Great Basin Naturalist Memoirs

Volume 9 A Utah Flora Article 4

1-1-1987 General References Stanley L. Welsh Monte L. Bean Life Science Museum and Department of and Range Science, Brigham Young University, Provo, Utah 84602

N. Duane Atwood USDA Forest Service, Intermountain Region, Ogden, Utah 84401

Sherel Goodrich Intermountain Forest and Range Experiment Station, Forest Service, U.S. Department of Agriculture, Ogden, Utah 84401

Larry C. Higgins , Department of Biology, West Texas State University, Canyon, Texas 79016

Follow this and additional works at: https://scholarsarchive.byu.edu/gbnm Part of the Anatomy Commons, Botany Commons, Physiology Commons, and the Zoology Commons

Recommended Citation Welsh, Stanley L.; Atwood, N. Duane; Goodrich, Sherel; and Higgins, Larry C. (1987) "General References," Great Basin Naturalist Memoirs: Vol. 9 , Article 4. Available at: https://scholarsarchive.byu.edu/gbnm/vol9/iss1/4

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This flora project, begun at Brigham Young University Kaye Thorne is here singled out for special thanks. Her in 1960, has a larger counterpart in the Intermountain duties at the herbarium of Brigham Young University, Flora Project of the New York Botanical Garden. Rather huge by any standard, were not an obstacle to helping than being competitive, although exclusive in personnel with the completion of the flora. She wrote two family and organization, the projects have been mutually collab- treatments, arranged for loans, provided illustrations of orative and beneficial. Drs. , Noel new taxa discovered, and did myriads of other tasks. Holmgren, Patricia Holmgren, James Reveal, and Rupert Many of the early collectors and avid botanists have C. Barneby have aided in identification and verification of passed from the scene. Included are Seville Flowers, critical material, have provided information on nomencla- Albert Osbun Garrett, Per Axel Rydberg, Marcus Eu- ture, have given locality data and information on new gene Jones, Sereno Watson, and John Charles Fremont. state records, allowed prior publication of nomenclatural Their contributions are here gratefully aknowledged. combinations and new taxa, and have given encourage- The writers are especially grateful to the new genera- ment and help at all times. Exchange specimens, both tion of collectors, including Gary Baird, Merton A. (Ben) those sent and those received, have been useful to both Franklin, Joel Tuhy, Betsy Neeley, and Ronald Kass. projects. We thank them for their help. Beverly Albee is here given special thanks for the loan of Specimens in critical groups have been routinely sent distribution maps showing specimens from the herbaria to specialists for verification and/or identification. Groups of the University of Utah and Utah State University. The and specialists are as follows: curators of the various herbaria are acknowledged for their help and suggestions. Compositae Arthur Cronquist Collaboration of officials of the various national parks Cirsium Gerald B. Ownbey and monuments, who so kindly granted collecting per- Boraginaceae Larry C. Higgins mits, is also acknowledged. The assistance of Larry Hays Leguminosae (general) Duane Isely at Zion National Park, Jim Holland, Vic Vierra, and Ron Astragalus {sind other Sutton at Glen Canyon National Recreation Area, and critical legumes) Rupert C. Barneby Lupintts David B. Dunn numerous people, including Superintendent Pete Perry Allium Dale B. McNeai of Canyonlands National Park was particularly helpful. Chrysothamnus Loran Anderson The former superintendent of Arches, Natural Bridges, Penstemon Noel Holmgren and Canyonlands, the late Bates Wilson, gave help and Umbelliferae Lincoln Constance encouragement during the early years of fieldwork lead- Various groups John Thomas Howell ing to this flora. Eleanor Inskip of the Canyonlands Natu- Eriogonum James L. Reveal ral History Association arranged for float trips through Loasaceae H. J. Thompson Cataract Canyon in collaboration with the National Park Gramineae Lois A. Amow Cruciferae Reed C. Rollins Service. Numerous federal and state agencies have col- Hydrophyllaceae N. Duane Atwood laborated in allowing and encouraging the collection of Cyperaceae Sherel Goodrich critical material. Donna House of the Navajo Heritage program aided in exploration of Navajo lands in Utah. Special thanks are hereby tendered to Drs. Dorald L. Thanks are given to Jody Chandler, subject matter Allred and Stephen L. Wood of the Life Science Museum librarian at the M. L. Bean Life Science Museum, branch at Brigham Young University for their help and encour- library, for her search of literature for chromosome num- agement. Drs. Bruce Smith and Jerran Flinders gave bers and for the citation of type locality information. financial assistance and allowed time for curation and Kathryn Taylor Mastin read the manuscript for typo- collection of materials. Many people have assisted with graphical errors and uniformity, especially of author ab- collection and identification of plant materials. Among breviations, and she assisted with preparation of the au- them are Mont Lewis, Kaye Thorne, Sherel Goodrich, thor list. The glossary was written by N. D. Atwood and Drs. Glen Moore, Joseph R. (Dick) Murdock, Bertrand supplemented by all other contributors. The index of F. Harrison, Walter P. Cottam, Arthur Holmgren, Bas- scientific names was formulated initially by N. D. Atwood sett Maguire, Elizabeth Neese, N. Duane Atwood, L. C. and S. Goodrich. Common names were extracted from Higgins, Steven L. Clark, and Richard Shaw. the text by Wendy Jefieries. GENERAL REFERENCES

References of a specific nature accompany many of the Atwood, N D and S L. Welsh 1985. New species ofTalinum families and genera in the taxonomic treatment. Many (Portulacaceae) from Utah, (ireat Basin Nat. 45: 485-487. other references have been used in a general way, and Bailey, L M 1951. Manual of cultivated plants. The MacMillan York. their use has contributed to the value of the flora. These (Company, New 1116 pp. 1976. (Revi.sed general references, many not cited in the text directly, Bailey, L M and K Z Bailey Hortus third. and expanded l)y the slaH of the Horto- are presented here. rium), MacMillan Fui)iisiiing C:o., Inc., New York. 12^X)

Ahnow, L, B. Albee, andA. Wyckoff. 1977, 1980. Flora ot the pp. central Wa.satch Front, Utah. Utah Mus. Nat. Hist., Salt Bahneby, R C and S, L. Welsh 1985. Ni'w species ofAstragalus Lake {>'ity. 662 pp. (Leguminosae) from southeastern Utah. (;reat Basin Nat. Atwood. N D.. S Goodhich. and S L Welsh. 1984. New A.s- 45:551-552. tragalus (Leguminosae) from the Goose Creek drainage, Ben.SON. L 1979. Plant classification. 2nd ed. D. C. Heath and Utah-Nevada. Great Basin Nat. 44: 263-264. Co., Lexington, Massachusett,s. 901 pp. 1987 Welsh et al; A Utah Flora, Introduction 11

Boh. N. L. 1960. The grasses of Burma, Ceylon, India, and Pak- 19.59. Va,scular plants of the Pacific Northwest. Part 4: istan. Pergamon Press, London. 767 pp. Ericaceae through Campanulaceae. Univ. Washington VV 1980. Publ. Bot. 17(4): 1-510. BROTHERSON. J D . L A SZYSKA. AN13 E EVENSON Poisonous plants of I'tah. Great Basin Nat. 40: 229-2.53. 1961. Va.scular plants of the Pacific Northwest. Part 3: 1982. Prelimi- Saxifragaceae to Ericaceae. Univ. Washington Publ. Bot. Chatterlky. M . B T Welsh, and S L Welsh 17(3): 1-614. nar\' inde.\ of authors of Utah plant names. Great Basin 1964. Vascular plants of the Pacific Northwest. Part 2: Nat. 42; 385-394. Salicaceae to Saxifragaceae. Univ. Washington Publ. Bot. ChRISTENSEN. E M 1967. Bibliography of Utah botany and wild- 17(2): 1-.597. land conservation. Brigham Young Univ. Sci. Bull. Biol 1969. Vascular plants of the Pacific Northwest. Univ. Ser. 9(1): 1-136. Washington Publ. Bot. 17(1): 1-914. CHRISTENSEN. E M andS L WELSH 19ft3. Pre.settleuicnt vege- Holmgren. .\ II 1948, 1957. Handbook ofthe va.scular plants of tation of the valleys of western Summit and Wasatch coun- the northern Wasatch. Lithotype Prk'css Company, San ties, Utah. Proc. Utah Acad. 40: 200-201. Francisco. 202 pp. CORRELL. D S AND M C JOHNSTON 1970. Manual of the vascu- 1960. Howell. J. T. E. McClintogk. and collaborators lar plants of Texas. Texas Research Foundation, Renner, Supplement, pp. 10.35-1076. In: Arizcma flora. University Texas. 1881 pp. of California Press, Berkeley and Los Angeles. Cronquist. a 1981. An integrated system of classification of Hubbakd. C E 1968. Grasses: A guide to their structure, identi- flowering plants. Columbia University Press, New York. fication, uses, and distribution in the British Isles. Revised 1262 pp. ed. Penguin Books). Harmondsworth. 463 pp. Cronquist. A.. A H. Holmgren. N H Holmgren, and J L Hulten. E 1964. The circumboreal plants. I. Almquist and Wik- Reveal. 1972. Intermountain flora. Vol. 1. Hafner Pub- sell. Stockholm. 275 pp. lishing Company, Inc., New York. 270 pp. Kass. R. ]. AND S L. Welsh. 1985. New species of Primula (Prim-

Cronquist. A . A H Holmgren. N H Holmgren. L Re J ulaceae) from Utah. Great Basin Nat. 45: .548-.5.50. veal, and p. K Holmgren 1977. Intermountain flora. Kearney, T H . R H Peebles, and collaborators 1969. Ari- Vol. 6. Columbia Univ. Press, New York. 584 pp. zona flora. University of California Press, Berkeley and DORN. R. 1977. Manual of the vascular plants of Wyoming. D Los Angeles. 1032 pp. Garland Publishing, Inc., New York. Vol. 1 and 2. 1498 Kingsbury, J M 1964. Poisonous plants ofthe and pp. Canada. Prentice-Hall, Inc., Englewood Cliffs, New Fremont, C. 1845. Report of exploring expedition to the Rocky J. Jersey. 626 pp. Mountains in the year 1843, and to Oregon and North Knobloch, I W 1963. Hybridization in the Gramineae. Dar- 1843-44. D.C. California in the years Washington winiana 12: 624-628. Garrett, A O 1936. Spring flora of the Wasatch region. Ed. 5. McVaugh. R 1983. Flora Novo-Galiciana. Vol. 14. Gramineae. Stevens & Wallis, Inc., Salt Lake City. 240 pp. Ed. W. R. Anderson. Univ. Mich. Press. Ann Arbor. 4.36 Gleason, H a and a Cronquist 1963. Manual of the vascular pp. plants of the northeastern United States and adjacent MUNZ. P A. 1968. Supplement to a California flora. University of Canada. Van Nostrand Reinhold Co., New York. 810 pp. California Press, Berkeley. 224 pp. Goodrich. S and S. L. Welsh. 1983. New variety of 1970. A California flora. University of California Press, Stephanomeria tenuifolia (Compositae) from Utah. Great Berkeley. 1681 pp. Nat. 43: 373-374. Basin MuRDOCK. J R and S L. Welsh 1971. Land use in Wah Wah Gould, F. W 1951. Grasses of the southwestern United States. and Pine vallevs, western Utah. Brigham Young Univ. Sci. Univ. Ariz.. Biol. Sci. BuU. 7: 1-352. Bull. Biol. Ser. 12(4): 1-25. 1975. The grasses of Texas. Texas A & M Press. College Neese, E. and S L. Welsh 1983. A new species oi Penstemon Station, Texas. 653 pp. (Scrophulariaceae) from the Uinta Basin, Utah. Great Basin Nat. 43: 373-374. Gould. F W , and R Moran 1981. The grasses of Baja Califor- nia, Mexico. San Diego Society of Natural History, San NiCORA, E. G 1978. Flora Patagonica, Part 3. Gramineae. Collec-

I. N. T. A., Buenos Aires, Argentina. Diego. 140 pp. cion Cientifica del Northstrom. E and S L Welsh 1970. Revision of the Gould, F. W and R. B. Shaw 1983. Grass systematics. 2d ed. T boreale complex. Great Basin Naturalist 30: Texas A & M Press, College Station. Texas. 397 pp. Hedysarwn 109-130. Graham. E H 1937. Botanical studies in the Uinta Basin of Utah Rehder, a 1949. Bibliography of cultivated trees and shrubs. and Colorado. Ann. Carnegie Mus. 26: 1-432. Arnold Arboretum of , Jamaica Plain, 1980. Halliday. P , R D Meikle, J Story, and H Wilkinson. Massachusetts. 825 pp. Draft index of author abbreviations compiled at the Scoggan. H 1978. The flora of Canada. Part 2. Natl. Museum herbarium. Royal Botanic Gardens, Kew. Her Majestys J. Canada, Ottawa. .545 pp. Stationer Office. 249 pp. Thornburg. a a 1982. Plant materials for use on surface-mined Hanson. A. A. 1972. Grass varieties in the United States. U.S. lands in arid and semiarid regions. U.S. Dept. Agric, Dept. Agric. Handbook No. 170: 1-124. SCS-TP-1.57. 88 pp. Harrington, H. D 19.54. Manual ofthe plants of Colorado. Sage TiDESTROM. I 192.5. Flora of Utah and Nevada. Contr. U.S. Natl. Books, Denver. 666 pp. Herb. 25: 1-665. Harrison. B F.. S L Welsh, and G Moore 1964. Plants of 1972. Evolu- TiDWELL. W. D . S R RUSHKOHTH. .AND D SIMPER. Arches National Monument. Brigham Y'oung Univ. Sci. tion of floras in the Intermountain Region, pp. 19-39. In: Bull. Biol. Ser. 5(1): 1-2.3. Intermountain Flora. Vol. 1. Hafner Publishing Com- Hintze, L F 1972. Geologic history of Utah. Brigham Young pany, New York. University, Dept. of Geology. Provo. 181 pp. 1964. to the woody Treshow. M , S L Welsh, and G Moore Guide Hitchcock, A S. 1935. Manual of the grasses of the United plants of Utah. Ed. 2. Purett Press, Boulder, Colorado. 160 pp. States. U.S. Dept. Agric. Misc. Publ. 200: 1-1040. Tsvele\'. N N 1984. Grasses of the Soviet Union. Parts 1 and 2. 1951. Manual ofthe grasses ofthe United States. 2nd ed., Russian Translation Series 8. A. A. Balkema, Rotterdam. 11% revised by Agnes Chase. U.S. Dept. Agric. Misc. Publ. pp.

200: 1-1051. TuTiN. T G . V H Heywood. N A Burges, D M Moore. D H Valentine. S Walters, and D. A. Webb 1964-1980. Hitchcock, C. L , A Cronquist, M Ownrey. and J W Thomp M SON 1955. Vascular plants ofthe Pacific Northwest. Part 5: Flora Europaea. Vol. 1-5. Cambridge University Press, Cam- Compositae. Univ. Washington Publ. Bot. 17(5): 1-343. bridge. 12 Great Basin Naturalist Memoirs No. 9

USDA Soil Conservation Service 1982. National list of scien- 1982c. A new species of Cnjptantha (Boraginaceae) dedicated tific plant names. SCS-TP-159. Vol. 1. U.S. Government to the memory of F. Creutzfeldt. Great Basin Nat. 42: 203-204. Printing Office. 416 pp. 1983. A bouquet of daisies (Erigeron, Compositae). Great Vallentine.J F 1961. Important Utah range grasses. Utah State Basin Nat. 43: 365-368. Univ. Ext. Circ. 281; 1-48. 1983b. New taxa in Thelespemxa and Townsendia (Compositae) VOSS. E G 1972. Michigan flora. Part 1. Cranbrook Institute of from Utah. Great Basin Nat. 43: 369-370. Science and University of Michigan Herbarium, Bloom- 1986. History of botanical exploration in Utah: The Fremont field Hills. 488 pp. era. Utah Acad. Sci. (in press). ed. 1983. International code of botanical nomenclature. Welsh, S L and N D Atwood 1977. An undescribed species of Adopted by the 13th International Botanical Congress, Thelypodiopsis (Brassicaceae) from the Uinta Basin, Utah. Sydney, August 1981. Bohn, Scheltema & Holkema, Great Basin Nat. 37: 95-96. Utrecht. 472 pp. Welsh, S L, N D Atwood, S Goodrich, E Neese, K H Thorne. Watson, S. 1871. Botany. In C. King, Report of the geological AND B Albee 1981. Preliminary index of Utah vascular plant exploration of the fortieth parallel. Vol 5. Washington: names. Great Basin Nat. 41: 1-108.

Government Printing Office. 525 pp. Welsh. S L , N D Atvvood, and J R Murdock 1978, Kaiparowits Weber, W A 1976. Rocky Mountain Flora. Colorado Assoc. flora. Great Basin Nat. .38: 125-179.

Univ. Press, Boulder. 479 pp. Welsh, S L, N D Atwood, and J L Reveal 1976. Endangered Welsh, S L 1957. An ecological survey of the vegetation of the threatened, extinct, endemic, and rare or restricted Utah Dinosaur National Monument, Utah. Unpublished The- plants. Great Basin Nat. .35: 327-376. sis, Brigham Young University. 86 pp. Welsh, S L, and R C Barneby 1981, A,s(raga/u,s lentiginosus (Fa- 1970a. New and unusual plants from Utah. Great Basin baceae) revisited — a unique new variety. Isleya 2(1): 1-2.

Nat. 30: 16-22. Welsh, S L , and S Goodrich 1980, Miscellaneous plant novelties 1970b. An undescribed species of Astragalus (Legumi- from Alaska, Nevada, and Utah, Great Basin Nat. 40; 78-88. nosae)from Utah. Rhodora72: 189-193. Welsh, S L, andG Moore 1968. Plants of Natural Bridges National 1971. Description of a new species of Da/ea(Leguminosae) Monument. Proc. Utah Acad. 45: 220-248. from Utah. Great Basin Nat. 31: 90-92. 1973. Utah plants: Tracheophyta. Brigham Young University 1974a. Anderson's flora of Alaska. Brigham Young Univ. Press, Provo, Utah. 473 pp.

Press, Provo, Utah. 724 pp. Welsh. S L , and E Neese 1983a. New species in Hymenoxys and 1974b. Utah plant novelties in Astragalus and Yucca. Perityle (Compositae). Great Basin Nat. 43: 369-.370. Great Basin Nat. 34: 305-310. 1983b. New variety ofOpuntia basilaris (Cactaceae) from Utah. 1975. Utah plant novelties in Cymopfenw and Penstemon. Great Basin Nat. 43: 700.

Great Basin Nat. 35: 377-378. Welsh. S L , and B W Olsen 1969. A living, prehistoric lumber 1978a. Problems of plant endemism on the Colorado tree. Proc. Utah Acad. 46: 149-159.

Plateau. Memoirs Great Basin Nat. 2: 191-195. Welsh, S L, and J L Re\Eal 1968. A new species of Townsendia 1978b. Endangered and threatened plants of Utah: a re- from Utah. Brittonia20: 375-.377.

evaluation. Great Basin Nat. 38: 1-18. Welsh, S L , and F G Smith 1983. New Haplopappus variety in 1981. New taxa of western plants— in tribute. Brittonia.33: Utah (Compositae). Great Basin Nat. 43: 371-372. 294-303. Welsh, S L, andK H Thorne 1978. Illustrated manual of proposed 1982a. Utah plant types— Historical perspective 1840 to endangered and threatened plants of Utah. U. S. Fish and 1981 —Annotated list and bibliography. Great Basin Nat. Wildlife Publication, Denver, Colordo. 318 pp.

42: 1-44. Welsh, S L , and C A Torr, 1981, Biotic communities of hanging

1982b. New taxa of thistles {Cirsium , Asteraceae) in Utah. gardens in southeastern Utah. National Geographic Society Great Basin Nat. 42: 199-202. Research Reports 13: 663-682.

GROUP TRACHEOPHYTA - VASCULAR PLANTS

Plants with a well developed vascular system (xylem .3(1). Plants fernlike and with broad leaves or free-floating aquat- and phloem) in the sporophyte generation, leaves (either ics with small overlapping leaves; reproduction by macrophylls or microphylls), and roots; reproduction by spores; flowers and woody cones lacking Polypodiophyta, 15 spores, or by microspores and megaspores, and by seeds p.

(in most), the latter borne in cones or flowers. — Plants neither fernlike nor free-floating aquatics (except in Lemnaceae); reproduction by .spores and seeds, these Key to the Divisions and Classes. borne in flowers or cones 4

I. Plants with small scalelike leaves, usually with a single 4(3). Seeds not borne enclosed by ripening caqiels, but vein (microphylls); reproduction by means of spores; naked and situated on the surface of a scale, these borne

flowers or woody cones lacking 2 crowded together on an iixis and forming a cone; flowers

not developed; leaves typically needle- or scalelike . . — Plants with large leaves, usually with more than a single Pinophyta, p. 24 vein (macrophylls), if scalclike, as occasionally, other- wise different from above; reproduction by spores or — Seeds borne in ripening carpels; plants with flowers;

seeds, the latter borne in flowers or cones 3 leaves mainly not needle- or scalelike (Magnoliophyta) . . 5

2(1). Stems jointed, fluted, and hollow in the internodes; 5(4). Flower parts mainly 4- or 5-merous; leaves typically leaves not green, reduced to a whorl of connate scales at net veined; stems increasing in diameter by means of a the nodes; plants neither grass- nor mosslike cambium between the xylem and phloem; cotyledons Etjuisetophyta, p. 9 typically 2 Magnoliopsida, p. .35

— Stems not jointed; leaves green and imbricated, not — Flower parts typically 3-merous; leaves typically paral-

whorled or forming a sheath at the nodes; plants either lel veined; stems usually lacking a cambium or, if aquatic and grasslike or terrestrial and mosslike present, producing entire vascular bundles; cotyledon I Lycopodiophyta, p. 13 Liliopsida, p. 647