The Early Stages of Pedaliodes poesia () in Eastern Ecuador (Lepidoptera: Satyrinae: Pronophilina) Author(s): Harold F. Greeney, Tomasz W. Pyrcz, Philip J. DeVries and Lee A. Dyer Source: Journal of Insect Science, 9(38):1-9. 2009. Published By: Entomological Society of America DOI: http://dx.doi.org/10.1673/031.009.3801 URL: http://www.bioone.org/doi/full/10.1673/031.009.3801
BioOne (www.bioone.org) is a nonprofit, online aggregation of core research in the biological, ecological, and environmental sciences. BioOne provides a sustainable online platform for over 170 journals and books published by nonprofit societies, associations, museums, institutions, and presses. Your use of this PDF, the BioOne Web site, and all posted and associated content indicates your acceptance of BioOne’s Terms of Use, available at www.bioone.org/page/terms_of_use. Usage of BioOne content is strictly limited to personal, educational, and non-commercial use. Commercial inquiries or rights and permissions requests should be directed to the individual publisher as copyright holder.
BioOne sees sustainable scholarly publishing as an inherently collaborative enterprise connecting authors, nonprofit publishers, academic institutions, research libraries, and research funders in the common goal of maximizing access to critical research. .SYVREPSJ-RWIGX7GMIRGI:SP`%VXMGPI +VIIRI]IXEP
8LIIEVP]WXEKIWSJ 4IHEPMSHIWTSIWME ,I[MXWSR MR IEWXIVR)GYEHSV 0ITMHSTXIVE7EX]VMREI4VSRSTLMPMRE ,EVSPH*+VIIRI]E8SQEW^;4]VG^F4LMPMT.(I:VMIWG ERH0II%(]IVH =ERE]EGY&MSPSKMGEP7XEXMSR 'IRXIVJSV'VIEXMZI7XYHMIW'SWERKE)GYEHSVGS*SGL]%QE^SREW5YMXS )GYEHSV >SSPSKMGEP1YWIYQSJXLI.EKMIPPSRMER9RMZIVWMX]-RKEVHIRE/VEOz[4SPERH 9RMZIVWMX]SJ2I[3VPIERW(ITEVXQIRXSJ&MSPSKMGEP7GMIRGIW2I[3VPIERW0% &MSPSK](ITEVXQIRX9RMZIVWMX]SJ2IZEHE 6IRS2IZEHE
%FWXVEGX ?MLM[KZQJM\PMQUUI\]ZM[\IOM[ 8MLITQWLM[XWM[QI 0M_Q\[WV NZWUVWZ\PMI[\MZV-K]ILWZ +P][Y]MI[KIVLMV[ 8WIKMIM *IUJ][WQLMIQ[\PMTIZ^ITNWWLXTIV\-OO[IZMTIQL[QVOTaWZQVXIQZ[WV\PMJW\\WU[QLMWNPW[\XTIV\TMI^M[ /I][SVHW %RHIWFEQFSS 'LYWUYIEGPSYHJSVIWXPEVZE4SEGIEITYTE 'SVVIWTSRHIRGI EVIZQQSWW$]ELSSGSQ FT]VG^XSQEW^$LSXQEMPGSQ GTHIZVMIW$YRSIHY HPH]IV$YRVIHY 6IGIMZIH %TVMP` %GGITXIH .YRI` 4YFPMWLIH .YRI %WWSGMEXI)HMXSV .MQ1MPPIV[EWIHMXSVSJXLMWTETIV 'ST]VMKLX 8LMWMWERSTIREGGIWWTETIV;IYWIXLI'VIEXMZI'SQQSRW%XXVMFYXMSRPMGIRWIXLEXTIVQMXWYRVIWXVMGXIHYWITVSZMHIHXLEX XLITETIVMWTVSTIVP]EXXVMFYXIH -772 `:SPYQI2YQFIV 'MXIXLMWTETIVEW +VIIRI],*4]VG^8;,YEPMRKYE01(I:VMIW4.(]IV0%8LIIEVP]WXEKIWSJ 4IHEPMSHIWTSIWME ,I[MXWSR MRIEWXIVR)GYEHSV 0ITMHSTXIVE7EX]VMREI4VSRSTLMPMRE TT .SYVREPSJ-RWIGX7GMIRGI EZEMPEFPISRPMRIMRWIGXWGMIRGISVK .SYVREPSJ-RWIGX7GMIRGI`[[[MRWIGXWGMIRGISVK .SYVREPSJ-RWIGX7GMIRGI:SP`%VXMGPI +VIIRI]IXEP -RXVSHYGXMSR [\I\][ QV XIZ\QK]TIZ \PM +PWK~ IVL -T .SYVREPSJ-RWIGX7GMIRGI`[[[MRWIGXWGMIRGISVK .SYVREPSJ-RWIGX7GMIRGI:SP`%VXMGPI +VIIRI]IXEP +IJI}I[ ;IV 1[QLZW IVL QVKT]LM[ IZW]VL PMK\IZM[ 8LMVHMRWXEV *MKYVIWKLOQ WN XZQUIZa KTW]L NWZM[\ JWZLMZML Ja KI\\TM XI[\]ZM IVL R ! FSH] PIRKXL ! XS QQ HIZIPSTQIRX W\PMZ LQ[\]ZJML PIJQ\I\[ C[MM /ZMMVMa M\ IT IVL XMQI!HE]W >ITMVKQI !! NWZ UWZM KWUXTM\M LM[KZQX\QWV[ WN \PM 0MIL IVL JWLa [QUQTIZ \W [MKWVL QV[\IZ J]\ QVLQ[\QVK\Ta [\]La IZMIE ?M KWTTMK\ML TIZ^IM I\ MTM^I\QWV[ ZIVOQVO [Y]IZM QV KZW[[[MK\QWV# JWLa KWTWZI\QWV W^MZITT UWZM aMT NZWU \W U IVL ZMIZML \PMU QV OTI[[ RIZ[ I\ \PM TW_Q[P# PMIL PWZV[ IVL KI]LIT \IQT[ [TQOP\Ta UWZM IUJQMV\ZM[MIZKPTIJTWKI\MLI\U XZWVW]VKML ?M ZMIZML [M^MZIT TIZ^IM NZWU \P QV[\IZ \W MKTW[QWV IVL *SYVXLMRWXEV *MKYVIWP EG \_W QVLQ^QL]IT[ NZWU MOO[ NW]VL QV \PM NQMTL \W IL]T\[ R ! FSH] PIRKXL ! XS QQ HIZIPST ?M ILLML NZM[P NWWL XTIV\ I[ VMMLML ZMUW^QVO NZI[[ IVL QIRXXMQI!¯HE]W WTL TMI^M[ LIQTa ?M UILM TIZ^IT UMI[]ZMUMV\[ \PM LIa 0MIL [QUQTIZ \W \PQZL QV[\IZ PWZV[ [TQOP\Ta UWZM XZW XZQWZ\WUWT\QVO VW]VKML TQOP\ KIZIUMTKWTWZML I\ UWT\QVO J]\ LIZSMVQVO \W WZIVOMJZW_V I[ TIZ^I UI\]ZM[ LIZSMZ IV\MZQWZTa .QO]ZM J# JWLa [QUQTIZ QV NWZU \W \PQZL QV[\IZ KI]LIT 6IWYPXW \IQT[ TWVOMZ LQ[\QVK\Ta [Y]IZM QV KZW[[[MK\QWV# OMVMZIT ZL )KK *MKYVIWEG JWLa KWTWZI\QWV VW\IJTa LQNNMZMV\ NZWU QV[\IZ R ! ETTVS\ QQ HMEQIXIV XMQI XS \PWZIKQK [MOUMV\[ _Q\P LQ[\QVK\ OZMMVQ[P PQOPTQOP\[ LIZS LEXGLMRKHE]W MVQVO IVL M`\MVLQVO \W ) TI\MZ TI\MZ QV \PQ[ [\ILQ]U# \MZ :W]VL _PQ\M IXXMIZQVO [UWW\P J]\ _Q\P UQV]\M QZZMO UQVIT [MOUMV\[ IVL ) ) IT[W _Q\P OZMMVQ[P PQOP ]TIZ ^MZ\QKITTa WZQMV\ML [\ZQI\QWV[ ^Q[QJTM ]VLMZ LQ[[MK\QVO TQOP\[# JWLa OZW]VL KWTWZ JMQOM _Q\P \PQV TWVOQ\]LQVIT \P [KWXM .QO]ZM I# TIQL [QVOTa V % WZ QV XIQZ[ V % # LWZ[WTI\MZIT IVL ^MV\ZWTI\MZIT _PQ\M [\ZQXM[# UI\]ZM XZQWZ \W MUMZOMVKM TIZ^IT PMIL KIX[]TM[ KTMIZTa ^Q[QJTM QV[\IZ _Q\P I \PQKS UQLLWZ[IT [\ZQXM Z]VVQVO MV\QZM JMTW_ KPWZQWV .QO]ZM[ JK# ]XWV MUMZOMVKM TIZ^IM TMVO\P WN JWLa OZMMVQ[P IV\MZQWZTa NILQVO \W ZMLLQ[P KWV[]UMMV\QZMMOO[PMTT KMV\ZITTa \PMV \W OZMMVQ[P IOIQV XW[\MZQWZTa MV\QZM JWLa _Q\P PQV\[ WN ZW[MKWTWZML JT][P# TI\MZ QV [\ILQ]U LM^MTWX *MVWXMRWXEV *MKYVIWGHM QVO XIQZ[ WN QVLQ[\QVK\ [UITT LIZS [XW\[ LWZ[WTI\MZITTa I\ R ! FSH] PIRKXL ! ¯ QQ HIZIPSTQIRX QV\MZ[MOUMV\IT[]\]ZM[WV)\PZW]OP) XMQI!¯HE]W 0MIL VMIZTa ZW]VL KIZIUMTKWTWZML UQV]\MTa ZM\QK] *MJXLMRWXEV *MKYVIWR HI TI\ML _Q\P [XIZ[M TWVO XITM [M\IM# JWLa I\ PI\KPQVO _PQ\M R ! FSH] PIRKXL ! XS QQ HIZIPSTQIRX .QO]ZM[ JK _QLM[\ I\ ) \IXMZQVO OZIL]ITTa XW[\MZQ XMQI!¯HE]WMRGPYHMRKTVITYTE WZTa IVL \MZUQVI\QVO QV I XIQZ WN XWWZTa LM^MTWXML KI]LIT 0MIL XITMZ \PIV QV XZM^QW][ QV[\IZ[ J]\ _Q\P LQ[\QVK\Ta \IQT[# [WN\ XITM [M\IM TI\MZITTa UW[\ KWVKMV\ZI\ML WV LIZSMZ IV\MZQWZ XWZ\QWV MXQKZIVQIT PWZV[ _MTT LMNQVML J]\ )!·)# XZW\PWZIKQK [PQMTL _MISTa [KTMZW\QbML KTMIZ IVL [\QTT [PWZ\ IVL ZW]VLML .QO]ZM V# JWLa \ZIXMbWQLIT QV JIZMTa VW\QKMIJTM .QO]ZM K# TI\MZ QV [\ILQ]U .QO]ZM L KZW[[[MK\QWV VIZZW_MZ LWZ[ITTa _QLM[\ IZW]VL ) TIZ^IM IZM OZMMVQ[P NZWU PW[\ XTIV\ UI\MZQIT ^Q[QJTM QV \PM OZW]VL KWTWZ XITM JMQOM _Q\P NQVM QVLQ[\QVK\ _I^a JMQOM O]\# JWLa ZW]VL QV KZW[[[MK\QWV IVL LM^MTWXQVO NQ^M XIQZ[ \W XQVSQ[P XI\\MZVQVO UQLLWZ[IT [\ZQXM _MTT LMNQVML WN QVLQ[\QVK\ TWVOQ\]LQVIT _PQ\M [\ZQXM[ JMOQVVQVO WV ) OZMMVQ[P \W ) IVL NZWU )!· ZMLLQ[P JZW_V KMV\ZITTa LIZSMVQVO XW[\MZQWZTa IVL \MZUQVI\QVO WV )!# R][\ XZQWZ OZMMVQ[P XWZ\QWV[ TQVML []JLWZ[ITTa _Q\P LIZS JZW_V \W UWT\QVO TIZ^IM LM^MTWX I LQ[\QVK\ ZMLJZW_V UQL JTIKS# [Q` XIQZ[ WN [UITT LQ[\QVK\ LIZS JZW_V LWZ[WTI\MZIT LWZ[IT [\ZQXM# KI]LIT \IQT[ JMKWUM UWZM ^Q[QJTM J]\ ZM [XW\[ NZWU <)# LWZ[WTI\MZIT _PQ\M [\ZQXM \PQKSMVQVO QV UIQV[PWZ\ \W QVLQ[\QVK\ KZM[KMV\[ WV KMV\ZIT IJLWUQVIT [MOUMV\[ IVL MV\QZM [\ZQXM \W )! []J\MVLML Ja LQ[\QVK\ J]\ QZZMO] 7IGSRHMRWXEV *MKYVIWIJL TIZ \PQV LIZS JZW_V [\ZQXM \PQ[ [\ZQXM \PQKSMZ M`\MVLQVO R ! FSH] PIRKXL ! XS QQ HIZIPSTQIRX ^MV\ZITTa QV\W [Y]IZM XI\KPM[ WV < IVL )# [XQZIKTM[ XMQI!¯HE]W LIZS# TI\M QV [\ILQ]U UW[\ XI\\MZV NILQVO [MM XZMX]XIT 0MIL ZW]OPTa Y]ILZI\M KIZIUMT JZW_V NQVMTa XQ\\ML IVL LM[KZQX\QWVM`KMX\NWZLIZSIZMI[WV<IVL)·) ZM\QK]TI\ML _Q\P UWLMZI\MTa LMV[M [PWZ\ XITM [M\IM MXQ KZIVQ]U JMIZQVO \_W [PWZ\ ZW]VLML PWZV[# JWLa ZW]VL 4VITYTE *MKYVIWI EF QV KZW[[[MK\QWV [QUQTIZ QV KWTWZI\QWV \W TI\M NQZ[\ QV[\IZ R ! PIRKXL ! ¯ QQ HIZIPSTQIRX XMQI M`KMX\ LWZ[WTI\MZIT IVL ^MV\ZWTI\MZIT _PQ\M TWVOQ\]LQVIT !¯HE]W [\ZQXM[ JMKWUQVO UWZM KWV\ZI[\QVO \PIV W\PMZ[# JWLa VW_ ;QUQTIZ \W TI\M NQN\P QV[\IZ J]\ JWLa ITUW[\ MV\QZMTa JMIZ[ UQV]\M OZIV]TI\QWV[ IVL [XIZ[M [PWZ\ XITM [M\IM \ZIV[T]KMV\_PQ\MXQVS\WJMQOM[QTSXIL_PQ\M\WJMQOM KI]LIT \IQT[ TWVOMZ IVL UWZM M^QLMV\ IVL UWZM _QLMTa [MXIZI\ML .SYVREPSJ-RWIGX7GMIRGI`[[[MRWIGXWGMIRGISVK .SYVREPSJ-RWIGX7GMIRGI:SP`%VXMGPI +VIIRI]IXEP *MKYVI -QQEXYVIWXEKIWSJ 4IHEPMSHIWTSIWME EX=&7E JVIWLP]PEMHIKKW FG VIGIRXP]LEXGLIH PEVZEERHTVILEXGLMRKIKK H TVIQSPXJMVWXMRWXEV I VIGIRXP]QSPXIHWIGSRHMRWXEV J TVIQSPX WIGSRHMRWXEV KLVIGIRXP]QSPXIHXLMVHMRWXEV M JMVWXMRWXEV N WIGSRHMRWXEV O XLMVHMRWXEV P JSYVXLMRWXEV Q XLMVHMRWXEV R JMJXLMRWXEV 4YTE *MKYVIW GJ R! PIRKXL ! ¯ IJLWUMV [QUQTIZ QV KWTWZ IVL WVTa [TQOP\Ta [K]TX\]ZML# QQ HIZIPSTQIRXXMQI!¯HE]W [M^MZIT LIa[ JMNWZM MKTW[QWV \PM MV\QZM X]XI JMKWUM[ 7^MZITT NWZU ZWJ][\ ZW]OPTa ZMK\IVO]TIZ J]\ _Q\P JT]V\ LIZSJZW_V.QO]ZM MLOM[ M`KMX\ NWZ QVLQ[\QVK\ ZW]VLML ZQLOM[ LWZ[WTI\MZITTa WV IJLWUMV IVL PMIL \IXMZQVO [TQOP\Ta IVL [Y]IZML WNN IV\MZQWZTa# OZW]VL KWTWZ XITM JMQOM _Q\P QZZMO]TIZ JZW_V (MWGYWWMSR NTMKSQVO IVL XI\\MZVQVO XI\\MZVQVO UW[\ LQ[\QVK\ IVL =VNWZ\]VI\MTa \PM XI]KQ\a WN X]JTQ[PML TQNM PQ[\WZa QV M`\MV[Q^M UQLLWZ[ITTa IVL LIZSM[\ QV \PZMM \ZQIVO]TIZ NWZUI\QWV WV 8MLITQWLM[ TMI^M[ ][ _Q\P NM_ [XMKQM[ NWZ XI\KPM[ \_W R][\ LWZ[IT WN _QVO XIL[ IVL WVM KMV\ZITTa KWUXIZQ[WV .SYVREPSJ-RWIGX7GMIRGI`[[[MRWIGXWGMIRGISVK .SYVREPSJ-RWIGX7GMIRGI:SP`%VXMGPI +VIIRI]IXEP *MKYVI -QQEXYVIWXEKIWSJ 4IHEPMSHIWTSIWME EX=&7E VIGIRXP]QSPXIHJSYVXLMRWXEV F JSYVXL MRWXEV G TVIQSPXJSYVXLMRWXEV H VIGIRXP]QSPXIHJMJXLMRWXEV I QEXYVIJMJXLMRWXEV 6W\IJTa \PQZL QV[\IZ[ IVL X]XIM \PM WVTa [\IOM[ QTT][ MTWVOI\M IVL IVO]TIZ .SYVREPSJ-RWIGX7GMIRGI`[[[MRWIGXWGMIRGISVK .SYVREPSJ-RWIGX7GMIRGI:SP`%VXMGPI +VIIRI]IXEP *MKYVI -QQEXYVIWXEKIWSJ 4IHEPMSHIWTSIWME EX=&7EF TVITYTEPPEVZE GJ TYTE []OOM[\ \PI\ 8 XWM[QI UIa \ISM I[ U]KP I[ I UWV\P TWVOMZ KWTWZI\QWV J]\ 8IZIXMLITQWLM[ M`PQJQ\[ I [PQVQVO JTIKS PMIL \W LM^MTWX NZWU MOO \W IL]T\ ;KP]T\bM LQL VW\ [XMKQNa \PM KIX[]TM TQSM \PI\ WN 8 bQVOIZI .SYVREPSJ-RWIGX7GMIRGI`[[[MRWIGXWGMIRGISVK .SYVREPSJ-RWIGX7GMIRGI:SP`%VXMGPI +VIIRI]IXEP *MKYVI %HYPX 4IHEPMSHIWTSIWME IQIVKMRKJVSQMXWTYTEEX=&7-RWIXWLS[WXLITYTENYWXTVMSV XSLEXGLMRK %GORS[PIHKQIRXW V]UJMZ ! WN \PM AIVIaIK] 6I\]ZIT 0Q[\WZa :M[MIZKP /ZW]X .WZ I[[Q[\IVKM QV \PM NQMTL _M \PIVS :INIMT /ZIVQbW IVL ?QTUMZ ;QUJI}I 0./ \PIVS[ :]\P )VV IVL 2WPV > )HMXSV´WRSXI 5WWZM \PM 8WX]TI\QWV *QWTWOa .W]VLI\QWV IVL 5I\\ 8IXMZ KWXQM[ WN \PQ[ IZ\QKTM _QTT JM LMXW[Q\ML QV \PM NWTTW_ 3IXTIV NWZ NQVIVKQIT []XXWZ\ .SYVREPSJ-RWIGX7GMIRGI`[[[MRWIGXWGMIRGISVK .SYVREPSJ-RWIGX7GMIRGI:SP`%VXMGPI +VIIRI]IXEP *MKYVI %HYPXWSJ 4IHEPMSHIWTSIWMEHSVWEPWYVJEGISRPIJXERHZIRXVEPWYVJEGISRVMKLX9TTIV WTIGMQIRƃ )GYEHSV2ETS&EI^E8IRERIEV%VVE]ERQ%TVMP%2IMPHPIK 1>9.0S[IVWTIGMQIRƂ )GYEHSV8YRKYVELYE&MWGE]E¯Q¯1E]%.EWM WOMPIK1Y^IYQ>SSPSKMG^RI9RM[IVW]XIXY.EKMIPPSRWOMIKS >SSPSKMGEP1YWIYQSJXLI.EKMIPPSRM ER9RMZIVWMX] .SYVREPSJ-RWIGX7GMIRGI`[[[MRWIGXWGMIRGISVK .SYVREPSJ-RWIGX7GMIRGI:SP`%VXMGPI +VIIRI]IXEP 6IJIVIRGIW 5QTTMZ 4, ! 0M_Q\[WV ?+ 7V 8ZWVWXPQTI I OMV][ WN LQ]ZVIT 4MXQLWX >QTWZQI )4 !! ;\]LQM[ WV \PM [a[\MUI\QK[ IVL JQWOMWOZIXPa WN \MZI" _Q\P NQO]ZM[ WN VM_ [XMKQM[ IVL ZMNMZMVKM \W ITT \PW[M [WUM UWV\IVM [I\aZQL J]\\MZNTQM[ 4MXQLWX\MZI 8P, LQ[[MZ\I\QWV _PQKP PI^M JMMV NQO]ZML WZ LM[KZQJML 4IUI[ / >QTWZQI )4 8aZKb QTWZQI )4 1V" +W[\I + >IVQV ;) 4WJW 25 5MTQK ) MLQ\WZ[ 4IUI[ - MLQ\WZ )\TI[ WN 6MW\ZWXQKIT 4MXQLWX\MZI +PMKS 4QUQ\IKQWVM[ Y]M WNZMKMV LQ[\QV\I[ QV\MZXZM\IKQWVM[ \I` TQ[\" 8IZ\ ) 0M[XMZWQLMI · 8IXQTQWVWQLMI )[[WKQI\QWV NWZ .SYVREPSJ-RWIGX7GMIRGI`[[[MRWIGXWGMIRGISVK