69

STUDIES ON THE FAUNA OF CURAÇAO AND OTHER

CARIBBEAN ISLANDS: No. 198

Additional notes on Caribbean tiger-beetles

of the genera Cicindela and Megacephala

by

P. Wagenaar Hummelinck

(Lab. Zool. Oecol. & Taxon., Utrecht)

Another small collection oftiger-beetles which was made by the author

notes during his visits to the Antilles may justify these additional to the papers of JONGE POERINK, on Cicindela (1953, this series vol. 4) and the present author, on Megacephala (1955, vol. 6).

Studies 4 Studies 6 Studies 64

1953 1955 present paper

page page page plate

Cicindela auraria 122 71 Ic

Cicindela trifasciata trifasciata 129 74 Id

Cicindela trifasciata ascendens 132 76 lie

Cicindela suturalis suturalis ; 133 76 lie, IVg-i

Cicindela suturalis hebraeai 135 78 la, IVc-e

Cicindela suturalis guadeloupensis 137 80 lid

Cicindela graphiptera graphiptera 138 80 Ila

Cicindela graphipterafulgidiceps 139 80 lb

Cicindela boops 81 lib

Cicindela carthagenajamaicana 83 Illa-c

Cicindela argentata 85 IVf

Cicindela marginata 86 IVa

Cicindela dorsalis media 88 IVb

Key to the Cicindelids 89

Megacephala acutipennis 91 106 Vc

Megacephala affinis affinis 93 106

Megacephala affinis gracilis 95 106 Vb

Megacephala carolina carolinaJ 97 107

Megacephala carolina carolina forma occidentalis 102 107 Vd-e

Megacephala sobrina sobrina forma typica . . . 105 108 Vla-b

Megacephala sobrina sobrina forma antiguana. 108 109 Vic

Megacephala sobrina sobrina forma bonaireana 110 109 Va

Megacephala sobrina sobrina infuscatai 112 111 VId

Key to the Megacephalas 112

Distribution 123

References 128 70

Fig. 26. Some body measurements taken in Cicindela.

auraria the Fig. 27. Elytrae ofa female specimen ofCicindela from Aruba,showing position

ofa number of small erect setae in which may be distinguished: a) a single row ofabout 15-

of about the 30 near the lateral margin, b) a scattered longitudinalrow (in principle) 10 on

dorsal scattered of small less the part, c) an irregularly group setae, more or bordering

proximal part of the dark area.

The author is indebted to Mrs. C. S. OLDENBURGER - EBBERS, Mrs. J. S.

DE LEEUW VAN WEENEN - DE HART and Mr. F. VAN DER HEIDE for

allowing him to include a number of measurements and other data,

produced during their student’s practical course in taxonomy at the Zoo-

logical Laboratory of the Utrecht University in 1967-1969. CARLA

OLDENBURGER (cf. Table 2, Figs. 27-34) and JEANNETTE DE LEEUW VAN

WEENEN (cf. Figs. 39-41) studiedthe greater part of the Cicindela material,

while VAN DER HEIDE (cf. Table 5, Figs. 42-46) examined most of the

Megacephala specimens.

Loans from the Science Museum ofthe Institute of Jamaica and from the British Museum

(Natural History) are gratefully acknowledged. Several interesting specimens collected by

Dr. Ir. R. H. COBBEN (Wageningen) and Dr. F. CHALUMEAU (Guadeloupe) have also been

studied, while a few specimens from the Zoölogisch Museum of Amsterdam and the

Rijksmuseum van Natuurlijke Historie at Leiden are included for comparison. Dr. 71

CHALUMEAU and MICHAEL A. IVIE (Columbus, Ohio) kindly informed me about their

activities. Furthermore I with Dr. THOMAS H. coleopterological greatlyenjoyed my contacts

FARR (Kingston, Jamaica) and Mr. C. M. C. BROUERIUS VAN NIDEK (Voorburg,

Netherlands).

A few illustration from the two preceding reports have been reproduced again (Figs. 14

and 20, Pis. I—II and V-VI) to make these additional notes a little more attractive to all

those who share the author's admiration for these most engaging elements of the Antillean

beetle fauna. I am indebted to the official artist of the Zoological Laboratory of Utrecht

Mr University, H. VAN KOOTEN for the excecution of the photographs.

If not otherwise stated, all measurements have been taken from specimens preserved in

alcohol.

The length of the elytron - considered to be a suitable measure for comparing relative

of - width of sizes body and legs has been measured from shoulder to apex. The an elytron

has been taken as being the half ofthe width of the abdomen; when measured separately its

maximum width should be about 20% (cJ}-25% (?) more (cf. Fig. 26, and Stud. 6 fig. 2). The

is defined the of and it is body-length by adding up length elytron, pronotum head, though

evident that a body-length without the protrudingparts of abdomen and labrum,will often

be quite different from the body sizes as given in literature.

Not much value be attached these which be taken ought to to measurements, hardly can

in - be of for mutual an exact way they may, however, importance comparison.

All new localities indicated by station numbers have been described in these Studies, vol.

63 (1981).

The greater part of the material has been presented to the Zoologisch Museum of

Amsterdam, while other specimens have been deposited at the Rijksmuseum van

NatuurlijkeHistorie, Leiden, the British Museum (N.H.), London, The Institute of Jamaica,

Kingston, and the Institut de Recherches Entomologiques de la Caraibe, Guadeloupe. It is

to be regretted that in some cases the poor state ofpreservation reflects accidental collecting

which undertaken with the of insects. duringtrips were not purpose collecting

Although the author admires the taxonomical work ofRIVALIER (and other coleopterolo-

gists), his nomenclature intentionallywas not used, as the present publication only aims at

additional information in which his giving some to two previous papers certainly more

satisfying classification was not used. The few data regardingthe penis should be considered

only as a stimulus to give more attention to the structure ofthe male copulatory organ when

studying the modest thoughinteresting cicindelid fauna of the Antilles.

Cicindela auraria Klug, 1834

[Pl. Ic; figs. 27-28, 35a-c, 39a, 40a, 41]

JONGE POERINK 13-14 from 1953, p. 122-128, figs. 27, 28a-c, pis. [Material Margarita, Bonaire, Klein Bonaire, Curacao, Aruba, Paraguana and La Goajira, Venezuela,

Colombia and Panama; synonymy.] 72

Cicindela FERNANDEZ YEPEZ & ROSALES 170 auraria, 1956, p. [Gran Roque.]

Dromochorus (Ellipsoptera) auraria, SCHILDER 1953, p. 560.

259. Habroscelimorpha auraria, RIVALIER 1954, p.

ARUBA: Spaans Lagoen, muddy area, IV. 1957, collected by R. H. Cobben (2,33 2$$).

Salinja Balashi near Spaans Lagoen, 22.X.1967 (1 9).

St. Jorisbaai SE 23.X.1968 8 NE corner, Curasao: , corner, mudflat, (4

St. Martha mudflat 4.VIII.1967 1 4 larvae from bur- Salinja , near saltpond, (1 (J ?,

rows). Salinja St. Kruis, N mudflat, 20.11.1970 (2 SS 1 $)•

KLEIN BONAIRE: Salinja Abau, salty mudflat, 25.111.1955 (2r?r? 19).

BONAIRE: Lagoen, NW part, hiding in cracks of salty mudflat, 2 & 9.III.1955 (3(JcJ

2?2); 7.XII.1963 19). Salinja di Cai, N, 9.IX.1967 (1 (J). Awa Lodo di Lac, only

a few specimens on a muddy limestone flat, 19.VIII.1967 (1 J). Isla di Pedro, Lac,

dried limestone mud with crusts ofblue algae, 7.III. 1970 (2 499)- Casdi Meeuchi,

Lac, soft whitish mudflat covered by felt-like algae, 9.III.1970 (19)- Witte Pan,

5.III.1970 (19)- Blauwe Pan, E, salty mudflat with flakes of tuffoid limestone,

9.III.1955 (1 9); N of crystallizers, disturbed saltflat, 14.111.1970 (222)- Salinja Mar-

tinus, S of Kralendijk, muddy shore of saltpond, 1.IV.1955 (8 JJ 629)-

MARGARITA: Punta de Piedras, Estacion de Investigaciones Marinas de Margarita,

Sta. 801, white sandy beach ofabout 10 m broad, separatedby a few low dunes from

a muddy sandflat; a single specimen among many C. graph. fulgidiceps, 13.1.1964

(1 $); Sta. 802, sandy mudflat near mangroves, 9.1.1964 (53??).

COLOMBIA: Santa Marta, salt lake, 23.11.1896 (1 §, RMNH).

width 3.9 Body length cJ 8.7-9.4-10.3 99.2-9.6-10.6 mm; Q4.1 mm; about 2.4 times as long as wide. (Measurements from 20 20 $$ from

Aruba, Bonaire and Margarita, see Table 2).

Labrum yellowish white, with 6-9 submarginal setae. Frontal margin more (?) or less (cJ) outwards curved with a small but distinct central tooth (Figs. 28, 35).

hairs. Width the Head glabrous, except for 2 supra-orbital across eyes about 1.85 pronotum-width = 0.45 elytron-length. Genae densely clothed with decumbent bristles.

Pronotum about 1.5 mm in length, <$ 2.2 $2.3 mm wide, about 1.5 times

furrows rather as broad as long. Median line distinct, transverse deeply impressed. Lateral upperparts broadly but sparsely clothed with decumb- ent bristles.

Penis about 4 mm long, slender, almost 8 times as long as wide. Flagel- lum that well 360° making a complete loop encompasses over (Figs. 39a,

40a). Six chitinous plates counted.

Legs (cf. Tables 3 and 4) of $$ only slightly longer than of $$. I, II and

III about 1.3, 1.6 and 2.0 times elytron-length, respectively, equalling 73

about 0.8,1.0 and 1.25 body-length. Femur and tibia III about 0.7 and 0.6

Trochanters I and with elytron-length, resp. II a single long seta.

Elytron 5.0-6.2-6.7 $ 5.0-6J-6.6 mm in length (cf. Table 2), about 3.2 times as long as wide. Lateral side weakly convex; the last 1/5 part rather

off the rather abruptly turning to apex, distinctly curved inwards, its margin finely serrated, ending into a rather sharply projected spine. The yellowish-white marking consists of a wide uninterrupted marginal area which extends from shoulder and is less to apex more or tri-lobed within, the apical lunular part only reaching the medial margin. See also Fig. 27.

The labra ofthe Margarita specimens are uniform in having 6 submarginal setae; half of the Bonaire specimens possess the samenumber, while the others have 7-8 setae; oneAruba animal has 6 sdl je, the other four 7-9. It may be noted that the fourth joint of the antenna is about 0.9 times as long as the third one, as in other species.

Description of larva (according to Mrs. DE LEEUW - VAN WEENEN; fig. 41)

Dorsal side of head dark ventral side of head and chestnut-brown. - brown; pronotum Diameter ofocellus 2 subequal to the distance between ocelli 1 and 2. Fronto-clypeal-lateral

broad. caudal of frons 3 - Antenna area as long as U-shaped ridge on part bearing setae. with the proximal and second segments subequal in length; the distal about two-fifth the

ofthe second. The of the its mesal length proximal segment galeabearing 4 setae on margin.

4 Ligula with fine setae arranged in a transverse row. Proximal end of labial palpus with 4 setae; distal segment with 1 seta. - Pronotum with the cephalo-lateralmargins extending as far cephalad as the mesal portion.

The larva of be included in the the C. auraria may key givenby HAMILTON (1925, p. 19) in following way:

25. Proximal segment of the galeawith four stout setae on the mesal margin; head and

pronotum brown a

Proximal segment ofthe galeawith three stout setae on the mesal margin; head and

pronotum not brown, metalic colored 27

a. Diameter of ocellus 2 distinctly less than the distance between ocelli 1 and 2; median

hooks 2 or 3 setae, inner hooks 4 setae 26

Diameter of ocellus 2 subequal to the distance between ocelli 1 and 2; median hooks

inner hooks little 1 stout seta, 2 setae auraria 74

Cicindela trifasciata trifasciata Fabricius, 1781

[Pl. Id; figs. 29-30, 35d-e, 39b, 40b]

JONGE POERINK 1953, p. 129-132, figs. 28d-f, 29c d, 30a-b, pis. 13 and 16 [Material from

Margarita, Antigua, St. Barts, St. Martin, Anguilla, St. Croix, St. Thomas, Puerto

Rico, Hispaniola, Jamaica, Gran Cayman, Little Cayman, Cuba and Cayenne;

synonymy.]

Cicindela trifasciata, LENG & MUTCHLER BEATTY 132 1914, p. 393; 1917, p. 194; 1944, p. [St.

WOLCOTT 227 MISKIMEN & Croix]; 1948, p. [Puerto Rico]; BOND 1970, p. 78 [St.

Croix]; BUTLIN 1976, p. 128 [Anegada];IVIE 1983, p. 197-198,fig. 2a [St. Thomas, St.

John, Tortola, Anegada, St. Croix.]

Dromochorus (Ellipsoptera) trifasciata, SCHILDER 1953, p. 560.

Cicindelidia RIVALIER 258. trifasciata trifasciata, 1954, p.

Cicindelidia BALAZUC & CHALUMEAU BOYD 14. trifasciata, 1978, p. 20-22, fig. 2; 1982, p.

Cicindela DAHL tortuosa Dej. var., 1892, p. 108 [Bermuda.]

TOBAGO: Point Rockley Bay, NE Red with C. sut. hebraea on dark sandy beach,

14.1.1955 (5 specimens, damaged).

GUADELOUPE: Pointe des Chateaux, 20.IV.1980,coll. F. Chalumeau (1 ?); 26.VI.1982,

F. Chalumeau (3 6 ??) [el. 1. 6.9 $ 7.0 mm; tibia III J0.7 §0.65, tars. (J 0.75 ? 0.7 el. 1.].

ANTIGUA: Deep Bay, sandy mudflat near saltpond, 17.VII.1955 (2 SS 2 99). St. John's

Harbour, the Cove, sandy beach, 1.VIII.1955 (5 SS 3 99)- Beach Hotel, sandy beach,

11.VI.1965, coll. D. C. Geijskes (1 9, RMNH). Dickinson Bay near McKinnons Salt Pond, with C. sut. suturalis on greyish sand, 25.VII.1967 (5 SS 3 $$).

BARBUDA: Codrington Village, at light, 5.VII.1955; 10.V.1982, F. Chalumeau (1 S

1 $). Low Pond, Sta. 674, 5.VII.1955 (7

Lagoon, S of Codrington Village, 4.VII.1955 SE mud sandy mud, (6

Village, with C. sut. suturalis on whitish sand, leeward of Lagoon (2SS 2$?). Billy

Point, near entrance ofGreat Lagoon, Sta. 832, with C. sut. suturalis on white sand

beach, 22.VII.1967 (299). Bull. Hole, Sta. 667, low limestone flat near brackish water

pond, 9.VII.1955 (1 S)- Beach on Barhuda, 20.VI.1965, coll. D. C.Geijskes (6SS 599,

RMNH).

ST. KITTS: Frigate Bay, near Sta. 677, muddy sand flat, 20.VII.1955 (7CJ

ST. MARTIN: AtweWs Pond, E of Philipsburg, sandy mudflat near saltpond,

2.VI.1955 (7CJCJ 1599); 3.VI.1955 (ASS 899); 26.IX.1963 (\1 SS 1299); 30.VII.1967

299)- Great white beach near Atwell's 1.IX.1963 (4SS Bay, sandy Pond, (1 C J);

26.IX.1963 (1 J). Great SaltpondNE, salty mudflat, 28.IX.1963 (11 SS 13 99). Oyster

Pond, sand bar near sea with cow faeces, 13.X.1963 (299)- Saline de Grand' Case,

sandy mudflat near saltpond, 22.VI.1973 (3 SS 2 99)- Baie aux Cailles, NE of Simson

Lagoon, Sta. 830, 28.VII.1967 (2SS 19)- Flamingo Pond, part of Simson Lagoon,

soft mudflat in drying lagoon, with 2 larvae hidingunder pieces of wood, 16.X.1963

1 (2 (J 9)- Simson Bay flat, part of drying Simson Lagoon, near Sta. 1130A, muddy

sandflat, 6.VI.1955 599); 16.X.1963 (6

lands, 21.IX. 1956, coll. R. H. Cobben (1 S 499). 75

ANGUILLA: Crocus Bay, near Sta. 1704, sandy beach, 3.VII.1973 (2rJrJ 1 $). Anguilla,

mudflat near school, 7.VII.1965, E.G.S.G. (1 ?).

CAICOS ISLANDS: South Caicos, VI.1957, coll. T. H. Farr (6 C?C? 299, Sri. Mus.

Jamaica). TURKS ISLANDS: Grand Turk, Waterloo, 26.IV.1954, coll. C. B. Lewis (1 (J); VI.1957,

T. H. Farr (8 c}

ANEGADA: Setting Point, beaches and saltponds, 9.VIII-21.IX.1975, R. K. Butlin

(3 CJCJ 499. Br. Mus.).

PUERTO RICO: Playa Luquillo, E of San Juan, Sta. 040, sandy beach with decay of

Syringodium, 1.V.1973 (2

coll. David JAMAICA: Hope River bed, St. Andrew, near August Town, 11.XII.1955,

Gregory (1 cJ). Port Henderson, St. Catherine, saline flats near sea, 21.1.1953, R. P.

Bengry (1 (J 1 9)- Portland Ridge, Clarendon, 1 mi E ofJackson Bay, 3.VIII.1953, C.

B. Lewis (19). Milk River beach, 10.1.1954, G. R. Proctor (19). Negril, Westmorel-

and, 24.111.1955, T. H. Farr (1 9)- Falmouth, Trelawney, 23.VIII.1955, T. H. Farr

(2 (JcJ). All specimens in Sci. Mus. Kingston. - Great Saltpondnear Port Henderson,

C. sand bar near Rhizophora, 8.V.1973 (3

Yallahs, Sta. 1676), muddy shore, 6.V.1973 (1 9). Mona nearKingston, at Sta. 026, a

few of sand freshwater 16.VI.1973 square metres greyish along a reservoir, (1 spec.).

LITTLE CAYMAN: South Hole Sound, near Sta. 1698, near Rhizophora, 5.VI.1973

(observed).

CAYMAN BRAC: South Bay Ligoon, near Sta. 1701, soft muddy area, 3.VI.1973

(observed).

SURINAME: Matappica, muddy sand near saltpan, 20.X.1940 (1 S). Coronie sand bar

with shells near shore, 15.111.1945 (19)- Bigisanti, at light in swamp near shore,

11.VII.1955 (1 (J). Paramaribo, at light, 3.IX.1958 (1

10.VIII.1959 (1 (J). Paloemeu, Swanilboto val, 2.IX.1959 (1 cJ). All specimens collected

by D. C. Geijskes, RMNH Leiden.

Body length $ 9.1-10.3-11.0 $ 10.2-70.5-11.1 mm; width about $4.1

$ 4.3 mm; about 2.5 times as long as wide. (Measurements of44 $$ 43 $$ from Antigua, Barbuda, St. Kitts, St. Martin, Turks & Caicos I. and

Jamaica, cf. Table 2).

Labrum yellowish-brown, with 8-13 submarginal setae. Frontal margin

tooth outwards curved, with a distinct central triangular (Figs. 29, 30).

Head, incl. genae, glabrous, except for 2 supra-orbital hairs. Width

across the eyes about 1.8 pronotum-width = 0.62 elytron-length.

Pronotum about 7.9 mm in length $2.2 $2.5 mm wide, about 1.15 as broad as long. Median grove distinct, transverse furrows deeply im-

with decumbent pressed. Lateral upperparts sparsely clothed bristles.

rather with truncated about 6 Penis about 3| mm long, plump a apex,

times as long as wide. Flagellum encompassing two complete loops (Figs.

39b, 40b). Six chitinous plates counted.

Legs (cf. Tables 3 and 4) of $$ somewhat longer of $9-1, II and III in 76

SS about 1.45, 1.65 and 2.15 times elytron-length, resp., equalling about

0.9, 1.0 and 1.3 body-length. I, II and III in ?? about 1.35, 1.45 and 2.0 times elytron-length, equalling about0.8,0.9 and 1.2 body-length. Femur and tibiaIII about 0.7 and 0.65 elytron-length. Trochanters I and II with a single long seta.

Elytron <$ 5.8-6.6-7.6 $ 6.1-6.7-7.7mm in length (cf. Table2), about 3.2 times as long as wide. Lateral side weakly convex with a shallow inward curve just above the middle, the last 1/5 part turning off to the apex, which is rather pointed in the rounded in the $, ending in a short acute

The consists spine. yellowish-white to brownish-yellow marking of: (1) a hoof-shaped humerallunula fromwhich the frontal part reaches the place at which the side of the pronotum meets the elytron, and the hind part

middle band ends in a more or less large spot; (2) a strongly S-shaped which is crenated and sometimes interrupted in the middle; (3) a hooked apical lunula; all these components joined by (4) a more or less wide marginal area.

Cicindela trifasciata ascendens LeConte, 1851

[Pl. IIc]

JONGE POERINK 15 from 1953, p. 132-133, figs. 28g-h, 29e, pi. [Material Hispaniola, Cuba, Bermuda, Mexico and U.S.A.; synonymy.]

Cicindela BEATTY 132 the flats of trifasciata tortuosa Dejean, 1944, p. [Taken on sandy

Krause lagoon, Oct. 1937; det. J. M. Valentine.]

Cicindelidia RIVALIER 258. trifasciata ascendens,, 1954, p.

Dromochorus (Ellipsoptera) trifasciata ascendens, SCHILDER 1953, p. 560

Cicindela suturalis suturalis Fabricius, 1798

[Pls. IIe, IVg-i; figs. 31, 36c-d, 40d]

Cicindela suturalis Fabricius (typical form), JONGE POERINK 1953, p. 133-135, figs. 28i-k, 29b, 30c-d, pis. 15-16 [Material from Antigua, St. Barts, St. Martin, St. Thomas and

Hispaniola; synonymy.] 77

Cicindela suturalis, LENG & MUTCHLER 1914, p. 393; 1917, p. 194; BUTLIN 1976, p. 128

IVIE 2c-e St. [Anegada]; 1983, p. 196-197, fig. [St. Thomas, John, Anegada.]

Cicindela suturalis MANDL 3-5. suturalis, 1958, p. 25, figs.

Dromochorus SCHILDER 560. (Ellipsoptera) suturalis, 1953, p.

RIVALIER BALAZUC & CHALUMEAU Cylindera (Plectographa) suturalis, 1954, p. 266; 1978, p.

4 "nous doutons fort de la ... Nous 23-25, fig. [St. Barts; presence en Guadeloupe.

n'avons de materiel de la pu ... nousprocurer Martinique."]

BOYD 16. Cylindera suturalis, 1982, p.

ANTIGUA: Deep Bay, white sand near saltpond, cf. Sta. 1393,17.VII.1955(4CJ

Fort Bay, white sand beach, 18.VII.1955 (2<$<$ 2??). Dickinson Bay N, white sand

beach, 19.VII.1967 (9CJ

C 25.VII.1967 Pond, single specimen among . trif. trifasciata, greyish sand, (19).

Antigua, A. D. Torlese (1 3$, Brit. Mus. 1932-148).

BARBUDA: Martello Tower beach, white sand, 8.VII.1955 (5 c?c? 4??). Palm Beach, W

of Codrington Village, white sand beach along bar bordering Great Lagoon,

12.VII.1955 (1 3 ??). Billy Point, Sta. 832, white sand beach near entrance ofGreat

Lagoon, 22.VII.1967 (7

ANEGADA: Setting Point, beach and saltponds,9.VIII-21.IX.1975, R. K. Butlin (1 9,

Brit. Mus.).

ST. THOMAS: Magens Bay, white sand beach, 20.VIJ955 (4c?c? 6 59).

Body length

$3.4 mm; about 2.5 times as long as wide. (Measurements from 13 cM

17?? from Antigua, Barbuda and St. Thomas, cf. Table 2).

Labrum yellowish-brown, with 8-10 submarginal setae. Frontal margin weakly outwards curved (?) or nearly straight (cJ) with a small central

tooth (Fig. 31).

incl. for 2 hairs and Head, genae, glabrous, except supra-orbital a row of decurrent bristles near the posterior margin. Width across the eyes about 1.3 pronotum-width = 0.43 elytron-length.

Pronotum <$ 1.4 ? 1.5 mm in length 1.75 $ 1.9 mm wide, about 1.25 as broad as long. Median line and transverse furrows rather distinct. Lateral

upperparts rather sparsely clothed with decumbent bristles.

Penis about 2\ mm long, rather plump, about 5-j times as long as wide.

Flagellum making a single loop well over 360° (Fig. 39d). Five chitinous plates counted.

Legs (cf. Tables 3 and 4) of only very slightly longer than of $$. I, II and about and 2.0 times III 1.25,1.5 elytron-length, resp., equalling about

0.75, 0.9 and 1.2 body-length. Femur and tibia III about 0.65 elytron- length. Trochanters I with a single long seta.

Elytron 5.0-5.2-5.5 $ 5.0-5.6-6.5mm length (cf. Table 2), about 3.3 78

wide. Lateral times as long as side weakly convex; the last 1/4 part turning off the truncated its in to slightly apex, margin finely serrated, ending a

sharply projecting spine. The yellowish-white marking is dominating to

such that there is left an extent only one elongate subtriangular sutural

mark of about 2/3 elytron length, which arises froma trapezium-like base

which is as wide as about 2/3 part of the width ofthe proximal part ofthe

The of this is elytron. top sutural mark abruptly widened and often partly enclosed by a narrow, more or less irregular line. More rarely this black

sutural mark shows further development (somewhat resembling subsp.

hebraea; cf. PI. Ia).

Cicindela suturalis hebraea Klug, 1834

[Pls. Ia, IVc-e; figs. 32a-c, 39d, 40e]

hebraea Cicindela suturalis var. Klug, JONGE POERINK 1953, p. 135-137, figs. 281-n, pi. 13 [Material from Trinidad, Barbados, St. Vincent, Guadeloupe, Hispaniola,

Colombia, Venezuela, Cayenne, Brasil and Peru; synonymy.]

Cicindela suturalis LENG & MUTCHLER WOLCOTT var. hebraea, 1914, p. 393; 1917, p. 194;

1948, p. 226.

Cicindela suturalis hebraea, MANDL 1958, p. 25, figs. 3-5 [St. Thomas, Hispaniola,

Colombia, Guyana, Cayenne, Brasil.]

Dromochorus SCHILDER 560. (Ellipsoptera) hebraea, 1953, p.

Cylindera (Plectographa) suturalis hebraea, RIVALIER 1954, p. 266.

suturalis BALAZUC & CHALUMEAU Cylindera (Plectographa) race hebraea, 1978, p. 25, fig. 5

["ne parait pas representee dans les Antilles frangaises".]

suturalis BOYD 17 R. Cylindera hebraea, 1982, p. [Guadeloupe, Dominicana.]

CURASAO: Emmastad, at light, X.1956, coll. R. H. Cobben (1 $); golf links, 26.X.1956,

Cobben (1 $). [The two specimens ofthis new island locality had partly been eaten

by insects after arrival in Utrecht.]

TRINIDAD: IDS Gallos Point, sandy beach with some debris, Sta. 790,16.1.1964 (8 SS

11 ??). Cocos Beach at Nariva Swamp bridge, rather white sand with some decay,

Sta. 789, 17.1.1964 (1 C? 1$). Maracas Bay, white sand beach, 29.IX.1956, R. H.

Cobben (6

TOBAGO: Rockly Bay, Scarborough, dark sand shore with rock debris and Sargas-

sum, Sta. 583, 20.1.1955 (3

14.1.1955 (2cJ

ST. VINCENT: St. Vincent, H. H. Smith 231 (19, Br. Mus.).

MARTINIQUE: Ste Aure, Roguet, 8.X.1981, F. Chalumeau (Zool. Mus. Amsterdam)

[el. 1. 4.9 mm].

PUERTO RICO: Playa Mediania Aha, E of San Juan, Sta. 038, sandy shore near

drainage from swampy region, 1.V.1973 (333 19)- 79

Body length 7.5-7.5-8.6 $ 7.7-SJ-8.8 mm; width about (J 2.9

times from ?3.25 mm; about 2.6 as long as wide. (Measurements \1 S6

10?$ from Trinidad and Tobago; cf. Table 2).

Labrum yellowish-brown, with 8-10 submarginal satae. Frontal

outwards curved with margin weakly ($) or nearly straight {<$) a small

central tooth (Figs. 32a-e).

Head, incl. genae, glabrous, except for 2 supra-orbital hairs and a row of decurrent bristles the Width the near posterior margin. across eyes about 1.3 0.43 pronotum-width = elytron-length.

Pronotum 1.4 1.5 mm in 7.7 1.85 mm about 1.2 as $ length, c J $ wide, broad Median distinct. Lateral as long. line and transverse furrows up- perparts rather sparsely clothed with decumbent bristles.

Penis about 2j mm long, rather plump, about 5| times as long as wide.

Flagellum almost encompassing two loops (Figs. 39e, 40d). Seven chit-

inous plates counted.

Legs (cf. Tables 3 and 4) of $<$ only very slightly longer than of $$. I, II and III about and 2.0 times about 1.25,1.5 elytron-length, resp., equalling

0.75, 0.9 and 1.2 body-length. Femur and tibia III about 0.65 elytron-

length. Trochanters I with a single long seta.

Elytron <$ 0.48-5.05-5.5 $ 5.2-5.55-5.8 mm in length (cf. Table2), about

3.4 times as long as wide. Lateral side weakly convex; the last 1/4 part

to turning off the slightly more (?) or less ((J) truncated apex, its margin

finely serrated, ending in a small spine. The yellowish-white marking

consists of: (1) a hoof-shaped humeral lunula, the hind part ending in a

more or less triangular spot; (2) a strongly S-shaped middle band; (3) a

hooked all these apical lunula; components are joined by (4) a more or

less wide marginal band - a design which shows a striking similarity to

" that of C. trifasciata trifasciata.

F. CHALUMEAU (in press) considers C. sut. hebraea to be a continental form which does not occur on the islands from Grenada to Cuba. According to him the populations of the Greater Antilles (and those from Trinidad and Curagao) need further investigation.

Considering differences in elytral design and punctation mainly, he distinguishes four sub- species: the nominal form (from Anegada to Antigua), (C. sut. guadeloupensis (Guadeloupe) and two new forms (Martinique, and Grenada + ? St. Vincent). 80

Cicindela suturalis guadeloupensis Fleutiaux & Sallé, 1889

[Pl. IId]

suturalis Fleutiaux & JONGE POERINK Cicindela var. guadeloupensis Salle, 1953, p. 137-138,

figs. 280-q, pi. 15 [Material from Guadeloupe,Cayenne (?) and Brasil; synonymy.]

Cicindela suturalis var. guadeloupensis, LENG & MUTCHLER 1914, p. 194.

Cylindera (Plectographa)suturalis race guadeloupensis, BALAZUC & CHALUMEAU 1978, p. 25-

26, fig. 6 [Guadeloupe.]

Cicindela suturalis MANDL 3-5 nocturna Steinheil, 1875, 1958, p. 26, figs. [STEINHEIL'S

" ofC. from Colombia does identification with description nocturna not encourage an

C. suturalis guadeloupensis.]

suturalis 17. Cylindera guadeloupensis, BOYD 1982, p.

Cicindela graphiptera graphiptera Dejean, 1831

[Pl. IIa]

Cicindela JONGE POERINK graphipteraDejean (typical form), 1953, p. 138-139, figs. 28r-t, pi.

15 [Material from Costa Rica and Colombia; synonymy.]

Dromochorus SCHILDER 561. (Ellipsoptera) graphiptera, 1953, p.

RIVALIER 261. Opilidia graphiptera, 1954, p.

Cicindela graphiptera fulgidiceps Putzeys, 1845

[Pl. Ib; figs. 32d-e, 36a-b, 39e, 40f]

Cicindela JONGE POERINK graphipteravar. fulgidiceps, 1953, p. 139-141, figs. 28u-w, 29f, pis.

13 and 17 [Materialfrom Margarita,Tortuga, Venezuela mainland and Colombia;

synonymy.]

Dromochorus (Ellipsoptera) fulgidiceps, SCHILDER 1953, p. 561.

Opilidia graphipterafulgidiceps, RIVALIER 1954, p. 261.

MARGARITA: Punta Mosquito, Sta. 797, sandy beach with debris, 13.1.1964 (6(J

4?$). Punta Mangle, Sta. 800, white sand beach, 10.1.1964 (1 2$$). Punta de

Piedras, near Sta. 802, white sand beach, 9.1.1964 (2

32??).

Body length $9.7-77.2-11.7mm; width $3.7 $4.2, about 2.6 times as long as wide. (Measurements of 11c? 13 $ from Marga- rita, cf. Table 2). 81

Labrum brownish yellow, with 6-8 submarginal setae. Frontal margin

with slightly outwards curved (9) or almost straight (rj) a small but

distinct central tooth (Figs. 32d-e).

Head, incl. genae, glabrous, except for 2 supra-orbital hairs. Width

across the eyes almost 1.2 pronotum-width = 0.4 elytron-length.

about 1.6 in 2.3 about 1.45 Pronotum mm length, 3 $2.5 mm wide, rj

$ 1.55 times as broad as long. Median line weak but distinct, transverse

furrows deeply impressed. Each side broadly and rather densely clothed

with decumbent central covered. Hindbor- bristles; upper parts sparsely

der with a fringe of rather long bristles.

Penis about 2j mm long, rather plump, about 5j times as long as wide.

full Flagellum not yet completing a loop (Figs. 39f, 40e).

Legs (cf. Tables 3 and 4) of conspicuous length, distinctly longer

than 9?. I, II and III in about 1.2, 1.6 and 2.55 times elytron-length,

about 1.0 and 1.55 II and III in resp., equalling 0.75, body-length. I, 99 about 1.0,1.3 and 2.05 times elytron-length, equalling about0.65,0.8 and

1.3 body-length. Femur III about S 1-05 90-9; tibia III cJ0.7 90.6 elytron-

length. Trochanters I and II without a long seta.

Elytron <$ 6.1-6.5-6.7 $ 7.0-7.7-7.9mm (cf. Table 2), about 3.6 times as

wide. Lateralside little almost the last long as very convex or straight; 1/4 part turning offto the more(?) or less (

serrated, ending into a small spine. The yellowish-white marking is

dominating to such an extent that from the dark parts there are only a few

character-like figures left in which sometimes a design tending to that of

the typical form (comparable with those of C. suturalis hebraea or C.

be trifasciata) can recognized.

Cicindela boops Dejean, 1831

[Pl. IIb; figs. 33, 37a-c, 39c, 40c]

Cicindela Mannerheim 258-260 viridi- Boops. DEJEAN, 1831, p. ["Subcylindrica, supra

laterali antice intus cupreo-aenea; elytris punctatis, margine subinterrupto dentato,

fasciaC.media subtransversa abbreviata, lunulaque apicis albis; ano testaceo. / Agilis Klug / C. Auraria Schonherr / Elle se trouve dans File de Saint-Domingue, et

l'ai de M. le de le lui ai conserve." - je re?u conte Mannerheim, sous nom que je 82

KLUG 27 C. from C. cf. JONGE Cicindela boops, 1834, p. [Separates auraria boops,

POERINK BATES 506 C. aurariae 1953, p. 122.]; 1890, p. ["C. boopi (Mann.) et (Klug)

proxima affines, differt inter alia, elytrorum margine albo postice profunde inden-

& MUTCHLER 393 HORN LENG & tata."]; LENG 1914, p. [Haiti, Cuba.]; 1915, p. 402;

MUTCHLER 1916, p. 691-692, fig. 2, pi. 12 fig. 4 [Puerto Rico; short descr.]; 1917, p.

194 HORN BEATTY 132 BLACKWELDER [boops]; 1926, p. 304; 1944, p. [St. Croix];

Rico alkali flats 1944, p. 17; WOLCOTT 1948, p. 226, fig. 1 [Puerto "only around and

near salt ponds of the southwestern corner of the island: at Faro de Cabo Rojo,

Parguera, Guanica and Ensenada."]; JONGE POERINK 1953, p. 122, 127, pi. 15

[Differences between C. auraria and C. boops; Cuba, Hispaniola, Puerto Rico,

Jamaica.];MISKIMEN & BOND 1970, p. 78; IVIE 1983, p. 194-196,fig. 2b [Anegada, St.

Croix].

Dromochorus SCHILDER 560. (Ellipsoptera) boops, 1953, p.

RIVALIER BOYD 15. Habroscelimorphaboops, 1954, p. 259, fig. 4b; 1982, p.

PUERTO RICO: Cabo Rojo, Salina Corozo, salty mudflat, 18.IX.1963 (1 $ 2$?). La

Parguera, E of Cabo Rojo, Salina Papayo, muddy salt flat, 13.IX.1963 (13(JcJ 3$?).

Isla Mayueyes, S of La Parguera, at light, 13.IX.1963 (1 §).

TURKS ISLANDS: Grand Turk VI. 1957, T. H. Farr (2 7$$). ,

CAICOS ISLANDS: South Caicos, central lowlands, 19.IV.1954, G. R. Proctor (1 CJ);

VI. 1957, T. H. Farr (1 rf).

JAMAICA: Port Henderson, salina, 30.1.1953, R. P. Bengry (1 2?$); 31.1.1953,Bengry

(6

Inst, of Jamaica, Kingston.

Body length 8.6-/0.0-11.0 $9.0-70.5-11.8 mm; width $3.7 $4.0 mm;

<$ 2.7 9 2.6 times as long as wide. (Measurements of 24 S3 20 9? from

Puerto Rico, Jamaica and Caicos & Turks Islands; see Table 2).

Labrum (Figs. 33, 37a-c) brownish, with 6 submarginal setae. Frontal

margin outwards curved, with a distinct triangular tooth.

Head, including genae, glabrous (except for 2 supra-orbital hairs).

Width across the eyes about pronotum-width = 3/7 elytron-length.

Pronotum about 7.6mm in length $2.0 92.7 mm wide, about as broad Median rather transverse furrows more as long. grove shallow,

deeply impressed. Lateral sides sparsely clothed with decumbentbristles.

times wide. Penis about 4j mm long, rather plump, about 5j as long as

and half Seven chit- Flagellum encompassing one a loop (Figs. 39c, 40c).

inous plates counted.

Legs (cf. Tables 3 and 4) of S3 distinctly longer than of 99, cupreous

in 1.65 2.0 times brown or testaceous. I, II and III SS about 1.35, and

elytron-length, resp., equalling about 0.85, 1.0 and 1.25 body-length. I, II and III in 99 about 1.2, 1.4 and 1.75 elytron-length, equalling about 0.75, 83

0.9 and 1.15 body-length. Femur III about $0.7 $0.65; tibia III $0.65

$0.6 elytron-length. Trochanters I and II with a single large seta.

Elytron <$ 5.5-6.5-1A $ 6.0-6.9-7.8mm in length, about

long as wide, i.e. more slender than in C. auraria, with lateral sides less

than in convex and apex a little more rounded; more roundedin the 9 the

with distinct without inward in the last $, a apical spine, an curve 1/5 part of the lateral side. Yellowish-white marking generally less wide and more distinct as in C. auraria, with a marginal area which is often interrupted by the dark hook in the Puerto Rico specimens.

Cicindela carthagena jamaicana (van Nidek, 1980)

[Pl. III a-c; figs. 34c-d, 37d-g]

Cicindelidia carthagenajamaicana BROUERIUS VAN NIDEK, 1980, p. 129 ["Jamaica, Duncans,

Try, 21-23.VIII.1966",3 6 $$: "The colour of the elytra... is brilliant blue-green

with dark blue punctures and a velvet-like patch in the middle of each elytron in the

colour ofthe elytra ofthe typical form. The medium elytral band is along the margin

connected with the marginal spot, situated between this band and the apical lunula.

into than in The apex of the elytra runs sharper the clear spine the typical form....

differs from the typical form by its markings and its colour, and from colossea (W.

Horn 1926), aside from colour and markings, by its size."]; BOYD 1982, p. 13.

1831: HORN 289. Dromochorus [Cicindela carthagena Dejean, 1915, p. 388; 1926, p. (Dromo-

560. Cicindelidia RIVALIER chorus) carthagena,SCHILDER, 1953, p. carthagena, 1954,

p. 257.]

JAMAICA: Port Morant (St. Thomas), 15.11.1956, R. P. Bengry (1 (J). Lyssons Beach

near Morant Bay, 6.IV.1953, Bengry (3 Two miles E of Morant Bay, 1.IV.1953,

Bengry (1(J). Bowden 28.VI.1954, T. H. Farr (2??). Palisadoes (St. Andrew), , 27.V.1947, G. B. Thompson (1

21.1.1953, Bengry

Green Bay, 27.111.1955, Bengry(1

All specimens in the Institute of Jamaica, Science Museum. - Duncans (N coast of

Trelawny), 21 .VIII.1966, Howden & Becker (1 5). - Great Saltpond entrance at Port

Clarence, near Port Henderson, sand bar near Rhizophora, with C. trif. trifasciata,

8.V.1973 (15). Last two specimens in the Zool. Museum ofAmsterdam; four of the

other onespresented to the R.M.N.H. Leiden.

Body length 9.5—70.3—11.5 $ 10.9-/7.2-12mm; width rj 3.9 $4.4 mm; c? 2.6 ?2.5 times as long as wide. (Measurements of 11c? 10? pinned 84

Mus. Table alcohol specimens from Sci. Jamaica; see 2.) [A single 9

specimen from Great Saltpond measured 12.1 mm from head to apex of elytron.]

Labrum (Figs. 34c -d, 37d-g) yellowish white, with 6 (exceptionally 5) regularly placed submarginal setae. Frontal margin only very weakly curved, with a minute central denticle in more distinctly outward

with rather tooth in curved, a well-developed triangular $?.

for of Head, including genae, glabrous (except one pair supra-orbital

hairs). Width across the eyes about lj pronotum-width = 3/7 elytron-

length.

Pronotum $ 1.9 $ 2.0 mm in length; rj 2.3 $ 2.5 mm wide; about 1.2 times

as broad as long. Median furrow weak but distinct; transverse furrows

deeply impressed. Upper flanks clothed with decumbent bristles; front,

mediodorsaland hindparts glabrous.

Legs (cf. Tables 3 and 4) of slightly longer than of $$, shining

cupreous-green.I, II and III in <$<$ about 1.4, 1.5 and 2.0 times elytron-

length, resp., equalling about 0.85,0.9 and 1.2 body-length. I, II and III in

99 about 1.3, 1.45 and 1.8 times elytron-length, equalling about 0.8, 0.9 and 1.1 body-length. Femur and tibia III about c?0.65 90.6 elytron-

length. Trochanters I and II without a long seta.

Elytron J 6.0-6.4-6.8 96.7-7.0-7.5 mm in length, about c? 3.4 93.1 as

long as wide. Lateral side weakly convex, at 4/5 ofits length turning off to

the apex; the last 1/5 part finely serrated, ending into a little spine. The

yellowish-white elytron marking consists of (see PI. Illa-c): 1) a hooked

humeral lunula without a distinct proximo-medial branch: 2) a S-shaped

middle band; 3) a hooked apical lunula; 4) a dotbetween the medial parts of the middle band and the apical lunula which may be narrowly con- nected with the proximal part of the apical lunula; 5) a marginal dot, situated between the lunula and the middle which be apical band, may narrowly connected with the latter, thus showing a tendency of forming an interrupted marginal area which connects the humeral and apical lunulae.

to of Though lam unable discover any characteristics which would justify the creation a

behalf ofthe Jamaican of Cicindela it be new taxon on specimens carthagenaDejean, may wise to maintain the subspecies jamaicana,i, for the present. The occurrence ofC. carthagena might be expected on other Greater Antilles, visa the ancient and probableerroneousrecord of C. rufiventris in Haiti. 85

Cicindela argentata Fabricius, 1801

[Pl. IVf; figs. 34e-g, 38a-c]

Cicindela 242 argentata FABRICIUS, 1801, p. ["C. capite thoracequeargentatis, elytris nigris

puncto striga undata lunulaque pallidis. Habitat in America meridionali. D. Smidt.

Mus. D. de Sehestedt."]

from be The specimens of C. argentata Guadeloupe may distinguished as var. pallipes

Fleutiaux & Salle, 1889, or as a subspecies.

Cicindela 215 de M. venant de la Guade- argentata, Dejean 1831, p. ["regu ... Chevrolat,

loupe . ..]; LENG & MUTCHLER 1914, p. 393; HORN 1915, p. 406; LENG & MUTCHLER

690 "Occurs on 194 1916, p. [var. pallipes only Guadeloupe."]; 1917, p. [var.

HORN BLACKWELDER 17. pallipes]; 1926, p. 308; 1944, p.

Dromochorus SCHILDER 561 (Cicindosa) argentata, 1953, p.

Brasiella RIVALIER BALAZUC & CHALUMEAU 3 argentata, 1954, p. 263; 1978, p. 22-23, fig.

[Several localities on Guadeloupe, Basse-Terre.].

Brasiella RIVALIER Id of the argentata pallipes, 1955, p. 80, fig. [Description type specimen

from Guadeloupe in the Chaudoir collection.]; BOYD 1982, p. 15.

R. H. CURASAO: Emmastad, golflinks, on brownish weathered diabase, 27.X.1956,

Cobben (3(J(J 3 9$); at light, X.1956, Cobben (2 $$). Rio Canario, 1956?,B. de Jong

(1?).

TRINIDAD: SI. Augustine, trench along road, IX.1956, R. H. Cobben (1 J 1 $).

SURINAME: Paramaribo, at light, 19.XI.1946, coll. D. C. Geijskes (1?); 2.XII.1958,

Geijskes(l (J); V.1959, P. H. van Doesburg, jr. (1 $); at light, 10.IV.1962,Broekhuizen

(1 cJ); Zorg en Hoop, 13.IX.1963, L. D. Brongersma (1?). Afobaka, 20.X.1964,

Geijskes (1 J). Kabel, 18.X.1958, van Doesburg (1 (J). Gansee, 10.XII.1963, Geijskes

(1 J). Rainville,28.VII.1977,A. van Assen (1 J). All Surinam specimens from RMNH

Leiden, labeled Brasiella venustula by BROUERIUS VAN NIDEK.

WEST INDIES, without data, possibly from Curasao or Trinidad (3 identification

doubtful).

6.5-7.4-8.0 width Body length $6.5-6.7-1.0 $ mm; $2.3 $2.5 mm; 2.9

times as long as wide. (Measurements of 2$$ 6$$ from Curasao and

Trinidad; see Table 2).

with 8 6 Labrum (Figs. 34e-g, 38a-c) yellowish white, (sometimes or 7) submarginal setae. Frontal margin slightly outwards curved, with a (very)

small central tooth.

Head, including genae, glabrous (except for 2 supra-orbital hairs); dark cupreous brownish-green. Width across eyes about lj pronotum-width

= 4 elytron-length.

Pronotum $ 1.3 $ 1.4 mm in length, $ 1.2 $ 1.4 mm wide, about as broad as long, sometimes slightly longer. Mediangroove and transverse furrows 86

rather indistinct. Upper lateral sides and front part near the median line

sparsely clothed with decumbent bristles.

Legs (cf. Tables 3 and 4) of

II and III in Surinam about 1.35,1.5 and 2 times elytron-length, resp.,

equalling about 0.85, 0.9 and 1.2 body-length. I, II and III in 99 from

Curasao and Suriname about 1.25, 1.4 and 1.8 times elytron-length,

equalling about 0.75,0.85 and 1.1 body-length. Femur and tibia III about

0.6 Trochanter with elytron-length. I a single large seta, II without.

Elytron <$ 4.1-4.3-4.6 94.4-4.9-6.3 mm in length, about $3.9 93.7 as

long as wide. Lateral side nearly straight, at 7/8 of its length rather

abruptly turning offto the pointed apex. Margin of the last 1/8 part finely

serrated, ending in a rather small (9) to minute (S) spine. Pits of sculpture

in field. The shining green a cupreous yellowish-white elytron marking

consists of: 1) a single dot in the middleof the proximal part; 2) a some-

times interrupted tortuous middle band; 3) a single dot in the middle of

the caudal part; 4) an apical lunula along the lateral margin, not reaching

the medial margin.

The specimens from Curasao are resembling FLEUTIAUX & SALLH'S var. pallipes from

~

- mentioned Guadeloupe. In contrast to the other cicindelids in this paper, C. argentata occurs in inland localities, obviously avoiding sea shores and other saline habitats.

Cicindela marginata Fabricius 1775

[Pl. IVa; figs. 34a-b, 38d-e, 39f, 40g]

Cicindela marginataFABRICIUS, 1775,1792,p. 176 ["C. viridis, elytris margine fascia undata - duobus albis. Habitat in LENG LENG & punctisque Virginia."] 1902, p. 162;

MUTCHLER 393 HORN LENG & MUTCHLER 1914, p. [Bahamas, Cuba.]; 1915, p. 394;

690 in Bahamas and Cuba. In the United States far 1916, p. [Occurs Florida, as

north as Maine, always on marshy sea coasts."]; 1917, p. 194; HORN 1926, p. 297;

BLACKWELDER 18. 1944, p.

Dromochorus (Ellipsoptera) marginata,SCHILDER 1953, p. 561.

RIVALIER BOYD 17. Ellipsoptera marginata, 1954, p. 267; 1982, p.

Florida: Virginia Key, sandy beach, 31.VIII.1963 (4c?(J 2$?). Key Biscayne near

Bear Cut, Sta. 690, white sand with decaying Thalassia, 7.IX.1963 (2 $$ 3 9?); sandy

beach with 1.IX.1963 Cocoa mangroves, (2

Beach, 23.VII.1966,R. D. & M. F. Ward (1 J,Zool. Mus. Amsterdam). Coral Gables,

shore, det. K. Mandl 1958, C. Blumenthal (1$, Zool. Mus. Amsterdam).

South Carolina: mudflat with Beaufort near Huntington Island, salty grasses,

29.VIII.1963 (1 (J 1$). 87

Body length $10.0-11.0-12.3 $ 10.5-77.0-11.3mm; width $4.3

$4.8 mm; 2.5 92.3 times as long as wide. (Measurements of 6$ 5$ alcohol specimens; see Table 2.).

Labrum (Figs. 34a-b, 38d—f) yellowish brown, with 9-13 submarginal

Frontal setae. margin only weakly and often irregularly curved, with a

small, more or less triangular tooth in $$.

Head sparsely covered with short decumbent bristles, the genae densely

covered. Width the across eyes about pronotum-width = 3/7 elytron-

length.

2.1 2.0 in 2.2 2.4 about 1 Pronotum $ $ mm length; $ $ mm wide; $ $ 1.2

times as broad as long. Median furrow rather weakly, transverse furrows

lateral the more deeply inpressed. Upper parts, front, hindpart along

subbasal transverse furrow, and the region along the median line, sparsely

covered with decumbentbristles.

about and about 4 times Penis 4| mm long, plump club-like, as long as

wide.Flagellum making three loops or more (Figs. 39d, 40f). Six chitinous

plates counted.

Tables 3 and of little than of Legs (cf. 4) SS only very longer $$. I, II and III in SS about 1.4, 1.7 and 2.2 times elytron-length, resp., equalling about 0.8, 1.0 and 1.3 body-length. I, II and III in 9? about 1.3, 1.6 and

2.05 times elytron-length, equalling about 0.8, 0.95 and 1.25 body-length.

Femur and tibia III about <$0.1 $0.65 elytron-length. Trochanters I and

II with a single long seta.

6.1-6.9-7.6 6.5-7.0-7.4 in about 3.1 Elytron

long as wide. Lateral side weakly convex, at 4/5 of its length turning off to

the truncated apex; the last 1/5 part finely serrated, ending into a rather

sharply projecting spine. The yellowish-white elytron marking consists of

(see PI. IVa): 1) a not well-definedhooked humeral lunula with a more or less isolated big dot which may be considered as its proximo-medial

and curved disto-medial branch, narrow, strongly branch; 2) a sharply tortuous middle band which is many times interrupted in its caudally directed part; 3) a hooked apical lunula; 4) an uninterrupted marginal area which broadly connects the humeral and apical lunulae. 88

Cicindela dorsalis media LeConte, 1856

[Pl. IVb; fig. 38f-g, 39g, 40h]

two suture Cicindela dorsalis SAY, 1817, p. 20 ["Brassy: elytra white; curved lines oneach,

and curved branch near the base green; lip and tail pail. Inhabitats New Jersey."];

13 5. - SCHAUP 91 93 1818, p. 415, pi. fig. 1884, p. 98, fig. mm],p.99, fig. [var.

13-14 in LENG 392 media, mm, Ga, S.C., N.C., Fla.]; 1902, p. 161; HORN 1915, p. &

CAZIER 293-294. 1908, p. 37, figs. 154-155; 1926, p. 295; 1954, p.

SCHILDER 560. Ellipsoptera dorsalis, 1953, p.

Habrosclerimorpha dorsalis, RIVALIER 1954, p. 158.

dorsalis BOYD 14. Habrosclerimorpha media, 1982, p.

SOUTH CAROLINA: Beaufort, HuntingtonIsland, oceancoast, hidingunder tree trunk

on white sand shore, 29.VIII.1963 (2

The specimens from Beaufort, S.C., belong to the "Florida east coast north to New Jersey population" described as C. dorsalis var. media LeConte, 1856.

about Body length 11.8-72.7-12.4mm; width 5.0 mm; 2.5 times as long

as wide (See Table 2).

Labrum (Fig. 38f-g) yellowish brown, with 6 submarginal setae.Frontal

less margin very slightly outwards curved or almost straight, more or

with small tooth which is less in undulated, a triangular developed <$<$.

Head including glabrous, for 2 supra-orbital hairs. , genae, except

about Width across the eyes about lyo pronotum-width = 3/7 elytron-

length.

Pronotum 2.2 mm in length, about 2.8 mm wide, about 1.25 times as broad as long. Median grove and transverse furrows distinct. Lateral

with decumbent parts and frontal and caudal areas densely covered

central covered. bristles; upper part more sparsely

Penis about4| mm long, slender, about 8 times as long as wide. Flagel-

lum making a single loop only. Seven chitinous plates counted.

Legs (cf. Tables 2 and 3) of distinctly longer than of ?$. I, II and III

in about 1.5 and 2.0 times about (J(J 1.25, elytron-length, resp., equalling

0.8, 1.0 and 1.3 body-length. I, II and III in 9? about 1.1, 1.3 and 1.8

elytron-length, equalling about 0.7, 0.85 and 1.2 body-length. Femur and

tibia III about

7.7 7.8 mm in about 3.2 times as as wide. Elytron c J $ length, long

Lateral side weakly convex; the proximal 1/5 part rather abruptly nar- rowing towards the shoulders; the distal 1/5 part finely serrated, turning 89

off to the somewhat truncated apex, ending into a minute spine. The

yellowish-white marking is dominating to such an extent that there are

only a few arabesque-like striae left (PI. IVb).

SIMPLIFIED KEY TO THE CICINDELIDS

BASED ON MATERIAL TREATED IN THIS PAPER

la Genae glabrous 3

lb Genae with decumbent bristles 2

2a Yellowish-white elytron marking consists of a wide, uninterrupted

fromshoulder to less trilobed marginal area extending apex, more or

within (PI. Ic) C. auraria

2b Yellowish-white elytron marking consists of a broad, uninterrupted

marginal area which connects the humeral and apical lunulae, and a

sharp tortuous middle band which is often interrupted in the caud-

ally directed medial part (PI. IVa) C. marginata

3a Yellowish-white elytron marking consists ofa rather wide, generally

uninterrupted marginal area extending from shoulder to apex, more

less trilobed within or distinctly (PI. lib) C. boops

3b Yellowish-white elytron marking consists of an often rather caprici-

ous design which may be strongly dominant 4

4a Yellowish-white marking dominant in such a way that only a few

arabesque-iike figures are left, including a longitudinal loop (PI. IVb)

C. dorsalis

4b Yellowish-white marking consists of a tortuous and often rather

capricious design which may be strongly dominant 5

5a Hindleg not very long, generally about body-length, its femur

about as long as the tibia, less than 3/4 elytron-length.... 7

5b Hindleg very long, generally about 1| body-length in males, its

femur much longer as the tibia, equalling elytron-length; first and

second trochanter without setae 6

6a Yellowish-white marking consists of a semicircular humeral lunula, 90

often middle a strongly tortuous and band, a semicircular apical

lunula, and a connecting marginal area (PI. Ha) .

C. graphiptera graphiptera

6b Yellowish-white strongly dominantof the dark parts are only a few

character-like left C. figures (PI. Ib) . . . graphiptera fulgidiceps

la Yellowish-white marking consisting of single dots in the middle of

the and distal proximal parts, a tortuous middle band, and an apical

lunula along the lateral margin; first trochanterwith a single seta (PL

IVf) C. argentata

lb Yellowish-white marking without single dots in the proximal and

distal parts 8

8a Yellowish-white design with a dot between middleband and apical

first and second trochanter without lunula; a single long seta (PI.

Illa-c) C. carthagena jamaicana

8b Yellowish-white design without a dot between middle band and

apical lunula; first trochanter with a seta 9 single long ....

9a Margin of labrum nearly straight, with very small central tooth;

pronotum about as wide as long, its medianline rather indistinct;

length of elytron mostly about mm 11

9b Margin of labrum curved, somewhat roof-shaped, with rather well-

developed central tooth; pronotum about as wide as long, its

median line distinct; length of elytron mostly about 65 mm. 10

10a Tortuous design striking, with an uninterrupted marginal area and a

distinct humeral lunula (PI. Id) C. trifasciata trifasciata

10b middle band Tortuous narrow, humeral lunula mostly consisting of

small a spot only (PI. lie) C. trifasciata ascendens

11a Yellowish-white few dark strongly dominant, leaving a parts only,

especially along the suture; no lunulae can be distinguished (PI. He)

C. suturalis suturalis lib Tortuous middle band and sharply hooked lunulaerather wide (PI.

IVc, e) C. suturalis hebraea

11c Tortuous middle band and lunulae rather narrow . (PL lid). . .

iC. suturalis guadeloupensis 91

28. Labrum and of of Cicindela from above: a from Marga- Fig. apex right elytron auraria, rita (Punta de Piedras); b-d from Bonaire (b-c Lagoen, d Salinja Martinus);e-f from Klein

Bonaire; g-i from Aruba (Spaans Lagoen). - Half of the specimens in C. auraria have 6

- males rule less submarginal setae, the other half 7 to 8, rarely 9. Margin of labrum in as a

protrudingthan in females. Apex of elytron in the female a little less pointed than in the

male.

Fig. 29. Labrum and apex of right elytron of Cicindela trifasciata trifasciata: a-b from St.

Martin (Simson Lagoon); c-d from Grand Turk; e-f from South Caicos; g-h from Jamaica

(Hope River). - The labrum of C. trifasciata has a variable number of submarginal setae, 8-

13. - Margin of labrum in males generally a little less protruding than in females. Apex of

elytron in the female less pointed than in the male. 92

30. Labrum and of of Cicindela a-b from Fig. apex right elytron trifasciata trifasciata:

Antigua (St. John’s); c-d from Barbuda (Great Lagoon); e-f from St. Kitts (Frigate Bay). - Cf. Fig. 29.

31. Labrum and of of Cicindela suturalis suturalis: a-b from St. Fig. apex right elytron

Thomas (Magens Bay); c-d from Barbuda (Martello Tower beach); e-ffrom Antigua (Deep

Bay). - The labrum of C. suturalis has commonly 8 submarginal setae, sometimes 9 or 10, more rarely 7. - Margin of labrum in males not protruding or only slightly curved outwards.

Apex of elytron especially in females distinctly truncated. 93

Fig.C.32. Labrum and apex of right elytron of Cicindela suturalis hebraea (a-c) and graphiptera fulgidiceps(d-e): a from Tobago (Red Point); b-c from Trinidad (b Los Gallos

Point, c Cocos Beach); d-e from Margarita (d Punta de Piedras, e Punta Mosquito). - The labrum of C. graphipterahas commonly 6 submarginalsetae, sometimes 7 or 8. - Margin of labrum in males generallynot protruding. Apex of elytron in females somewhat truncated.

Fig. 33. Labrum and apex of right elytron of Cicindela boops: a-b from Jamaica (Port

Henderson); c-d from South Caicos; e-f from Puerto Rico Saliña Papayo). - The labrum of - C. boops has 6 submarginal setae. Margin of labrum in males generally less protruding

than in females. Apex of elytron in the female less pointed than in the male. 94

Fig. 34. Labrum and apex of right elytron of Cicindela marginata (a-b), C. carthagena jamaicana(c-d) and C. argentata (e-g). - The labrum of C. marginatashowed 9-11 submar-

C. and C. 8. - of C. ginal setae; carthagena 6, argentata commonly Margin marginata more

or less irregularly curved, of C. carthagena and C. argentata, generally less protruding in

males than in females. Apex of elytron as a rule less pointed in the females.

Fig. 35. Labrum and mandibles of Cicindela auraria (a-c) and C. trifasciata trifasciata (d-e):

a-c from Curaçao (St. Jorisbaai); d from Barbuda (S ofCodrington Village); e from Antigua

(Dickinson Bay). 95

Fig. 36. Labrum and mandibles of Cicindela graphipterafulgidiceps (a-b) and C. suturalis

suturalis (c—d): a-b from Margarita (Pta Piedras); c-d from Antigua (Dickinson Bay).

Fig. 37. Labrum and mandibles of Cicindela boops (a-c) and C. carthagenajamaicana(d-g):

a-c from Puerto Rico (a-b Sal. Papayo, c Sal. Corozo); d-g from Jamaica (d Green Bay, e

Port Morant, f-g Port Henderson). 96

Labrum and mandibles of Cicindela C. and C. Fig. 38. argentata (a-c), marginata (d—e)

dorsalis (f-g): a-b from Curaçao, c from unknown Caribbean locality; d-e from Florida (d

Virginia Key, e Key Biscayne); f-g from South Carolina (Beaufort) - with details of

bifurcated tip of right ￿ mandible in two specimens of C. marginata and C. dorsalis.

39. from the male of: Cicindela auraria,b,C. Fig. Flagellum copulatory organ a, trifasciata

C. suturalis C. C. trifasciata; c, boops; d, C. hebraea; e, graphiptera fulgidiceps; f, marginata; g,

C. dorsalis media. 97

40. Male of: Cicindela auraria; b, C. trifasciata Fig. copulatory organ (penis, aedoeagus) a,

trifasciata; c, C. boops; d, C. suturalis suturalis; e, C. suturalis hebraea; f, C. graphiptera

fulgidiceps; g, C. marginata; h, C. dorsalis media. 98

Fig. 41. Larva ofCicindela auraria from the mudflats ofSanta Martha, Curaçao: a, head; b,

fifth abdominal d, nine - pronotum; c, segment; segments to ten all dorsal aspect; e, antenna;

f, maxilla; g, metathoracic leg. 99

TABLE 2

MEASUREMENTS IN Cicindela, IN MM

by CARLA OLDENBURGER, in which a few unpublished data from

+ included. JONGE POERINK’S paper ( ) are

Elytron Pronotum Head Body El. Pron. Head Body

length width 1. w. 1. w. 1. w. l./w. w./l. w./l. l./w.

C. auraria

Aruba

$ 2 6.7 - 6.8 2.0 1.6 2.5 1.7 3.1 10.3 4.0 3.4 1.6 1.7 2.5

? 3 6.6-7.0-7.4 2.2 1.6 2.5 1.8 3.4 10.6 4.4 3.2 1.5 1.9 2.2

+ 6 6.2 2.0 1.5 2.3 9.5 4.0 3.1 1.4 2.4

+ 9 6 6.7 2.1 1.6 2.5 10.0 4.2 3.2 1.5 2.4

Bonaire

13 5.2-6.2-6.8 1.9 1.5 2.2 1.5 2.8 9.5 3.9 3.3 U 1.8 2.4

9 13 5.7-6J-7.0 2.0 L5 2.3 1.6 3.0 9.6 4.2 3.2 1.5 1.9 2.3

Klein Bonaire

+ tj 17 6.3 2.0 1.5 2.2 9.5 3.9 3.2 1.4 2.4

+ $ 26 6.5 2.1 1.5 2.3 9.6 4.1 3.1 1.5 2.3

Margarita

? 4 5.0-6.0-6.4 1.9 1J 2.1 1.6 2.7 9.2 3.8 3.1 1.4 1.8 2.4

La Goajira (P. Lopez)

S 8 6.3 2.0 1.6 23 9.7 4.0 3.2 1.4 2.4

? 13 6.5 2.1 1.6 2.4 9.8 4.3 3.0 1.4 2.3

C. trifasciata trifasciata

Antigua

? 5 6.1-6.4-6.6 2.1 1.9 2.2 1.8 3.2 10.3 4.3 3.1 1.1 1.7 2.4

Barbuda

cJ 5 6.5-6.(5-6.8 2.2 2.1 2.3 1.6 3.2 10.4 4.3 3.0 1.0 2.0 2.4

? 7 6.1-6.7-7.5 2.3 1.9 2.4 1.7 3.2 10.3 4.6 2.9 1.2 1.9 2.2

St. Kitts

cj 6 5.9-6.4-7.1 2.0 1.8 2.1 1.6 3.0 9.7 4.3 3.2 1.1 1.8 2.4

? 6 6.1-6.6-7.7 2.3 1.9 2.3 1.8 3.1 10.2 4.5 2.9 1.2 1.7 2.2

St. Martin

c? 6 6.5-6.7-7.3 2.1 1.9 2.2 1.7 3.0 10.5 4.2 3.2 1.1 1.7 2.5

? 6 6.3-6.7-7.4 2.2 1.9 2.2 1.7 3.2 10.2 4.3 3.0 1.1 1.9 2.4

+ ? 16 7.3 2.4 2.1 2.5 11.3 4.7 3.1 1.2 2.4 100

Table 2 (cont.)

Elytron Pronotum Head Body El. Pron. Head Body

length width 1. w. 1. w. L w. l./w. w./l. w./l. l./w.

Grand Turk (dry specimens)

cJ 7 6.6-7.0-7.3 2.0 2.0 2.4 2.1 3.2 11.0 4.0 5.5 1.2 1.5 2.7

9 4 6.7-7.0-7.5 2.1 2.0 2.5 2.1 3.4 11.1 4.2 3.3 1.2 1.6 2.6

South Caicos (dry specimens)

6 6.0-6.7-7.6 2.0 2.0 2.3 2.1 3.2 10.7 4.0 3.3 1.1 1.5 2.7

9 2 6.5 - 7.3 1.9 2.0 2.4 2.3 3.2 10.8 3.7 3.6 1.2 1.4 2.9

Jamaica (dry specimens)

S 7 6.0-6.6-7.0 2.0 1.9 2.2 1.8 3.0 10.0 3.9 3.3 1.1 1.6 2.5

9 1 6.8 2.0 2.0 2.3 2.2 3.1 10.7 4.0 3.4 1.1 1.4 2.6

C. suturalis suturalis

Antigua

tJ 5 5.0-5.2-5.4 1.6 1.5 1.8 1.3 2.3 7.9 3.2 3.2 1.2 1.7 2.4

9 5 5.0-5.6-5.9 1.7 1.5 2.0 1.3 2.3 8.3 3.4 3.3 1.3 1.7 2.4

Barbuda

$ 4 5.1-5.2-5.5 1.5 1.4 1.8 1.3 2.3 8.0 3.1 3.4 13 1.8 2.6

9 6 5.4-5.9-6.5 1.8 1.6 1.9 1.4 2.5 8.7 3.6 3.3 1.2 1.8 2.4

St. Thomas

(J 4 5.0-5./-5.2 1.5 1.3 1.6 1.3 2.2 7.8 3.1 3.4 1.2 1.7 2.5

9 6 5.1-5.4-5.6 1.6 1.4 1.8 1.3 2.4 8.1 3.2 3.4 1.2 1.8 2.5

C. suturalis hebraea

Trinidad

S 12 4.9-5.7-5.5 1.4 1.4 1.7 1.3 2.2 7.8 2.8 3.6 1.2 1.7 2.7

9 6 5.4-5.6-5.7 1.7 1.5 1.8 1.4 2.4 8.4 3.3 3.3 1.2 1.7 2.5

Tobago

cJ 5 4.8-5.0-5.3 1.5 1.4 1.7 1.2 2.2 7.7 3.0 3.3 1.2 1.8 2.6

9 4 5.2-5.5-5.8 1.5 U 1.9 1.4 2.4 82 3.2 3.6 1.3 1.7 2.6

C. graphipterafulgidiceps

Margarita

9 13 7.0-7.7-7.9 2.1 1.6 2.5 1.8 2.9 112 4.2 3.6 1.5 1.6 2.6

La Goajira (P. Lopez)

$ <5.5 1.8 1.6 2.4 9.9 3.5 3.6 U 2.8

+ 9 10 7.4 1.9 1.7 2.6 11.4 3.8 3.9 1.5 3.0

La Goajira (R. Hacha)

+ cj 4 6.0 1.5 1.5 2.2 9.4 3.1 3.9 1.4 3.0

+ 9 5 7.4 1.9 1.7 2.5 11.4 3.9 3.8 1.5 3.0 101

Table 2 (cont.)

Elytron Pronotum Head Body El. Pron. Head Body

length width 1. w. 1. w. 1. w. L/w. w./l. w./l. l./w.

C. boops

S. Caicos & Gr. Turk (dry specimens)

cJ 3 6.5-6.S-7.1 1.9 1.6 2.2 1.9 2.9 10.4 3.8 3.6 1.4 1.5 2.7

? 7 6.6-7.3-7.6 2.0 1.7 2.2 2.1 3.1 11.0 4.0 3.6 13 1.5 2.7

Puerto Rico

14 6.4-6.S-7.4 1.9 1.6 2.2 1.6 3.0 10.3 3.9 3.6 1.4 1.9 2.7

? 6 6.7-7.7-7.8 22 1.6 2.4 1.8 32 11.0 4.4 33 1.5 1.8 2.5

Jamaica (dry specimens) tJ 7 5.5-6.0-6.7 1.6 13 1.8 1.7 2.6 9.3 3.4 3.8 1.2 1.5 2.7

? 7 6.0-6.5-6.7 1.8 13 1.9 1.7 2.6 9.6 3.6 33 1.3 1.5 2.6

C. carthagena jamaicana

Jamaica (dry specimens) s 11 6.0-6.4-6.8 1.9 1.9 2.3 1.8 2.7 10.3 3.9 3.4 1.2 1.5 2.6

$ 10 6.7-7.0-7.5 2.2 2.0 2.5 2.2 3.0 11.2 4.4 3.1 1.2 1.3 2.5

C. argentata

Curasao (dry specimens)

cJ 1 4.6 1.1 1.4 1.2 1.1 2.0 7.0 2.2 4.2 0.8 1.8 3.2

$ 5 4.4-5.0-6.3 1.3 1.4 1.4 1.5 1.9 7.5 2.5 3.8 1.0 1.2 3.0

Trinidad (dry specimens)

cJ 1 4.1 1.2 1.2 1.2 1.2 1.8 6.5 2.4 3.4 1.0 1.5 2.7

? 1 4.5 1.2 1.4 1.5 1.4 2.0 7.2 2.4 3.7 1.0 1.4 3.0

C. marginata

Key Biscayne cJ 6 6.1-6.9-7.5 2.2 2.1 2.2 1.8 3.1 11.0 4.3 3.1 1.0 1.7 2.5

? 5 6.5-7.0-7.4 2.4 2.0 2.4 2.0 3.3 11.0 4.8 2.9 1.2 1.6 2.3 102

TABLE 3

AVERAGED MEASUREMENTSIN CICINDELA LEGS, IN MM.

length of III II I

fem. tib. fem. tib. Elytron body tar. leg tar. leg fern. tib. tar. leg

C. auraria

Curacao

3 5 6.7 10.7 4.95 4.2 4.7 13J8 3.8 3.3 3.75 10.9 3.1 2.5 3.0 8.7

9 8 6.6 10.5 4.75 4.1 4.1 13.0 3.7 3.3 3.3 10.3 3.1 2.5 2.8 8.6

Bonaire

9 10 6.65 10.5 4.55 3.9 3.9 12.5 3.55 3.05 3.05 9.8 3.05 2.35 2.8 8.2

C. trifasciata trifasciata Antigua

c? 6 6.95 11.6 4.9 4.9 5.35 /5./ 3.6 3.7 4.0 11.0 3.25 2.7 4.0 /0.0

Barbuda

c? 9 6.5 70.9 4.6 4.5 5.05 14.1 3.5 3.5 3.8 10.7 3.2 2.5 3.8 9.9

9 7 6.7 11.1 4.6 4.35 4.5 3.4 3.2 3.2 9.1 3.1 2.5 3.3 8.9

St. Martin

$ 10 6.95 11.6 4.9 4.7 5.2 14.8 3.75 3.6 4.05 11.4 3.5 2.7 4.0 10.1

9 10 7.3 12.3 5.0 4.8 5.0 14.7 3.8 3.65 3.8 1U 3.45 2.7 3.65 9.8

Anegada

t? 3 6.9 11.0 4.85 4.75 5.1 14.7 3.7 3.6 4.0 113 3.15 2.55 3.85 9.6

9 4 6.85 11.1 4.85 4.55 4.85 14.3 3.9 3.45 3.5 10.9 3.2 2.5 3.35 9.1

C. suturalis suturalis

Antigua & Barbuda

9 7 5.45 8.8 3.5 3.45 3.7 /0.« 2.7 2.35 2.75 7.8 2.3 1.8 2.45

C. suturalis hebraea

Trinidad & Tobago

9 5 5.4 8.7 3.6 3.5 3.5 /O.0 2.65 2.45 2.75 7.

C. graphipterafulgidiceps

Margarita tJ 10 6.7 M 7.0 4.85 4.95 170 3.75 3.0 3.8 10.5 2.9 205 3.05 8.0

9 10 8.0 12.60 7.0 4.75 4.8 16.4 3.8 3.0 3.45 10.3 2.95 2.05 2.85 7.9

C. boops

Puerto Rico cJ 10 7.15 11.5 5.05 4.6 4.85 14.5 4.0 3.75 3.9 11.7 3.35 2.7 3.7 9.7

9 5 7.6 11.6 4.9 4.5 4.05 13.4 3.7 3.6 3.35 10.6 3.15 2.7 3.0 8.9

Jamaica (dry specimens)

9 2 7.95 12.2 4.5 4.3 4.0 12.8 3.5 3.4 3.2 10.1 3.1 2.65 3.0 8.8 103

length of III II I

Elytron body fem. tib. tar. leg fem. tib. tar. leg fem. tib. tar. leg

C. carthagena jamaicana

Jamaica (dry specimens)

$ 4 6.25 10.3 4.15 4.05 4.15 12.5 3.15 2.9 3.5 9J 3.0 2.35 3.5 8.9

9 2 7.0 11.3 4.4 — — — 3.45 3.25 3.4 10.1 3.2 2.6 3.3 9.1

Jamaica (paratype dry)

9 1 6.8 — 4.0 4.2 3.8 12.0 3.2 3.0 3.25 9.4 3.0 2.3 3.2 8.5

Jamaica (Saltpond)

9 1 7.2 11.6 4.4 4.25 4.2 12.8 3.4 3.2 3.35 9.8 3.1 2.55 3.25 9.1

C. argentata

Curacao (dry specimens)

9 2 4.55 7J5 2.45 2.75 3.1 8.3 2.9 2.0 — — 1.9 1.5 2.2 5J

Curasao?

9 3 5.2 8.7 3.0 3.1 3.35 9.4 2.4 2.2 2.7 7.3 2.15 1.8 2.55 6.5

Suriname (dry specimens) cJ 3 5.0 8.15 3.1 3.2 3.5 9.9 2.3 2.2 2.9 7.4 2.15 1.85 2.9 6.8

9 3 5.1 8.3 3.0 3.0 3.0 9.0 2.4 2.1 2.45 7.0 2.25 1.8 2.5 6.5

C. marginata

Key Biscayne

9 4 7.6 12.6 5.2 4.95 5.8 15.8 4.0 3.75 4.6 121 3.3 2.7 4.0 10.1

C. dorsalis

S. Carolina (Beaufort)

9 2 7.75 12.1 5.8 4.35 4.05 14.2 3.9 3.1 3.05 10.1 3.15 2.45 2.85 8.4 104

TABLE 4

AVERAGED MEASUREMENTS IN CICINDELA LEGS

EXPRESSED IN PERCENTAGES OF THE ELYTRON LENGTH

Elytron III II I

- length

in mm fern. tib. tar. leg fem. tib. tar. leg fem. tib. tar. leg

C. auraria

Curacao

3 5 6.7 73.9 62.7 80.1 206 56.7 49.3 60.0 162 45.3 37.3 44.8 750

9 8 6.6 71.7 62.1 62.1 196 56.1 50.0 50.0 156 47.7 38.2 43.2 130

Bonaire

3 10 6.3 72.5 63.5 69.8 206 59.0 52.0 59.7 167 49.1 37.6 49.7 138

C. trifasciata trifasciata

Antigua

3 6 6.95 70.7 70.7 77.0 218 52.1 53.4 57.7 159 46.9 39.0 57.7 AM

Barbuda

3 9 <5.5 70.8 69.2 77.7 2/7 53.8 53.9 58.8 165 49.2 38.2 58.5 752

$ 7 (5.7 68.7 64.9 67.2 203 50.7 47.8 47.9 135 46.3 37.8 48.8 133

St. Martin

3 10 6.95 70.6 67.7 74.9 213 53.7 51.9 58.3 164 50.0 38.9 57.7 146

$ 10 7.3 68.5 65.5 67.9 201 52.1 50.0 52.0 153 47.3 37.0 50.0 134

Anegada

3 3 6.9 69.7 68.5 73.3 212 53.4 52.0 57.43 762.6 45.5 37.1 55.7 7JS

$ 4 6.S5 71.1 66.9 70.7 208 57.0 49.9 50.97 /5SJ 47.2 36.9 49.0 133

C. suturalis suturalis

Antigua& Barbuda

3 7 5.2 65.4 65.6 72.1 202 50.5 45.7 54.4 150 42.5 35.0 50.4 128

9 7 5.45 65.0 63.5 69.0 198 49.8 43.6 50.6 145 43.0 34.0 45.0 122

C. suturalis hebraea

Trinidad & Tobago

9 5 5.4 65.7 64.3 67.5 195 49.1 45.8 51.0 145 42.7 34.0 44.1 121

C. graphipterafulgidiceps Margarita

3 10 6.7 104.5 72.1 73.6 254 55.8 44.8 56.7 157 43.3 30.6 45.6 119

9 10 8.0 87.5 59.4 59.8 205 47.5 37.5 42.9 128 37.1 25.8 35.6 99

C. boops

Puerto Rico

3 10 7.15 70.4 64.4 67.8 203 56.0 52.7 54.6 164 46.9 38.0 51.5 136

9 5 7.6 64.5 59.2 53.3 176 48.7 47.4 44.0 139 41.7 39.5 39.5 117 105

Elytron III II I

- length

in fern. tib. fern. tib. tar. fern. tib. tar. mm tar. leg leg leg

C. carthagena jamaicana Jamaica (dry spec.)

S 4 6.25 66.5 64.9 66.4 201 50.5 46.5 56.1 152 48.1 37.9 56.1 142

5 2 7.0 63.0 — — — 50.0 46.0 49.0 145 46.0 37.0 47.5 130

Jamaica (dry, paratype)

? 1 6.8 58.8 61.8 55.9 176 47.0 44.1 47.2 138 44.4 32.3 48.5 125

(Saltpond)

? 1 7.2 61.0 59.0 58.2 178 47.2 44.5 46.5 136 43.0 35.4 45.0 126

C. argentata

Curasao (dry spec.)

S 2 4.55 53.4 60.1 67.2 /«/ 40.0 41.0 — — 42.1 33.3 47.2 121

Curapao?

$ 3 5.2 57.7 59.8 64.4 180 46.0 42.7 51.9 140 48.2 32.7 48.0 125

Suriname (dry spec.)

? 3 5.1 59.5 58.1 59.3 176 47.7 40.8 48.1 /i(5 44.0 35.3 49.1 128

C. marginata

Key Biscayne

$ 4 7.(5 68.2 64.8 75.7 207 52.5 49.2 60.4 161 43.3 35.4 52.5 132

C. dorsalis

S. Carolina

9 2 7.75 74.5 56.5 52.2 183 50.2 40.0 39.3 130 40.7 31.5 36.5 108 106

Megacephala acutipennis Dejean, 1825

[Pl. Vc; fig. 47a-b]

WAGENAAR HUMMELINCK 1955, p. 91-93, figs. 3-5, pi. 6 [Material from Hispaniola and

Cuba; synonymy.]

13 Megacephala AcutipennisDejean, 1825, p. ["Obscuro-aenea; ore antennis, ano, pedibus

elytromrumque apicis macula obliqua testaceis; elytris obliqua testaceis; elytris

punctatis, aculeatis... Saint-Domingue"]. - Megacephala acutipennis, LENG &

MUTCHLER 393 St. Thomas and Porto BOYD 1914, p. [Haiti,Cuba; not Rico.]; 1982,

4 ?Puerto p. ["Cuba, Hispaniola, Rico".]

Leg measurements in a few specimens from Haiti - already studied before (1955, p. 91) - proved that the male M. acutipennis - being on the average somewhat smaller than the female - has comparatively longer

legs (cf. Tables 6 and 7).

Megacephala affinis affinis Dejean, 1825

[Fig. 47c-d]

WAGENAAR HUMMELINCK 7 from Trinidad and 1955, p. 93-95, figs. 6-7,9-10, pi. [Material

Hispaniola, Venezuela, Guiana, Brasil, Colombia and Honduras.]

12-13 Megacephala Affinis DEJEAN, 1825, p. ["Viridi-obscura; ore, antennis, ano, pedibus elytrorumque apicibus macula communi cordata late emargjnata testaceis; elytris

subrugosus; geniculis obscuris. ... Cayenne."]

A few indicate that the of the measurements legs male specimens are

slightly longer than those of the female individuals (cf. Tables 6 and 7).

Megacephala affinis gracilis Reiche, 1842

[Pl. Vb]

WAGENAAR HUMMELINCK 1955, p. 95-97, figs. 8-10, pi. 7 [Material from Curasao and

Colombia; synonymy.] 107

Megacephala Carolina Carolina (Linnaeus, 1767)

[Fig. 47g-i]

WAGENAAR HUMMELINCK 14 and 6 from U.S.A. 1955, p. 97-102, figs. 11, 16, pi. [Material and Mexico.]

Measurements of a few specimens from various localities - for the

greaterpart already studied before- do not show appreciable differences:

the size of the females somewhat than that of the average being larger males, while the legs of the latter may be slightly longer (cf. Tables 6 and

7).

Megacephala Carolina Carolina L. forma occidentalis Klug, 1829

[Pl. Vd-e; fig. 47j]

WAGENAAR HUMMELINCK 1955, p. 102-103,figs. 12-13 and 15, pi. 6 [Materialfrom Cuba;

synonymy.]

carolina BOYD 4 Bahamas Megacephala occidentalis, 1982, p. [Cuba, (Watling Is.).]

JAMAICA: Half Way Tree, 25.X.1952, R. P. Bengry (1

16.XI.1952, Bengry (1

Bengry (2

Lynn (2 SS 299); Hope River bed behind U.C.W.E., 19.IX.1952, W. G. Lynn (1 9).

Beacon-Pedro Plains, in hole in ground, clear weeded corn, terra rossa soil,

21.X.1953, W. B. Dixon (1 V). Yallahs River at Morant Bay Road, 31.X.1954,T. H.

Farr (1 cj). Kingston 10, 60 Renfield, 26.IX.1980, C. Christie (1

from Sci. Mus. Jamaica.

GRAND CAYMAN: June 1962, T.H.Farr (lcJ; lenght of elytron 9.5 mm, of penis 4.5 mm).

The material from Jamaica does not differfrom that of Cuba described in 1955 (cf. Table 5). The Cayman specimen shows the distinguishing colour characteristics mentioned LENG & MUTCHLER by (1916, p. 687). 108

Megacephala sobrina Dejean, 1831

103-105 the delimitation of WAGENAAR HUMMELINCK 1955, p. ["Admitting very vague the

lower a certain subdivision of the sobrina in and categories, ... species subspecies

which into distribution varieties has been maintained, may give some insight its and

differentiation on the West-Indian islands."]

Megacephala sobrina, FERNANDEZ YEPEZ & ROSALES 1956, p. 170 [Gran Roque.]

In most specimens the length of the elytron is mm, of the body

head without (= elytron + pronotum + labrum) mm; legs I, II and III being a little over 1, about lj, and almost 2 times elytron-length, resp. Though the female specimens, when averaged, are about 5% larger than the male individuals, their legs may be 5-10% shorter (cf. Tables 5-

7).

Measurements of the following specimens from RMNH Leiden - labelled M. sobrina

confusa Chd. by Mr. C. M. C. BROUERIUS VAN NIPEK - are included in Tables 6 and 7:

SURINAME: Paramaribo, Zorg en Hoop, 16.V.1955,D. C. Geijskes (1 (J); 12.VI.1957,

P. H. van Doesburg, jr. (2 2$); Combe,26.XII.1964, Geijskes (1

Coropina Kreek, at light, 15—17. VII.1977, A. van Assen (1 ?); 9.VII.1978, van Assen

"As the first description of the species prevents any classification ofthe type-specimens in

minor systematic categories,... material... from ... Margarita and Los Testigos ... has been

considered the form of the sobrina.” If as representing typical subspecies (cf. 1955, p. 105)

further restriction might be desirable, those specimens collected in the Patio of Hotel

Sta. be considered material Central, 155, Porlamar, 25.V.1936, may as lecto-type (1955, p.

105, figs. 17a, c, f, h, j and 20a, c-e, pi. 8).

Megacephala sobrina sobrina Dejean, 1831 (forma typica)

[Pl. VIa-b; figs. 42a-b, e-f, 48a-d, 50a-d]

WAGENAAR HUMMELINCK 8 from 1955, p. 105-108, figs. 17-20, pi. [Material Margarita,

Cubagua, Los Testigos and Patos.]

Punta MARGARITA: Mosquito beach debris on sandy shore, Sta. 797, 13.1.1964 ,

(3

Piedras, sandy beach debris near lagoon entrance, Sta. 802, 9.1.1964 (10 7 $?)■

The newly collected material agrees with that described before, the female individuals being slightly but distinctly larger than the male speci- 109

802: 9.85 15.3 mens, e.g. at Sta. average elytron-length J 9.1 $ mm,body

$ 16.5 mm (cf. Table 5).

Megacephala sobrina sobrina, forma antiguana Leng & Mutchler, 1916

[Pl. VIc; figs. 44-45, 49a-h, 51a-d]

WAGENAAR HUMMELINCK 9 from 1955, p. 108-110,figs. 17s-v, 20f-i, pi. [Material Antigua,

St. Barts, St. Martin and Hispaniola; synonymy.]

BALAZUC & CHALUMEAU 1 Megacephala(Tetracha) sobrina, 1978, p. 18-20, fig. [Pro parte:

St. Barts, also St. Martin and Tintamarre.]

sobrina BOYD 4. Megacephala confusa Chaudoir, 1982, p.

Megacephala sp., BUTLIN 1976, p. 128 [Anegada.]

BARBUDA: Darby's Cave, sink-hole in limestone terrace, Sta. 600, 10.VII.1955

(elytron). Codrington Village, limestone flat near lagoon, Sta. 603, 5.VII.1955

(elytron). Low Pond, brackish cattle pond near lagoon N ofCodrington Village, Sta.

674a, 21.V1I.1967 (61 21 ??).

ST. KITTS: Frigate Bay, near saltpond, Sta. 677, 20.VII.1955 (1$).

ST. MARTIN: Point Blanche, below cow faeces near shore, 25.VII. 1955 (3

Great Saltpond, NE shore, debris on salty mud, Sta. (710), 29.IX.1963 (1 CJ). Oyster

Pond SE shore, below cow faeces on 13.X.1963 (9r?c? 15 $$). Cul-de-Sac, , grassy area, 3.XI.1978, coll. F. Chalumeau (2??).

ANEGADA: Setting Point,9.VIII-21.IX.1975, R. K. Butlin (1?, Br. Mus.).

The material collected on Barbuda perfectly agrees with that from St.

that the from Low Pond Martin, except specimens are notably more

robust than the Oyster Pond individuals (cf. Tables 5-7). In a single case

the design ofthe elytrae resembled that of M. sobrina infuscata, the black

the dorsal far about spot superseding area as as 9/10 sutural-length.

Megacephala sobrina sobrina, forma bonaireana Humlnk, 1955

[Pl. Va; figs. 42c-d, g-i, 43, 47f, 48e-o, 50e-h]

WAGENAAR HUMMELINCK 7 from Klein 1955, p. 110-111, fig. 17-20, pi. [Material Bonaire, Bonaire, Curasao and Aruba; synonymy.] 110

BONAIRE: DOS Pos, below rock debris near fresh-water well NE ofGoto, 5.XII.1963

(18

7.XII.1963 (1 ?). Pos Frances, low limestone flat with brackish mud S of Kralendijk,

1.IV.1955 (1 cj dead).

KLEIN BONAIRE: POS Calbas, nearpuddle ofbrackish water on low limestone terrace,

Sta. 63A, 3.XII.1963 (1 $).

CURASAO: Boca Labadera, N coast of Sta. Catarina, 29.111.1970 (1 (J).

ARUBA: Salinja Master, near Savaneta, among debris near abandoned saltpan,

Rooi rock 29.IV.1955 (19). Bringamosa, among plant decay and non-calcareous

brackish brooklet in central Sta. 5.XI.1963 debris near Aruba, (103B), 89$);

(103d), 20.X. 1967 (1

If it would be necessary giving this forma a more restricted area, it

could be: type locality Bonaire, type lot (4 SS 2 ?$) from Deenterra, Sta.

186, 25.111.1937(cf. 1955, p. Ill; figs. 17k-m, o-p, r; 18a, d; 19e, and 201;

pl. 9, ?).

The new material agrees with that described before (cf. Tables 5-7).

The female specimen of Salinja Master, Aruba, is exceptionally large:

11 total 18 width of of elytron-length mm, length mm; body 6, pronotum

4.3, of head 4.7 mm.

When comparing the Aruba materialwith the other sobrina's from the

Leeward Group, F. VAN DER HEIDE observed the following differences.

MARGARITA ARUBA BONAIRE

specimens: specimens: specimens:

Head

usually a glossy green usually a glossy green purple colour

with a more or less with a more or less dominating in central

reddish shine. distinct purple shine. part, from without

there often with green.

Pronotum

with with colours a glossy green a a glossy green a purple

yellowish red to yellowish red to dominating in central

purple shine. purple shine. part; lateral parts

green. 111

Elytron dorsal area a bright dorsal area a bright dorsal area a bright

often intensive yellowish red, often purple, with a purple, most with a purple touch; yellow touch; near the suture; latero-dorsal area a latero-dorsal area a latero-dorsal area a

with blue. brilliant green with a brilliant green a brilliant greenish metallic shine, as a metallic shine, as a rule enclosing base rule enclosing base and both lateral sides and both lateral sides of the black spot, of the black spot, covering or nearly covering or nearly covering the entire covering the entire base of the elytron. base of the elytron.

Megacephala sobrina infuscata Mannerheim, 1837

[Pl. VId; figs. 46c-e, i-k, 49i-l, 51e-h]

WAGENAAR HUMMELINCK 9 from St. St. 1955, p. 112-114, figs. 21-22, pi. [Material Croix,

John, Puerto Rico, Hispaniola and Cuba; synonymy.]

Megacephala sobrina var. infuscata, WOLCOTT 1948, p. 225, fig.

Tetracha sobrina infuscata, BEATTY 1944, p. 131 [St. Croix.]

IVIE 1 St. Megacephala sobrina, 1983, p. 192-194, fig. [St. Thomas, Anegada, Croix.]

BALAZUC & 1 Megacephala (Tetracha) sobrina, CHALUMEAU, 1978, p. 18-20, fig. ["La race

forme infuscata ... ne peut etre tenue pour une insulaire, puisqu'elle se retrouve en

Guyane ... et en Colombie."]

Megacephala(Tetracha) rutilans Thomson = M. sobrina infuscata. MISKIMEN & BOND 1970,

78 St. p. ["Very uncommon on Croix."]

BOYD 4. Megacephala sobrina infuscata, 1982, p.

PUERTO RICO: Saliiia Papayo, near La Parguera, hiding under timber on salty

mudflat, Sta. (699), 13.IX.1963 (1 J 1 $). Saliiia Corozo, near Cabo Rojo, debris on

mudflat near saltlake, 18.IX.1963 (4

The newly collected material agrees with that described before (cf.

Tables 5-7). 112

SIMPLIFIED KEY TO THE MEGACEPHALAS

BASED ON MATERIAL TREATED IN PRESENT & PREVIOUS PAPERS

M. 1a Elytron acuminate at apex (PI. Vc) acutipennis

2 lb Elytron rounded at apex

2a Right mandible with 3 apical teeth. Lateral teeth of labrum larger

than central ones. Plainly coloured, blue or green predominating,

without black spot 3

2b Right mandible with 4 apical teeth. Lateral teeth usually not larger

than central ones, often weakly developed. In most cases vividly

brilliant with and with coloured, green predominating, usually purple,

black 4 a spot

with brownish about 3a Generally blue, often greenish or shades. Elytron

4 times as long as wide M. affinis affinis

3b Generally green, often with bluish sides. Elytron about 3f as long as

wide (PI. Vb) M. affinis gracilis

4a Apical part of elytron without traces of imbricated granules; hind

margin smooth. Hind part of head only occasionally with irregular,

small wrinkles. Margin of labrum as a rule slightly protruding before

curving backwards into the side margin. Apical lunula rounded at

base and sharply sinuated on its inner edge, abruptly narrowing to-

carolina carolina wards the suture M. ... 5

4b of with imbricated hind micro- Apical part elytron granules; margin

scopically serrated. Hind part of head with many, irregular, small

wrinkles. Margin of labrum as a rule not protruding before curving

backwards into the side margin. Apical lunula usually more or less

flattened at base and only weakly sinuated on its inner edge, slightly

narrowing towards the suture 6

of hind femora blackish. about 5a Outer part rarely Elytron 3| as long as

wide M. carolina carolinaf. typica

5b Outer part ofhind femora, towards the knees, usually distinctly black-

ish. Elytron about as long as wide (PI. Vd-e)

M. carolina carolina f. occidentalis 113

6a Reddish dorsalarea well separated from the black spot. Elytron about

7 as long as wide

6b Reddish dorsal area fused with or superseded by the much larger

black spot. Elytron about 3| as long as wide (PI. VId)

M. sobrina infuscata

la Dorsal area as a rule yellowish-red, sometimes touching the black

spot. Black spot usually extended as far as about f sutural length (PI.

Vla-b) M. sobrina sobrina f. typica

lb Dorsal area purplish, sometimes touching the black spot. Black spot

usually extended as far as about f sutural length

M. sobrina sobrina f. antiguana

7c Dorsal area purplish, generally touching the black spot. Black spot

extended far usually as as } sutural length (PI. Va)

M sobrina sobrina f. bonaireana

Fig. 42. Labrum and mandibles of Megacephala sobrina sobrina: a-b, e-f from Margarita

(Pta Mangle); c-d, g-i from Bonaire (Dos Pos). - All specimens with 4, often somewhat

reduced setae. 114

Fig. 43. Labrum and mandibles of Megacephala sobrina sobrina: from Aruba (a-g Rooi

Bringamosa; h Salinja Master). - As a rule with 4, rarely with 6 setae.

Fig. 44. Labrum and mandibles of Megacephala sobrina sobrina: from St. Martin (a-f, h-i

Point Blanche - With 4 which be often reduced. Oyster Pond; g flat). setae may

Fig. 45. Labrum and mandibles of Megacephalasobrina sobrina: from St. Kitts (a Frigate

Bay) and Barbuda (b-e Low Pond). - Mostly with 4 setae. 115

Fig. 46. Labrum and mandibles ofMegacephala carolina carolina (a-b, f-h) and M. sobrina

infuscata(c-e, i-k): from Jamaica (a-b, f-g Hope River, h Halfway Tree) and Puerto Rico (c-

- of small extra tooth. 4 Setae e, i, k Saliña Corozo, j Sal. Papayo). Left mandible e showing a

which be somewhat reduced. may

Fig. 47. Labrum and mandibles of Megacephala acutipennis (a-b from Hispaniola), Af. affinis affinis (c-d from Trinidad), M. sobrina sobrina (e from St. Martin,ffrom Curaçao), M. carolina carolina (g from S. Carolina, h Georgia, i Texas and j Cuba). [From Studies 6,

1955]. 116

Fig. 48. Variation in dorsal design of right elytron in Megacephalasobrina sobrina typical form (a-d) and forma bonaireana (e-o): from Margarita (a Pta Piedras, b and d Pta Mos- quito, c Pta Mangle), Bonaire (e-h Dos Pos), Klein Bonaire (i), Curaçao (j, Boca Labadera),

and Aruba (k-n R. Bringamosa, o Sal. Master). 117

Fig. 49. Variation in dorsal design in Megacephala sobrina sobrina forma antiguana (a-h),

M. sobrina infuscata (i-l) and M. carolina carolina forma occidentalis (m-o): from Barbuda

(a-d Low Pond), St. Martin (e-h Oyster Pond), Puerto Rico (i-j Sal. Corozo, k-l Sal.

Papayo), and Jamaica (m-o Hope R.). 118

Fig. 50. Colour scheme of Megacephalasobrina sobrina f. typica (a—d) and f. bonaireana(e-

from - h): Margarita and Los Testigos (a-d), and Bonaire (e-h). Dorsal area of elytron a bright yellowish red (intypical form)or purple (inf. bonaireana);laterodorsal area a brilliant

green; lateral area a brilliant greenish blue. [From Studies 6, 1955]. 119

Fig. 51. Colour scheme of Megacephala sobrina sobrina f. antiguana(a-d) and M. sobrina infuscata (e—h): from Antigua, St. Barts and St. Martin (a-d), and St. Croix, Puerto Rico,

Hispaniola and Cuba. - Dorsal area of elytron in antiguana a bright yellowish purple, laterodorsal area a brilliant green; lateral area a brilliant greenish blue. Dorsal area in infuscata fused with, or possibly superseded by black area, the remnant a bright purple; laterodorsal area narrow, a brilliant green which may become dominated by the blue ofthe

lateral area. [From Studies 6, 1955]. 120

TABLE 5

MEASUREMENTS IN Megacephala, in mm

by F. VAN DER HEIDE, in which a few data obtained by the present author (x) are included.

Elytron Pronotum Head Body El. Pron. Head Body

length width 1. w. 1. w. 1. w. l./w. w./l. w./l. l./w.

M. sobrina sobrina

Aruba

cj 6 8.3-S.9- 9.3 2.55 3.25 3.55 2.65 3.9 15.0 5.05 3.5 1.1 1.45 2.95

9 9 8.3-9J-10.3 2.75 3.35 3.8 2.85 4.1 15.5 5.45 3.4 1.15 1.45 2.8

+ Aruba (R. Bringamosa)

98 — ____ 15.7 _ 3.5 l.i 1J5 3.1

Bonaire

9 12 8.9-9.5-9.9 2.8 3.5 3.95 2.9 4.25 16.0 5.65 3.4 1.15 1.45 2.8

+ Bonaire (Dos Pos)

9 3 9.6-9.9-10.0 2.85 3.7 4.0 3.0 4.2 16.6 5.7 3.5 1.1 1.4 2.9

Margarita

9 4 9.0-9.7-9.4 2.65 3.25 3.75 2.85 4.1 15.2 5.3 3.4 1.15 1.45 2.85

+ Margarita (Pta Piedras)

10 8.3-9./- 9.6 2.55 3.4 4.0 2.8 4.2 15.3 5.1 3.55 1.15 1.55 3.0

9 7 9.2-9.S-10.0 2.8 3.7 4.1 2.9 4.4 16.5 5.6 3.2 1.1 1.5 2.95

+ Barbuda (Low Pond) tJ 16 8.3-9.4-10.5 2.65 3.6 3.95 3.1 4.15 16.7 5.3 3.55 1.15 1.4 3.0

9 15 8.2-9.5-10.0 2.75 3.6 3.9 3.15 4.15 16.2 5.5 3.45 1.1 1.4 3.0

St. Martin

S 13 7.3-S.2-8.9 2.35 3.1 3.5 2.6 3.8 13.9 4.75 3.45 1.1 1.45 2.9

9 17 2.6 3.2 3.75 2.8 3.95 14.7 5.15 3.35 1.15 1.45 2.8

+ St. Martin (Oyster Pond)

$ 7 7.5-S.J-9.0 2.4 3.35 3.7 2.85 3.85 14.5 4.8 3.55 1.15 1.35 3.0

9 7 8.6-9.2-9.5 2.6 3.5 3.9 2.95 4.1 15.3 5.2 3.55 1.1 1.35 3.0

M. sobrina infuscata

Puerto Rico

cj 5 8.1-S.S-9.1 2.5 3.2 3.55 2.65 3.85 14.65 5.0 3.5 1.1 1.45 2.9

6 9 8.4-9.0-9.3 2.65 3.25 3.70 2.75 4.05 15.0 5J 3.4 1.15 1J 2.8

Puerto Rico (Sal. Corozo)

$ 4 8.2-9.4-9.8 2.2 — — — — 15.2 4.4 3.45 1.15 1.4 2.9

9 5 9.4-9.6-9.8 2.25 15.6 4.5 3.4 1.15 1.4 2.8

M. carolina carolina

Jamaica (dry spec.)

cJ 7 7.4-7.9-8.6 2.7 2.65 3.1 2.4 3.55 12.9 4.5 2.9 1.15 1J 2.9

9 9 8.0-SJ-8.7 3.0 2.75 3.3 2.5 3.7 13.55 4.85 2.75 1.2 1.5 2.8 121

TABLE 6

AVERAGED MEASUREMENTS IN MEGACEPHALA LEGS, in mm.

length of III II I

Elytron body fern. tib. tar. leg fem. tib. tar. leg fern. tib. tar. leg

M. sobrina sobrina

Aruba

9 4 9.6 16.4 5.8 5.5 6.25 17.3 4.75 4.3 4.55 13.6 4.0 3.0 3.45 10.5

Curacao eJ 1 9.0 15.3 5.7 5.6 6.2 17.3 4.8 4.2 4.6 73.(5 3.9 3.0 3.7 10.6

Bonaire

9 5 9.9 76.5 6.05 5.75 6.3 18.1 4.9 4.55 4.6 14.1 4.05 3.05 3.15 70.2

Margarita a* 5 9.25 A5.7 5.85 5.7 6.4 17.9 4.9 4.45 4.7 14.0 4.1 3.15 3.75 11.0

9 5 9.8 76.5 6.2 5.85 6.25 18.3 5.0 4.3 4.45 13.7 4.0 3.05 3.5 10.5

Barbuda

S 8 9.2 15.6 5.65 5.55 6.45 17.7 4.65 4.2 4.6 13.4 3.9 2.9 3.5 10.3

9 8 9.25 15.6 5.35 5.15 5.8 /<5.5 4.45 3.95 4.2 12.5 3.8 2.8 3.2 9.8

St. Martin

3 5 8.45 /4.5 5.35 5.2 6.15 16.7 4.3 4.0 4.3 12.6 3.85 2.95 3.45 10.3

9 5 8.95 /5.2 5.7 5.3 6.1 77./ 4.55 4.1 4.25 12.9 3.85 2.9 3.25 10.0

M. sobrina infuscata

Puerto Rico

S 2 8.65 14.6 5.4 5.15 6.0 76.5 4.45 3.9 4.4 12.8 3.75 2.95 3.35 10.1

9 2 9.3 15.8 5.85 5.4 6.0 17.2 4.55 4.05 4.2 72.7 3.9 2.9 3.1 9.9

M. sobrina

Suriname (dry spec.) tJ 3 9.1 15.2 6.0 5.7 6.35 18.0 4.95 4.55 4.6 14.1 4.15 3.2 3.9 11.2

5 3 9.2 /5J 6.3 5.7 6.15 17.9 4.6 4.2 4.35 13.2 4.0 3.0 3.2 10.3

M. carolina carolina

Cuba (dry spec.)

S 2 9.4 ISJ 6.0 5.95 6.0 17.8 4.7 4.45 4.45 13.6 4.0 3.15 3.25 10.4

9 4 9.6 16.0 5.8 5.3 5.8 /<5.# 4.7 4.2 4.3 13.2 3.85 2.9 3.2 10.0

Florida (dry spec.) c? 2 9.7 /ff.0 5.85 5.65 6.4 17.9 4.8 4.45 4.6 13.9 4.05 3.05 3.5 10.6

Carolina (dry spec.)

9 3 10.0 17.0 6.15 5.75 5.9 17.7 4.75 4.45 4.45 13.7 4.0 3.1 3.4 10.6

Oaxaca (dry spec.)

6" 3 9.7 16.0 6.0 5.9 6.2 18.1 4.9 4.6 4.8 14.4 4.1 3.2 4.0 11.3

M. acutipennis

Hispaniola (dry spec.) 8.9 S 3 7.3 13.0 4.35 4.15 4.5 M.0 3.85 3.45 3.7 11.0 3.25 2.55 3.05

$3 8.1 14.0 4.85 4.25 4.35 HJ 3.9 3.5 3.25 10.6 3.25 2.55 2.55 8.4

M. affinis affinis

Colombia & Venezuela (dry spec.) 13.3 3.8 3.0 4.0 10.8 3 2 9.6 16.0 5.3 5.3 5.8 15.6 4.4 4.3 4.6

9 3 10.3 17.0 5.9 5.7 6.1 17.8 4.6 4.4 4.3 13.2 4.0 2.9 3.35 70.5 122

TABLE 7

AVERAGED MEASUREMENTS IN MEGACEPHALA LEGS

expressed in percentages of the elytron length

Elytron III II I - length

in fern. mm tib. tar. leg fern. tib. tar. leg fern. tib. tar. leg

M. sobrina sobrina

Aruba

9 4 9.6 60.3 57.4 65.0 180 49.4 44.7 47.3 /« 42.6 31.2 35.9 109

Curacao

6* 1 9.0 63.0 62.0 69.0 /92 53.0 47.0 51.0 151 43.0 33.0 41.0 118

Bonaire

c? 5 9.6 58.1 62.0 72.2 203 52.4 48.9 51.6 /5j 44.1 33.3 39.9 117

9 5 9.9 61.1 58.1 63.6 183 49.4 46.1 46.2 M2 40.9 30.8 31.8 103

Margarita

6" 5 9.25 63.1 61.3 68.9 /9i 52.9 48.2 50.1 152 44.3 34.0 40.3 //«

9 5 9.8 63.3 59.5 63.5 186 50.9 44.0 45.4 140 40.7 31.2 35.6 /07

Barbuda

o* 8 9.2 61.7 60.5 70.0 192 50.1 45.7 49.8 146 42.2 31.6 38.2 112

9 8 9.25 58.1 55.7 62.6 179 48.1 42.7 45.4 136 40.8 30.5 34.6 106

St. Martin

o* 5 8.45 63.3 61.8 72.9 /9« 51.2 47.2 51.2 150 45.5 34.8 41.0 121

9 5 8.95 63.8 59.5 68.0 /9/ 51.0 45.9 47.9 145 43.0 32.4 36.2 ///

M. sobrina infuscata

Puerto Rico

cJ 2 S.65 62.5 59.8 69.2 192 51.5 45.0 50.5 148 43.5 33.9 38.8 116

9 2 9J 62.1 58.3 64.9 187 49.5 43.7 45.0 138 42.5 31.4 33.8 107

M. sobrina

Suriname (dry spec.)

6" 3 9.1 66.2 63.0 70.0 199 54.4 50.0 50.8 /56 45.7 35.3 43.0 123

9 3 9.2 68.7 62.2 67.0 195 50.3 45.9 47.2 M5 44.5 32.7 34.9 112

M. carolina carolina

Cuba (dry spec.)

o* 2 9.4 63.7 63.2 63.7 189 49.9 47.5 47.5 145 42.5 33.5 34.6 ///

9 4 9.6 59.8 54.9 59.9 175 48.8 43.3 44.4 136 40.0 30.1 33.1 103

Florida (dry spec.)

o* 2 9.7 60.3 58.2 66.0 185 49.4 45.8 47.3 143 41.8 31.3 36.1 /OP

Carolina (dry spec.)

9 3 10.0 61.8 57.5 59.2 177 47.8 44.8 44.8 137 40.1 31.1 34.1 106

Oaxaca (dry spec.)

S 3 9.7 62.0 61.0 64.2 187 50.7 47.6 49.7 149 42A 33.1 41.4 117

M. acutipennis

Hispaniola (dry spec.) 122

M. affinis affinis

Colombia & Venezuela 113 o» 2 9.6 56.0 52.1 61.3 173 46.1 45.0 48.2 139 39.8 31.4 41.9 102 9 3 /0J 57.2 55.1 59.7 173 44.8 41.9 42.2 128 38.9 28.2 33.0 123

DISTRIBUTION

When studying the geographical distributionof Cicindelids in the West

Indies, LENG & MUTCHLER (1916) produced a table which "indicates a

of the Cicindela from the United dispersal genus states to Cuba, Porto

Rico and Hispaniola, with a decreasing number of species as we proceed

southward; accompanied, however, by a dispersal of one species (sutur- alis) in the opposite direction, from South America northward. The

southward dispersal apparently ceases at Guadeloupe, the northward reaches Porto Rico; but additional data may alter these statements." In

this table- see below - attention is paid to the West Indian - Floridian

and the isolated of C. in Guade- relationship to occurrence argentata

loupe, a species which in many varieties abounds in South America.

Florida trifasciata marginata & 17 more species

Bahamas trifasciata marginata

Cuba trifasciata marginata boops & 4 more species

Grand Cayman trifasciata

Jamaica trifasciata

Hispaniola trifasciata boops suturalis

Porto Rico trifasciata boops suturalis

St. Thomas trifasciata suturalis

St. Barts suturalis

Barbuda trifasciata suturalis

Antigua trifasciata suturalis

suturalis Guadeloupe trifasciata argentata

Martinique suturalis

St. Vincent suturalis

Barbados suturalis

Grenada suturalis

Table 8 establish that the has Comparing we may general picture not

changed considerably after sixty years.

Our knowledge ofthe Virgin Islands fauna has increased, and Jamaica

has got two more species, including a not uncommon one (C. carthagena jamaicana) only recently described and possibly also to be expected in other of parts the Greater Antilles. We may now consider C. boops as a

characteristic species ofthe Greater Antilles, also occurring on the Virgin

Islands and the Caicos & Turks Islands north of Hispaniola.

Cicindela trifasciata and C. suturalis remained the commonest two

the first one shows distributional south of Guade- species, though a gap 124

loupe and the second one has not (yet?) been reported from Jamaica,

Cuba and the Bahamas. C. trifasciata generally occurs on mud flats,

of salt and along the margin ponds, near shallow mangrove lagoons (its occurrence along the border of an artificial freshwater lake at Mona,

Jamaica, was quite a surprise); C. suturalis, on the contrary, prefers sandy beaches. In all four cases in which C. suturalis suturalis and C. trifasciata trifasciata were collected in the same locality, this habitat-preference could be observed, while in a few cases when single individuals were found in a "wrong" habitat, this clearly was by accident.

Though subspecific categories in C. suturalis are justified by differences in elytral design, there remain cases of doubt, making interpretation of

in literature data as given often uncertain. In the present material, the

of hebraea from Rico looked with specimens Puerto may be at some suspicion. Further study of the C. suturalis-complex will possibly yield several new and interesting taxonomic data with regard to its Antillean representatives.

Generally speaking, the impression was obtained that typical suturalis

(with a strongly dominant yellowish-white design) prefers white-sandy beaches, while the subspecies hebraea (with a yellowish-white tortuous middle band and hooked lunula) occurs on sandy beaches which are coloured by mud, or slightly polluted. In the subspecies guadeloupensis the same dark elytral design is still more developed, resembling that of c.

trifasciata trifasciata.

LENG & observed that "hebraea and MUTCHIJER (1916, p. 694) already suturalis have never been found together. C. trifasciata has, however, been found on the same beach little further back from the sea." running ... a

On Tobago, trifasciata trifasciata and suturalis hebraea have been collec- ted in the same locality, on dark sand.

The identificationof a Jamaican cicindelid, not uncommon in coastal and riverine habitats, as a subspecies ofa well-known continentalspecies,

C. carthagena, was rather a surprise. C. carthagena jamaicana was found

sandbar at the entrance ofGreat on a Saltpond, near mangroves, among several C. trifasciata trifasciata. This locality is situated quite near the salinaof Port Henderson in which R. P. BENGRY found C. boops, a species which preferably occurs in saline muddy areas. LENG & MUTCHLER (1916,

of "Alkali flat p. 691) give a picture an at SantaRita, Porto Rico", which 125

shows a low muddy section in which C. boops occurred, while C. trifas- ciata was found alongside a small brook.

In the most northern part of the Caribbean, C. marginata inhabits

well sandy beaches as as marshy coastal areas.

Following those authors who consider the West Indies ceasing to exist beyond the island of Grenada, LENG & MUTCHLER ignore Trinidad &

Tobago, and all islands of the Leeward Group, including those (west of

Margarita) the faunaof which has a remarkable, endemic character. With regard to the cicindelids, however, the species obtained do not differfrom those occurring on the adjacent mainland.

From Trinidad & Tobago C. trifasciata trifasciata and C. suturalis hebraea are recorded from haline coastal areas, and C. argentata from non-haline inland localities.

On the islands of the Leeward Group (west of Trinidad) C. trifasciata

to appears be replaced by C. auraria, and C. suturalis by C. graphiptera.

However, in Margarita C. auraria occurs together with C. trifasciata trifasciata. Cicindela graphiptera fulgidiceps has not been recorded from

Curasao, Aruba and Bonaire, though it is known from the shores of the opposite mainland, where it has been found together with C. auraria (cf.

JONGE POERINK 1953, p. 139, fig. 27).

Recent finds of C. suturalis hebraea and C. argentata on Curasao (not recorded of before), once more illustrate our still scanty knowledge the beetle fauna of this small and densely populated island.

- The ofdistribution in in Table - pattern Megacephala as expressed 8 is taking shape only ifwe are subdividing the species Megacephala sobrina into few a lower taxonomical categories, difficult to distinguish and almost unrecognizable among the profusion of comparable mainland races withoutknowing an exact find-spot. In this way we may distinguish a subspecies infuscata inhabiting the Greater Antilles and Virgin Islands; a forma antiguana occurring on the northern islands of the Windward

Group as far south as Antigua; a forma bonaireana on the Netherlands

Antillean islands of the Leeward Group, and a typical form from the

Venezuelan islands of Margarita and Los Testigos.

The occurrence of Megacephala carolina and M. acutipennis on the

Greater Antilles clearly points to a Floridianrelationship, while M. affinis 126

continental fauna element which represents a may prove to be more diversified after a more thorough research of the non-coastal island areas.

Hiding places of the nocturnal Megacephala sobrina were found in

fresh brackish-water various habitats, near or wells, in gardens, pastures with cow faeces, and on muddy areas near lagoons and saltlakes. TABLE 8 127

GEOGRAPHICAL DISTRIBUTION OF THE CICINDELIDS

treated in this paper occidentalis antiguana typica bonaireana f. f. jamaicana ascendens trifasciata fulgidiceps f. f. suturalis hebraea guadeloupensis carolina infuscata sobrina media sobrina sobrina affinis gracilis Cicindela dorsalis marginata carthagena boops trifasciata trifasciata suturalis suturalis suturalis argentata auraria graphiptera Megacephala carolina acutipennis sobrina sobrina sobrina sobrina affinis affinis

Southeast USA O O x ?

Bahamas x x

Cuba xxx xxx

Cayman Islands x x ?

Jamaica O x O x

Hispaniola xxxxx xx? ?

Puerto Rico O O O O

St. Thomas x O

St. John

Tortola

Anegada x x x

St. Croix x x

Anguilla O

St. Martin Ox ? O

St. Barts x x ? x

St. Kitts O O

Barbuda O O O

O O Antigua x

Guadeloupe x x

La Desirade

Martinique

St. Vincent x

Barbados x

Grenada

Tobago O O

Ox Trinidad x

Patos x

Los Testigos O

Margarita O O O O

Cubagua O

Tortuga O

Bonaire O O

Klein Bonaire O O

Curasao x x O Ox

Aruba O O

S. Amer. mainland x xxxOO ??xx

O according to material collected by the author

x according to specimens studied from other sources

—from literature.

? doubtful record. 128

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PLATE I

Cicindela - a suturalis hebraea from Brasil, ￿. b C. graphipterafulgidiceps from La Goajira,

Colombia, ￿.- cC. auraria from Aruba, ￿. - d C. trifasciata trifasciata from St. Martin, ￿. 131/II

PLATE II

Cicindela graphiptera graphiptera from Costa Rica, ￿. - b C. boops. - c C. trifasciata ascendens from U.S.A., ￿. - d C. suturalis guadeloupensis, ￿. - e C. suturalis suturalis, ￿. 132/III

PLATE III

a Cicindela carthagenajamaicanafrom Jamaica,Saltpond, ￿ [7.2 mm el.l.]. - b C. carthagena jama icana from Jamaica,paratype ￿, Mus. Amsterdam [6.9 mm el.l.]. - c Same paratype. - d

C. carthagena from Mexico, Playa Camaron, ￿, Mus. Amsterdam [7.4 mm el.l.]. 133/IV

PLATE IV a Cicindela marginatafrom Key Biscayne. Fla., ￿ [7.0 mm el.l.]. - b C. dorsalis media from

South Carolina,Beaufort, ￿ [7.7 mm el.l.]. - c C. suturalis hebraea from Puerto Rico, Playa

Mediania Alta, ￿. - dC. suturalis hebraea from Trinidad, Los Gallos Point, ￿. - e C. sut.

- f from Rio Mus. Amster- hebraea from same locality, ￿. C. argentata Curaçao, Canario,￿,

- variation dam [4.9 mm el.l.]. g-i C. suturalis suturalis from Antigua, Deep Bay, showing of

elytral design in one sample [5.2, 5.1 and 5.5 mm in length]. 134/V

PLATE V a Megacephala sobrina sobrina f. bonaireana from Curaçao, ￿. - b M. affinis affinis from - Curaçao, ￿. - c M. acutipennis from Hispaniola, Haiti, ￿. d-e M. carolina carolina f.

occidentalis from Cuba,￿ & ￿. 135/VI

PLATE VI a-b Megacephalasobrina sobrina, typical form, from Margarita, ￿ & ￿. c M. sobrina sobrina f. antiguana from St. Martin, ￿. - d M. sobrina infuscata from Puerto Rico, ￿.