) Nujiang Lisu Aut. Pref., Gaoligong Shan E pass 20 km NW Liuku, 3000 m (creek valley with devast. prim. forest, ferns, litter, moss, sift.), 25°58'49''N, 98°41'48''E, 3.IX.2009, D.W.Wrase [25] / Sammlung M. Schülke, Berlin", 2 exs. (cSch, cKoc); "CHINA: Yunnan, Nujiang Lisu Pref., Gaoligong Shan, E pass 20 km NW Liuku, 25°58'49''N, 98°41'48''E, 3000 m, creek valley with devast. prim. forest, litter & moss sifted, 3.IX.2009, leg. M. Schülke [CH09-25]", 1 ex. (cSch); "CHINA (Yunnan) Nujiang Lisu Aut. Pref., Gaoligong Shan, "Cloud Pass" 21 km NW Liuku, 25°58'21''N, 98°41'01''E, 3150 m, shrubs & bamboo, litter sifted, 2.IX.2009, leg. M.Schülke [CH09-22] / Sammlung M. Schülke, Berlin", 5 exs. (cSch, cKoc); "CHINA (Yunnan) Nujiang Lisu Aut. Pref., Gaoligong Shan, "Cloud Pass" 3150 m, 21 km NW Liuku (shrubs, Vaccinum, bamboo, litter sifted) 25°58'21''N/ 98°41'01''E, 2.IX.2009, D. W. Wrase [22A] / Sammlung M. Schülke, Berlin", 1 ex. (cSch). Distribution. Northwestern Yunnan (Fig. 8). The species is widely distributed along the northern and central parts of the Gaoligong Shan range. One specimen was collected at the opposite side of the Salween river in the Nu Shan range. Remarks. Ischnosoma zipapertum sp. n. is the second species of the I. spelaeum species group (Kocian, 1997) recorded from China. It is distinguished from all other Chinese species, except I. taibaiense, by the long ocular seta (as long and stout as the setae of the pronotum), small eyes (shorter than temples), a broader head in relation to pronotum, shorter elytra, and generally smaller body. The modifications of the male sternites VII and VIII are unique, among the Chinese species of Ischnosoma including the only described Chinese species of the I. spelaeum species group: I. taibaiense (see Zhou et al. 2005: 810, figure 3, 4).

Ischnosoma vicinum sp. n. Figs 2, 9–12, 54.

Description. Body (Fig. 2) blackish-brown. Posterior margins of pronotum, elytra and tergites slightly paler, reddish-brown. Legs, palpi and antenna brownish-yellow, with the middle antennomeres brownish. Pronotum, elytra and abdomen with blueish hue. Head 0.65–0.73 mm broad; 0.52–0.58 times as broad as pronotum, broader than long. Eyes slightly longer than temples. Surface with distinct reticulate and undulate microsculpture. Antenna slender, 1.69–1.99 mm long. Antennomere X as broad as long or slightly longer than broad. Pronotum 1.03–1.31 mm broad; 0.93–1.03 mm long; 1.2–1.3 times as broad as long, broadest in posterior half. Surface with distinct transverse microsculpture, finely and sparsely punctate. Posterior angles distinct. Lateral margin near lateral macropunctures with some additional small punctures and setae. Elytra quadrangular, very slightly widened posteriad; along suture 0.86–0.98 mm long; 1.02–1.11 times as broad as pronotum; 0.9–0.95 times as long as pronotum. Surface with very fine transverse microsculpture. Sutural row with 9–13 punctures, discal row with 11–13 punctures, and lateral row with 11–14 punctures. Wings reduced. Abdomen slightly tapering towards apex, densely puntate. Tergite VII without palisade fringe. Surface smooth with very fine transverse microsculpture. Length of forebody 2.53–2.78 mm; total body length 5.0–5.8 mm.

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 207 FIGURES 9–11. Ischnosoma vicinum sp. n. Posterior portion of male sternite VII (9); posterior portion of male sternite VIII (10); aedeagus in ventral (left) and lateral view (11). Scale bars: 0.1 mm: a (Figs 9–10), b (Fig. 11).

FIGURE 12. Distribution of Ischnosoma vicinum sp. n. (square), Ischnosoma involutum sp. n. (triangle) and Ischnosoma gemellum sp. n. (circles) in China.

208 · Zootaxa 4105 (3) © 2016 Magnolia Press KOCIAN & SCHÜLKE Male. Sternite VII in the middle of posterior margin broadly concave, without modified pubescence (Fig. 9). Sternite VIII with posterior margin deeply triangularly concave in the middle, without setae in posterior portion, only anteriorly and in posterior angles with modified, thick and flattened setae. Medial area very sparsely punctate with short thin setae (Fig. 10). Paramere and aedeagus see Fig. 11. Etymology. The specific name (Latin, adjective: near, neighboring) alludes to the similarity of this species with I. farkaci. Type material. Holotype, male: "P.R.CHINA, Yunnan, E slope Cangshan at Dali, N25°40'13.2'', E100°07'54.8'', 11.V.2010, 2728 m , sifting05, V. Grebennikov" (CAS). Paratypes (3): "CHINA: NW-Yunnan, Bailakou pass, 3750 m, 28.V.–7.VI.2006, S.Murzin & I.Shokhin / Sammlung M. Schülke, Berlin", 1 ex. (cKoc); "CHINA: Yunnan (CH07-03), Dali Bai Auton. Pref., Diancang Shan W Dali, 25°41'49''N, 100°06'24''E, 2970 m, sifted at rock edges and under small shrubs, 28.V.2007, M. Schülke / Sammlung M. Schülke, Berlin", 2 exs. (cSch). Distribution. Northwestern Yunnan (Fig. 12): Diancang Shan and Haba Shan ranges. Remarks. I. vicinum sp. n. is very closely related to I. farkaci Kocian. Both species differ from other species of the I. pictum group by the combination of the following characters: body dark coloured, blackish-brown to blackish, pronotum broad with conspicuous posterior angles, broadest in posterior fourth, elytra shorter than pronotum, humeral angles marked, head and pronotum with very conspicuous microsculpture. Lateral pronotal margins of both species often with some additional short setae besides long lateral setae. Posterior margin of male sternite VIII in both species very deeply triangularly concave. I. vicinum sp. n. differs from I. farkaci by generally smaller body, paler coloration and the modifications of the male sternite VIII (modified palisade setae only anteriorly and posteriorly), while in I. farkaci these setae are also present laterally.

Ischnosoma foedum sp. n. Figs 13, 16–19, 46, 55.

Description. Body (Fig. 13) reddish-brown to brownish. Head (except mouthparts), central portion of elytra, and anterior portions of tergites (especially tergite V to VII) darker, balckish. Pronotum in anterior and median portions often with darker spot. Expansion of dark colour is variable. Legs, antennae and palpi yellow to brownish-yellow, antennae often darker, reddish-brown. Elytra with a slight greenish hue. Head 0.51–0.58 mm broad; 0.55–0.60 times as broad as pronotum. Surface smooth without microsculpture, rarely with very inconspicuous microsculpture. Antenna slender, 1.26–1.46 mm long. Antennomere X as long as broad or slightly longer than broad. Pronotum 0.88–1.01 mm broad; 0.73–0.83 mm long; 1.2–1.27 times as broad as long. Surface shiny with very fine, more or less indistinct microsculpture, finely and sparsely punctate. Elytra distinctly widened posteriad, along suture 0.68–0.79 mm long; 1.09–1.2 times as broad as pronotum; 0.93–0.98 times as long as pronotum. Surface with very fine transverse microsculpture. Sutural row with 8 to 10 punctures, discal row with 7 to 10 punctures, and lateral row with 7 to 10 punctures. Hind wings reduced. Abdomen gradually tapering towards apex, very sparsely puntate. Latero-tergites and tergites apically with long, stout, dark setae. Tergite VII without palisade fringe. Surface smooth without distinct microsculpture. Length of forebody 1.87–2.27 mm; total body length 3.6–4.8 mm. Male. Sternite VII with posterior margin slightly concave, along central part of posterior margin with thin long setae (Fig. 16). Sternite VIII with areas of palisade and beard-like setae forming three to four rows; and with area of medial dense and posteriorly flattened and pointed setae. Setae in antero-lateral parts of sternite VIII longer, thin and diagonally postero-laterad (Fig. 17). Paramere basally stout. Internal structure of aedeagus see Fig. 18. Etymology. The specific epithet (Latin adjective: ugly, ghastly) alludes to the indistinctive habitus of this species. Type material. Holotype, male: "P.R. CHINA, Yunnan, E slope Cangshan at Dali, N25°40'15.5'', E100°07'45.4'', 18.V.2010, 2657 m, sifting 17, V. Grebennikov / coll. Smetana (CAS)". Paratypes (52): same data as the holotype (4 exs.) (cKoc, cSch, cSme); "P.R. CHINA, Yunnan, E slope Cangshan at Dali, N25°40'15.1'', E100°07'39.9'', 10.V.2010, 2711 m, sifting 04, V. Grebennikov", 3 exs. (CNC, cKoc); "P.R. CHINA, Yunnan, E slope Cangshan at Dali, N25°40'13.2'', E100°07'54.8'', 11.V.2010, 2728 m, sifting

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 209 05, V. Grebennikov", 11 exs. (CNC, cKoc); "CHINA: N-Yunnan Dali Bai Nat. Aut. Pref. Diancang Shan 3 km E Dali, 25°41'N, 100°07'E, 29.VIII.2003, 2700 m, A. Smetana [C138]", 1 ex. (cSme); "CHINA: N-Yunnan Dali Bai Nat. Aut. Pref. Diancang Shan 5 km SSW Dali, 25°38.7'N, 100°08.3'E, 26.VIII.2003, 2800 m, A. Smetana [C136]", 1 ex. (cSme); "CHINA: Yunnan, above Dali, 2700–2900 m, 14.IV.1999, leg. W. Schawaller", 2 exs. (SMNS); "CHINA: Yunnan, above Dali, 2500–2700 m, 8.–18.IV.1999, leg. W. Schawaller", 4 exs. (SMNS, cKoc); "CHINA (N-Yunnan), Dali Bai Nat. Aut. Pref., Diancang Shan, 5 km SSW Dali old town, creek valley above cablecar, 2800 m, 25°38.7'N, 100°08.3'E (shrubs, bamboo, moss, old flood debries), 26.VIII.2003, Wrase [15] / Sammlung M. Schülke, Berlin", 5 exs. (cSch, cKoc); "CHINA (N-Yunnan), [C03-15] Dali Bai Nat. Aut. Pref., Diancang Shan, 5 km SSW Dali old town, creek valley above cablecar, 2800 m, 25°38.7'N, 100°08.3'E (shrubs, bamboo, moss, old flood debries), 26.VIII.2003, M. Schülke / Sammlung M. Schülke, Berlin", 2 exs. (cSch); "CHINA (N-Yunnan), [CH07-03] Dali Bai Nat. Aut. Pref., Diancang Shan W Dali, 25°41'49''N, 100°06'24''E, 2970 m, sifted at rock edges and under small shrubs, 28.V.2007, M. Schülke / Sammlung M. Schülke, Berlin", 1 ex. (cSch); "CHINA (N-Yunnan), [CH07-05] Dali Bai Nat. Aut. Pref., Diancang Shan W Dali, 25°41'33''N, 100°06'36''E, 2927 m, litter at moist escarpment sifted, under stones, 28.V.2007, M.Schülke / Sammlung M. Schülke, Berlin", 3 exs. (cSch, cKoc); "CHINA: Yunnan, Dali Bai Aut. Pref., Zhemo Shan, 7 km SW Xiaguan, 25°32–33'N, 100°10–11'E, 2870–2970 m, scrub with bamboo, oaks & Rhododendr., litter sifted, 18.IX.2009, leg. M.Schülke [CH09-60] / Sammlung M. Schülke, Berlin", 6 exs. (cSch, cKoc); "CHINA: Yunnan, Dali Bai Aut. Pref., Zhemo Shan, 7 km SW Xiaguan, 2870–2970 m (scrubs with bamboo, oaks, Rhododendr., litter sifted), 25°32–33'N, 100°10–11'E, 18.IX.2009, D.W.Wrase [60] / Sammlung M. Schülke, Berlin", 8 exs. (cSch, cKoc); "CHINA: Yunnan, Dali Bai Aut. Pref., Jizu Shan, summit plateau, 37 km NE Dali, 3150 m, (mixed forest, sifted from litter, moss), 25°58'30''N, 100°21'36''E, 5.IX.2009, D.W.Wrase [28] / Sammlung M. Schülke, Berlin", 1 ex. (cSch).

FIGURES 13–15. Habitus of Ischnosoma foedum sp. n. (13); Ischnosoma gemellum sp. n. (14); Ischnosoma involutum sp. n. (15).

210 · Zootaxa 4105 (3) © 2016 Magnolia Press KOCIAN & SCHÜLKE FIGURES 16–18. Ischnosoma foedum sp. n. Male sternite VII (16); male sternite VIII (17); apical portion of aedeagus in ventral (left) and aedeagus in lateral view (18). Scale bars: 0.1 mm: a (Figs 16–17), b (Fig. 18).

FIGURE 19. Distribution of Ischnosoma foedum sp. n. (diamonds), Ischnosoma abstrusum sp. n. (circles) and Ischnosoma yunnanum sp. n. (squares) in China.

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 211 Distribution. Northwestern Yunnan (Fig. 19): mountain ranges arround Er Hai Lake (Diancang Shan, Zhemo Shan, Jizu Shan). Remarks. I. foedum sp. n. belongs to the I. pictum species group. It differs from the other Chinese species with a similar chaetotaxy of the male sternite VIII by shorter, posteriorly distinctly widened elytra with less than 10 discal punctures, completely reduced hind wings, shorter antennae, a generally shinier body surface, and the shape of the large internal sclerite of the aedeagus (broad in lateral view and blunt apically; Fig. 18).

Ischnosoma involutum sp. n. Figs 12, 15, 20–22, 51, 60.

Description. Pronotum blackish-brown with lighter postero-lateral and posterior margins; elytra pale brown with blackish-brown area in the middle; head and abdomen blackish; posterior margins of tergites brownish-yellow; mouthparts, palpi, legs and basal two antennomeres yellow, antennomere III–XI blackish. Darker parts of pronotum, elytra, and tergites with weak blueish hue (Fig. 15). Head 0.59 mm broad; 0.65 times as broad as pronotum, broader than long, narrower posteriad, broadest in anterior half. Eyes very large and bulbous, longer than temples. Surface smooth without microsculpture. Antenna stout and long, antennomere X distinctly longer than broad. Pronotum 0.91 mm broad; 0.75 mm long; 1.2 times as broad as long, broadest in the middle. Surface with very fine transverse microsculpture, finely and sparsely punctate.

FIGURES 20–22. Ischnosoma. involutum sp. n. Male sternite VII (20); male sternite VIII (21); aedeagus in ventral (left) and lateral view (22). Scale bar: 0.1 mm: a (Figs 20–21), b (Fig. 22).

212 · Zootaxa 4105 (3) © 2016 Magnolia Press KOCIAN & SCHÜLKE Elytra distinctly widened posteriad; along suture 1.06 mm long; 1.33 times as broad as pronotum; 1.4 times as long as pronotum. Surface with very fine transverse microsculpture. Sutural row with 17 punctures, discal row with 17 punctures and lateral row with 18 punctures. Hind wings fully developed. Abdomen gradually tapering towards apex, sparsely puntate. Tergite VII with palisade fringe. Surface smooth with very fine microsculpture. Length of forebody 2.47 mm; total body length 4.15 mm (up to abdominal segment VI). Male. Sternite VII at posterior margin slightly concave, along central part of posterior margin with fine long setae (Fig. 20). Sternite VIII with areas of palisade and beard-like setae forming three rows; with central acute seta; and with area of medial dense setae (Fig. 21). Paramere and aedeagus: Fig. 22. Etymology. The specific epithet (Latin adjective: intricate, obscure) alludes to the extreme similarity of the habituswith some other species of this species group. Type material. Holotype, male: "CHINA N Yunnan, Xue Shan nr. Zhongdian 3800 m 26.VI.1996 27°49N 99°34E C43 / collected by A. Smetana, J. Farkač and P. Kabátek" (cSch). Distribution. Northwestern Yunnan (Fig. 12): Xue Shan range. Remarks. I. involutum sp. n. belongs to the species the I. pictum group, which is characterized by the following character combination: male eyes protuberant and bulbous; head widened anteriad; antenna long; antennomeres VIII to XI distinctly longer than wide; pronotum narrow, in posterior half with parallel lateral margins; elytra long and conspicuously widened posteriad, with numerous discal punctures. The new species differs from all the similar species by the shape and chaetotaxy of the male sternite VIII (with single pointed modified seta in the middle of the posterior margin between the clusters of beard-like and palisade setae.

Ischnosoma yunnanum sp. n. Figs 3, 19, 23–26, 53, 58.

Description. Body (Fig. 3) blackish, anterior margin of pronotum, humeral spot and posterior margin of elytra, posterior margins of tergites, legs, palpi and basal two antennomeres brownish-yellow. Pronotum, elytra, and abdomen with distinct blue-goldish hue. Head 0.65–0.68 mm broad; 0.62–0.65 times as broad as pronotum, distinctly broader than long, narrower posteriad, broadest in the middle. Eyes very large and bulbous, more than twice as long as temples. Surface smooth without microsculpture. Antenna stout and very long (1.84–2.12 mm). Antennomere X distinctly longer than broad. Pronotum 1.03–1.04 mm broad; 0.88 mm long; 1.17–1.18 times as broad as long. Posterior half of pronotum parallel-sided, posterior angles acute. Surface with very fine transverse microsculpture, finely and sparsely punctate. Elytra distinctly widened posteriad; along suture 1.21 mm long; 1.3–1.41 times as broad as pronotum; 1.37 times as long as pronotum. Surface with very fine transverse microsculpture. Sutural row with 16–18 punctures, discal row with 17–18 punctures and lateral row with 17–18 punctures. Hind wings fully developed. Abdomen gradually tapering towards apex, sparsely puntate. Tergite VII with palisade fringe. Surface smooth with very fine microsculpture. Length of forebody 2.9 mm; total body length 5.0–5.6 mm. Male. Sternite VII at posterior margin distinctly and broadly concave, along central part of posterior margin with thin long setae (Fig. 23). Sternite VIII with areas of palisade and beard-like setae forming six rows, these rows contiguous in the middle, and with medial area of very dense setae (Fig. 24). Paramere and aedeagus: Figs 25–26. Etymology. The specific epitheth is an adjective derived from Yunnan, the Chinese province where this species was collected. Type material. Holotype, male: "CHINA (Yunnan) Nujiang Lisu Pref., Gaoligong Shan, "Cloud pass" 3150 m, 21 km NW Liuku (shrubs, Vaccinium, bamboo, litter sifted), 25°58'21''N 98°41'01''E, 2.IX.2009, D.W.Wrase (22A) / Ischnosoma cf. rosti det. M. Schülke 2012 / Sammlung M. Schülke, Berlin" (cSch). Paratypes (2): "P.R.China, Yunnan, E slope N Gaoligongshan, N27°47'22.1'' E098°32'17.7'', 24.V.2010, 3027 m, sifting 21, V. Grebennikov", 1 ex. (CNC). "CHINA: N-Yunnan [C2005-16] Nujiang Lisu Aut. Pref., Gongshan Co., Gaoligong Shan, side valley, 3000–3050 m, 27°47.90'N, 98°30.19'E / conif. forest with Rhododendron, broad

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 213 leaved bushes, litter, moss, dead wood sifted along creek and snowfields, 21.VI.2005, leg. M. Schülke [C2005- 16]", 1 ex. (cKoc). Distribution. Northwestern Yunnan (Fig. 19): northern and central parts of the Gaoligong Shan range. Remarks. I. yunnanum sp. n. differs from all similar Chinese species of the I. pictum group by the chaetotaxy of the male sternite VIII. It is the only species with contiguous areas of beard-like and palisade setae.

FIGURES 23–26. Ischnosoma yunnanum sp. n. Male sternite VII (23); male sternite VIII (24); apical portion of aedeagus in ventral (left) and lateral view (25); extruded inner structure of aedeagus in lateral view (26). Scale bars: 0.1 mm: a (Figs 23–24), b (Figs 25–26).

Ischnosoma abstrusum sp. n. Figs 19, 27, 30–32, 52, 59.

Description. Body (Fig. 27) dark brownish, head blackish, anterior margin of pronotum, humeral spot, posterior margin of elytra, posterior margins of tergites, legs, palpi and first two antennomeres brownish-yellow. Pronotum, elytra, and abdomen with distinct bluish hue. Head 0.56–0.61 mm broad; 0.56–0.59 times as broad as pronotum, slightly broader than long. Eyes big, more than twice as long as temples. Temples behind eyes parallel. Surface smooth without microsculpture. Antenna relatively fine, 1.44–1.74 mm long. Antennomere X slightly longer than broad. Pronotum 0.96–1.1 mm broad; 0.75–0.92 mm long; 1.15–1.23 times as broad as long, broadest in posterior third. Surface with very fine transverse microsculpture, finely and sparsely punctate. Elytra slightly widened posteriad; along suture 0.98–1.18 mm long; 1.17–1.26 times as broad as pronotum; 1.19–1.30 times as long as pronotum. Surface with very fine transverse microsculpture. Sutural row with 12–16 punctures, discal row with 11–17 punctures and lateral row with 12–17 punctures. Wings fully developed. Abdomen gradually tapering towards apex, sparsely puntate. Tergite VII with palisade fringe. Surface smooth with very fine microsculpture. Length of forebody 2.9 mm; total body length 5.0–5.6 mm. Male. Sternite VII in the middle of posterior margin deeply concave, with area of very dense thin setae (Fig. 30). Sternite VIII with relatively long areas of palisade and beard-like setae forming six rows, and with median area of very dense setae (Fig. 31). Paramere and aedeagus: Fig. 32.

214 · Zootaxa 4105 (3) © 2016 Magnolia Press KOCIAN & SCHÜLKE Etymology. The specific epithet (Latin adjective: hidden, secret, concealed) alludes to the impossibility to distinguish this species from it's relatives based only on habitus characters.

FIGURES 27–29. Habitus of Ischnosoma abstrusum sp. n. (27); Ischnosoma duplicatum (Sharp), female (28); Ischnosoma duplicatum (Sharp), male (29).

Type material. Holotype, male: "CHINA: Yunnan, Lincang Pref., Xue Shan, 48 km N Lincang, 2070 m, 24°19'03''N, 100°07'13''E, forest remnant, N-slope, litter & mushrooms sifted, 12.IX.2009, leg. M. Schülke [CH09- 45] / Sammlung M. Schülke, Berlin" (cSch). Paratypes (22): "CHINA: Yunnan, Lincang Pref., Bangma Shan, E pass, 17 km NW Lincang, 23°57'31''N, 99°56'13''E, 2040 m , secondary pine forest, litter, dead wood & mushrooms sifted, 9.IX.2009, leg. M.Schülke (CH09-36) / Sammlung M. Schülke, Berlin", 1 ex. (cSch); "CHINA: Yunnan (CH07-19) Dehong Dai Aut. Pref., mountain range 31 km E Luxi, 2280 m, 24°29'31''N, 98°52'58''E, secnd. pine forest with old decid. Trees, litter sifted, 3.VI.2007, A. Pütz", 1 ex. (cPüt); "P.R.CHINA, Sichuan, Emei Shan, N29°33.6' E103°20.6', 27.VI.–5.VII.2009, 1800–2400 m, sifting 11–17, V.Grebennikov", 1 ex. (CNC); "CHINA [6]—Yunnan, NE Kunming, 25°08'35''N, 102°53'49''E, 2320 m, mixed forest sifted, 13.VIII.2014, V. Assing", 2 exs. (cAss, cKoc); "CHINA [10]—Yunnan mt. NW Wuding, mixed forest, 25°36'53''N, 102°18'59''E, 2190 m, 17.VIII.2014, V. Assing", 1 ex. (cAss); "CHINA [11]—Yunnan, Wuding, Lion Mountain, 2200 m, 25°31'59''N, 102°22'36''E, sifted, 17.VIII.2014, V. Assing", 1 ex. (cAss); "CHINA [9b]—Yunnan, mt. W Xundian, 2300 m, mixed for., 25°34'58''N; 103°08'42''E, sifted, 16.VIII.2014, V. Assing", 5 exs. (cAss, cSch, cKoc); "CHINA [4]—Yunnan, NE Kunming, 25°09'07''N, 102°53'46''E, 2280 m, sec. pine for. with alder 11.VIII.2014, V. Assing", 1 ex. (cSch); "CHINA [5]—Yunnan, NE Kunming, 25°08'40''N, 102'53'48''E, 2290 m, mixed forest, sifted, 11.VIII.2014, V. Assing", 1 ex. (cAss); "CHINA [9]—Yunnan, mt. W Xundian, 2300 m, mixed for., 25°34'58''N, 103°08'42''E, sifted, 15.VIII.2014, V. Assing", 1 ex. (cAss). TAIWAN: "Tengchih (1400 m) Kaohsiung, Taiwan, 25.–30.IV.2001, Sugaya H. leg. (FIT) [flight intercept

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 215 trap] / Ischnosoma spec. nov. det. M. Schülke 2002", 1 ex. (KUM); "Taiwan: Kaohsiung, Tengchih, 2100 m, 19.–23.IV.2001, leg. H. Sugaya", 4 exs. (KUM, cKoc, cSch); "Taiwan: Kaohsiung, Tengchih, 1400 m, 29–30.IV.2001, leg. H. Sugaya", 1 ex. (KUM); "Taiwan: Nantou, Tehuashe, 900 m, 4.–5.V.2001, leg. H. Sugaya", 1 ex. (KUM). Distribution. Widely distributed from Western Yunnan and Sichuan to Taiwan (Fig. 19). Remarks. I. abstrusum sp. n. differs from similar species of the I. pictum group by the shape of the male sternite VII (posterior margin in the middle deeply concave and bordered by a cluster of very dense modified, thin long setae. The only other species with a similarly shaped male sternite VII are I. discoidale discoidale (Sharp), I. discoidale taiwanum Kocian, and I. ruth Kocian, but these species are distinguished by smaller body size, fewer and sparser modified setae on the male sternite VII, and the different chaetotaxy of the male sternite VIII.

FIGURES 30–32. Ischnosoma abstrusum sp. n. Male sternite VII (30); male sternite VIII (31); aedeagus in ventral (left) and lateral view (32). Scale bars: 0.1 mm: a (Figs 30–31), b (Fig. 32).

Ischnosoma gemellum sp. n. Figs 12, 14, 33–35, 40, 57.

Description. Body (Fig. 14) distinctly two-colored, blackish and redish-yellow. Pronotum in posterior part laterally with reddish-yellow spot. Humeral spot, posterior margin of elytra, posterior margins of tergites, legs, palpi and first two antonnomers reddish-yellow. Elytra and abdomen with slight bluish hue. Head 0.55–0.56 mm broad; 0.57–0.59 times as broad as pronotum, as broad as long. Eyes big, distinctly longer than temples. Surface with distinct sparse microsculpture composed of transverse and obligue waves. Antenna thin, 1.51–1.61 mm long. Antennomere X slightly longer than broad.

216 · Zootaxa 4105 (3) © 2016 Magnolia Press KOCIAN & SCHÜLKE Pronotum 0.96 mm broad; 0.82–0.88 mm long; 1.08–1.16 times as broad as long, broadest behind middle. Surface with very fine transverse microsculpture, finely and sparsely punctate. Elytra slightly widened posteriad; along suture 1.03–1.06 mm long; 1.23–1.26 times as broad as pronotum; 1.2–1.26 times as long as pronotum. Surface with very fine transverse microsculpture. Sutural row with 12–18 punctures, discal row with 12–16 punctures and lateral row with 15–17 punctures. Wings fully developed. Abdomen gradually tapering towards apex, sparsely puntate. Tergite VII with palisade fringe. Surface smooth with very fine transverse microsculpture. Length of forebody 2.47–2.73 mm; total body length 4.5–5.1 mm. Male. Sternite VII in the middle of posterior margin broadly concave, with sparse thin setae (Fig. 33). Sternite VIII near posterior margin with area without pubescence, bordered by suture. Non-pubescent area at posterior margin enframed by beard-like setae forming two rows. Anterior to non-pubescent area with median cluster of short, dense, and pointed setae (Fig. 34). Paramere and aedeagus: Fig. 35. Etymology. The specific epithet (Latin, adjective: twin-born) alludes to the similarity of this species with I. bolitobioides. Type material. Holotype, male: "CHINA: Yunnan (CH07-14A), Baoshan, Pref. Gaoligong Shan, 33 km SE Tengchong, 2100–2200 m, 24°51'22''N, 98°45'36''E, decid forest, litter, wood, fungi sifted, 4.VI.2007, M.Schülke / Sammlung M. Schülke, Berlin" (cSch). Paratypes (2): "CHINA: N-Yunnan (C2005-07A), Diqing Tibet Aut. Pref., Deqin Co., Meili Xue Shan, E-side, 12 km SW Deqin, 2890 m, 28°25'30''N, 98°48'47''E / small creek valley, mixed forest with bamboo, leaf litter, moss, dead wood, sifted, 13.6.2005, leg. M.Schülke (C2005-07A) / Sammlung M. Schülke, Berlin", 1 ex. (cSme); "CHINA, Sichuan, Gongga Shan, Lake abv. Camp 2, 2750 m, 25.VII.1994, A. Smetana (C23)", 1 ex. (cKoc).

FIGURES 33–35. Ischnosoma gemellum sp. n. Male sternite VII (33); male sternite VIII (34); aedeagus in ventral (left) and lateral view (35). Scale bars: 0.1 mm: a (Figs 33–34), b (Fig. 35).

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 217 FIGURES 36–45. Male sternite VIII with position of dark setae, beard-like palisade setae and median dense setae, schematic. Ischnosoma evae Kocian (36); I. bohaci Kocian (37); I. fusciventre (Tikhomirova) (38); I. bolitobioides (Bernhauer) (39); I. gemellum sp. n. (40); I. absalom Kocian (41); I. farkaci Kocian (42); I. ruth Kocian (43); I. discoidale discoidale (Sharp)(44); I. discoidale taiwanum Kocian (45). Scale bar: 0.1 mm.

218 · Zootaxa 4105 (3) © 2016 Magnolia Press KOCIAN & SCHÜLKE FIGURES 46–53. Male sternite VIII with position of dark setae, beard-like palisade setae and median dense setae, schematic. Ischnosoma foedum sp. n. (46); I. noemi Kocian (47); I. splendidum (Gravenhorst) (48); I. maderi (Bernhauer) (49); I. duplicatum (Sharp) (50); I. involutum sp. n. (51); I. abstrusum sp. n. (52); I. yunnanum sp. n. (53). Scale bar: 0.1 mm.

Distribution. Western Yunnan (Gaoligong Shan, Meili Xue Shan) and Western Sichuan (Gongga Shan) (Fig. 12). Remarks. I. gemellum sp. n. is very closely related to I. bolitobioides, from which it differs by the chaetotaxy of the male sternite VIII (median cluster of modified setae larger and formed by two types of setae: broadly flattened setae bordering anterior margin of median non-pubescent area and short pointed setae anterior to these flattened setae. In I. bolitobioides, there are only broadly flattened setae arranged in some irregular rows anterior to non-pubescent area.

Unnamed and presumably undescribed species

Besides some other single female specimens of doubtful identity, the material studied contained two distinctive undescribed species, which are easily distinguished from all the named Chinese species, but which are not described for want of males.

Ischnosoma spec. 1

"CHINA: Yunnan, Baoshan Prefecture, mountain range 14 km E Tengchong, 1850 m, 25°00'28''N, 98°38'07''E, secondary mixed forest, litter sifted, 1.VI.2007, M. Schülke [CH07-16]", 3 ♀♀ (cSch). This brachypterous species resembles I. chinense regarding the irregular pubescence of the elytra, but is easily distinguished by shorter elytra with much finer and denser pubescence, and by the absence of an ocular seta.

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 219 FIGURES 54–57. Forebody. Ischnosoma vicinum sp. n. (54); I. foedum sp. n. (55); I. zipapertum sp. n. (56); I. gemellum sp. n. (57). Scale bar: 1 mm.

Ischnosoma spec. 2

"CHINA: N-Yunnan, Dali Bai Nat. Aut. Pref., Diancang Shan, 4 km W Dali old town, 25°41.4' N, 100°06.7' E, 2900–3000m, E slope with devastated forest and old pine forest, litter and mushrooms sifted, 31.VIII.2003, leg. M. Schülke [CH03-20]", 1♀ (cSch). This brachypterous species is characterized by small eyes, the ocular seta inserting in the median portion at some distance behind the eyes, the inner anterior seta of the pronotum inserting at some distance from the anterior margin of the pronotum, elytra with very distinct microsculpture, the absence of a carina at the anterior margin of the pronotum. These character combination suggests that this species is most likely not a member of the I. pictum group, but probably represents an undescribed species group.

New records of described species

Ischnosoma absalom Kocian, 2003 Figs 41, 61.

CHINA: Shaanxi: S-Shaanxi, SW Zhouzhi, Qingling Shan, 33°44'02''N, 107°58'06''E, 1900 m, NE-slope, stream

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 221 valley, mixed forest, litter and soil sifted, 25.VII.2012, leg. V. Assing, 2 exs. (cAss, cSch); same data, but "leg. D.W. Wrase [1], 1 ex. (cSch); S-Shaanxi, SW Meixian, Qingling Shan, 34°01'31''N, 107°24'13''E, 1870 m, N-slope, secondary deciduous forest, near stream, litter and grass sifted, 26.VII.2012, leg. V. Assing, 2 exs. (cAss, cSch); same data, but "leg. D.W. Wrase [2]", 4 exs. (cSch); same data, but "leg. M. Schülke [CH12-02]", 2 exs. (cSch); Qinling Shan, pass at road Zhouzhi–Foping, 105 km SW Xian, N-Slope, 1990 m, 33°44'N, 107°59'E, 4.VII.2001, leg. A. Smetana [C93], 3 exs. (cSme); same data, but "2.VII.2001 [C89]", 4 exs. (cSme, cKoc); same data, but "1880 m, 33°44'N, 107°58'E, 4.VII.2001 [C92]", 1 ex. (cSme); S-Shaanxi, Qinling Shan, mountain range W pass on road Xian–Shagoujie, 45 km SSW Xian, 33°52'N, 108°46'E, 2600 m, 25.VII.2001, leg. A. Smetana [C118], 1 ex. (cSme); Yunnan: Lijiang, Naxi Autonomous County, E Yulonxue Shan, 30 km N Lijiang, 27°09.0'N, 100°14.9'E, 2800–2900 m, 13.VIII.2003, leg. A. Smetana [C120], 1 ex. (cSme). Remarks. Ischnosoma absalom was described from the central parts of the Qinling Shan range, where the species is widely distributed and relatively common. The above specimen from Yunnan represents a new province record (Fig. 61).

FIGURE 61. Distribution of Ischnosoma absalom Kocian (circles) and I. chinense (Bernhauer) (triangles) in China (material examined and literature records).

Ischnosoma bohaci Kocian, 2003 Figs 37, 62.

CHINA: Gansu: S-Gansu, N Chengxian, W-Qingling Shan, 34°08'16''N, 105°46'42''E, 1760 m, N-slope, secondary deciduous forest margin, sifted, 28.VII.2012, leg. V. Assing, 23 exs. (cAss, cSch); same data, but "leg. M. Schülke [CH12-04], 3 exs. (cSch); S-Gansu, N Chengxian, W-Qingling Shan, 34°10'20''N, 105°42'19''E, 1830 m, stream valley with secondary deciduous forest, moist litter sifted, 29.VII.2012, leg. V. Assing, 4 exs. (cAss, cSch); same data, but "leg. M. Schülke [CH12-06], 4 exs. (cSch); S-Gansu, N Chengxian, W-Qingling Shan, 34°08'16''N, 105°46'42''E, 1760 m, heap of rotting plants, sifted, 28.VII.2012, leg. V. Assing, 1 ex. (cAss); S- Gansu, N Chengxian, W-Qingling Shan, 34°10'17''N, 105°42'56''E, 1850 m, mixed secondary forest margin, litter sifted, 29.VII.2012, leg. V. Assing, 2 exs. (cAss, cSch); S-Gansu, mountains 36 km SE Longnan, 33°13'20''N, 105°15'10''E, 2170 m, N-slope with shrubs and scattered coniferous trees, litter & mushrooms sifted, 31.VII.2012, leg. M. Schülke [CH12-07], 3 exs. (cSch); S-Gansu, W-Qinling Shan, 101 km NW Longnan, 34°03'14''N, 104°10'00''E, 2200 m, SW-slope with shrubs, litter sifted, 1.VIII.2012, leg. M. Schülke & D.W. Wrase [CH12-09], 2 exs. (cSch); S-Gansu, W Qingling Shan, 125 km NW Longnan, Lazikou pass, S-side, Zhuli valley, 34°07'57''N, 103°56'15''E, 2260 m, N-slope, mixed forest with oak and pine near creek, litter and dead wood sifted, 3.VIII.2012,

222 · Zootaxa 4105 (3) © 2016 Magnolia Press KOCIAN & SCHÜLKE leg. D.W. Wrase [CH12.11], 1 ex. (Sch); Shaanxi: S-Shaanxi, SW Meixian, Qingling Shan, 34°01'31''N, 107°24'13''E, 1870 m, N-slope, secondary deciduous forest, near stream, litter and grass sifted, 26.VII.2012, leg. V. Assing, 1 ex. (cAss); same data, but "leg. M. Schülke [CH12-02], 3 exs. (cSch); Taiwan: Kaohsiung Hsien, Kuanshan trail, st. Kaunshanchi Riv., 2400 m, 20.IV.1992, leg. A. Smetana [T94], 1 ex. (cSme). Remarks. Ischnosoma bohaci is widely distributed in central and south-eastern China including Taiwan (Fig. 62).

FIGURE 62. Distribution of Ischnosoma bohaci Kocian in China (material examined and literature records).

Ischnosoma bolitobioides (Bernhauer, 1923) Figs 39, 63.

CHINA: Gansu: S-Gansu, N Chengxian, W-Qinling Shan, 34°08'16''N, 105°46'42''E, 1760 m, N-slope, secondary deciduous forest margin, sifted, 28.VII.2012, leg. V. Assing, 3 exs. (cAss, cSch); Hubei: SW-Hubei, 30 km NE Hefeng, Mulinzi, 30.1N, 110.2E, 23.–24.V.2004, leg. J. Turna, 3 exs. (NHMW, cSch); W-Hubei, Muyuping S env., 31.45'N, 110.4°E, 1300 m, 18.V.2004, leg. J. Turna, 1 ex. (NHMW); Shaanxi: S-Shaanxi, SW Zhouzhi, Qinling Shan, 33°44'02''N, 107°58'06''E, 1900 m, NE-slope, stream valley, mixed forest, litter and soil sifted, 25.VII.2012, leg. V. Assing, 1 ex. (cAss); Qinling Shan, pass between Zhouzhi and Foping, 105 km SW Xian, N-slope, 1990 m, 33°44'N, 107°59'E, 2./4.VII.2001, leg. M. Schülke, 2 exs. (cSch); S-Shaanxi, SW Zhouzhi, Qinling Shan, 33°44'02''N, 107°58'06''E, 1900 m, 25.VII.2012, leg. V. Assing, 1 ex. (cAss); S-Shaanxi [CH12-01] Qinling Shan, 52 km SSW Zhouzhi, 33°44'02''N, 107°58'06''E, 1900 m, NE-slope, creek valley, mixed forest, litter and soil sifted, 25.VII.2012, leg. M. Schülke, 2 exs. (cSch). Sichuan: 160 km S Chengdu, Emei Shan, 1400 m, 28.VI.1991, leg. C. Holzschuh & Ji, 8 exs. (NHMW, cKoc); Emei Shan, 160 km SSW Chengdu, 1000–1700 m, 23.VI.1994, leg. H. Schillhammer, 5 exs. (NHMW, cKoc); same data, but "1530–1700 m, 22.VI.1994, leg. C. Holzschuh", 1 ex. (NHMW); S-Sichuan, S Xichang, Luoji Shan, 2300–2500 m, litter sifted, 16.–24.VII.1996, leg. S. Kurbatov, 1 ex. (MHNG); Yunnan: Nujiang Lisu Aut. Prefecture, Gongshan, Gaoligong Shan, 2500 m, 27°45.404'N, 98°35.749'E, litter and dibris sifted at snowfield edge during heavy rain, 19.VI.2005, leg. M. Schülke, 1 ex. (cSch); Nujiang Lisu Aut. Prefecture, Gaoligong Shan, E pass 20 km NW Liuku, 3000 m, creek valley with devastated primary forest, 25°58'49''N, 98°41'48''E, 3.IX.2009, leg. D.W. Wrase, 1 ex. (cSch); Nujiang Lisu Aut. Prefecture, Gaoligong Shan, side valley, 18 km NW Liuku, 2500 m, 25°58'10''N; 98°42'27''E, devastated primary forest, 9./10.VI.2007, leg. D. W. Wrase, 1 ex. (cSch); Nujiang Lisu Aut. Prefecture, Nu Shan, 7 km NNW Coajian, 25°43'29''N, 99°07'57''E, secondary pine forest, litter and bark sifted, 11.VI.2007, leg. M. Schülke, 1 ex. (cSch); Nujiang Lisu Aut. Prefecture, Gaoligong Shan, creek valley 20 km NW Liuku, 3000 m, 25°58'49''N, 98°41'48''E, bamboo and litter

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 223 sifted, 9.VI.2007, leg. M. Schülke, 1 ex. (cSch); Nujiang Lisu Aut. Prefecture, Gaoligong Shan, side valley, 19 km NW Liuku, 2730 m, 25°59'02''N, 98°42'23''E, devastated primary forest, litter sifted, 9.VI.2007, leg. M. Schülke, 1 ex. (cSch); Dali Bai Autonomous Prefecture, Diancang Shan W Dali, 25°41'47''N, 100°06'32''E, 3016 m, moist escarpment, litter sifted, 28.V.2007, leg. M. Schülke, 2 exs. (cSch); N-Yunnan, E-slope N Gaoligong Shan, 27°59'11.6''N, 98°33'25.3''E, 3086 m, sifting 24, leg. V. Grebennikov, 1 ex. (CNC); Dali, E-slope Cangshan, 25°40'13.2''N; 100°07'54.8''E, 2728 m, sifting 01, 9.V.2010, leg. V. Grebennikov, 2 exs. (CNC); Dali, E-slope Cangshan, 25°40'15.1''N; 100°07'39.9''E, 2728 m, sifting 04, 2711 m, 10.V.2010, leg. V. Grebennikov, 1 ex. (CNC); Lincang Prefecture, Wuliang Shan, old pass road, W side, 24°42'58.6''N, 100°29'52.0''E, 2200 m, small creek valley with primary forest remnant, litter and debris sifted, 12.IX.2009, leg. M. Schülke [CH09-47], 1 ex. (cSch). TAIWAN: Ilan Hsien: 17 km 100 Logging road, 1650 m, flight intercept trap, 22.VII.2003, leg. Chun Lin Li, 1 ex. (MNB); Kaohsiung Hsien: Peinantashan trail, 2500 m, 3.V.1995, leg. A. Smetana [T171], 1 ex. (cSme); Nantou Hsien: Meifeng, 2100m, 6.V.2001, leg. H. Sugaya, 4 exs. (KUM, cSch); Meifeng, 2100m, 26.X.2001, leg. H. Sugaya, 3 exs. (KUM, cSch); Pingtung Hsien: Peitawushan, Kuai-Ku Hut, 2130 m, 27.IV.1992, leg. A. Smetana [T101], 2 exs. (cSme, cSch); Taichung Hsien: Anmashan, 2150 m, 13.V.1992, leg. A. Smetana [T129], 1 ex. (cSme); Taoyuan Hsien: Chianshan, 1200m, 20.XI.2000, leg. H. Sugaya, 1 ex. (KUM); Kaohsiung Hsien: Tengchih, 19.IV.2001, leg. H. Sugaya, 1 ex. (KUM); Tengchih, 19.–23.IV.2001, leg. H. Sugaya, 13 exs. (KUM, cSch). Remarks. Ischnosoma bolitobioides, originally described from Japan (Bernhauer, 1923) and subsequently recorded from Taiwan (Cameron, 1949), was recently recorded also from mainland China (Fujian, Sichuan) and Thailand by Kocian (2003). The records presented above suggest that the species is widely distributed in the central and southern parts of mainland China (Fig. 63). The records from Gansu, Hubei, Shaanxi and Yunnan are new province records.

FIGURE 63. Distribution of Ischnosoma bolitobioides (Bernhauer) in China (material examined and literature records).

Ischnosoma chinense (Bernhauer, 1939) Fig. 61.

CHINA: Zhejiang: West Tianmu Shan N. R., way to peak of immortals, 30°20'34''N, 119°25'51''E, 1100–1200 m, primary mixed forest, litter and moss sifted, 15.VI.2007, leg. A. Pütz [CH07-36], 3 exs. (cPüt, cKoc); same data, but "leg. M. Schülke", 1 ex. (cSch); same data, but "leg. D.W. Wrase [36], 2 exx (cSch); Hangzhou Pref., Tianmu Shan, S pass 41 km WNW Linan, NE valley, 350 m, 30°21'34''N, 119°19'01''E, plant refuse near creek, 17.VI.2007

224 · Zootaxa 4105 (3) © 2016 Magnolia Press KOCIAN & SCHÜLKE leg. D.W.Wrase [38], 1 ex. (cSch); Hangzhou Pref., Tianmu Shan, 40 km WNW Linan, water reservoir, 300 m, 30°20'56''N, 119°18'42''E, litter from rock edges, sifted, 17.VI.2007 leg. D.W.Wrase [39], 18 exs. (cSch). Remarks. All recent records (Kocian, 2003 and this paper) are in the immediate vicinity of the type locality.

Ischnosoma convexum (Sharp, 1888) Fig. 64.

CHINA: Fujian: Wuji Shan Nat. Res., Sangan env., 900 m, 30.V.–12.VI.2001, leg. P. Hlaváč & J. Cooter, 1 ex. (cSch); Sichuan: S-Sichuan, env. Xichang, 1600 m, 28.VII.1996, leg. S. Kurbatov, 1 ex. (MHNG); W-Sichuan, Aba Tibet. Aut. Pref., Weizhou Co., Quionglai Shan, Wolong valley, 20 km WNW Dujiangyan, 1100 m, 31°05'N, 103°26'E, brook bank, 14.VII.1999, leg. D.W. Wrase, 1 ex. (cSch); Yunnan: Dali Bai Aut. Pref., 36 km N Dali, ruderal pasture with pines and shrubs, 26°01'20''N, 100°08'14''E, 2158 m, litter sifted under pines and shrubs, 24.VIII.2009, leg. M. Schülke [CH09-04], 1 ex. (cSch); N-Yunnan, Nujiang Lisu Aut. Pref., Gongshan Co., Gaoligong Shan, small creek valley 17 km N Gongshan, 27°54.41'N, 98°39.55'E, 1525–1600 m along creek under stones / gravel / leaf litter sifted, 20.VI.2005, leg. M. Schülke [C2005-15], 1 ex. (cSch); Zheijiang: Hangzhou Pref., Tianmu Shan, 40 km WNW Linan, water reservoir, 30°20'56''N, 119°18'42''E, 300 m, plant refuse, litter from rock edges, 17.VI.2007, leg. M. Schülke [CH07-39], 3 exs. (cSch, cKoc); Tianmu Shan, pass 25 km NNW Linan, 620–820 m, 30°25'40''N/ 119°35'30''E, creek valley, bamboo, mixed forest, litter sifted, 16.VI.2007 leg. D.W. Wrase [37], 1 ex. (cSch); Remarks. Ischnosoma convexum is widely distributed in Asia, from India eastwards to Japan, Indonesia and the Solomon Islands. In China, the distribution seems to be restricted to the southern half of the country (Fig. 64). The above records from Fujian, Sichuan, and Zhejiang are new province records.

FIGURE 64. Distribution of Ischnosoma convexum (Sharp) in China (material examined and literature records).

Ischnosoma discoidale discoidale (Sharp 1888) Figs 44, 65.

CHINA: Gansu: S-Gansu [CH12-06] W-Qinling Shan, 47 km N Chengxian, 34°10'20''N, 105°42'19''E, 1830 m, creek valley with secondary deciduous forest, moist litter sifted, 29.VII.2012, leg. M. Schülke, 2 exs. (cSch); Hubei: Daba Shan, Bachtal 11 km NW Muyuping, 31°30'N, 110°22'E, 1960 m, 18.VII.2001, Mischwaldgesiebe, leg. M. Schülke, 1 ex. (cSch); Daba Shan, Pass E mt. Da Shennonggija, 12 km NW Muyuping, 31°38'N, 110°21'E,

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 225 16.–22.VII.2001, 1950–2050 m, mixed forest, litter, moss, dead wood and mushrooms sifted, leg. M. Schülke & D.W. Wrase, 6 exs. (cSch); Shaanxi: Qinling Shan, pass at road Zhouzhi > Foping, 105 km SW Xian, N-slope, 1700 m, 33°46'N, 107°58'E, 3.VII.2001, leg. A. Smetana [C91], 3 ex. (cSme, cKoc); Shaanxi/Chongqing: border Shaanxi-Sichuan [now Shaanxi-Chongqing], Daba Shan, pass 20 km SSE Zhanping, 1700–1800 m, 31°44'N, 109°35'E, 12.VII.2001, leg. A. Smetana [C101], 1 ex. (cSme); Sichuan: Emei Shan, 29°33.6'N, 103°20.6'E, 1800–2400 m, sifting 11–17, 27.VI.–5.VII.2009, leg. V. Grebennikov, 1 ex. (CNC). Remarks. Ischnosoma discoidale discoidale, originally described from and common in Japan, is recorded from parts of central China for the first time (Fig. 65). The species was doubtfully recorded also by Li (1993).

FIGURE 65. Distribution of Ischnosoma discoidale discoidale (Sharp) (squares), I. discoidale taiwanum Kocian (circles) and I. ruth Kocian (triangle) in China (material examined and literature records).

Ischnosoma discoidale taiwanum Kocian, 2003 Figs 45, 65.

TAIWAN: Ilan Hsien: 76 km N Crossing Highway, 1180 m, flight intercept trap, 3.–4.IX.2002, leg. Chun Lin Li, 1 ex. (MNB); Kaohsiung Hsien: Peinantashan trail, 2450 m, 2.V.1995, leg. A. Smetana [T170], 1 ex. (cSch); Peinantashan trail, 2500 m, 3.V.1995, leg. A. Smetana [T171], 1 ex. (cSme); Taichung Hsien: Anmashan, 2700 m, litter, 8. IV. 2002, leg. H. Sugaya, 6 exs. (KUM, cSchü). Remarks. This brachypterous subspecies of Ischnosoma discoidale is restricted to Taiwan (Fig. 65).

Ischnosoma duplicatum (Sharp, 1888) Figs 28–29, 50, 66.

CHINA: Fujian: Wuyi Shan Nat. R., Sangan env., 900 m, 30.V.–12.VI.2001, leg. P. Hlaváč & J. Cooter, 1 ex. (cSch); Hubei: Daba Shan, Pass E Da Shennongjia, 12 km NW Muyuping, 31°36'N, 110°21'E, creek valley, 1950–2050 m, deciduous forest, dead wood, moss and mushrooms sifted, 19.VII.2001, leg. M. Schülke, 1 ex (cSch); Sichuan: Ya'an Prefecture, Shimian County, Xiaoxiang Ling, side valley above Nanya Cun near Caluo, 11 km S Shimian, 1250 m, 7.VII.1999, leg. D. Wrase, 1 ex. (cSch); same data, but "leg. A. Pütz", 1 ex (cPüt); Ya'an Prefecture, Tianquan Co., W Erlang Shan pass, 2780 m, 29.51.27N, 102,15.47E, sifted, 21.VI.1999, leg. A. Pütz, 1 ex. (cPüt); S-Sichuan, pass between Wupo and Jinyang, Corylus-Rhododendron-Quercus shrubs, 15.–21.VI.2004, leg. R. Fabbri, 1 ex. (cSch); Xiling Shan, 1600–2400 m, litter sifted, 30.VII.–4.VIII.1996, leg. S. Kurbatov, 1 ex.

226 · Zootaxa 4105 (3) © 2016 Magnolia Press KOCIAN & SCHÜLKE (MHNG); Yunnan: Yunnan, above Dali, 2700–2900 m, 14.IV.1999, leg. W. Schawaller, 1 Ex. (SMNS); Baoshan Prefecture, Gaoligong Shan, 65 km NNE Tengchong, 1750 m, 25°35'20''N, 98°40'21''E, secondary mixed forest, litter and moss sifted, scree, 27./31.VIII.2009, leg. M. Schülke, 2 exs. (cSch); Baoshan Prefecture, Gaoligong Shan, 33 km SE Tengchong, 2150 m, 24°51'22''N, 98°45'36''E, devastated primary forest, 28.VIII.2009, leg. M. Schülke, 1 ex. (cSch); Baoshan Prefecture, Gaoligong Shan, W Paß, 35 km SE Tengchong, 2100 m, 24°50'18''N, 98°45'43''E, devastated primary forest, 28.VIII.2009, leg. M. Schülke & D. W. Wrase, 2 exs. (cSch); Baoshan Prefecture, Gaoligong Shan, near Xiaoheishan N.R., 35 km SE Tengchong, 2110 m, 24°50'16''N, 98°45'43''E, deciduous forest, litter sifted, 30.V.2007, leg. M. Schülke, 1 ex. (cSch); Baoshan Prefecture, Gaoligong Shan, 29 km ESE Tengchong, 24°55'37''N, 98°45'09''E, 2350 m, devastated deciduous forest, dead wood, moss and mushrooms sifted, 1.VI.2007, leg. M. Schülke, 1 ex. (cSch); Baoshan Prefecture, Gaoligong Shan, 32 km SE Tengchong, 2150–2250 m, 24°51–53'N, 98°45'E, devastated primary and secondary forest, litter sifted, 26.VIII.2009, leg. M. Schülke, 2 exs. (cSch); Baoshan Prefecture, Gaoligong Shan, W pass 32 km SE Tengchong, 1600 m, cleft with devastated primary forest, litter sifted, 24°51'11''N; 98°44'27''E, 28.VIII.2009, leg. D. W. Wrase, 1 ex. (cSch); Baoshan Prefecture, mountains 25 km S Tengchong, 1900 m, 24°48'21''N, 98°32'05''E, cleft with devastated primary forest, litter sifted, 30.VIII.2009, leg. M. Schülke, 1 ex. (cSch); Nujiang Lisu Autonomous Prefecture, Gongshan County, Gaoligong Shan, 2500 m, 27°45.404'N, 98°35.749'E debris on snowfield during havy rain, sifted, 19.VI.2005, leg. M. Schülke, 1 ex. (cSch); Dali Bai Autonomous Prefecture; Diancang Shan, 5 km SSW Dali old town, creek valley above cableway, 25°38.7' N, 100°06.3' E, shrubs, Bamboo, litter, moos and debris sifted, 2800 m, 26.VIII.2003, leg. M. Schülke, 1 ex. (cSch); Dali Bai Autonomous Prefecture, Diancang Shan, 4 km W Dali old town, E-slope, 2900–3000 m, 25°41.4' N, 100°06.7' E, former quarry, 31.VIII.2003, leg. D.W. Wrase, 1 ex. (cSch); Dali Bai Autonomous Prefecture, Diancang Shan, 3 km W Dali old town, pine forest at "Cloud road", 25°41.1'N, 100°06.8'E, 2650–2750 m, litter from deep trenches and mushrooms sifted, 3.IX.2003, leg. M. Schülke, 1 ex. (cSch); Dali Bai Autonomous Prefecture, Diancang Shan W Dali, 25°41'09''N, 100°06'32''E, 3000–3200 m, cleft in devastated mixed forest, litter and debris sifted, 27.V.2007, leg. M. Schülke, 1 ex. (cSch); Dali Bai Autonomous Prefecture, Diancang Shan 43 km NW Dali, 3078 m, 25°59'35''N, 99°52'06''E, W pass, Quercus, Rhododendron, Bamboo, litter sifted, 29.V.2007, leg. M. Schülke, 3 exs. (cSch); Yunnan, Dali Bai Autonomous Prefecture, Diancang Shan 45 km NW Dali, 2730 m, 26°01'20''N, 99°53'17''E, creek valley, pines, ferns, litter sifted, 29.V.2007, leg. M. Schülke [CH07-09], 1 ex. (cSch); Dali Bai Autonomous Prefecture, Diancang Shan, pass 43 km NW Dali, 25°59'33.5''N, 99°52'12.5''E, pasture with shrubs, litter and moss sifted, 23.VIII.2009, leg. M. Schülke & D. W. Wrase, 3 exs. (cSch); Dali Bai Autonomous Prefecture, Wuliang Shan, 9 km SW Weishan, 2450–2500 m, 25°10'14''N, 100°14'22''E, W-slope, Quercus and Pinus, litter sifted, 13.VI.2007, leg. M. Schülke and A. Pütz, 4 exs. (cSch, cPütz); Dali Bai Autonomous Prefecture, Wuliang Shan, 9 km SW Weishan, 2480 m, 25°10'15.5''N, 100°14'21.8''E, W-slope, Quercus and Pinus, litter sifted, 14.IX.2009, leg. M. Schülke, 3 exs. (cSch); Dali Bai Autonomous Prefecture, Zhemo Shan, 7 km SW Xiaguan, 2870–2970 m, shrubs with bamboo and Rhododendron, litter sifted, 25°32–33'N, 100°10–11'E, 18.IX.2009, leg. M. Schülke & D. W. Wrase, 4 exs. (cSch); Dehong Dai Autonomous Prefecture, mountain range 31 km E Luxi, 2280 m, 24°29'31''N, 98°52'58''E, secondary pine forest with single old deciduous trees, litter sifted, 3.VI.2007, leg. M. Schülke, D. W. Wrase & A. Pütz, 11 exs. (cSch, cPütz); Nujiang Lisu Autonomous Prefecture, Gongshan County, Gaoligong Shan, side valley, 3000–3050 m, 27°47.90'N, 98°30.19'E, coniferous forest with Rhododendron and shrubs, litter, moss and dead wood sifted along creek valley and snowfieds, 21.VI.2005, leg. M. Schülke, 8 exs. (cSch); Nujiang Lisu Autonomous Prefecture, Gaoligong Shan, valley 18 km W Gongshan, 3020 m, 27°47'54''N, 98°30'13''E, mixed forest, litter, moss and dead wood sifted, 7.VI.2007, leg. M. Schülke, 2 exs. (cSch); Nujiang Lisu Autonomous Prefecture, Nu Shan, 7 km NNW Coajian, 25°43'29''N, 99°07'57''E, 2420 m, secondary pine forest with shrubs, litter and bark sifted, 11.VI.2007, leg. M. Schülke, 3 exs. (cSch); Nujiang Lisu Autonomous Prefecture, Gaoligong Shan, side valley 19 km NW Liuku, 25°59'02''N, 98°42'23''E, 2730 m, devastated primary forest, litter sifted, 9./ 10.VI.2007, leg. M. Schülke, 7 exs. (cSch); Nujiang Lisu Autonomous Prefecture, Gaoligong Shan, E pass 20 km NW Liuku, 3000 m, creek valley with primary forest, fern roots, litter and moss sifted, 25°58'49''N, 98°41'48''E, 3.IX.2009, leg. D.W. Wrase, 1 ex. (cSch); Shanzhi, Jizu Shan, 2180–2580 m, along path to summit, mixed forest (Pinus, Quercus, Rhododendron), litter sifted, 27°58'N, 100°22'E, 22.–24.VI.2007, leg. J. Hájek & J. Růžička, 1 Ex. (cSch); Pu'er Prefecture, Ailao Shan, 37 km NW Jingdong, 24°45'12''N, 100°41'24.5''E, 2300 m, devastated forest remnant, litter and dead wood sifted, 13.IX.2009, leg. M. Schülke, 1 Ex. (cSch); Lincang Prefecture, Bangma Shan, E pass 17 km NW Lincang, 23°57'31''N, 99°56'12''E, secondary pine forest, litter sifted, 9.IX.2009,

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 227 leg. M. Schülke, 1 ex. (cSch); Lincang Prefecture, Bangma Shan, 33 km SSW Lincang, 2150 m, 23°35'41''N, 100°00'27''E, deciduous forest remnant, N-slope, litter and dead wood sifted, 11.IX.2009, leg. M. Schülke, 5 exs. (cSch); Lincang Prefecture, Bangma Shan, 20 km NW Lincang, 23°58'25''N, 99°54'36''E, water reservoir, devastated forest with ferns, litter and fern roots sifted, 9.IX.2009, leg. M. Schülke, 3 exs. (cSch); Lincang Prefecture, Laobie Shan, Wei Bo Shan pass, 24°08'16''N, 99°42'53''E, creek valley with secondary mixed forest, litter and moss sifted, 8.IX.2009, leg. M. Schülke, 4 exs. (cSch); Lincang Prefecture, Xue Shan, 48 km N Lincang, 2070 m, 24°19'03''N, 100°07'13''E, forest remnant, N-slope, litter sifted, 12.IX.2009, leg. M. Schülke, 1 ex. (cSch); Lincang Prefecture, Xue Shan, 11 km ENE Lincang, 2510 m, 23°55'01''N, 100°11'17.5''E, secondary pine forest with Rhododendron, 10.IX.2009, leg. D. W. Wrase, 3 exs. (cSch); Lincang/Dali Prefecture, Wuliang Shan, old pass road, N pass, 24°45'16.4''N, 100°29'50.3''E, 2350 m, forest remnant and tea plantation, litter sifted, 16.IX.2009, leg. M. Schülke, 5 exs. (cSch); N-Yunnan, Nujiang Lisu Aut. Prefecture, Gongshan Co., Gaoligong Shan, valley at 3000–3050 m, 27°47.90'N, 98°30.19'E, 21.VI.2005, leg. A. Smetana [C 169], 2 exs. (cSme); N-Yunnan, Diqing Tibet Aut. Prefecture, Deqin Co., Meili Xue Shan, E-side, 12 km SW Deqin, 28°25.30'N, 98°48.47'E, 2890 m, 13.VI.2005, leg. A. Smetana [C160], 1 ex. (cSme); N-Yunnan, W slope N Gaoligong Shan, 27°53.626'N, 98°24.168'E, 2500 m, 8.VI.2009, sifting 01, leg. V. Grebennikov, 1 ex. (CNC); N-Yunnan, E slope N Gaoligong Shan, 27°46.8'N, 98°33.1'E, 2000–3000 m, 12.–15.VI.2009, sifting 1–7, leg. V. Grebennikov, 18 exs. (CNC, cSch); N-Yunnan, E slope N Gaoligong Shan, 27°59'01''N, 98°32'56.9''E, 3018 m, 27.V.2010, sifting 23, leg. V. Grebennikov, 3 exs. (CNC); N-Yunnan, E slope N Gaoligong Shan, 27°47'22''N, 98°32'17.7''E, 3027 m, 24.V.2010, sifting 20, leg. V. Grebennikov, 11 exs. (CNC, cSch); N-Yunnan, E slope N Gaoligong Shan, 27°45'40.8''N, 98°36'03.2''E, 2536 m, 3.VI.2010, sifting 27, leg. V. Grebennikov, 6 exs. (CNC, cSch); N-Yunnan, E slope N Gaoligong Shan, 27°59'11.5''N, 98°33'25.3''E, 3086 m, 28.V.2010, sifting 24, leg. V. Grebennikov, 3 exs. (CNC); N-Yunnan, E slope N Gaoligong Shan, 27°45'27.1''N, 98°35'34.5''E, 2600 m, 2.VI.2010, sifting 26, leg. V. Grebennikov, 11 exs. (CNC, cSch); W-Yunnan, E slope Cangshan at Dali, 25°40'13.2''N, 100°07'54.8''E, 2728 m, sifting 06, 11.V.2010, leg. V. Grebennikov, 1 ex. (CNC); Dali Bai Nat. Aut. Prefecture, Diancang Shan W Dali, 25°41'53''N, 100°06'45''E, 2860 m, pine forest, litter, moss and dead wood sifted, 28.V.2007, leg. A. Pütz [CH07- 06], 1 ex. (cPüt); Yunnan [6], NE Kunming, 25°08'35''N, 102°53'49''E, 2320 m, mixed forest with alder, oak, and pine, litter and mushrooms sifted, 13.VIII.2014, leg. V. Assing, 1 ex. (cAss); Yunnan [9], mountain W Xundian, 25°34'58''N, 103°08'42''E, 2300 m, mixed forest with alder, pine, shrub undergrowth, litter, twigs, and roots of herbs sifted, 15.VIII.2014, leg. V. Assing, 1 ex. (cAss); Yunnan [10], mountain NW Wuding, 25°36'53''N, 102°18'59''E, 2190 m, degraded mixed forest with alder, oak, and pine, litter, mushrooms, and dead wood sifted, 17.VIII.2014, leg. V. Assing, 3 exs. (cAss); Yunnan [12], mountain WNW Wuding, 25°38'45''N, 102°06'55''E, 2390 m, mixed forest margin with alder and pine, litter sifted, 18.VIII.2014, leg. V. Assing, 6 exs. (cAss); same data, but "leg. M. Schülke", 2 exs. (cSch); Yunnan [12a], mountain WNW Wuding, 25°38'45''N, 102°06'55''E, 2390 m, mixed forest margin with alder and pine, litter sifted, 1.IX.2014, leg. V. Assing, 9 exs. (cAss); same data, but "leg. M. Schülke", 8 exs. (cSch); Yunnan [22], SE Pingbian, Dawei Shan, 22°54'31''N, 103°41'44''E, 2100 m, primary subtropical broad-leaved forest, litter sifted, 27.VIII.2014, leg. V. Assing, 2 exs. (cAss); Yunnan [22a], SE Pingbian, Dawei Shan, 22°54'31''N, 103°41'44''E, 2100 m, primary subtropical broad-leaved forest, litter sifted, 28.VIII.2014, leg. V. Assing, 1 ex. (cAss); Yunnan [23], mountain W Yuxi, 24°27'11''N, 102°29'58''E, 2250 m, secondary mixed forest, litter, roots, and moss sifted, 31.VIII.2014, leg. V. Assing, 1 ex. (cAss); Zhejiang: West Tianmu Shan N. R., path to "Peak of immortals", 30°29'34''N, 119°25'51''E, 1100–1200 m, primary mixed forest, litter and moss sifted, 15.VI.2007, leg. A. Pütz, 1 Ex. (cSch). TAIWAN: Chiayi Hsien: Fengchifu, 12.–13.XI.2000, leg. H. Sugaya, 1 ex. (cSch); Kaohsiung Hsien: Tengchih, 1400 m, 21.–23.VII.2000, 20.–22.IV.2001, leg. H. Sugaya, 4 exs. (KUM, cSch); Tengchih, 2100 m, 19.–23.IV.2001, leg. H. Sugaya, 5 exs. (KUM, cSch); Peinantashan trail, 2450 m, 2.V.1995, leg. A. Smetana [T170], 1 ex. (cSme); Road above Tona Forset station, km 16–17, 1700–1800 m, 28.IV.1998, leg. A. Smetana [T190], 1 ex. (cSme); Nantou Hsien: Meifeng, 2100m, 26.X.2001, leg. H. Sugaya, 1 ex. (KUM); Meifeng, 2100 m, 6.V.2001, leg. H. Sugaya, 9 exs. (KUM, cSch); Meimu, 1600 m, flight intercept trap, 5.–10.IV.2002, leg. H. Sugaya, 33 exs. (KUM, cSch); Meimu, 1600 m, litter sifted, 30.III.2002, leg. H. Sugaya, 4 exs. (KUM, cSch); Taichung Hsien: "40 km 200 Logging road", 2000 m, 26.–27.VI.2002, flight intercept trap, leg. Chun Lin Li, 4 exs. (NMB, cSch); Anmashan, 2200m, 8.IV.2002, litter, leg. H. Sugaya, 1 ex. (KUM); Anmashan, 2 km, 24.–26.VI.2003, flight intercept trap, leg. Chun Lin Li, 9 exs. (MNB, cSch). Remarks. Ischnosoma duplicatum is widely distributed from the Russian Far East and Japan southwards to

228 · Zootaxa 4105 (3) © 2016 Magnolia Press KOCIAN & SCHÜLKE Thailand, Nepal, and northern India. In China (Fig. 66) its distribution seems to be confined to the southern half of the country, where the species is very common. Records from Hubei, Sichuan and Zhejiang are new province records.

FIGURE 66. Distribution of Ischnosoma duplicatum (Sharp) in China (material examined and literature records).

Ischnosoma evae Kocian, 2003 Figs 36, 67.

CHINA: Gansu: S-Gansu, W-Qingling Shan, 132 km NW Longnan, Lazikou pass, 34°14'32''N, 103°54'29''E, 3000 m, N-slope, pasture with shrubs, litter sifted, 2.VIII.2012, leg. M. Schülke [CH12-10], 3 exs. (cSch); Dagcanglhamo (= Langmusi), 34°04.6–05.1'N, 102°37.7–38.1'E, 3464–3644 m, wet coniferous forest (Picea, Abies, Rhododendron), N-slope, detrius and wet moss sifted, 25.VI.2995, leg. J. Hájek, D. Král & J. Růžička, 1 ex. (cSch); Qinghai: Lang Shi Dang Jing Qu (park) 75 km NW Honggu, 36°54'08.8''N, 102°21'16.9''E, 2896 m, creek valley on overgrown scree, Populus and Betula forest, litter sifted between rocks, 28.VI.2011, leg. M. Schülke [CH11-11], 7 exs. (cSch); same data, but "36°54'05.2''N, 102°21'07.1''E, 2925 m [CH11-11a]", 1 ex. (cSch); road 301 km 180, 43 km ESE Men Yuan, 37°09'32.6''N, 102°02'06.0''E, 2704 m, creek valley with Picea, Salix, Populus and Betula, litter and moss sifted, 5.VII.2011, leg. M. Schülke [CH11-19], 11 exs. (cSch); same data, but "leg. D.W. Wrase [19]", 1 ex. (cSch); Shaanxi: S-Shaanxi, Qinling Shan, mountain range W pass on road Xian–Shagoujie, 45 km SW Xian, 33°52'N, 108°46'E, 2675 m, 26.VII.2001, leg. A. Smetana [C119], 3 exx (cSme); same data, but "25.VII.2001 [C118], 6 exs. (cSme); S-Shaanxi, Daba Shan, mountain range N pass 22 km NW Zhenping, N-slope, 2685 m, 32°01'N, 109°21'E, 13.VII.2001, leg. A. Smetana [C102], 1 ex. (cSme); same data, but "14.VII.2001 [C103]", 1 ex. (cSme); Sichuan: N-Sichuan, N Songpan, road S 301, km 118, N Gongangling pass, 33°03'15''N, 103°43'36''E, 3390 m, spruce forest with shrubs, litter, moss, and mushrooms sifted, 9.VIII.2012, leg. V. Assing [19], 4 exs. (cAss, cSch); same data, but "leg. M. Schülke [CH12-19]", 1 ex. (cSch); N-Sichuan pass NW Songpan, 32°55'32''N, 103°25'56''E, 3600 m, moist N-slope with Salix and other shrubs, litter, grass roots, and moss sifted, 11.VIII.2012, leg. V. Assing [24], 21 exs. (cAss); same data, but "leg. M. Schülke [CH12-24]", 10 exs. (cSch); same data, but "leg. D.W. Wrase [24]", 5 exs. (cSch); N-Sichuan, 70 km N Songpan, road S 301, above Gan lake, 33°15'26''N, 103°46'03''E, 2700 m, spruce forest with birch, litter, mushrooms, moss, and dead wood sifted, 12.VIII.2012, leg. M. Schülke [CH12-26], 6 exs. (cSch); same data, but "leg. D.W. Wrase [26]", 1 ex. (cSch); W- Sichuan, Ganzi Tibetian Autonomous Prefecture, Daxue Shan, 10 km S Kangding, 3150 m, 29°59'N, 101°55'E, 28.VI.1999, leg. A. Pütz, 1 ex. (cPüt); W-Sichuan, Yaan Prefecture, Tianquan County, E Erlang Shan pass, 2900 m, 29°52.36'N, 102°17.82'E, 22.VI.1999, leg. A. Pütz, 2 exs. (cPüt, cKoc); W-Sichuan, Ganzi Tibetian Autonomous

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 229 Prefecture, Weizhou County, Qionglai Shan, Wolong valley, 69 km WSW Guanxian, 3500 m, 38°53.57'N, 102°58.63'E, mixed forest, 15.VII.1999, leg. A. Pütz, 1 ex. (cPüt); W-Sichuan, Ganzi Tibetian Autonomous Prefecture, Daxue Shan, 15 km W Kangding, Mugecuo, below lowest lake, 30°09.18'N, 101°51.16'E, 3300 m, 27.VI.1999, leg. A. Pütz, 1 ex (cPüt); road Pingwu–Nanping, Maoniu Shan, N-side, 2950–3000 m, 29.V.–5.VI.2006, leg. A. Puchner, 1 ex. (NHMW); W-Sichuan, Qionglai Shan, Mou Pi Shan, 3100–3700 m, Barkam, 15 km S Zhuokaji, mixed and relict coniferous forest, 10.–30.VI.2004, leg. R. Fabbri, 2 exs. (cSch); NW- Sichuan, Maelma, 40 km SE of Aba, 3250 m, shrubs, 9.–29.VI.2004, leg. R. Fabbri, 1 ex. (cSch); NW-Sichuan, 20 km W of Luhua, 2750–3200 m, mixed forest, 8.–28.VI.2004, leg. R. Fabbri, 1 ex. (cSch); Jitiang, 3.–14.VII.2001, leg. E. Kučera, 1 ex. (cSch); W-Sichuan, Jintiang (Liang-ch-ke), 6.–14.VI.2002, leg. E. Kučera, 3 exs. (cSch); Yunnan: Dali, Cangshan, E-slope, 25°40'25.1''N, 100°05'57.6''E, 3806 m, sifting 15, 17.V.2010, leg. V. Grebennikov, 3 exs. (CNC, cKoc); same data, but "25°40'14.7''N; 100°06'12.0''E, 3827 m, sifting 16", 9 exs. (cSme, cKoc); same data, but "25°40'07.6''N; 100°06'12.9''E, 3887 m, sifting 18, 19.V.2010", 8 exx (cSme); same data, but "25°39'54.4''N, 100°05'53.0''E, 3991 m, sifting 14, 15.V.2010", 2 exx (cSme); sama data, but "25°39'54.7''N, 100°06'04.5''E, 3815 m, sifting 19, 19.V.2010", 2 exx (cSme); N-Yunnan, Diqing Tibetian Autonomous Prefecture, Zhongdian County, Xue Shan, 23 km S Zhongdian, 27°38.3'N, 99°41.5'E, 3675–3725 m, 2.VI.2005, leg. A. Smetana [C149], 3 exs. (cSme, cKoc); N-Yunnan, Zhongdian County, Xue Shan, 10 km SW Zhongdian, 27°46.5'N, 99°36.5'E, 3800 m, 20.VIII.2003, leg. A. Smetana [C129], 18 exs. (cSme, cKoc); N- Yunnan, Diqing Tibetian Autonomous Prefecture, Zhongdian County, Xue Shan, near lake 23 km S Zhongdian, 27°37.1'N, 99°36.5'E, 3850 m, 6.VI.2005, leg. A. Smetana [C153b], 6 exs. (cSme, cKoc); same data, but "5.VI.2005 [C152]", 3 exs. (cSme); same data, but "27°37.1'N, 99°36.5'E, 15.VI.2005 [C161]", 3 exx (cSme); N- Yunnan, Zhongdian County, pass 28 km ESE Zhongdian, 27°43.9'N, 99°58.2'E, 3700–3750 m, 22.VIII.2003, leg. A. Smetana [C131], 1 ex. (cSme); N-Yunnan, Diqing Tibetian Autonomous Prefecture, Bitai Hai öake area, 29 km ESE Zhongdian, 27°43.65'N, 99°58.97'E, 3540 m, 1.VI.2005, leg. A. Smetana [C147], 1 ex (cSme); N-Yunnan, Zhongdian County, Xue Shan, 10 km SW Zhongdian, 3700–3800m, 27°46.5'N, 99°36.5'E, primary mixed forest, leaf litter sifted, 20.VIII.2003 leg. D.W. Wrase [10A], 37 exs. (cSch); N-Yunnan, Zhongdian County, pass 28 km ESE Zhongdian, devastated primary forest with young Abies, Larix, Betula, Rhododendron, 27°43.9' N, 99°58.2' E, 3700–3750m, 22.VIII.2003, leg. M. Schülke [CH03-12], 2 exs. (cSch); N-Yunnan, Diqing Tibetian Autonomous Prefecture, Zhongdian County, Bitai Hai Lake area, 29 km ESE Zhongdian, 27°43.65'N, 99°58.97' E, 3540m, creek valley, devastated mixed forest, litter, moss and dead wood sifted, 1.VI.2005, leg. M. Schülke [CH05-01], 3 exs. (cSch); N-Yunnan, Diqing Tibetian Autonomous Prefecture, Zhongdian County, Xue Shan, 23 km S Zhongdian, 3675–3725m, 27°36.3' N, 99°41.5' E, devastated mixed forest, sifted from leaf litter and dead wood, 2.VI.2005, leg. M. Schülke [CH05-02], 5 exs. (cSch); N-Yunnan, Diqing Tibetian Autonomous Prefecture, Zhongdian County, 35 km ESE Zhongdian, 3450m, 27°41.00'N, 100°01.47'E, devastaded mixed forest near small creek, sifted from litter, moss and dead wood, 3.VI.2005, leg. D.W.Wrase [03], 1 ex. (cSch); N-Yunnan, Diqing Tibetian Autonomous Prefecture, Zhongdian County, Xue Shan near lake 23 km S Zhongdian, 3895 m, 27°37.1' N, 99°38.5' E, devastated mixed forest, meadows, lake border, leaf litter and dead wood sifted, 5.VI.2005, leg. M. Schülke [CH05-05], 7 exs. (cSch); same data, but "6.VI.2005 [CH05-05A]", 12 exs. (cSch); same data, but "15.VI.2005 [CH05-05B]", 1 ex. (cSch); same data, but "15.VI.2005, leg. D.W. Wrase [5B]", 3 exs. (cSch); same data, but "pitfall traps (vinegar), 6.–15.VI.2005 [CH05-5F], 4 exs. (cSch). Remarks. The distribution of Ischnosoma evae (Fig. 67) is restricted to the eastern slopes of the Tibetian plateau and adjacent mountains such as the Qinling Shan and the Daba Shan ranges. The records from Qinghai are a new province record.

Ischnosoma farkaci Kocian, 2003 Figs 42, 68.

CHINA: Yunnan: NW-Yunnan, Bailakou pass, 3750 m, 28.V.–7.VI.2006, leg. S. Murzin & L. Shokin, 1 ex. (cSch); CHINA: N-Yunnan, Diqing Tibet. Aut. Prefecture, Zhongdian Co., Bitai Hai Lake area, 29 km ESE Zhongdian, 27°43.65'N, 99°58.97' E, 3540m, creek valley, devast. mixed forest, litter, moss, dead wood, 1.VI.2005, leg. M. Schülke [C2005-01], 1 ex. (cSch); N-Yunnan, 32 km N Lijiang, Maoniuping, Yak meadows, 27°09.9'N, 100°14.5'E, 3540 m, steep slope, wet mixed forest (with Pinus, Abies, Rhododendron), detrius, leaves

230 · Zootaxa 4105 (3) © 2016 Magnolia Press KOCIAN & SCHÜLKE and moss sifted, 16.VI.2007, leg. J. Hájek & J. Růžička [CH28], 1 ex. (cSch); N-Yunnan, Deqing Tibetian Autonomous Prefecture, Bitai Hai lake area, 29 km ESE Zhondian, 27°43.55'N, 99°58.97'E, 1.VI.2005, leg. A. Smetana [C147], 1 ex. (cSme); N-Yunnan, Zhongdian County, 51 km SSE Zhongdian, 27°55.3'N, 99°56.5'E, 2970 m, 16.–19.VIII.2003, leg. A. Smetana [C124], 1 ex. (cSme); N-Yunnan, 32 km N Lijiang, Maoniuping, Yak meadows, 27°09.9'N, 100°14.5'E, 3540 m, steep slope, wet mixed forest (with Pinus, Abies, Rhododendron), detrius, leaves and moss sifted, 21.VI.2007, leg. J. Hájek & J. Růžička [CH41], 1 ex. (cKoc). Remarks. Ischnosoma farkaci was described from the surroundings of Zhongdian (Kocian, 2003). All the recent records are from the same area in northern Yunnan (Fig. 68).

FIGURE 67. Distribution of Ischnosoma evae Kocian in China (material examined and literature records).

Ischnosoma fasciatocolle (Champion, 1922) Fig. 68.

CHINA: Yunnan: Nujiang Lisu Autonomous Prefecture, Nu Shan, 7 km NNW Coajian, 25°43'29''N, 99°07'57''E, 2420 m, secondary pine forest with shrubs, litter and bark sifted, 11.VI.2007, leg. A. Pütz, 1 ex. (cPüt); Cangshan near Dali, E slope, 25°40'15.5''N, 100°07'45.4''E, 2657 m, sifting 17, 18.V.2010, leg. V. Grebennikov, 1 ex. (cSch); N Yunnan, Nujiang Lisu Autonomous Prefecture, Gongshan County, Gaoligong Shan, snowfield at 2500 m, 27°45.404'N, 98°35.749'E, 19.VI.2005, leg. A. Smetana [C185], 1 ex. (cSme). Remarks. Ischnosoma fasciatocolle was previously known from northern India and Nepal. The material represents the first records from China (northwestern Yunnan) (Fig. 68).

Ischnosoma fusciventre (Tikhomirova, 1973) Figs 38, 68.

CHINA: Beijing: Yan Shan, Dongling Mts., Xiaolongmen, 1400 m, 15.–16.VI.2001, leg. P. Hlaváč & J. Cooter, 14 exs. (cSch); Gansu: Xinlong Shan, 70 km S Lanzhou, 2225–2380 m, 7.VIII.1994, leg. A. Smetana [C32], 1 ex. (cSme); Hebei/Nei Mongol: pass Chengde–Chifeng, 41.6°N, 118.2°E, 30.–31.V.2002, leg. J. Turna, 1 ex. (NHMW); Hubei: W-Hubei, Daba Shan, pass E of mt. Da Shennongjia, 12 km NW Muyuping, 31°30'N, 110°21'E, 1950 m, leg. A. Smetana [C117], 1 ex. (cSme); Shaanxi: Qinling Shan, mountain W pass at autoroute km 70, 47 km S Xian, 2500–2600 m, 26.–27.VIII.1995, leg. A. Pütz, 1 ex. (cSch). Remarks. Ischnosoma fusciventre is widely distributed from Far East Russia and Japan southwards to central China (Fig. 68). Records from Beijing, Gansu, Hebei/Nei Mongol and Hubei are new province records.

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 231 FIGURE 68. Distribution of Ischnosoma farkaci Kocian (squares), I. fasciatocolle (Champion) (diamonds), I. fusciventre (Tikhomirova) (circles) and I. hieroboam Kocian (triangle) in China (material examined and literature records).

Ischnosoma hieroboam Kocian, 2003 Fig. 68.

CHINA: Yunnan: NNE Pingbian, broad-leaved subtropical forest, 23°00'39''N, 103°42'10''E, 1500 m, 26.VIII.2014, leg. V. Assing, 4 exx (cAss, cSch). Remarks. Ischnosoma hieroboam was described from Myanmar (Kocian, 2003) and is here recorded from China for the first time (Fig. 68).

Ischnosoma noemi Kocian, 2003 Figs 47, 69.

CHINA: Yunnan: Dali Bai Autonomous Prefecture, Jizu Shan, path to cable car, 37 km NE Dali, 25°58'N, 100°23'E, 2450 m, mixed forest, sifted from litter, moss & pine apples, 5.IX.2009, leg. M. Schülke [CH09-29], 3 exs. (cSch, cKoc); N-Yunnan, Diqing Tibetian Autonomous Prefecture, Deqin County, Meili Xue Shan, E-side, 12 km SW Deqin, 28°25.30'N, 98°48.47'E, 2890 m, 9.VI.2005, leg. A. Smetana [C156], 1 ex. (cSme); N-Yunnan, Zhongdian Co., 48 km N Zhongdian, 28°16.6' N, 99°45.7' E, 3220m, creek valley, devastated primary forest, dead wood, moss and mushrooms sifted, 21.VIII.2003, leg. M. Schülke [C03-11], 1 ex. (cSch); N-Yunnan, Zhongdian Co. 51 km SSE Zhongdian, 2970m, 27°25.3'N, 99°56.5'E, creek valley, mixed coniferous forest with shrubby vegetation, bamboo, 16.VIII.2003 leg. D.W. Wrase [04], 1 ex. (cSch); N-Yunnan, Diqing Tibetian Autonomous Prefecture, Deqin County, Meili Xue Shan, E-side, 12 km SW Deqin, 2890 m, 28°25.30'N, 98°48.47'E, small creek valley, mixed forest with bamboo, leaf litter, moss and dead wood sifted, 9.VI.2005, leg. M. Schülke [C2005-07], 1 ex. (cSch). Remarks. The distribution of this species is restricted to the northern and western parts of Yunnan (Fig. 69).

232 · Zootaxa 4105 (3) © 2016 Magnolia Press KOCIAN & SCHÜLKE FIGURE 69. Distribution of Ischnosoma noemi Kocian (diamonds), I. rosti (Bernhauer) (circles), I. maderi (Bernhauer) (triangles) and I. mandschuricum (Bernhauer) (square) in China (material examined and literature records).

Ischnosoma quadriguttatum quadriguttatum (Champion, 1923) Fig. 70.

CHINA: Fujian: Kuatun [Guadun] (2300 m) [elevation wrong], 27°40'N, 117°40'E, 4.III.1938, 6.III.1938, 27.III.1938, 7.IV.1938, 24.V.1938, leg. J. Klapperich, 5 exs. (MAKB, cSch); Kuatun [Guadun], 26.III.1946, leg. Tschung sen., 2 exs. (NHMW); Wuji Shan Nat. Res., Sangan env., 900 m, 30.V.–12.VI.2001, leg. P. Hlaváč & J. Cooter, 6 exs. (cSch, cKoc); Gansu: S-Gansu, N Chengxian, W-Qingling Shan, 34°10'20''N, 105°42'19''E, 1830 m, stream valley with secondary deciduous forest, moist litter sifted, 29.VII.2012, leg. V. Assing, 3 exs. (cAss, cSch); S-Gansu, N Chengxian, W-Qingling Shan, 34°10'17''N, 105°42'56''E, 1850 m, mixed secondary forest margin, litter sifted, 29.VII.2012, leg. V. Assing, 6 exs. (cAss, cSch); S-Gansu, mountains SE Longnan, 33°13'20''N, 105°15'10''E, 2170 m, N-slope with shrubs and scattered coniferous trees, litter sifted, 31.VII.2012, leg. V. Assing, 3 exs. (cAss, cSch); S-Gansu, S Longnan, Min Shan, 33°05'24''N, 104°45'13''E, 1500 m, N-slope, macchia, litter and moss sifted, 6.VIII.2012, leg. V. Assing, 2 exs. (cAss); S-Gansu, N Chengxian, W-Qingling Shan, 34°08'16''N, 105°46'42''E, 1760 m, N-slope, secondary deciduous forest margin, sifted, 28.VII.2012, leg. V. Assing, 1 ex. (cAss); S-Gansu [CH12-04] W Qinling Shan, 43 km N Chengxian, 34°08'16''N, 105°46'42''E, 1760 m, N-slope, secondary deciduous forest margin, sifted, 28.VII.2012, leg. M. Schülke, 4 exs. (cSch); S-Gansu [CH12-04c] W Qinling Shan, 43 km N Chengxian, 34°08'16''N, 105°46'42''E, 1760 m, S-slope, secondary deciduous forest margin, litter between rocks sifted, 28.VII.2012, leg. M. Schülke, 3 exs. (cSch); S-Gansu [CH12-05] W Qinling Shan, 47 km N Chengxian, 34°10'17''N, 105°42'56''E, 1850 m, mixed secondary forest margin, litter sifted, 29.VII.2012, leg. M. Schülke, 3 exs. (cSch); S-Gansu [CH12-06] W Qinling Shan, 47 km N Chengxian, 34°10'20''N, 105°42'19''E, 1830 m, creek valley with secondary deciduous forest, moist litter sifted, 29.VII.2012, leg. M. Schülke, 1 ex. (cSch); S-Gansu [CH12-17] Min Shan, 38 km SSW Longnan, 33°05'24''N, 104°45'13''E, 1500 m, N-slope, macchia, litter and moss sifted, 6.VIII.2012, leg. M. Schülke, 1 ex. (cSch); S-Gansu [CH12-08]

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 233 Mts. 38 km SE Longnan, 33°11'20''N, 105°14'24''E, 2030 m, N-slope with scree, moss, fern roots and litter sifted, 31.VII.2012, leg. M. Schülke, 1 ex. (cSch); S-Gansu Mts. 38 km SE Longnan, 2130 m, 33°11'16''N, 105°14'24''E W-slope with scree, scrubs and trees, litter, soil and moss sifted, 7.VIII.2012, leg. D.W. Wrase [18B], 1 ex. (cSch); Guizhou: Leishan Co., SE Kaili, NE Leishan, Leigong Shan, E-slope, 2.5 km E of pass, 26°23.39'N, 108°13.33'E, 1600 m, 23./24.VI.2001, leg. H. Schillhammer [17a], 1 ex. (NHMW); Hainan: Bawangling Nat. Forest Park, 12,3 km SE of Baotie, 19°5.20'N, 109°11.80'E, 1050 m, 8.V.2011, sifting moist accumulation of leaf litter along stream in primary forest, leg. M. Fikáček, 1 ex. (NMP); Hubei: SE-Hubei, Mufu Shan, Jiugongshan forest park, 29.4N, 114.6E, 1000 m, 3./5./18.VI.2002, leg. J. Turna, 1 ex. (NHMW); W-Hubei, Daba Shan, pass E of mt. Da Shennongjia, 12 km MW Muyuping, 31°30'N, 110°21'E, creek valley, 1950–2000 m, mixed deciduous forest, moss, dead wood and mushrooms sifted, 19.VII.2001, leg. M. Schülke [CH01-13c], 2 exs. (cSch); W-Hubei, Daba Shan, creek valley 8 km NW Muyuping, 31°28'N, 110°22'E, 1540 m, edge of small creek, 18.VII.2001, leg. D.W. Wrase, 2 exs. (cSch); same data, but "deciduous forest, sifted from moss, leg. M. Schülke [CH01-16a]", 2 exs. (cSch); Shaanxi: Qinling Shan, Taibai Shan, above Houzhenzi, 1300–1700 m, 9.VI.–3.VII.1998, leg. P. Jäger & J. Martens, 1 ex. (SMNS); Qinling Shan, pass at road Zhouzhi–Foping, 105 km SW Xian, N-slope, 1880 m, 33°44'N, 107°58'E, 4.VII.2001, leg. A. Smetana [C92], 2 exs. (cSme); same data, but "1700 m, 33°46'N, 107°58''E, 3.VII.2001 [C91]", 1 ex. (cSme); same data, but "1990 m, small creek valley, mixed deciduous forest with bamboo and small meadows, 2./4.VII.2001, leg. D.W. Wrase [01]", 1 ex. (cSch); same data but: "litter, dead wood and mushrooms sifted, 2./4.VII.2001, leg. M. Schülke [CH01-01]", 2 exs. (cSch); S-Shaanxi [CH12-31], Micang Shan, 40 km SW Hanzhong, 32°52'25''N, 106°37'11''E, 1530 m, N-slope, mixed secondary forest, litter and moss sifted, 16.VIII.2012, leg. M. Schülke, 4 exs. (cSch); S-Shaanxi [CH12-30], Micang Shan, 33 km S Hanzhong, 32°44'44''N, 106°52'46''E, 1360 m, stream valley, forest margin, litter and soil sifted, 15.VIII.2012, leg. M. Schülke, 2 exs. (cSch); S-Shaanxi [CH12-32], Micang Shan, 42 km S Hanzhong, 32°40'43''N, 106°48'33''E, 1090 m, stream valley, shady S-slope, secondary mixed forest, litter, grass, and herbs along path sifted, 17.VIII.2012, leg. M. Schülke, 2 exs. (cSch); S-Shaanxi [CH12-28] Micang Shan, 34 km S Hanzhong, 32°44'22''N, 106°51'55''E, 1460 m, W-slope, deciduous forest margin with bamboo, litter, grass, and moss sifted, 14.VIII.2012, leg. M. Schülke, 1 ex. (cSch); S-Shaanxi, Micang Shan 42 km S Hanzhong, 1090 m, 32°40'52''N/106°49'16''E, NW-slope, margin of mixed forest with rocks, litter, moss and soil sifted) 14.VIII.2012, leg. D.W. Wrase [27], 1 ex. (cSch); S- Shaanxi, Micang Shan 40 km SW Hanzhong,1530 m, 32°52'25''N/106°37'11''E, N-slope, mixed secondary forest, litter and moss sifted, 16.VIII.2012, leg. D.W. Wrase [31], 1 ex. (cSch); S-Shaanxi, Qinling Shan, 42 km SW Meixian, 1875 m, 34°01'32''N/107°24'13''E, N-slope, secondary deciduous forest near creek, litter & grass sifted, 26.VII.2012, leg. D.W. Wrase [02A], 1 ex. (cSch); Micang Shan, 33 km S Hanzhong, 1360 m, 32°44'44''N/ 106°52'46''E, stream valley, shady brookside with bamboo/decidious shrubs, litter/moss/soil sifted, 15.VIII.2012, leg. D.W.Wrase [30B], 1 ex. (cSch); S-Shaanxi Micang Shan, 42 km S Hanzhong, 32°40'43''N, 106°48'33''E, 1090 m, stream valley, shady S-slope, secondary mixed forest, litter, grass, and herbs along path sifted, 17.VIII.2012, leg. V. Assing, 4 exs. (cAss, cSch); S-Shaanxi, Micang Shan, 34 km S Hanzhong, 32°44'22''N, 106°51'55''E, 1460 m, W-slope, deciduous forest margin with bamboo, litter, grass, and moss sifted, 14.VIII.2012, leg. V. Assing, 1 ex. (cAss); S-Shaanxi, Micang Shan, 42 km S Hanzhong, 32°40'52''N, 106°49'16''E, 1090 m, NW-slope, mixed forest margin with rocks, litter, grass, and moss sifted, 14.VIII.2012, leg. V. Assing, 5 exs. (cAss, cSch); S-Shaanxi Micang Shan, 30 km S Hanzhong, 32°45'56''N, 106°53'57''E, 1070 m, stream valley, litter and soil sifted, 15.VIII.2012,, leg. V. Assing, 2 exs. (cAss); Shaanxi/Chongqing: Shaanxi/Chongqing border, pass 20 km SSE Zhenping, 1700–1800 m, 31°44'N, 109°35'E, small creek valley, young mixed forest, litter and moss sifted, 9./ 12.VII.2001, leg. D.W. Wrase [07], 3 exs. (cSch); same data, but "leg. M. Schülke [CH01-07], 1 ex.; same data, but "leg. A. Smetana [C101], 1 ex. (cSme); Sichuan: Emei Shan, 29°33.6'N, 103°20.6'E, 1800–2400 m, sifting 11–17, 27.VI.–5.VII.2009, leg. V. Grebennikov, 32 exs. (CNC, cSch); Emei Shan, 29°32'56''N, 103°20'28''E, 2310 m, sifting 38, 20.VI.2010, leg. V. Grebennikov, 2 exs. (CNC); Emei Shan, Leidongping, 2500 m, 29°32'N, 103°21'E, 18.VII.1996, leg. A. Smetana, J. Farkač & P. Kabátek, 1 ex. (cSme); Emei Shan, Wannian, 1050 m, 19.–30.III.1999, leg. W. Schawaller, 1 ex. (SMNS); C-Sichuan, Wenjiang Distr., Dujiangyan County, Qingcheng Shan, 56 km NW Chengdu, 30°54'N, 103°33'E, 975 m, creek bank, forest remnant, 18.VI.1999, leg. M. Schülke, 1 ex. (cSch); same data, but "field ridge, leg. D.W. Wrase", 1 ex. (cSch); same data, but "leg. A. Pütz", 2 exs. (cPüt); Qingcheng Shan 65 km NW Chengdu, 37°53'N, 103°53'E, 18.V./3.–4.VI.1997, leg. A. Pütz, 4 exs. (cPüt); W- Sichuan, Ganzi Tibet. Auton. Prefecture, Luding, W Erlang Shan pass, 2600 m, 7 km SSE Luding, 29°51'N, 102°15'E, litter and mushrooms sifted, 29.VI.1999, leg. M. Schülke, 1 ex. (cSch); same data, but 20.–29.VI.1999,

234 · Zootaxa 4105 (3) © 2016 Magnolia Press KOCIAN & SCHÜLKE leg. D.W. Wrase", 1 ex. (cSch); W-Sichuan, Ya'an Prefecture, Shimian County, Daxue Shan, road between Anshunchang and Wanba, 20 km WSW Shimian, 1500 m, 9.VII.1999, 3 exs. (cPüt); same data, but " 12 km W Shimian, 1300 m", 1 ex. (cPüt); S-Sichuan, S Xichang, Luoji Shan, 2300–2500 m, litter sifted, 16.–24.VII.1996, leg. S. Kurbatov, 1 ex. (MHNG); Xiling Shan, 1600–2400 m, litter sifted, 30.VII.–4.VIII.1996, leg. S. Kurbatov, 1 ex. (MHNG); Yunnan: SW-Yunnan, Rehai hot springs, 6 km SW Tengchong, 24°57.1'N, 98°26.2'E, 1400 m, 5.VI.2007, sifted detrius and leaves in grove margin above hot stream, along old path, leg. J. Hájek & J. Růžička [CH11], 1 ex. (cSch); SW-Yunnan, Gaoligong Shan, 90 km W Baoshan, 26.–28.V.1995, leg. S. Becvar, 1 ex. (NHMW); W-Yunnan, above Dali, 2000–2200 m, 4.–17.IV.1999, leg. W. Schawaller, 1 ex. (SMNS); NW-Yunnan, E slope N Gaoligong Shan, 27°46.8'N; 98°33.1'E, 2000–3000 m, sifting 1–7, 12.–15.VI.2009, leg. V. Grebennikov, 3 exs. (CNC); NW-Yunnan, E slope N Gaoligong Shan, 27°45'27.1''N; 98°35'34.5''E, 2600 m, sifting 26, 2.VI.2010, leg. V. Grebennikov, 3 exs. (CNC); NW-Yunnan, E slope N Gaoligong Shan, 27°45'40.8''N; 98°36'03.2''E, 2536 m, sifting 27, 3.VI.2010, leg. V. Grebennikov, 1 ex. (CNC); W-Yunnan, Nujiang Lisu Aut.Pref., Nu Shan, 7 km NNW Coajian, 25°43'29''N, 99°07'57''E, 2420 m, secondary pine forest with shrubs, litter and bark sifted, 11.VI.2007, leg. M. Schülke [CH07-30], 5 exs. (cSch); same data, but "leg. D.W. Wrase [30]", 2 exs. (cSch); same data, but "leg. A. Pütz", 2 exs. (cPüt, cKoc); W-Yunnan, Baoshan Pref., mountain range 25 km S Tengchong, 1900 m, 24°48'28''N, 98°32'03''E, devastated primary deciduous forest, litter and fungi sifted, 2.VI.2007, leg. M. Schülke [CH07-17], 2 exs. (cSch); same data, but "leg. D.W. Wrase [17]", 1 ex. (cSch); W- Yunnan, Nujiang Lisu Aut. Pref., Salween side valley 5 km S Fugong, road SS228 km 223, under stones, moist embarkment, 8.VI.2007, leg. A. Pütz [CH07-25], 1 ex. (cPüt); W-Yunnan, Baoshan Pref., mountain range 22 km S Tengchong, 1750 m, 24°49'29''N, 98°29'27''E, secondary forest, litter and dead wood sifted, 2.VI.2007, leg. A. Pütz [CH07-18], 1 ex. (cPüt); W-Yunnan, Baoshan Pref., Gaoligong Shan, 33 km SE Tengchong, 2100–2200 m, 24°51'22''N, 98°45'36''E, deciduous forest, litter, wood and fungi sifted, 4.VI.2007, leg. A. Pütz [CH07-14A], 1 ex. (cPüt); same data, but "pitfall trap, 31.V.–4.VI.2007, leg. D.W. Wrase, 1 ex. (cSch); W-Yunnan, Baoshan Pref., mountain range 14 km E Tengchong, 1850 m, 25°00'28''N, 98°38'07''E, secondary mixed forest, litter sifted, 1.VI.2007, leg. A. Pütz [CH07-16], 1 ex. (cPüt); W-Yunnan, Baoshan Pref., Gaoligong Shan, 29 km ESE Tengchong, 24°55'37''N, 98°45'09''E, 2350 m, devastated deciduous forest, litter, wood and fungi sifted, 1.VI.2007, leg. A. Pütz [CH07-15], 4 exs. (cPüt); same data, but "leg. D.W.Wrase", 1 ex. (cSch); Yunnan, Baoshan Pref., Gaoligong Shan, W Pass 35 km SE Tengchong, 2100 m, 24°50'18''N, 98°45'43''E, devastated primary deciduous forest, litter, wood and mushrooms sifted, 25.VIII.2009, leg. M. Schülke [CH09-06], 2 exs. (cSch, cKoc); S- Yunnan, Pu'er Pref., Ailao Shan, 37 km NW Jingdong, 24°45'12''N, 100°41'24.5''N, 2300 m, devastated forest remnant, litter and dead wood sifted, 13.IX.2009, leg. M. Schülke [CH09-48], 3 exs. (cSch); W-Yunnan, Baoshan Pref., Gaoligong Shan, 33 km SE Tengchong, 2150 m, 24°51'22''N, 98°45'36''E, devastated primery deciduous forest, litter, wood and mushrooms sifted, 6.VIII.2009, leg. M. Schülke [CH09-08], 1 ex. (cSch); S-Yunnan, Lincang Pref., Xue Shan, 11 km ENE Lincang, 2510 m, 23°55'01''N, 100°11'17.5''E, secondary pine forest with Rhododendron, small cleft with water, litter & mushrooms sifted, 10.IX.2009, leg. M. Schülke [CH09-39], 2 exs. (cSch); S-Yunnan, Lincang/Dali Pref., Wuliang Shan, old pass road, N pass, 24°45'16.4''N, 100°29'50.3''E, 2350 m, forest remnant & tea plantation litter, mushrooms and grass sifted, 16.IX.2009, leg. M. Schülke [CH09-55], 1 ex. (cSch); S-Yunnan, Lincang Pref., Wuliang Shan, old pass road, W side, 24°42'58.6''N, 100°29'52.0''E, 2200 m, small creek valley with primary forest remnant, litter sifted, 16.IX. 2009, leg. M. Schülke [CH09-47a], 1 ex. (cSch); S-Yunnan, Lincang Pref., Bangma Shan, E pass, 17 km NW Lincang, 2040 m, 23°57'31''N / 99°56'13''E, secondary pine forest with ferns, litter sifted, 9.IX.2009, leg. D.W.Wrase [36], 1 ex. (cSch); C-Yunnan [23], mountain W Yuxi, 24°27'11''N, 102°29'58''E, 2250 m, secondary mixed forest, litter, roots, and moss sifted, 31.VIII.2014, leg. V. Assing [23], 1 ex. (cAss); C-Yunnan [6], NE Kunming, 25°08'35''N, 102°53'49''E, 2320 m, mixed forest with alder, oak, and pine, litter and mushrooms sifted, 13.VIII.2014, leg. V. Assing [06], 1 ex. (cAss); C-Yunnan [5], NE Kunming, 25°08'40''N, 102°53'48''E, 2290 m, mixed deciduous forest with scattered pine trees, litter and mushrooms sifted, 11.VIII.2014, leg. V. Assing [05], 1 ex. (cAss); S-Yunnan [22/22a], SE Pingbian, 22°54'31''N, 103°41'44''E, 2100 m, primary subtropical broad-leaved forest, litter sifted, 27./28.VIII.2014, leg. V. Assing, 14 exs. (cAss, cSch); Yunnan [2a], mountains NE Dongchuan, 26°14'10''N, 103°12'31''E, 2670 m, scree slope with bushes and herbs, litter sifted, 9.VIII.2014, leg. V. Assing, 1 ex. (cAss); N-Yunnan, Nujiang Lisu Autonomous Prefecture, Gongshan County, N Gaoligong Shan, 2500m, 27°45.404'N, 98°35.749'E, litter & debries at snowfield sifted during rain, 19.VI.2005, leg. M. Schülke [C2005-12], 2 exs. (cSch); S-Yunnan, Xishuangbanna, 23 km NW Jinghong, vic. Na Ban (NNNR), 22°09.49'N, 100°39.92'E, forest EKL, 26.III.2009, leg. L. Mengl, 1 ex.

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 235 (NME); S-Yunnan, Xishuangbanna, 20 km NW Jinghong, near Man Dian (NNNR), pitfall trap, 22°07.80'N, 100°40.05'E, 740 m, rubber plantation, 28.VI.2008, leg. A. Weigel, 1 ex. (NME); Zheijiang: Tianmu Shan, pass 25 km NNW Linan, 30°25'40''N, 119°35'30''E, 620–820 m, creek valley with bamboo and mixed forest, litter sifted, 16.VI.2007, leg. A. Pütz [CH07-37], 5 exs. (cPüt); same data, but "leg. D.W. Wrase", 3 exs. (cSch); West Tianmu Shan N.R. way to peak of immortals, 30°20'34''N, 119°25'51''E, 1100–1200 m, primary mixed forest, litter and moss sifted, 15.VI.2007, leg. A. Pütz [CH07-36], 2 exs. (cPüt); Tianmu Shan, pass 25 km NNW Linan, 620–820 m, 30°25'40''N, 119°35'30''E, creek valley with bamboo and mixed forest, litter, sifted, 16.VI.2007, leg. M. Schülke [CH07-37], 5 exs. (cSch); Zheijiang/Anhui: Zheijiang/Anhui border, pass W Linan, 30°17'59''N, 118°52'41''E, 625 m, walnut plantation, plant refuse and litter sifted, 19.VI.2007, leg. M. Schülke, 1 ex. (cSch);

FIGURE 70. Distribution of Ischnosoma quadriguttatum quadriguttatum (Champion) in China (material examined and literature records).

TAIWAN: Ilan Hsien: 75 km , N crossing highway, 1100 m, 22.V.2003, flight intercept trap, leg. Chun Lin Li, 2 exs. (NMB); Shen-Mi Lake, 24°22'43''N, 121°44'12''E, 1110 m, 10.V.1995, leg. A. Smetana [T177], 1 ex. (cSme); Shen-Mi Lake, 24°22'43''N, 121°44'12''E, 1110 m, 9.V.1995, leg. A. Smetana [T175], 4 exs. (cSme, cSch); Kaohsiung Hsien, Tengchih, 1400 m, 21.–23. VII. 2000, leg. H. Sugaya, 2 exs. (KUM, cSchü); Tengchih, 1400 m, 25.–30. IV. 2001, leg. H. Sugaya, 3 exs. (KUM, cSchü); Tengchih, 2100 m, 19.–23. IV. 2001, leg. H. Sugaya, 6 exs. (KUM, cSchü); Tengchih, 1700 m, 2.–3. IV. 2002, litter, leg. H. Sugaya, 2 exs. (KUM); Tona Forest Station, 1050 m, 27.–30.IV.1998 (flight intercept trap), leg. A. Smetana & L. Robillard [T193], 1 ex. (cSch); Road above Tona Forest Station, km 16–17, 1700–1800 m, 28.IV.1998, leg. A. Smetana [T190]. 2 exs. (cSme); Nantou Hsien: Meifeng, 2100 m, 6. V. 2001, leg. H. Sugaya, 20 exs. (KUM, cSchü); Meimu, 1600 m, litter, 30. III. 2002, leg. H. Sugaya, 5 exs. (KUM); Tehuashe, 900 m, 4.–5. V. 2001, leg. H. Sugaya, 1 ex. (KUM); Nanshanchi, 800 m, 27. X. 2000, leg. H. Sugaya, 2 exs. (KUM, cSchü); Meifeng, 2130 m, 4.V.1998, leg. A. Smetana [T199], 1 ex. (cSch); Pintung Hsien: Chialoshui, 100 m, 11. III. 2000, leg. M. Maruyama, 1 ex. (KUM); Taichung Hsien: Anmashan, 2150 m, 13.V.1992, leg. A. Smetana [T129], 1 ex. (cSme); Anmashan, 2225 m, 2.V.1990, leg. A. Smetana [T37], 1 ex (cSme); Anmashan, 2000 m, 24.–26.VI.2003, flight intercept trap, leg. Chun Lin Li, 4 exs. (NMB); Chipen- Wenchuan, 400 m, 6.–8. XI. 2000, leg. H. Sugaya, 3 exs. (KUM, cSchü); Chipen-Wenchuan, 400 m, 24.–27.IV.2001, leg. H. Sugaya, 5 exs. (KUM, cSchü); Taitung Hsien: road no. 20, after Li-Tao (Lidau), km 174, 1300 m, mountain forest litter, 8.IV.2007, leg. S. Vít, 2 exs. (cSch); W-Taitung, road no. 24, Jhihben, forest recreation area, 400 m, litter sifted, 12.IV.2007, leg. S. Vít, 1 ex. (cSch); Taipeh: Noi Dong logging road, 850 m, 19.II.2004, flight intercept trap, leg. Chun Lin Li, 3 exs. (MNB);

236 · Zootaxa 4105 (3) © 2016 Magnolia Press KOCIAN & SCHÜLKE Remarks. Ischnosoma quadriguttatum is one of the most widespread species of the genus, its distribution ranging from Pakistan to Indonesia. In China the species is very common in the central and southern parts of the country (Fig. 70). Records from Fujian, Gansu, Hainan, Hubei and Shaanxi are new province records.

Ischnosoma rosti (Bernhauer, 1923) Fig. 69.

CHINA: Beijing: Yan Shan, Dongling Mts., Xiaolongmen, 1400 m, 15.–16.VI.2001, leg. P. Hlaváč & J. Cooter, 2 exs. (cSch); Beijing, Badaling, 1.VII.1992, leg. M. Jäch, 1 ex. (NHMW); Gansu: S-Gansu, W-Qingling Shan, NW Longnan, 34°03'14''N, 104°10'00''E, 2200 m, SW-slope with shrubs, litter sifted, 1.VIII.2012, leg. V. Assing, 3 exs. (cAss, cSch); S-Gansu [CH12-15] Min Shan, 30 km W Longnan, 33°26'19''N, 104°36'17''E, 1470 m, deep cleft with scree, shrubs at N-slope, litter sifted, 5.VIII.2012, leg. M. Schülke, 1 ex. (cSch); S-Gansu, W.Qinling Shan, 101 km NW Longnan, 2200 m, 34°03'14''N, 104°10'00''E, SW-slope with shrubs, litter sifted, 1.VIII.2012, leg. D.W. Wrase [09], 3 exs. (cSch); Sichuan: Nanping, 6.–9.VI.2001, leg. E. Kučera, 1 ex. (cSch). Remarks. Ischnosoma rosti was described from Japan and later also recorded from Far East Russia (Kocian, 2003). The species is here recorded from China for the first time (Fig. 69).

FIGURE 71. Distribution of Ischnosoma splendidum (Gravenhorst) in China (material examined and literature records).

Ischnosoma splendidum (Gravenhorst, 1806) Figs 48, 71.

CHINA: Qinghai: Daban Shan, 62 km NNW Honggu, creek valley, Picea, Populus, Betula forest, 36°51'15–28''N, 102°36'34''–37'07''E, 2236–2350 m, creek valley, litter, dead wood and moss sifted, 26.VI.2011, leg. M. Schülke [CH11-09], 3 exs. (cSch, cKoc); Daban Shan, 25 km ESE Menyuan, 37°16'21.6''N, 101°52'37.4''E, 2795 m, loamy field edges, 6.VII.2011, leg. M. Schülke [CH11-22], 2 exs. (cSch); road from Honggu to Men Yuan, 37°08'46''N, 102°14'58.7''E, 2473 m, river valley, forest edge, semi-open pasture, sifted from mushrooms, 3.VII.2011, leg. M. Schülke [CH11-16b], 1 ex. (cSch); Lang Shi Dang Jing Qu (park), 75 km NW Honggu, 36°54'05.2''N, 102°21'07.1''E, 2925 m, creek valley on overgrown scree, Populus and Betula forest with Rhododendron, litter

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 237 sifted between rocks, 28.VI.2011, leg. M. Schülke [CH11-11a], 3 exs. (cSch); Sichuan: N-Sichuan, pass 35 km NNW Songpan, 32°55'32''N, 103°25'56''E, 3600 m, moist N-slope with Salix and other shrubs, litter, grass roots and moss sifted, 11.VIII.2012, leg. M. Schülke [CH12-24], 2 exs. (cSch). Remarks. Ischnosoma splendidum, a species of Holarctic distribution, is also distributed in the northern parts of China, southwards to the northernmost part of Sichuan province (Fig. 71). The records from Qinghai and Sichuan are new province records.

TABLE 1. Catalogue of the Ischnosoma species of China Species Distribution in China Distribution outside China absalom Kocian, 2003 Shaanxi, Yunnan (Fig. 61) – abstrusum sp. n. Sichuan, Taiwan, Yunnan (Fig. 19) – bergrothi (Hellén, 1925) Heilongjiang Northern and Eastern Europe, Siberia, Mongolia, Russian Far East bohaci Kocian, 2003 Fujian, Gansu, Shaanxi, Taiwan (Fig. 62) – bolitobioides (Bernhauer, 1923) Fujian, Gansu, Hubei, Sichuan, Shaanxi, Japan, Thailand Taiwan, Yunnan (Fig. 63) chinense (Bernhauer, 1939) Zhejiang (Fig. 61) – convexum (Sharp, 1888) Fujian, Hong Kong, Sichuan, Taiwan, Japan, India, Southeast Asia, Yunnan, Zhejiang (Fig. 64) Solomon Islands discoidale discoidale (Sharp, 1888) Gansu, Hubei (Fig. 65) Japan, Thailand discoidale taiwanum Kocian, 2003 Taiwan (Fig. 65) – duplicatum (Sharp, 1888) Fujian, Hubei, Sichuan, Taiwan, Yunnan, Japan, Nepal, India, Thailand Zhejiang (Fig. 66) evae Kocian, 2003 Gansu, Qinghai, Sichuan, Shaanxi, Yunnan – (Fig. 67) farkaci Kocian, 2003 Yunnan (Fig. 68) – fasciatocolle (Champion, 1922) Yunnan (Fig. 68) Nepal, India foedum sp. n. Yunnan (Fig. 19) – fusciventre (Tikhomirova, 1973) Beijing, Gansu, Hebei, Nei Mongol, Hubei, Japan, Russian Far East Shaanxi (Fig. 68) gemellum sp. n. Yunnan (Fig. 12) – hieroboam Kocian, 2003 Yunnan (Fig. 68) Myanmar involutum sp. n. Yunnan (Fig. 12) – maderi (Bernhauer, 1943) Beijing, Sichuan, Yunnan (Fig. 69) – mandschuricum (Bernhauer, 1923) Liaoning (Fig. 69) – noemi Kocian, 2003 Yunnan (Fig. 69) – quadriguttatum quadriguttatum Anhui, Fujian, Chongqing, Gansu, Guizhou, Nepal, Pakistan, India, Southeast (Champion, 1923) Hainan, Hong Kong, Hubei, Sichuan, Asia Shaanxi, Taiwan, Zhejiang (Fig. 70) rosti (Bernhauer, 1923) Beijing, Gansu, Shanxi (Fig. 69) Russian Far East ruth Kocian, 2003 Yunnan (Fig. 65) – splendidum (Gravenhorst, 1806) Hebei, Qinghai, Sichuan (Fig. 71) Europe, Middle Asia, Siberia, Japan, Canada, USA taibaiense Zhu et al., 2005 Shaanxi (Fig. 8) – thubal Kocian, 2003 Taiwan Thailand vicinum sp. n. Yunnan (Fig.12) – yunnanum sp. n. Yunnan (Fig. 19) – zipapertum sp. n. Yunnan (Fig. 8) –

CHINA: Shaanxi: S-Shaanxi, Qinling Shan, pass on road Zhouzhi–Foping, 105 km SW Xian, N-slope, 1990 m, 33°44'N, 107°59'E, small creek valley, mixed deciduous forest, bamboo, small meadows, litter, dead wood and mushrooms sifted, 2./4.VII.2001, leg. M. Schülke [CH01-01], 8 exs. (cSch); same data, but "leg. D.W. Wrase [01]", 1 ex. (cSch). Remarks. Ischnosoma taibaiense is known only from the surroundings of the Taibai Shan, the highest peak in the Qinling Shan range in southern Shaanxi (Fig. 8). All the specimens listed above were collected near the type locality.

Key to the Ischnosoma species of China and Taiwan

Ischnosoma mandschuricum, most likely a synonym of I. fusciventre and known only from the female holotype, is omitted in the key.

1 Anterior margin of pronotum without carina; exceptionally such a carina may be present only in lateral portions of anterior margin; best seen with opposite light reflection...... 2 - Anterior margin of pronotum with carina along all its length...... pictum species group. 7 2 Ocular seta as long and thick as setae of pronotum (Fig. 56)...... spelaeum species group. 3 - Ocular seta very short or missing...... I. convexum species group. 4 3 Larger, 3.1–3,6 mm (forebody 1.39–1.69 mm), elytra only with sutural, discal and lateral rows of punctures. Male sternite VII in the middle of posterior margin broadly and deeply concave, with median dent bearing some short setae (Fig. 5). Sternite VIII with posterior margin in the middle very deeply concave, with 6–7 rows of modified setae similar to opened zip (Fig. 6). Yunnan...... zipapertum sp. n. - Smaller, 2.5–3.1 mm (forebody 1.11–1.28 mm), elytra with additional non-setiferous punctation. Male sternites VI and VII only with slight emargination at posterior margin, both with large triangular fields of dense setae (Zhu et al., 2005: figures 3–4). Shaanxi...... taibaiense Zhu et al. 4 Elytra besides sutural, lateral, and discal rows of punctures with fine punctation and pubescence on whole surface. Zhejiang...... chinense (Bernhauer) - Elytra besides sutural, lateral and discal row of punctures without additional punctation and pubescence...... 5 5 Pale fine pubescence in the middle of male sternite VIII not reaching posterior margin of sternite (Kocian 2003: figures 5, 9, 12). Base of inner structure of aedeagus at most with one oblong finger-shaped structure (Kocian 2003: figure 6). Body generally darker, pronotum and anterior part of tergites III–V brown. Widely distributed in China, East and Southeast Asia...... convexum (Sharp) - Pale fine pubescence in the middle of male sternite VIII reaching posterior margin of sternite. At least one pair of setae situated at the margin (Kocian 2003: figures 42, 46, 61, 180). Base of inner structure of aedeagus with group of finger-shaped structures (Kocian 2003: figures 58, 185). Body generally paler, pronotum (sometimes with darker spot in anterior part) and tergites III–V reddish-yellow...... 6 6 Dark setae at posterior margin and in the middle of male sternite VII not reaching margin (Kocian 2003: figure 179). Two pairs of dark setae in the middle of male sternite VIII (Kocian 2003: figure 180). Macropterous species. Thailand, Taiwan...... thubal Kocian - Dark setae at posterior margin and in the middle of male sternite VII reaching margin (Kocian 2003: figure 45). Dark setae on male sternite VIII only in posterior and lateral parts of sternite (Kocian 2003: figure 46). Wing-dimorphic species, brachypterous or apterous specimens known only from Taiwan and Nepal. Widely distributed in South, East and Southeast Asia...... quadriguttatum quadriguttaum (Champion) 7 Elytra short, at suture shorter, as short as, or slightly longer than pronotum...... 8 - Elytra long, at suture distinctly longer (at least 1.1 times) than pronotum...... 16 8 Dorsal surface of head shiny, without microsculpture...... 12 - Dorsal surface of head with conspicuous microsculpture...... 9 9 Posterior margin of male sternite VII only slightly concave (Fig. 16). Sternite VIII: Fig 17, aedeagus: Fig. 18. Known from mountain ranges around Er Hai Lake in western Yunnan...... foedum sp. n. - Posterior margin of male sternite VII deeply concave in the middle (Kocian 2003: figures 391, 400, 407)...... 10 10 Setae in last row of median area of male sternite VIII flattened and pointed. Male sternite VIII as in Fig. 43. Aedeagus: Kocian (2003: figure 408). Yunnan...... ruth Kocian - Setae in last row of median area of male sternite VIII thin and fine, similar to other setae in this area. Male sternite VIII as in Figs 44 and 45. Aedeagus: Kocian (2003: figures 393, 396, 397, 402)...... 11 11 Median area of dense setae of male sternite VIII larger (Fig. 44; Kocian (2003: figure 392)). Japan, central China, Thailand...... discoidale discoidale (Sharp) (brachypterous morph)

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 239 - Median area of dense setae of male sternite VIII smaller (Fig. 45; Kocian (2003: figure 401)). Taiwan...... discoidale taiwanum Kocian 12 Smaller species, 3.8–5.0 mm (forebody 2.0–2.3 mm). Body uniformly reddish. Posterior angles of pronotum very well developed, angular. Male sternite VIII in posterior half with bald area, bordered by bow-like suture, and with median area of dense pointed and stouter setae (Kocian 2003: figure 440). Aedeagus: Kocian (2003: figures 441–442). Shaanxi and Yunnan...... absalom Kocian - Larger species, 5.0–6.0 mm (forebody 2.3–3.0 mm. Body coloration different. Male sternite VIII and aedeagus different. . 13 13 Body paler, reddish-brown to brownish, head, central part of elytra and anterior parts of tergites often darker, blackish. Prono- tum, elytra and abdomen with distinct bluish-golden hue. Pronotum narrower, posterior angles less marked. Apical margin of male sternite VIII broadly triangularly concave (Fig. 36), posterior half without bald area (Kocian 2003: figure 427). Aedea- gus: Kocian (2003: figure 428). Widely distributed from Qinghai southwards to Yunnan...... evae Kocian - Coloration different. Pronotum wider, posterior angles more distinct angular. Posterior half of male sternite VIII with distinct bald area...... 14 14 Pronotum pale reddish, posterior margin of elytra broadly pale. Posterior margin of male sternite VIII only with small triangu- lary emargination (Kocian 1997, Fig. 63). Aedeagus: Kocian (1997: figure 64). Northern and eastern Europe, Siberia, Mongo- lia, Russian Far East, in China only recorded from Heilongjiang...... bergrothi Hellén - Head, pronotum and elytra unicolorous, dark brown to black. Posterior margin of male sternite VIII very deeply triangularly concave (Figs 10, 42)...... 15 15 Bald apical area of male sternite VIII bordered by thick modified setae anteriorly, laterally, and posteriorly (Fig. 42). Aedea- gus: Kocian (2003: figure 455). Northern Yunnan...... farkaci Kocian - Bald apical area of male sternite VIII bordered by thick modified setae only anteriorly and posteriorly (Fig. 10). Aedeagus as in Fig. 11. Yunnan. (Diancang Shan)...... vicinum sp. n. 16 Elytra with two irregular discal rows of punctures (Kocian 2003: figure 316). Male sternites VII and VIII, and aedeagus: Kocian (2003: figures 308–312). Russian Far East, Japan, China, Thailand, Myanmar, Nepal and northern India...... duplicatum (Sharp) - Elytra with one discal row of punctures...... 17 17 Discal row of elytra with less than 10 punctures...... 18 - Discal row of elytra with more than 10 punctures...... 19 18 Smaller. Pronotum and elytra uniformly yellowish-red, elytra sometimes darker around scutellum. Head surface mostly with unconspicuous microsculpture. Male sternite VIII as in Fig. 48; aedeagus: Kocian (1997: figure 73). Holarctic species, in China known from Hebei, Qinghai, and Sichuan...... splendidum (Gravenhorst) - Larger. Elytra dark with paler humeral spot and posterior margin. Head surface shiny, without microsculpture. Male sternite VIII as in Fig. 38, aedeagus: Kocian (2003: figure 421). Russian Far East, Japan, northern and central China...... fusciventre (Tichomirova) 19 Posterior half of male sternite VIII with distinct bald area...... 20 - Posterior half of male sternite VIII without distinct bald area...... 21 20 Colorful species, head, pronotum and elytra black, only two transverse spots at the median part of margins of pronotum, humeral portions and posterior margins of elytra brightly yellowish. Anterior margin of bald area on male sternite VIII bordered with modified short and stout flattened (beard-like) setae (Fig. 34) and median area of dense pointed setae. Aedeagus as in Fig. 35. Yunnan...... gemellum sp. n. - Less colorful species, head black, pronotum entirely pale reddish, sometimes with indistinctly darkened anterior part. Elytra pale brown, variable darkened in the middle. Anterior margin of bald area on male sternite VIII bordered only with modified short and thick flattened (beard-like) setae (Kocian 2003: figure 443). Aedeagus as in Kocian (2003: figure 444). Japan, China, northern Thailand...... bolitobioides (Bernhauer) 21 Head of male with large protuberant eyes (Figs 58, 60). Width of head across eyes dinstincly larger than across temples. Prono- tum less transverse, smaller than elytra at shoulders. Species with distinct sexual dimorphism of eye size, shape of head, length of elytra and size of basal antennomeres...... 22 - Head of males with smaller, not protuberant eyes (Figs 57, 59). Width of head across eyes smaller than, or equal to width across temples. Pronotum more transverse. Species without distinct sexual dimorphism of eye size, shape of head, length of elytra and size of basal antennomeres...... 26 22 Areas of palisade and beard-like setae of male sternite VIII larger, covering one third or more of median length of sternite (Fig. 24)...... 23 - Areas of palisade and beard-like setae of male sternite VIII smaller, covering one quarter or less of median length of sternite (Fig. 21)...... 25 23 Areas of palisade and beard-like setae of male sternite VIII larger, forming sixs rows, in the middle not separated from each other, bald median area small and closed (Fig. 24). Yunnan...... yunnanum sp. n. - Areas of palisade and beard-like setae of male sternite VIII smaller, forming three to five rows, in the middle separated from each other, bald median area posteriorly opened...... 24 24 Shape of median bald area behind median area of dense setae more rectangular (Fig. 47). Areas of palisade and beard-like setae of male sternite VIII larger, forming five rows. Area of median setae smaller, posterior rows of setae flattened and pointed (Kocian 2003: figure 355). Aedeagus as in Kocian (2003: figure 356). Yunnan...... noemi Kocian - Bald area behind median area of dense setae not distinctly rectangular (Fig. 49). Areas of palisade and beard-like setae of male sternite VIII smaller, forming three or four rows. Area of median setae larger and denser, median dense setae thin and not dis-

240 · Zootaxa 4105 (3) © 2016 Magnolia Press KOCIAN & SCHÜLKE tinctly pointed (Kocian 2003: figure 330). Aedeagus as in Kocian (2003: figures 331, 336). Beijing, Sichuan, Yunnan...... maderi (Bernhauer) 25 Middle of posterior margin of male sternite VIII between areas of palisade and beard-like setae connected by pointed modified seta. Bald area behind median area of dense setae larger, median setae sparser (Fig. 21). Yunnan...... involutum sp. n. - Middle of posterior margin of male sternite VIII between areas of palisade and beard-like setae without modified setae. Bald area behind median area of dense setae smaller, median setae denser (Kocian 2003: figure 324). Russian Far East, Japan, Northern China (Beijing, Gansu, northern Sichuan)...... rosti (Bernhauer) 26 Head surface mostly with distinct microsculpture. Elytra relatively short, at suture about 1.2 times longer than pronotum. Pos- terior margin of male sternite VII very broadly concave, bordered by very sparse long thin setae (Fig. 37; Kocian (2003: figure 435)). Aedeagus as in Kocian (2003: figures 437, 438). Gansu, Shaanxi, Fujian, Taiwan...... bohaci Kocian - Head surface shiny, without microsculpture...... 27 27 Posterior margin of male sternite VII very broadly concave, bordered by very sparse long thin setae (as in Fig. 33) or denser thin setae (Kocian 2003: figure 344)...... 28 - Posterior margin of male sternite VII only in the middle broadly and deeply concave and bordered by dense very thin and long setae (as in Fig. 30)...... 29 28 Smaller species with shorter elytra (at suture about 1.2 times as long as pronotum). Coloration of head and pronotum uniformly pale. Posterior margin of sternite VIII bordered in median part by dense thin setae (Kocian 2003: figure 344). Aedeagus: Kocian (2003: figure 346). Yunnan, Myanmar...... hieroboam Kocian - Larger species with longer elytra (at suture more than 1.3 times as long as pronotum). Posterior margin of sternite VIII in the middle bordered by sparse thin setae (Kocian 2003: figure 410). Aedeagus as in Kocian (2003: figure 412). Northern India, Nepal, Yunnan...... fasciatocolle (Champion) 29 Larger species with long elytra (at suture more than 1.2 times as long as pronotum). Colour darker, head, pronotum and elytra brown, with paler humeral spot. Bald median area between areas of palisade and beard-like setae of sternite VIII smaller (Figs 31, 52). Aedeagus: Fig. 32...... abstrusum sp. n. - Smaller species of variable elytral length (at suture 0.9–1.2 times as long as pronotum). Colour paler, pronotum and elytra yellowish-red, head and elytra in the middle often darker. Bald median area between areas of palisade and beard-like setae of male sternite VIII larger (Figs 44; Kocian 2003: figure 392). Aedeagus: Kocian (2003: figures 393, 396, 397). Japan, central China, Thailand...... discoidale discoidale (Sharp) (macropterous morph)

Acknowledgments

We are indebted to the colleagues indicated in the material section for the loan of material under their care, especially D.W. Wrase, who accompanied the second author on almost all of the ten collecting trips to China between 1995 and 2013 and who donated all his staphylinid material to the second authors collection. Special thanks also go to Aleš Smetana (Ottawa) for the generous gift of some holotypes and other unique specimens. The second author thanks also Bernd Jaeger and Joachim Willers (MNB) for the opportunity to prepare habitus photographs and Volker Assing (Hannover) for a linguistic revision of the English manuscript.

References

Bernhauer, M. (1923 “1922”) Neue Staphyliniden der palaearktischen Fauna. Koleopterologische Rundschau, 10, 122–128. Bernhauer, M. (1939) Neuheiten der chinesischen Staphylinidenfauna (Col.). Mitteilungen der Münchner Entomologischen Gesellschaft, 29, 585–602. Bernhauer, M. (1943) Neue Staphyliniden der paläarktischen Fauna. Koleopterologische Rundschau, 29, 71–76. Cameron, M. (1949) New species and records of staphylinid beetles from Formosa, Japan, and South China. Proceedings of the United States National Museum, 99, 455–477. http://dx.doi.org/10.5479/si.00963801.3247.455 Campbell, J.M. (1991) A revision of the genera Mycetoporus Mannerheim and Ischnosoma Stephens (Coleoptera: Staphylinidae: Tachyporinae) of North and Central America. Memoirs of the Entomological Society of Canada, 156, 1–169. http://dx.doi.org/10.4039/entm123156fv Champion, G.C. (1922) Some Indian Coleoptera (7). The Entomologist's Monthly Magazine, 58, 31–34. Champion, G.C. (1923) Some Indian Coleoptera (10). The Entomologist's Monthly Magazine, 59, 43–53, 77–80. Ganglbauer, L. (1895) Die Käfer von Mitteleuropa. Die Käfer der österreichisch-ungarischen Monarchie, Deutschland, der Schweiz, sowie des französischen und italienischen Alpengebietes. Zweiter Band. Familienreihe Staphylinoidea. 1. Theil: Staphylinidae, Pselaphidae. Carl Gerold's Sohn, Wien, 881 pp. Gravenhorst, J.L.C. (1806) Monographia Coleopterorum Micropterorum. Henricus Dieterich, Gottingae, 236 + [12] pp. Hellén, W. (1925) Die nordeuropäischen Arten der Gattung Mycetoporus Mann. (Col.). Notulae Entomologicae, 5, 33–47.

ISCHNOSOMA FROM CHINA Zootaxa 4105 (3) © 2016 Magnolia Press · 241 Hansen, V. (1952) Danmarks Fauna. Bd. 58. Biller XVI. Rovbiller 2. Del. G. E. C. Gads Vorlag, København, 251 pp. Kocian, M. (1997 “1996”) A revision of Western Palearctic species of the genus Ischnosoma Stephens (Coleoptera, Staphylinidae: Tachyporinae). Acta Universitatis Carolinae. Biologica, 40, 241–299. Kocian, M. (2003) Monograph of the world species of the genus Ischnosoma (Coleoptera: Staphylinidae). Acta Universitatis Carolinae. Biologica, 47, 3–151. Kraatz, G. (1857) Naturgeschichte der Insecten Deutschlands. Erste Abtheilung Coleoptera. Zweiter Band. Lieferung 3–6. Nicolai, Berlin, pp. 377–1080. Li, J.-K. & Chen, P. (1993) [Studies on fauna and ecogeography of soil animal]. Northeast Normal University Press, Changchun, ii + 265 pp. [in Chinese with English title] Li, L.-Z., Zhao, M.-J. & Sakai, M. (2000) A new genus of the subfamily Tachyporinae (Coleoptera, Staphylinidae) with description of a new species from Japan. Japanese Journal of Systematic Entomology, 6, 11–18. Luze, G. (1901) Bolitobiini. Revision der paläarktischen Arten der Staphyliniden-Gattung Bryocharis Boisd. et Lac., Bolitobius Mannh., Bryoporus Kraatz und Mycetoporus Mannh. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, 51, 662–746. Palm, T. (1966) Svensk Insektfauna. 9. Häfte 4. Skalbaggar. Coleoptera. Kortvingar: Fam. Staphylinidae. Undefam. Habrocerinae, Trichophyinae, Tachyporinae. Entomologiska Föreningen i Stockholm, Stockholm, 91 + 2 pp. Rey, C. (1883) Tribu des brévipennes. Deuxieme Rameau Bolitobiates. Quatorzième et Quinzième familles: Tachyporiens et Trichophyens. Annales de la Société Linnéenne de Lyon, Series 2, 29, 13–125. Schülke, M. (1997–1998) Eine neue Art der Gattung Ischnosoma Stephens aus Griechenland (Insecta: Coleoptera: Staphylinidae: Tachyporinae). Reichenbachia, 32, 231–234. Schülke, M. (2001–2002) Eine neue Art und neue Funde von Arten der Gattung Ischnosoma Stephens, 1829 (Insecta: Coleoptera: Staphylinidae: Tachyporinae). Reichenbachia, 34, 127–135. Schülke, M. (2003) Eine neue Art und neue Funde von Arten der Ischnosoma spelaeum-Gruppe aus Anatolien (Insecta: Coleoptera, Staphylinidae, Tachyporinae). Linzer biologische Beiträge, 35, 453–460. Schülke, M. (2007) On the Turkish species of the genus Ischnosoma (Insecta: Coleoptera: Staphylinidae: Tachyporinae). Entomological Problems, 37 (1–2), 21–30. Sharp, D.S. (1888) The Staphylinidae of Japan. The Annals and Magazine of Natural History, Series 6, 2, 277–295, 369–387, 451–464. http://dx.doi.org/10.1080/00222938809460928 Stephens, J.F. (1829) The nomenclature of British insects; being a compendious list of such species as are contained in the Systematic Catalogue of British Insects, and forming a guide to their classification, &c. &c. Baldwin & Cradock, London, 4 + 68 pp. Tikhomirova, A.L. (1973) [new taxa]. In: Kryzhanovskii, O., Tikhomirova, A. & Filatova, L. (Eds.), Stafilinidy (Coleoptera, Staphylinidae) Yuzhnogo Primoria. In: Giliarov, M. (Ed.), Ekologiia pochvennykh bespozvonochnykh. Izdatelstvo Nauka, Moskva, pp. 144–173. Zhu, J.-W., Li, L.-Z. & Zhao, M.-J. (2005a) A new species of the genus Ischnosoma from China (Coleoptera, Staphylinidae, Tachyporinae). Acta Zootaxonomica Sinica, 30 (4), 809–811. Zhu, J.-W., Li, L.-Z. & Zhao, M.-J. (2005b) Ischnosoma bergrothi (Hellén) new to China (Coleoptera, Staphylinidae). Journal of Shanghai Normal University (Natural Sciences), 34 (4), 83–85.