'DJtsoddo dsqnsowauoqluorE prreDa.raalnd'dsqns oopunu T '(DSouISnspspwDLrr^) sorJcdsqnso,,'\l tlllt\\ DttD^ g'optwlt g.'suaJSatl|pueDfisnd'ouuac'ddsqns 'suuqns 'ourocrstods 'osstwalaotd '(Daiqdrod Dtsn7uoqnsg 7 g g pue sumrr,rrld) sarcedsqns o,rl qlr,lt J'r1t1rJ?.ltldg '(suaJof-tr n pse oll[qdoaeoqd) sarcadsqnsor'l tllrt* o11&qdouaoqdg 'opunlt.t8tu g'DtuaLlJsarJnuE'D.arqdourypa fl'sqoryaw fl'Dloa]nl g'Dpt^ n'D KqdolsryX 'Dt!o{tpqaq g'slsuednqbua^u dsqnsttanpnS g'(otaJo.tdnrpreDsuap)selcedsqns o,n1q1r,n zslap g 'Dtodst.tr g'Dlou^t1c n'(upaltQod puersoiltrlrr) sercadsqnso,n1q1r,n rsolldzJ A'DJJa D'dsqns DaDutlJD.tofl'otrpqD to.rseun luu or.llroJ clcq poqulscpserredsqns puu sercedsArou Joqlo

'{uBr crJrcadsol potE^ole eleq nxpt .roJscuruu ,rcu ,{iusscf,cLrc.ru (,{1c1u1g o.utw{xo sttr'DJunpal g =) reddoH ry ro{ooJg 'Je^ oprtop[ 7 pue ('q)uag DtoltldouDpu.t DJunpat g =).reddog ry :elooJ1DaDaqJon A ,{Io{BIS 'Gf DlstiSuDqnsr?,i DJtolpd.r .loJ uorluurquoc .rcu s sr JcddoH T Je{oolg (dlo{glg) olsnSuoqns g ',{tclulg 'n 'sasuapuowsaO ? Lrxptp,trNstsuapuout^trp sorJedselSrns erp.;o Surlsrsuor seuesqnspue '(1uu: crlrcedso1 pcsr?i e.roq sraqto eerql puu pcqr:csop,4y,tr.cu or? qJrq,r Jo 0Z) scrJcds,Z EursuduroJ 'azLundsl^a7 sxttosqus- aDLu.tadsua'js rossn4f,pJnfl ur pcqsrlqulscclc sorJesqns,rou o.\{J

'qJlucscrurnrjeqroq pJBpuels puu'csnoqssqE cql ur pcsrsrsBurlpees 'serpnls plcrJuo pcspq sr uorsrAcJ srql 'rent]qrS,Ju pat g t'lsl\fttlx-^sn popnlJur flsnorAJ:d srclluru uouuoc 'fl)lcl;pl.:uqntq '(dlo1,r:lgoopup/r\sntdtlrrrg) lsro^cspuu'(uoprut{ Ltaup.toS 3ur)tulscql urnBelrq,rro 'lloqlncqr'd oopuu,,'lSurcnpo.rd-,{cuoq p uu :c(1urr1 luulodtur cq1cpnlcur puB uErltrISnVurolso.&\ oqlJo scoltlrrupuu slclllrru 'soc.rldunru csuduo t aow.tadsuaT crtrT posrAer sr ort?r'1snv uJclsod! ur Jrruopuo '(I66I) 11uscrccds gglo scr:cslcurlsrp ,(l1ucrur oLtoxel c'aDLutadst ta1 sortcssuldtlntng 6SI-I :(I)S 'ullsJtsnV ?rsf ,(nN urq so16ur (oucceg.{1,r,1- uxptctrN)Dwrads^aTsouos sn (pcnn) sorJcdspclllspu? '.rcddo11pue DJunpa.tA' oopu Dtt4.rr/r/,{/irJrtJouorsr^cl3r ruouoxol V q ucqdclg H I'tr{'ta{oorg

lJDJlsqv

'oorruuPrl 9909 "rlDltsnv rrrls.li\ I9 Yog o d '.4u.:) rtr[cs.t{ cJrtplri\Aurr|r]snv ru.tsry' lu3(u.3euBr\tpuB'I puI, uonrN.suo3 Jo tucrxusd.( z '€jrcquEJ '009I 'OdtSJ 'urnuEqrrH l09Z trroti.r.r.J I"lrdEJ uDrt"rlsnv xo{ O d tluorllN uerrerlsnV r

'H "reddo116 ueqdel5prru rre{oorg I'I I

BllsJtsnv uralsaM u! (aeaJe ,{I{ - uapw\N aDuradsutaTsaltas snldtlncng) 'DJunparg 'oopunL salJadspalllB puB sntdqDJng lo uoJsl^al JlruouoxBl y

(r66r)68r-r ,(rs "'sr{nN NuyrsiaVol.8, No. I (1991)

Introduction

atractiveness Theeucalypts of WesternAustralia are renowned for boththeir diversity and their in colourtreatments asornamentai;arden and street hees. Their popular appeal is reflected 1979; gi;inunu-;"tof recentbooks(e'g. Kelly 1969,1t78; Chippendale 19?3;Wrigley andFagg ilolliclay andWatton 1980; Elliot andJones 1986; Brooker and Kleinig 1990)'

eucalypts Despitethe burgeoning popular literature devoted to them,many Western Australian insufficient are diff]cult to iclentify.Because of the existenceof many undescribedtaxa and work andmany knowledgeof morphologicalvariation in somegroups, most availableteys fail to Chippendale Lotanicaidescriptionsare inadequate (e.g' Blackall and Grieve 1954; Blakely 1965; and the isi:, rsss). This applieseuen ior sonri of the mostcommon mallees of the wheatbelt goldfields.

of the C.A. Gardner,{he Governmcnt Botanist from 1928to 1960,proposed a full treatment volumewas State'seucalypts in thesecond volume of theFlora of WesternAustralia However'this neverpublished.Aseriesofleaf]etsonwesternAuStralialleucalyptswaspublishedbyGardnerfrom butthe lg51-ig66. Thesewere subsequcntly published posthumously in bookform (Gardner1979), work coveredless than half tnespecies now knownin theflora and provided no keysor iaxonomic overview.

PryorandJohnson(1971)andChippenclale(1988)mademajorcontributionsi|rheatmentsofall eucaliptsthen known, placing named Westem Australian taxa in a broadertaxonomic context' However,bothworks are now considerably outdated as they omit many recently recognisedWestern Ausiraliansperics (cf. Brookerand Kleinig 1990)'

wehavecompletedtaxonomic research on manywestem Australian eucalypts ovel thepasttwo 1978; decades(Brooker 1g12, 1973, 1g14, 1976,1979, 1981, 1986' 1988; Brooker and Blaxell 1989; Brookerand Done 1986;Brooker and Edgecombe 1986; Brooker and Hopper 1982' 1986' Hopper1982).

we arecurrently aware of the existcnceof over 100undescribed taxa in westeln Austlalian eucalyptS.Thosenottreatedherewillbedescriberlinforthcomingpublicationsbyourselves,by f. Craytingand Brooker, or by L.A.S. Johnsonand colleagues of the New SouthWales National tterbariumle.g.Johnson andHill 1991;Hill andJohnson1991). Many ofthese undescribed taxa were illustratedin the recentlypublished field guideof Brookerand Kleinig (1990)'

The major unresolvedtaxonomic problems in westem Austlalian eucalyptsa.re found in " Symphyoiyrtul',the largestinformal subgenus of hyor andJohnson (1971), and to which theseries and tiriiptr^a, belongs. @rom hereoninformal taxa of Pryor and Johnsonand of Johnson colleagues(e.g. Johirson and Hill 1991),not formallypublished under the rules of theIntemational is Code of BotanicalNomenclatule, are refened to within invertedcommas.) This subgenus chalacterisedbytwoopelcula(|heouterofwhichinmostspeciesdehiscesearlyinbuddevelopment more andleaves apersistentnecrotic scarat thejunction ofthe inneroperculum and the ), thantwo veiticalrows of ovulesper placenta, unbranched axillary inflorescences, or if apparently comooundand terminal then the anthers are adnate. 'palsralro (orzrpr.4 rtr"q ruo(l'pnq u^1or8ltnJ trF"a ut ts€elt"'rurod eql'aqnl 'tprruor -y,{ter eqlrJ"qiaroLupuuBuotsr rrr^r1'Sururqs'snorruuor rlrH^\E urntnrrrdo:t"3upultfc aqnt-xtlBr :?uotqrur Boq€ 'p.8pa o^\1'p"u"u"Usatronprd'parrdotIZI-9'Ae[rx€roI alpl rtaqun:(rnolor luaralrp eJourSr€uleqlpue qup(uaql) $prs 'otEullunru 'a]BIoeruRI'al"uralle 'priu ',{qqruqs,, qtoq uo snornEIS i tollad € olul pardojl?u1?rqe Itnbrul l9q^\auos se^eal

:s,$olloJsB ue^rB sr l€ur8uo oqllo (8I61) ueprel,qdq peqsrlqnduon?lsuer1 V

's9u0s ,rurouoxsla,t.rlf,rnlsrp uol Suriuoyeqsu uorlruSoce::rcq1 puc rxel snouEAeql Jo luerulee4 luanbosqns 'g eql SuueprsuocuitlNr Drtnpa.t jo enSololordaq1 uodn 1p'rp ol lnJesnsr l[ pue uoqueluocur 'ocurs lcq,rourosuooq seq ou.ros Jo snte1s eqI poqsrJqnduoo qe^e\DJunp g Joserleuu,r pue serceds pelclerIuro^eS tt8I rlr.,cBuclssreld octuBld,, s, uuBurqo-I ur reneq3S dq poqsgqnd.sum'arwadsb.aT 'ns swd(1otngoqlut senadspeumu lsrg aql'nunpaJsntdtlDtng ocunpatglo anSoplotdaq1

lJols-rHJ!urouoxsJ

'(066I'1, l, rcddoH)B.roljl pcrcAucpuA pcr?lJcC Jo tsll s,clstSeql uo tueus:u1drraql Eurpnlcur 'sc,rrlsrllurlucueSnuctu puB uouu^JcsuoJ olslrllJBJ ol scruBuoJrnbor pw'elqDreuln,r ro peroSuepuc 'ejl]r 'tL6I eq,{f,Luuxrl pcqlr3scpuncsoql Jo roqunue oslv (886I olepuoddrqJ:I16I uosuqoJ pu?lo,{Jd:996I .(lc{eJg:St6I rcupr?g)sroqlnn snor^crd ,{q scrcods c^rJ u?ql croru ou ol pqnqrr € uorq pnqlnql [xEl pcqu3scpunJo,{uJls clqelrcuo] ? urc'luocol pcsruSoJoJ,rou sr lr osngJoqpu€ 'sercods '(s:clceruqc lucuodtrr .{lleJrurouoJc sureluoc 1r pnqpuu pcas .{q fllnrcsdso) ,{lJsorSoloqdloru pcquJsun3jrc.{1ruclcsr ll 'op8Jops re^o pclclduoj cq plnoJ s,{o^rnsplcq e,rrsueqorduroc croq,rerlullsnv ulclso,r-qlnos ol 3ruropucsr 1rcsn?Jcq uorsr^or roJ uosoqss?,r sorJosaqJ

'(sror.lto pre D$nSuDqISg'Dptultt 1'DllttldouaDqdg'Dpt^DL[ n'DaDuLlcoto g 8t'JpnltuoJcq) Jeneqos D3unpatg suol px.r.lo.lcJ,{lsnorAard soollnru uoruruoJ lf,]o^os puD (uoprBJ,'{ r.{rrp.rz3 g)1e11euenlq '(oopuoft 'llcqlecq,r n) wrfl a1rq,tro oopue,nBurpntsur uorprlsnVuJctsold oql Jo saalpu pu?scoll '( u^\orlIJscqoqt Jo curos srrr 0fLroJ soucs slqJ I l,6I uosuqolpu? ro^Jd.. aD)unpaU,,sel.Jas snJdtlDJtrfl 'atoH lsrnJoJrrr=) tnprcl,NaDLu.ladst\al $uxs sn4tpJna'Jo uorsr^crIlnJ E o{suopun o,r

'(I66I IlrH pueuosuqol),?ptDpuuv,,puc (,,sarq pS,, =),{lo{el8 aDJ.toluoJ'txsg puB'(686 [ roddoHpuu JaIooJg) IoddoHZ' rc{ooJgsrirrFrU ros A'(086IueCpu? 'QzDuDtuuDwqa1')cs'a'( '616I rrBS)reJ s 4'JJ0J t 86I r0)oorg)rali]]rgsa,ll,fl^o l3s g'(gg6l Je).oorg',,aDpunJaoll,,=)qluiLa1DtltuuoJ ras n uo s].ro,rpoqsrTqnd ,(tsnor,lord ur peldopesu,tr. qceo:ddusrq; Ic^olsoucsqns Jo souos oql lB sorpnls,{reuorsr,ter f,q epeureq osleIIr^\ suo[nquluoJ '(/86I '986I 't86I luBrrJruSrslcqt roprsuor o,r'ourrl €rrss aql tV lr.,asa6rpB.I) slddtBJouour eql '(€86 urqlr,nsdnorS lo pnB t scurldunHpw saSrpE-I),,?Jcuc8qns,,lddprne IIe go serpnls ur peldopz ',{Jrnbuo ,tlluoJorrlJeoJddp pJuelunu pell?lopoql .{q uow8qso^ur8ur^rosop Jo splegSurSualpqc ol? sor.rospu? suorl3ososeql Jo rrollnlo^apuu sdrqsuorlele:eqJ (116I) uosuqolpuB rofud dq seuJs0t prresuorlJcs , ur,{lpurloJurpocEld cro,r,,snuaEqns,, srrllJo sJaquelu u?{prlsnv tlJelso,lv!

'slarql)u?lLsa "qluJg '\anIN'gDtpupJJow o{ g purez,xJnlryntdDrtal !, xc fl'rm)T ueJ otDLnwnatuoJ t'uuurpeals T, ,{leltzlgDUuDpo\.tg' looHDdrDcorJowg' VluxgDrsaDlg'A'e 's.ro,\\oU 'A) InJJnolo33S.IBIqlr,r slstueuuulou,rou)l -lla,r du?urpuu'(.(1a1e1g ooprr,1.r oopu?.r'(J( .g) '(uaptetrq '( 'd t)lDydarorldwoS ;run1 ra1tol1u3g) aftug lt'olle. llantr^l lol octs.taqpswdtpcng) urel sBsee-rl Jeqrr-Ill uplpIlsnv rrJelsoldluErodurr,(ylecrurouoJe qrns sepnlJur, ,snufuuottldu^S,,

a"erE!,4m - uapret{,rlarads tvt s.uas s tdqDrng'raddoH O uaqdars2y re{oorg HII/{ NuytsiaVol. 8, No. I (1991)

IninteriordistrictsbetweenKingoeorge,ssoundandYofk,inthemonthofoctoberwilhneaflyadultbuds(tlertr.Preiss. and 24?' see-FiSure 96); in grav€llysterile land on the hill No.234); Februaryand March wrtx youngi"at Ol"t Z+S 95)' K;;;;;;p, "*.t"p"Riche' fruitins in November(No 232' seeFigure

Ashrubafathomhigh'branchletsangled,withthepetioleandlhemarginandtheriboftheleavesr€ddish-yellow.Theleaves Uuicontinuouswilh it andthe same length, an inch long)' 3_4inches long,6_ l2lin", U.o"O.C"ry*+oo""i*itt it'" faicet ofthe :-celled Fruitsumbellate' oblong-clavate' with the measuringabout 3 linesin treua*, opercor"uiri-i ii'i"s long'cupsule " valveserec!' touching the mouth of thecalyx{ube pdicel 6-?lines long, above 21, lines broact' t[rncate; included'

arelikely to be involved'particulady With hindsight,it is not hardto infer thatseveral entities "between localityas King rvnenine .peciJ, is basedon four syntypes'three with such.animprec^ise informationgiven concernsthe C"org";, Sou"Oand York". fne onty Oistinctiuemorphological "redunco" Wereferto theseobservations "ulro"*fy"-i.of operculumwith thetip (i'e'iumeddown)' below.

Sllbsequent|reatmentsofE.reduncaandthediscoveryofrelatedspecies.Bentham(1867)gave whilerefering to oldfield and considerableattent ion to E. redunca.He begansomewhat obscurely- "tn form' a shrubor smallree with Maxwell collections(i'e. not thetypes) by s'tating, ttreoriginal "Shrubby" "shrub" for the Preisstype a smoothwhite bar*" when the i,oiotogoe states and to wandoo'yet he separatelydistinguished this in collections.The treetbrm seemsto be ana'ilusion "large yat. he describedas a tree". ii, i"ilr*i,,g list of va,rietiesas E . redunca e/arawhich

var' melanophloiaBenth' Two morevaricties were published by Bentham,namely' E redunca "Leaves more prominentlyveined' - which is diagnoseclby larger-a (i'e than 3-4 inches.long)' bark"' andE reduncavar' Murchisonaird SouttrHutt rivers, small tree with smoothblack protologueof whichTurczanin-ow angustifolia,A sytrorymfor E. xanthonenaTnrcz'(1847) in the "w. is a Drummondspecimen ("3rd gi""., ,i," "'itt"ri,ru iocality of Australia"' However'this Riche(Erickson 1969)' The collection")and must have been collected between Bolgart and Cape distinctive'e'g' Ieaves o.GinuiO.r.tiption ofE xanthonemaincludes features that are leasonably ..narrow-linearlanceolate,',peduncles finally..very much deflected',,..Filaments remarkablefortheir "habitussingularis". Deflected reddishorange colour", and the quiteambiguoui but unqualified speciesas we know it and are not evidentin the tvpe ;;;;;;;t ar"enot a characteristicof this specimens.

Bentham,splacemelltofE.reduncainhiscomprehensiveclassificationisofnosignificance gto Eucalyptusseies N ormnles today,other tha; hisrecognition of thespecies as be6ngin thelarye of theNoflMl esinto Benth.which is diagnosedby characteristicsofthe stamens' Thefurtherdivision Muell E asperaF'Muetl" a subseriesto includ eE.reduncawilh E. patensBenth.,E.diversicolorF. " to Pryorand Johnson andE. phoeticea1.Muell., each belonging to a clifferentsubgenus according (1971)indicates the heterogeneity of this subseries(ftclzsae)'

by the 1870smust have Mueller'slater ftcatment (1879) of E. '"edunca cortibnted little to what E mntho,nema-wil\ beena groupofspecies frequently encountered in thefiekl' Muellersynonymised wandoo)which colld be E. redunca andreferredto E. redunca pincipally as a tree (presumably observationthat the reducedto sh b tbrm overconstderabl; stret;hes of"poor ground"' Mueller's whitecolourationoftho..balkcomesoff...... onfriction''probablydoesnotrefertowandoobutmost in thearea where wandoo would l\kelyto E . acc edensw. v . Fitzg., the dom inant powder-bark species lroqs .{lJ^rtul.r sq1- ,{1c1u1gaDsowno se\tes sudQoJnE Ltr sfuapuowsap g pccctd flcTulg 'ssuapuowsap .!I pLwD)rlrlpat gjo drqsuo4rlcr csolc eq1Sunr:eruor reupreg;o elonb s.,{1e1e1g punueprrl,,q clrdsc6 slucurclrJlnururBls eqlJo uorxauurlcrt:ud oql pu?.,spnqpcduqs-uroq,, aql ere sor:oss.,{1a1e1g ro1 srsouSBrpcql ul scrnlucglttu:tltu8ts eq1 f,luot:d seq qctq \ aDundsaaT sel.J.as 'aoJnu.roJqng 'zr^ 'seues s td|lD)trg s,ucpfuJr'\lol oruereJer lnoq:J^\ scrtossntdf,pcnA Jltuouox?l clc.rJsrp? olur scrjads lm^clor cql Jo rcqunu e dno:? ol :oqlne lsrrJ oql s?.,|{(lg6t) ftslets

'([Z6I) s]aqnrcJo uofetrJlsslrlo slq ul pal?erl sD,^( peu:acuocscrccds cqlJocuoN sSurlpeesJouortf,JrJrssslJ srtl ut pcpnlxurloxem ssuapuousap fl pue DwauoLlltlDxg ..aDpastguorsl^rp ?,, leurolut e8rul xr\'lur uaupnS A p[e Dpp're^ 'DJuttp 'I 'cauonbcs 'suopel,{loJ otunpal n ).r poJutdcq uor e scpue .,pelJasrq,,cql zi{'snuc8cqluI 'uonu3lJrss[[3 scrccds,{uuur ol uoururoJrolauJllqJ B ol ccucututo:do.teB (196I ) uopIuI,{ JalB[3 uI

'q3neso.r srqJo .3nlssrqlloscrJJdso,rtcscrltJopcos,rnsaqt[ql.{lclrluns!l!(tz6 l pol]oltoJlsrJ) s!suapuowsapg 'uoll?rrJlsselJ pu?(ZZ6l pclJollo3 1ssr1)r.tau p.toS g 1osuorldu3sep [uurBuo ur uc.t r3 s1n1cp oql uoj{ l[lnll]ucurrclrorrqJJrtsouSurpSuollsc-(IU6I/,l, puotoq:Zl6I rc)toorg)snucS cql utcnbtun urrol ',{3o1our pooso lco,lor nlrol lruorsuourp-7ur o1pcpnllc ,{1ca.!lDlucl fluo'srclf,nmqJ oscql l€ql u,lnoqs o^l]qptno,r pJcscsJql qtrm.{)r:ur1ru.tt1c]o] I durnldpus qlootus',,rrln3rqrool ole^o,,dlprluossc o.rnpJJS cql luql solrJrpurscuJs^ oql .rcJuorldrrrsop cq; 4s suasD)unp|.r'flJo peos^lBs dlFnlJo 'A 'scrJcds oq luql .{1c1r1trnst Y tol )^uasoruttpa"r clSursoqt pccnldcq qctqn ut sctccdssnidflzrzg .fospccs oql Jo uortnJr.lrssul,slq tryam.u.radsytal sotscs sndqDinn tylpolciro uoprshl'9Z6I uI

'odroqlsuclcg ruou puou;scq uo.rg fluo 'sc^col ';^nocnnlS'lcpucls 'A u,1oux lrrql qlr.^dqnJqs osonxoU e .{lqclousn^! lr lP\l PUEDJanpal ,,o1 csolo^lc^,, su,r sctcods oq l lnql rcuprcC V JJo sclonbscpnlcul onSololord ctll stsuapuowsap'A 'uorssn3srp srql ot lutl^clcr scrcods.,'rou Joqlous poqlrJscp (9Z6l) ,{lolutq puu uople]^[

'ttcpty1,t1 'luolxo sua3u14soU lollBru u,rorqJo lBql ouros 'pctqucsoj 'u^roJq ol ruu.l cql tDql3ur.{ns su polorb s?,14rcuprtrg VJ (pg arn8rd) ,,luo8uulss f,Je^ ' 'lli:r.ns,, 'oopu?,{\ -rca.uso1 ,(e r8-rc,r.Is scWU u rql ur palBcllroJcpqJrq,^A lruq cql roJalqulou oslP s?,,!\ 1I ruo{ tr pJln.rcdcs.{lrpuoJ lnrll c8rrloj rlsrnlqflaureJtxo rll t/{ lelle]Ule'uaup.rD8 ir su tZ6I uI uopIEII ,{q pcqrJJsopst..\\ .rts sua:'DJunpa.rl01 peln1c:dlesolc cq ot lcptsuoJ ,,'{ou3,^\ qctq,tr setceds y

'(^\oleq ccs) cscqt uo posBq.qe rrp^ .trsue qsrlqnd o1s]?.t! (t€6I) dle{Blg ',,Drunpat g ol clqulelaJ,, uoeq elpq,{uu qolq,tr pJoJslnux pu? urpJcpunJ ruor; su:utt:ads ol pJ.ueJet eq 'suossoJ 'luotxolsls lso8rp etuBsoql uI luerc]Jrp loJ ,{lquqoJdlnq uorslJopslq qlIA JnJuoJcd\ slql ap?uequoq,rs?,r ocunpat g loldatuoJ s.ucprctrAllBq,,!\utauer lou aJeeL' I{8rLoq|le(munpat Jp-A) 'rel?d'JcIJrql,, DJanpat A oJ$EJluoJur..sc^uol olcloeJu?l aIBJI€J €roui sllJo slseqeql uo luer cr;neds o1Dwau oqtuDr n pe]0$)r puB DllotusnquD m^ DJunpa| A Jo ldoJuor slq posr^oJueplq I'(€Z6I) 'pa^Jnc'3uo1 rc1e1 ,,ulncredoelEurLUnoe ,&oA dra,r,,sql ,{q,{lcSJBIposuelc?.l"Wetlqaulrl'o4ltaLro 'm^ 'uorlrppu 'r4o1qdoao1au 'DJDq Dranp n, paqsllqnd eq uI pue n11o{psn8uo sm^ DcunparA 'exel coJqteql pu" , tuupat g postu?oceJeq e r'tueu]l"erl s.ueqluog ps,\\olloJ(8I6I) uepl"l I

'f:ep,r,r.od ']I?d ,{pqfrls aq uBJ lpuorl€N rnonscl pasodo.rde\l Lrr'Idruexe roJ'(Dan^fid 'dsqnsaopr.r,{ !,) oopuu.^\lsouluroqUou cqlJo IrBq eqJ eBue11Buqlreqeql'3'c'luepunquuaaqa,req

'raddoH rrcrwf!,,I - uapr"trr,t,rlz radshzj sauassndqDrt C u.qd.rs ?t raloorg HItt{ Nuylsia Vol. 8, No. 1 (1991)

operculumofE. desmondensismay haveobscured its naturalaffinity withE. red.uncq.seed, znd cotyledonsdo not supportBlakely's assignmentof E. desmond,ensis(Brooker 1972),but these characterswere not refencdto by him in the series'digest.

Blately describedone new variety,E.redunca var. subangusta,based on specimensfrom cunderdinand Knutsford which Maiden (1918) had refenecl to typicalE. redu.nca.E. red.uncayar. elatawas raised to spccies,iz. E. wandoo.

Gardner(1945), in a commenkryon thespecies concept in Eucalyptus,gayeprominence to the difficulty of definingspecies in theE. reduncagrotp. without resolvingthi taxonomicproblems, he highlightedthe varianlsof bothE. wqndooand E. gardneri Ihatocatr awayfrom rhelocalities whcretlpical formsoccur, and tentatively suggested that all relovanttaxa be placed in thoone sDccies (E.reduncal.

Pryor(1962), discussing the series con ceptin E uc alyptus, ernphasiscd the value of thccotyrcdon shapein broadlyindicating natural atfinities. He recognised that a largenumber of spccies,inciuding thoseof Bl:rkely'sz ucalyptus saries Subcor t ae,hndincommonbisccted coryledons. and rhushe concurredwith theconcept of Maidcn(1931).

Pryor ( _ andJohnson 1971) largely follow B lakcryin thcir informar"Eucaryptus scrics Red.uncae" butcorrected Blakcly's anomalous placemcnt of E. tlesmond,ensis.This species was included "E. in lhe scriesbut in a monotypicsubserics, subsericsDesnorrl ensinae" . Thcycxpressed som e doubt ^bolttE. xanthonemdand statcd rhat it requiredfurth.' investigation.They aniicipatcd subspccies yus sfatusfor E. redunca subangustaand, melanophloia. E. reduncq var, oxymitrawas givcn informallyas a synonymofE. gardneri,as wassuggested earlier by Gardncr(1945).

pryor chippendale(1973, 1988) Iargely followcd andJohnson's rreatmcnr of lhc seriesbut rcinstatedMaiden's name Eucalyptus ser. Levispernae. E. redunca vars s ubengustuand melanophloia weresynonymised by chippcndalc(1988) under c. retluncawith a notcthat furlher r"ra..i ,ry demonsfratethat subspccific or specificrank wouldbe appropriatefor theser^xa. E. reduncayar. oxymitrdwas fotmally synonymiscdun dcr E. gardneriby chippendarc(19gg) without comment.

The four syntypesof E. redunca. From our field and herbariumstudies, wc havefound the applicationofmostofthe abovenamcs relatively easy. we wereuncertain, howcver, as ro which form thename E reduncavtr- reduncamight refcr and weassumed this would berevealed by examination of the syntypes.

From researchin manyherbaria, we havebeen able to rocateand examinc orly lhreeof thefour (iso-)syntypes, vrz.Preiss 232 (MEL),245 (LD), 247(LD) (Figures95, 96). We haveno reasonro bclievethat themissing syntype (prei.x234) does not bclong in thesame faxonomrc seriesas the other three.If it is subsequentryfound to be unrclated,it w not affectthc proccdure adoptecl below by our choiceof Preiss232 aslectotvne. sltx?puous2P 8z

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Brookton)' Fi}nte L EucalJplrs wctndoo s]bsp. wandoo illttstrating tree habit (road between Pingelly and

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'erudaql se esrro:dulr '$eaqs urnuuqreq alerudcs .g eqluo ue^rEsuollnqr4srp crll qlr.,r\spJoJJs uotsnlJuoJ srql puE oopuot{ g 1o1e4dfi am (ocunpat 'oopuDflt'g go sed$u.(s)LVZ'ZVZ ssDtd suaulJodseql l?ql e^allaqe/[ sepnl.ur1eq1 seneds ;o dnorEaq1 sr aseql;oauo (I16I uosuqofpuB to.&d flsras) sorJodsJedns l?u]roJur se osoql ,{\oqs o.r I elq€Jur pu? saSBourJ olqs^Jesqo [?Je^es eJ? eJoql l?ql lsaSSns aouttadsr taI sarJosur serpnls JnO

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l:agute2. Eucaltptus clili6old, a rypicat maller (AerodromeRoad, NW of Ravensthorpe).

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'serJads eql ro uo[euJoJeql ol sloJol{ol?plx u]el eql Joqloq,rJ?elJ lou srlr s? 'sle1clq1 snonBrqurusl lueuolelssrqJ'..rso^rltu eql Jo ,{copet[, olqerleuadrulpuu osuop Surturog,, '(tggl) .sn :llo^rxetrI ruo{ etou3ur,,'to11o; eql sellB'snd&Dld A Jo tuetut?ollslq ul ru?qluog ot 'Joll€ur u {ouI uuoloql ot socuoJeJeJlsarFaa oql ur ueos srqluo ,{ln8rquru ueeq s(e,np s;q oreql

'snd&Dld E se rlxrssarrads loJ osues^lo:ruu 1tul8r.lo s1r ur dpo slddpcne;o lrqsqoqlJo suouduJsop ur posn tseq sl.,{JolJEur,, teW epnlruoJ pue'suoDEJrIdde snor,ra:d snouu.L slr pale8psa,rure,req a m'aowadsltaT et4yo saroads o1po11dde uoeq s€q.,Icolrcu,, uret sql osn?reg

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Figarc 3, Eucalyptus subangusta slbsp. subangusa, ilustraring rhe ercct mallee habit (Canna).

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.. lrqBr{olqBulJepu reql qlr^l sltolJPru 'dsqns scpesruSorar eq lq8ru sroqlootuos pup otpuou,ts g'nllottqto g "fiJ[lJ ,sl3nn g'DSSotBg sduq:a6'sdeleql ur urreleql asn lou opea puepleu eql uI uotlluSocarf,see.rol,trol1elou ssop {oolmu eqlur $uo8ol?Jlrqeq ooJql Jo uorsnliuroql raqnlou8qe 3ur1ce1,(ypasoddns pue surals enb4qo ro 'pJstuSoJJt tJJr:lJuusrp Inoqlr^\ JJJI ll8rus ro eJIlpu|qruqscsn33a:o,(tt,t ^ltsuJsset Jt{l sI srqJ.. '(/ petousmqlnE eseql d) oJaqr,reslg,.eol IIErrIsro ealpursnoreqnlou8ll-uou esnJJe us FoIr?uI,, '.,rsqntrou611e uonrurJepeql p€sodord(ZZ,:066 3rure14pue:e1oo.rg 8ur-)l3q1nq 1e,rel punorS reau.ro1e Sursue seqf,u?rq IIB qtr^\ quqs peruuelseISuIs e,, s? IJolJEtu poulJep (86 :686I) II111

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Figljrc 4. Eucalrptkr pluricaulis s|'bsp.porpht 'ea, illuslraliDg lhe effuse mallee habit (SandalwoodRoad, N ol the Stirling Ranee).

t,- 'suaJs"r^'dsqns '.eer .dsqns D$n8uDqns A G) :otDdsuTq G) iDuptot g (p\ aft\evt oopuD,n ooFntD,tt fl (.) :StItde' oopuht diqns oopuo,rl g (9):opart, dsqns DaDul.trDto g (E) iDaradst^4 ,ras sadttD"nfl u sed,{t )tleg .9 ernSrC

'roddoH 'O 'H eBa:EuAlN - ttapft I aoutadst^ll sauas sntd.tlornq uaqdats ? raloorg I I{ NuytsiaVol.8, No. I (t99r)

which we find indistinguishabtein the two formsofE./avida. Adult andseedling leaf characters alsosupport the conceptof a singlespecies in thiscase.

shedding ^.The of thebark in predominantlysmooth-barked species varies from thedecortication of iregular slabs in E. wandoo(Fi.grre 5c) and,E. cap rosato thecharacteristic partly shedcurrs of deadbark in someof themallets, e.g. E . gard.neri(Figve sd). rn E. crispara(Figurese) such curls areprominent anddense, particularly on thebutt. In seyeralspecies (e.g. E. sribingusm,figure 51, thebark is shedannually leaving completely smooth stems.

Bark colour varieswith the time of exposureof new surfacesrevealed after the shedding of the oldestbarklayer. Thepersistentweathered surfaces are originally pale or darkand on sheddin!reveal new surfaces which arebright yellow, orangeor coppery.These colours subsequently facle. The colourof new barkin autumn-winteris particularlyusefur in distinguishingE. wanioo (yelow) from E. capillosa(orange) wherethe two speciesco-occur (e.g. betwein Kellirberrin anOborrigin;. ln E. wandoosrbsp. p ulverea,the smoothbark is notable in beingpowderylike themrerated E. ic idens (powderbarkwandoo)..

Bark on the lower trunkof malletssuch as E. crivicora,E. densaandE. praeterrnrssa often has horizontal scarsup to 90 mm longpresumably due to insectattack (e.g. Figure 79).

c.otyledons'. Thelaminaof thecotyredon in all speciesis deeplydivided ancl forms a y shapewirh thepetiole (Figure 6).

''@t

Figurc 6. Seedling ofr. relarcd with y-shaped corytedons.

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'(opb.o\! X u ueafi durqspus sarcadspeluler '3'o) pueoopuDt A u\sr, oJAoJe se^eol oql JnoloJJuol puv'(89 ern?rdslpqnr' A ut lsorAorreupu? '79 'g, ern8rgpueyI en?rg'olodsltc gpue'DaJa^Uddsqns oopuD^r ul lsopuolqsr JBet oql) odBqs '(,,DJunpaJ,,sarJadsJadns ;eay8uqpaes aq1 pueDplltuJ A ur opou?IulqUrJ eql loJ uJJg ol dn 6'e) scpour4uryo q13ue[e|{| eJB soljos eql ul sdHsuouelor6urssesss roJ lnJosn sJIlslJolJEJeqc raqlO

'seuas eqlJo senadsloqlo ur JnJcolou op souoqJuJ '( 'sJed'seEpzl) u-ruroJ srnq Euperpergoqs qlr^r spu?l8Io lueaJ€l.ju,eem DaDuqJrJ, 3,Jo souroqJr.q 'lsEJluor '(t86I eql uI se8rp?a o^oq?oe{ pus lue8le^rypue q1?ue14eql sprql-o.4\llnoq? roJ 'spuelE snonEDuoJeJDslleJ aq1 polua8rauaaqlJoslleJ d?J pe:red;o uorte8uola lBculoulur,{s oqlJo esnmaq(8arn8rg)swdtpcngtlanbwnerus8ullpeeseqluo,,srmq,,eso\L'opuntuSrugptmng4l'g 'DSolltdn g Jo (Lam8rd)sauoqclq eql srsouos oql urs8urlpeos oqlJo amleeJ SurIrIS tsou eql

'se^Bol llnpe ernpoJdesuqd srql puofeq poJu?^psEueld(sa^Ealllnp" u?ql Jep?orq,{1pnsn) eyua,r.nferE se^sal oql IIsl luJ97,{leleurrxo:dde sl lu?ldeql llun luaurdolo^epapou?.qu! Jo lesuo aq1 uorg'(SulEurEp) sopouelureq1;o uoqr8uole,(q seseqelotedeqllepelsJ8des ouroJeq se^eal'JeusareqJ'os?qd Sullpoos eqt e'r'sepou9-ZroJSurpees pols8uoleeql ur olrsoddouEureJ sn?d oql pu? xed€ looqs eql ls ,{loFssncoppocnpord ere selesl eq1 'sSu1peos 'seyas eql uI eq1ur sarceds1p Jo al3,{J eJll eql lnoq8norql e1e1o4ed eru sc.^Ea-I'sSuqpa aS

- 'raddoH .O .H.I.IAJ LI anawSN uaprcI^{ dtalads^rj seues sqdtlotllA usqdas T ra{oorg NuyBia vol.8, No. I (1991)

FiSure ?. Scabrid seedlinglea! of E. capillosa snbsp.coplllora (lef!) comparedwith a glabrousleaf ot E wandoo subsp.

Figure 8. llairs on the surfaceof thc leaf of 8. .dpirord (electronmicrognph counesy of P.Y. Ladiges). -\

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'suro,\oprs oql qll^\ uoJlcrJossEoJqlsrA fue (snorounu,{reA lnoqll,r pue'oEJel.{le^llulor puE osucp orpspuelSgo eq1'souoz lslsocretur Il?proquoql 'ourTlno eloq.\\ore salooJD oql eraq.rsyl/qrs g uI ur JBlnSoJrfltensn ols pusJtro1 s18urs e ur ezrs '(6 ur olqBw^ aje ^oqJ arn8rd)salooJu eql ur solpoq..putrtsr,, oq ol Jo sloluro^eql Jo suollJcslolur 'souos oql ts pal?nlrsoq ol JPodds,{eql oql ur sorJodsI[€ Jo so^931oql ul rnJso spuBt8IrO

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'arn:sqoput,{rc1ueu8ur,; 's! drc^ sr'lueserdueq^r'Surura^ ,{Juu:alen} tqns A prmDsuap t'E a'tuasq€.ro,fut1ueur8er; uaql sr lr eJoq,rsa^Bol lse,roJJBU oql lnq IIp ul lJupsrpsr Euruls,ldffn.rol (suro^aprs oql Jo ,{luapuedepurpeuJoJ sr se^ucl llnp€ ur ueosuro^ lEur8Jeurej,lur eql lurll pc.roqs ('lrJ JoD'lDpJmJ selJodsroqlo ouros Jo;) uro,r pur8reuu4ul eql seu^rou{ ,(ltuuorluo^uoc sr l€r.l,rlx.lo; pue lucururord ssolele seqcJeeq q8noqtp (6 ernErd)se^Bal llnpu og ul mllurrslllelog:adns sr rucllcdeql

'ur8ruu Jselaql oprsurlsnf uees,{1ree1c uelgo ere 1nq ,ne,; sru spue181o pueluosqs 'ure,r ro {so,r sr ql,ror8 Jo o8?lssrql l? Sururel,ftepre1 purEreurerudro{uo,r puorosD rujoJ ol dn {rrl ul?8epue urAJeu oql spJs^\ol puelxa slolura^ JoqUnJ (9g6I lDJa Jm)) .jle^Juur8JsruBrsd srql Jo eprqnO'soqJleJoseueseururEreuraqlJouoqs[[a,rpeurofAuruloJoqpuefplsrpSur^mJ'spJp,rlno qrjpru eqt uo{ puelxesura^ apls 'o'l'GL6I felcrH) snoujoJpoprqcoJq,tllpJrd,4l sr uorlDue^aql se^?clEullpaos eql uI'Qtlqlpns pa twsuDttl.ltt sa^Dalqsa{ut uaassD) spuDlS lto puDuoltouatrtDa.I

',{doueoaq1 aprsur stalqJueJq Joplo 'se,,ree1 eq1uo sa,realuaarS dssolS Suuo^oc ueerS-en1qIInp Jo u^rol3snoncrdsuoJ oql toJ elq4ou sr maaqJso.tJrufl'e?e se^eelllnpe eql se ssaurssolS pue rnoloJ ul luslsuoreJe selceds lsour apq16

'u,\\oJJeql eprsur o,1{ueoJ8 ,{ssolE ol ornl?u lnq qsruaar8pw 11npsru,{eq lseolle Jeof,lsJrJ eql u'Drunpal gu\pw qsmlqpueltnpere 'ueel8 se^eeleqluaupn?gu1 f,sso13arl-se^EeletgDplww ApwDlltlldouamld'g'DaouLlJDID'EuI '.{er8 qsmlqol uaer6-en1quo:;.{re,r pue IInp ers sa^Bal llnpu aql sorJods palBIeJ puE oopuDt fl uI

- 'rddoH a"5rs'u,(tl uapf"I/,t,n{?Jadn^4 s u.s sktclq,rnq C ueqdats ? raloorg H.I.t,{ Nuylsia Vol. 8, No. I (1991)

Figure 9. Leafvenation and oil glandsof E. flaida.

b

Figu re 10.Transverse section s of budsjust abovethe junction of the operculumand hypanlhium showing the number of whorl s of slamensin (aI f. /hvida Nd ltt E. Bardnzri.

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.,{ll3lsIp 'os 'pouolleu uapr,rrpue ,(pcuqsrpsorJeds euos uI leq^roruoss,(e,trp are selcunpad eq7 'salcunpa4

'srsoqlu?pue uoualrur pnq me^Ueq(srse,( g-Z) pouod 3uo1 aql s! seuasaql Jo orq?eJenbrun lou q8noqlelq?tou V

- 'raddoH .H.I asarqr,{y,{ uapiet{ ,r'rzradsl^v7 saues ffitdqDr,1a ( ueqdols T rdtoorg hl NuytliaVol. 8, No. I (1991)

a

b

c

(fight row) of buds wilh lhe Figure 11. staminal amngement as seenin longitudinalviews (left row) and longitudinal sections t6rs are 5 mm' !rop-erculum removed in (a) E.llavida, b) E. desno densis, (c) E, nicroschetM' Vertical scale .s.r..(qder'odo1pue",o,,"",",,,n;3;;;11fiilH:Jil:i;Tg"""lli,:|;l;lilij:;,f:::;:i JoueSuoJoelleur slr uro.gl3upslp selJods e s? pasruSocors,[ lnuprlS A toltuu eql'oldu?xo rod '(,ie)'I aas)&rlrqerlarelnlosq?Jo srlrqEq l8ql Sulfldullnoqllm (uorlelndodeur uuoJ tuBuruiop eql .e.r) sJolJeEqJllquq pe^reJred ,(Up?or ol tq8re,rue,,rr8 dlpreus8 e^Bq e,r 'snlelssercads Suru8rsse uI

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qdaJuoJ)!urouoxel

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'(6961:eddo11) a1oq.n e se '(gI snue8eql ur pussolJos eelleu Jeqlopleles u osp ueasruqled s e:n8rg)1cr4sp afuoq1sue.re1 aql ol lsea aErea Euy1.l15oql uo4 lleql?eq,ruJeqlnos 6ql ur lsou pele.4uoJuoJete BxuJ

'(Dpt^Dtl g'Dpun$t81xt!') uosac euolrr^ l?org 'pu?lul aqlse re; sesa8uer saues eql (€I emArC),tueqly ol uollossnguo{ sBaru1p;urer lseq8rq aql ,{[uoSurpro^s'lsBoc eql ol spJe.,[q]noseom uJeqp>I oql uro{ spuelxe aqutads!^aj'ns snldKpcng

,{qde$oa3o1Xq4

- .C ee.3eufl^I uaprcl{ rzu DdsrLq sauas sntdqorrz leddoH uoqdals T, ra{oorg H.I.,^I Nuyrsiavol. 8, No. I (1991)

" b N-'^wt#@ ffi ,* WW%f, Figure12.seedof(a)S.redunca(MIJI.Brook?r8658),and(b)tfllbd'Srlasubsprxbd'8rdra(MlllR'7914)'

"' \

WESTERNAUSTRAL A THE SOUTHWEST

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11

ser' tuvirper'rdr in southem FiCDrc13. Map showins lhe dislriburion and dcnsiry in l' x l" sqsaresof taxa of E 'dbpl'/s -.l

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'r3ddoH aearw(t,{ - uapplN anuladu,el sauas ntd\ocng C uaqd.ls ? ra{oorg H l hl Nuytsia vol. 8, No. I (1991)

Ourtaxonomic method is basedexplicitly on thewholeliYing plantwith somecmphasis upon its environment.We do not hold to thenotion that genetic fixity andexpression is necessarilyshown adequatelyon herbariumspecimens,label information included, particularly for treesin contrastto herbs(which can often bc preservedwhole in herbaria).

"Great Davisand Heywood (1963) stated that caremust be talen not to assume, ab initio,lhat spccieswe have recognisedin the herbarium(often from limitcd samplesthat are not even contemporancouswith oneanother) are valid representativcs of naturalpopulations in thegenepool sense", "limited We extendthe idca of samples"to limited information,as speciationin Eucalyptus inyolvessitc adaplation, natural colour ( segUJrderE.xantho nema),bark chancter istics, lcaf category, leafdispositionetc., all ofwhichcannotbe expressed in a dricdherbzrium specimen and, to be useful, requirea standardisedglossary of termsin thecollector's notcs.

Theseries Lsvispsmde , until ourresearch, is a cauli(xtaryexample of taxonomybased on limil.ed data.

Descriptions

Eucallptus seies LevispermaeMaiden Crit. Rev. Gcn. Eucalyptus7: 144 (1925).E. scrics "Koy "Class. SubcornutaeBlakely, Eucalypts"llI (1934).E. series"Reduncae"Pryorand Johnson, Eucalypts"44 (1971).

'Iype: E. reduncaSchaucr.

Trces,mallets or mallccs,wilh or withoutlignotubcrs. Bark wholly smoolh,wilh parlly decorticatcdIIakcs, or roughon lowerpafl of trunkor sl.cms.Branchlcts with or withoutpith glands, glaucousor not. Cotylcdonsbisected. Scedling lcaves pctidatc, oppositcfor 2-6 pairs,elliplic to ovato,green orglaucous, glabrous orhairy. Juvcnile leaves petiolate, alternating, lanccolatc, clliplic, deltoidor cordalc,grecn or glaucous,glabrous or hairy. Adult lcavcspcliolatc, alternaling, linear to broadlylanceolate, concolorous, grecn, bluish grccn or glaucous,dull or glossy.Irflorescences axillary,unbranchcd. Pcdunclcs always flattened, broadening towards thc top. Budspcdiccllatc or sessile,fusiform to doublc-conic,sometimes curved, outer operculum lost carly in bud formation, inneroperculum acutely or obtuselyconical or horn-shaped,sometimes uncinate. Outer filaments crcctwith anthersheld upwiuds, orerect for mostoftheirlcngth and finally dcflexcd near the top with antherspendulous, inncr ones erect for halftheirlength, then inflexed with anthcrspcndulous towards thc basoof the operculumcavityi rarely with all filamcntsinIlcxed. Anthers versatile, dorsifixed, narrow-oblong,opening byparallcl longitudinal slits. Filaments white, cream, oryellow. Ovary sunk into hypanthium,of 3-4 loculi; ovulesin 4 veflicalrows on an elongaledplacenta. Style bng, to undcrsidcofoperculum. Fruitpedicellate orrarely sessile, cupular, cylindrical orbarrel-shaped. Seed subsphericalto spherical, or sometimessomewhaf cuboid, rarely ovoid, smooth, whitish to lightgrcy- brown:hilum ventral. -r

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ae$su,{W - uaptyl.r'{,r,a ladsl^"1 saues sntdqDrn4 !6ddoH C uaqdals ?t.tatoorg H l-W NuytsiaVol. 8, No. 1 (1991)

The anomaly in this arangementis the Levispermae,which includesmost specieswith conspicuouspith glands,one without, and somewhere pith glandsare present but rareand often 'ft difficult to detect(a similaranomaly applies to theseries crassatae"in section"D umaria").This factor,and the absenceof sharedderived characters in commonwith otherseries, suggest that the seiesLevispermae is taxonomicallyisolated.

It is ofinterestto notethat rarc inter-series natural hybrids with membersoftheLevispermaehave been found (seeend of taxonomictreatment below), involving speciesof the "Loxophlebae" (E. loxophlebaxwadoo),theAccedentes (8.accedensxpluricaulis),the"Erythronemae" (E. recondita x mnthonema), the Cornutae s.lat. (8. spathulataxwandoo)the"Grossae"(E. grossaxhistophylla), "Dumosqe" lhe (E. conglobatax wandoo,E. arachnaeax obtusiflora), lhe "lncrdssatae" (E. desmandensisx incrassata, E. intassata x medialis,E. incrassatax pluricaulis,E. incrassatax phaenophyllastbsp. phaenophylla). Several of thesehybrids represent an intersectional cross, as the "Dumosae" seies and"Incrassatae"ate ln section"Dumaria".

Twenty-eightspecies arerecognised in thescries. E. desmondenstsstands apart from all othertaxa, andis placedin themonotypic subseries Desmondenses which is distinguishedin thefollowing key.

Key to subseri€so[ E. ser,Levisperntae

1. Stemsslender with sparsesometimes drooping crown; branchlets glaucous, lackingpith glands;operculum broadly conical, g equalto hypanthrum; all stamenscompletely inflexed subseriesDesmondenses

1. Stemsthick with crowndense, erect or spreading;branchlets glaucous in few taxa,pith glandspresentl operculum narrowly conical to attenuate, usuallylonger than hypanthium; not all stamenscompletely inflexed exceplin E. microschema,...... slbseriesLevispermne

Twelveinformal superspecics are recognisablewithin theE. subserieslevjspermae (Table 1). The "wa'doo" superspecies is characterisedby thetree or tree-malleehabit, large, often hairy seedling leaves,dull adultleaves more than 1.5 cm wide and compact spindle-shaped buds 1-2 cm long. It comprisesa groupof parapatricallyreplaced taxa, with E. pdndo, distinctin its glabrousseedlings from E. capillosa,E. livida andE. nigrifunda,all of whichhave hairy seedlings.

Thesuperspecies'p haenophylla" comprises agroupof ma.llees with glossyadultleaves more than 1 cm wide andbuds more than 1.5 cm long,often with theoperculum conspicuously nanower than thehypanthium at theirjunction.Again, thisis a widespreadparapatrica[y replaced group of taxa. Theeastelnmost species, E. tumidand E. histophylla,havemore allantoid buds than their western congeners.E. arachnaeastands apart from all othermembers of thesuperspecies in its roughbark andvery slenderoften uncinate opercula.

"Jubangusta" The superspecies includesthree species ofsmooth-barked mallees (orrarely small mallets)whose adult leaves are usually dull andnarrow (less than 1 cm wide)and with smallbuds lessthan 1 cm long. Thetaxa in thissuperspecies replace each other parapatrically or allopatrically. E. subtilishas the narowestseedling and adult leavesin the serieslevisp ermae, urd is isolated geographicallyand morphologically from theother two speciesin the superspecres. -".l'

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'reddoH ee.os!,{N - uaprcW aa4rads^"1 sau.s snrdttpnlg ( usqdels ?9re)toorg H I I NuytsiaVol,8, No. I (t991)

5. Branchlehnot glaucous Basalbark rough; buds to 15 x 3 mm; lower CorackerupCreek and Pallinup River areas; E of theStirling Range...... 21.E. melanophitra 8. Basalba* smoothor with scattered partly-detachedflakes; buds to 26 x 5 mm 9. Operculummore than 10 mm long;crown intenselyblue-green, new growthusually purplish 10. Adult leavesmore than 15mm wide; usuallyon lateriticridges 11. Operculummore than 15 mm long; west centralwheatbelt - Cadouxarea ...... ,,.,,, 23a. E. gardnerisubsp. gard,neri 11. Operculumless than 15 mm long; RavensthorpeRange...... 23b. E. gardneristbsp. ravensthorpensis 10. Adult leavesless than 12 mm wide; usuallyin depressionsor flat country;southern wheatbelt from Ongerupeast to Ravensthorpeand no h-east to Hyden...... 24a.E. densasubso. densa 9. Operculumless than 10 mm long;crown grey-green to blue-green;new growthnot purplish 12. Budsto 7 x 2 mm;fruit to 3 x 3 mm;small mallet to 5 m tau; northernwheatbelt, particularly Mullewato PaynesFind ...... l6b. E. subangustaslbsp. pusilla 12. Budsmore than 7 x 2 mrfl 13. Mallet or smallto medium-sizedhee maturing to morethan 8 m; new barkcreamy grey, pinkishor yellow;adult leaves green or blue-grey; juvenileleaves not hairy 14. Mallet;bark cream, grey or pinkish,often with looselyheld non-decorticated dead fragments held on the lower 1 m; blackhorizontal insect scars oftenpressnt on lowertrunk; buds to 14 mm long;BeaufoIn1et.....,...... 22.E.nraetermissa 14. Smallto medium-sizedtree; bark white,grey or yellow,often with scatteredloosely held darkrough-barked fragments; insect scars absent; budsto 19 mm long;Darling Range and westemwheatbelt, south-east to StirlingRange andlower Pallinup River .....,...... ,.. la. E. wandooslbsp. wandoo 13. Smalltree to 7 m; newbark orange;adult leaves conspicuouslygrey; juvenile leaves hairy; westem and southemgoldfields 3.E.livida 'dsqns """" """" wsnSuoqns A'p9I tl$qlller?N $ou pue'ooroql?ldpue SuluuBuruet { uoa,rtoqlleqleoq^r :6uol rurx 0I ot s6lJunped 'tZ :epl,i{ulttl 0I ol s0^B0lllnpu :l.llooursryBq l€ssg olodsttt'g'91 "" ""'uoJeBqqeoug-D[oouepu?l :8uol u x 9I ol solJunpad:epr^r uu SI ol se^eelllnpe :se{pu '€Z Eurtrnrpal?rruorop dll€Fred sD luetslsred ]J?q IEs?g 'ZZ Suotrxr I u8qlssol spng spnqe:n1eu uo uro!1e urnrqluud,(qse qlpr,r olxBsunlnc:edg '91 'A'eg "" """'lloql?oq,r DaDUIJDJD'dsqnsDaDuqJDro I€IuoJpu€ ureqlou :sr6lsnl,droprds u1 3uo1 uLr 8I ot spnq 'ltere :opr^ruu SI ol ,(nusuluoJdlou se^sal:plaq fllqErl Uuq Iesuq IZ owoclsnds'E'L '''''"''"""" " " " """ " "aomdncourq3 a1u1-dm8ur4 :6uoluur 0l ot spnq:opJ^\ruru 0I usqlssol 'IZ lreJesa^€ol:ptoq dlosool rysq lEseq '6I q8nor{mq ps?g 'dsqns """"" suarD[raiu! o tuldouaa$ fl'q6 e6ueged:oqlsuo^?U ot ls?e sltrg poo,4\xogprlu dnBunuru::a1 :f l88urlssuals :urnlqlu?d,{q uaqlra/uol?u .tPqE11sdluo "uJEIp '02 urnln.rado uruit ot spnq 'dsqrs '?6 "' Dlltqdouanld o1\&ldouaoqdg lloql€oq,rureqtnos puE Isquoc :lqSlsJtsT suols:unrqluud.tq ueqlro/t{oJJeu {lsnoncrdsuoc '02 unlnrJedo'uierpujur S'Z ol spng qloorxs{rDq ps?q 6I spnqemle ul uo uloflB unlqluEd,4quuql Jo,roJJeu unlnJJodo '8I uaerEol uear6qslnlq'JInp so^?ol '11 olruo^nf:8uolur"u SI u€rgssal unlncradg "" """' Dp!^Dtl'9" elFooEle;,1Jog pu?N:ruru 9 x 0I ol qtn{ :,,!\ollo,t$o,,r\ou 'fssolS :u6oJE se^Belollua^nf:6uo[ rurx SI uEqloJoru unlnc:edo 41 (u^roJceplsu! se^?ol mo,( puoJos uo tuople eq ,{luo ',{sso13 '91 ,(Bu.rscllsualrelpqi aJnl?u) uee$ se,,iee111npy 'sI srolcsJBqsJo uollEurquoJ0^oq8 eql lnoqtl]{A 'g s$uapuowsap 8z oAUBUofu oqlsue^Bu :unlqluEd.tq 'llnp'urc ol pnbe +'pcluoJ rxnlnJredo:.(eJE-enlq g'g x 0I 'snoJn?l8 ol se^B3lllnpB :spu8l8 qlrd lnoqlr/'\ slolqJu8Jq 'ru :ut\oJJ6uld6e,{\ seurleuros asruds qlr,,ir.lepuels ? ol suJals gI 'I (suonrurgap]lqeq ees) eelpw

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22. Budsmore than 1 cm long 24. Leavesconspicuously erect 25. Budsmore than 15 mm long;between FraserRange and Balladonia ...... 12.E. histophylla 25. Budsless than 10 mm long;east and south-eastof Newdegate...... L8,E. mitoschema 24. Leavesheld at variousangles 26. Basalba* smooth;adult leayes to 20 mm wide; juvenileleaves to 6 cm wide 27. Adult leavesto 11 x 2 cm; maturebuds allantoid.to 20 x 3 mm:E andNE of Ravensthorpeto IsraeliteBay ll. E. tumida 27. Adult leavesto l0 x 1.5cm; maturebuds fusiform. to 15 x 4 mm: Jenamungupand Boxwood Hills eastto RavensthorpeRange 9b,E. phaenophyllastbsp. interjacens 26. Basalbark loose, rough; adult leaves to 12 mm wide;juvenile leaves to 3 cm wide;E of Hyden,90 Mile Tank area 10.E. luteola 16, Adult leavesdull throughoutthe crown, blue-green or green 28. Leavesconspicuously erect 29. Budsmore than 15 mm long;between FraserRange and Balladonia ...... 12. E. histophylla 29. Budsless than 10 mm long;E andSE of Newdegate,...... ,...... 18.E. microschema 28. Leavesheld at variousangles 30. Leavesless than 10 mm wide 31. Basalbark rough; buds to 30 x 4 mm; salineor swampyground; DalyupRiver-Gibson...... ,.26b. E. variastbsp. salsuginosa 31. Basalbark smooth 32. Crownblue-green; buds to 20 x 4 mm 33. Adult leavesless than 9 mm wide, new growthoften purplish; maximum height3 mi Ravenstholpe-Jerramungup iirea...... 24a.E. densastbsp, improcera q8nor'esoo1lruq pseg'17 'Euo1 '97 lunlq ro el8nuolle uur 91 ueqlssal urnlncradg po^inmr uelJoxede unlncrodo '9€ :8uolruru ZI u?qleJour spnq eJnlen """"" DlDdsrtJl'gI ?eJeBqqeeug-3)loouepue^ :sd?^\s{Barq Juualq uo :sFnc '6€ pel€cluocepdlued ur es?ql? Ir?g DlstrSuaqns'dsqns Dtsn?uoqns 'A 'E9I """"""""' llJql?eq^ruequou aql ut uruldpuus '69 ,ro e,(uo ,{lqslou:qtoorus ryBg lleql?aqaJe4uec pue uroq ou:esnlqo '8t urnlncredo:snocrul8 lou slelqcwJg 'dsqns "" ou1nc nlsn?uDqnsg'J9l lleqleoq.\lIDIUoJ urols€o reJ :elnJ? ueuo unln edo:snocnelEslelqcuerg 35 'r€ uju g x 0I ol spnq:uiu g x t u?qlerou lFrg o111snd'dsqnsolsn8uDqns 'g'q9l slllH w8uol[ ol qlnos puld seu,&dpu? ?,rollnl^I 'rg :uru Z x rot spnq:uu € x € ol trnjC dp le peyrnc1ou urnlncrodo '9t lEuolr.uu.r 71 u?qlssel spnq elnlsy{ '0€ epl.^l\uur 0I u?qloJorx so^Bo'I 'wuesjoNJo slvqns fl'Ll s ol rysI et!fiI06 :opr^{ruur 9 ol se^80[llnpE 'l?cluoc :dn lE poloorllou unlncredo 'tg :6uol rurx, ol'loJrpadot Suuodqspng DllsoddD'dsqns DuauolltuDx 'g 'e6l """"""" " """"' eEueaEuyFrt5 's€ :epld\ruru 9I-6 se^?e'I owauoqlunx'dsqns m)tauolguox 'g 'e6I dnEunu€lel ot sur?lll4[ Jo AS :epld\tutu, u?qlssal sa^uo'I 'Sg 'elenueoe dp le pe{ooq uego urnlnorodo :EuolururyI ol 'elello3rpedFllrullslp spng 'tg 'zg uru € x rI ol spnq:u6e.rau^rorJ 'e9Z otrD^'dsqntsDltD^ fl (Je^eI ssuroqJ ol ls"o re,ra dnfp6) unldpues ocueredsg :u L lq8requnmx?ur :u6eJ3-aollo,{ 'tg ql^\or8,Y\ou 'ep!1r\ urur gI-6 se^selllnpv

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42. Thin stemmed(to 10 cm diam.); leavesto 15 mm wide;buds to 14 mm long:Kalgan Plains, StirlingRange 20.E. medialis 42. Thick stemmed(to 20 cm diam.); leavesto 18 mm wide; budsto 17 mm longl Jitamingto Tincurrinand Dryandra ...... 5. E. hebetifulia 41. Basalbark smooth 43. Branchletsglaucous; newly exposed barkbright orange;central eastem wheatbelt,...... ,... 2b.E. capillosaslbsp. polyclada 43. Branchletsnot glaucous 44. Leavesalways dull 45. Leavesgreen; newly exposed barknot orange:buds fusiform: operculumacute; seedling leaves glabrous;Stirling Range... l9b. E. mnthonemassbsp, apposita 45. Leavesconspicuously grey; newly exposedbark bright orange; buds allantoidor fusiformioperculum blunt; seedlingleaves hairy; Coolgardiedistrict south to Kumarl- PeakCharles ...... 3,E.livida 44. Leavesmaturing green and slightly glossy,at first blue-green;buds fusiform; operculumacute; W of Dandaragan...... ,..... ,,., 6, E. abdita 40. Operculummore than 12 mm long,attenuate 46. Basalbark rough; saline or swampy ground;DalyupRiver-Gibson 26b.E.variasubsp.sdhrgilara 46. Basalba* smooth;slopes, plains or lateritichills 47. Stragglyshrub; maturg adult leaves purplish;Tadn Rocksouth to StirlingRange and Fitzgerald River NationalPark...... 25b.E. pluricaulis slbsp.porphyrea 47. Erect-stemmedmalleesi mature adult leaves green or blue-green 48. Leavesgreen; coastal from CapeRiche to FitzgeraldRivgr NationalPark and RavensthorpeRange ...... 27. E. redunca 48. Leavesblue-green -I I I I

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'raddoH 3p'rr"utrr,^,r- uoprey,r 2,a t2.tsl^?j sauassntdqDrrg C saqdars ? ra{oorg HIt{ NuyrsiaVot. 8, No. 1 (1991)

Nole'r' subsp ]rdrdoo diffcrsfrom subsp.purvereainthe absence ofpowder bz'k, its consistently yellow new bark,irs smallerseedling leaves,ancr its non-gtaucousurancttets. The two subspecies a^pp_earto rntergrade noflh of Cingin,c.g. M.I.H. Brooker 9889 (AD, CANB, MEf_,nSW, pi:niiij rs lrom a population with very slightpowder bark and palc yellow new bark.

The Twinc Rcservedisjunct population , eastof Narcmbcen (M.l.l!.'recognition. Brooker 10541)has glossy adult lcaves within the canopyand may warranttaxonomic This popufoti"oni,, presumablva relict from wcttertimes when thedistribution of E. wanioo extend"d"";t'i"";;.ry ; thecastcrnwheatbelt. Likc thcJilakin Rock population ofjanal, anJttrecoddard,s soar population of E rridr, theTwine Rcserve stand of E. iaidoo occuiswett'tottreeastorthe main range o[the spccrcs'and is confincd10 favourabre soilsreccivi'g high runoff from a lz'ge graniteoutJrop.

lb. Eucallptus wandooBl|tkely stbsp.pulvereaBrookcr & Hopper,subsp. nov. (Figurc14, l5)

A subsp^ccicryphr slatura parviore, , corticepulvcreo, saepe palrido-aurantraco ubinovo glaucisdiffcrl. etramulis

Typus: 8.7 km N ol Wathcroo,Wcstcm Australia,3 Fcbruary' lggg, M.1.,. Brooker9gg5 & C. ^larurrc.rs(holo: PERTH; iso: AD, CANB,MEL, NSW).

Trcc10 r 5 n lrt wirhpowcrcrv Ixrk oftcnpllc orangcwhcn frcsh agcirg 10 whilc or grcy, graucous branchlcls,and juvcnilc lcavcs up ro l3 x lb crn.

Specinensexanined. WESTERNAUSTRALIA: c. I milc SE ot.Mt Lesucur,6 Jan. M I II Brookar235l(AD, 1970, CANB,PERTH);7 milcs N ofThrc"Sp.i,W, iE ap.il f Gu yNEof 9.l0,M.1.. Brooker ?:19!PERTH); Mrpcror,30"06,5I15. 12,8, irvrr?r, rcs:,,Vr rr. Brooker B00l (CANB,NSw, PERTI{);6.1km NE, of ArrinorowarO, rvrura*o, iSJi:,s I t 5. 38,8,I Nov.1984, M.LII.Brooker 8733 (CANB, NSW, MEL, pERTH); 6.9km Se oiArrrn towardsThrce Springs, 29" 28'S I 15"40'E ,3 lunc 1985,M.t tr. Bt.ookergl32 (CANgtEi;,\sw, pERTH);7.4 km N of Coombcrdafc, 30. 24'S I 16.03'E, 27 Jan. 1986,U t.U n,",otril',)t69(CANB, MEL, NSw, PERTH);5.2 krn N of Moora. 30"i5.S 1r 6. 01,8,1 Feb. i S8i,i.i.ri. u _ ot "r 9876(AD, MEL.Nsw, 'ERTH); 12 km CANB, N of Moora.:0" :z;s r ro. oiln, i"r."i. , *s, ,y r a . Brooker9886 (AD,CANB, MEL, NSW.PERTH);5.7 mitcsN of ThrceSpit,-Ur, iS b.t. 1966,G.M. C hippendole 38 (CANB,PERTH): proposed Ml Lesucur Rescrv",e of irlt f"._, i0" 06,5 I 15. 13,E,21 May t981, t.A. GriJJin3ts2 (pERTH).

Disu'ibutionancr habitar. Northern whcatbclt from thecahby areato Morawadistrict (Figure r4). The subspcciesoccurs in more scarercd srandsthan typicarE. wandoo.but occursin simirar vcgelati0nslruclurc and soils. c-o'setration st,,tu' widcsprcadin disjuncfpopuhtions.often Iocallyabundantbutpoorryrcprcsented rn conseryaflonrcseryes. -T t

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'rsddoH aeacsu,{l - uaplel I,trz.tadsr^4 sayassnYl&tung ( uaqdals ? raloorg HIhl Nuytsiavol. 8, No. 1 (1991)

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Figurc 16. Distribution of E. capillosa snbsp. capilloso (.) and E. capillose stbsp. polfclada (o\, ndbuds, fruiB, adutr Ieaves and silhouelte of a fifth node seedlinSleaf of E. cdpillosa st'bsp.capillosa (scatebar = I cm). '(HJr{gJ '996I 'sso.rJ rpJ W 9 S A ;IIV'rIL uosltA4I d IudV ZI ' '€Iuolsela 'gNVJ) ureqlnosJo A\ ru)t0t :(HJugd) lrr{rs J w l LI6I dtnl LZ :(Hldgd 9198 artoll e'u'696I'rEW9Z'eopru) S tu Z€:(HJUgd) 6669 ar{oy gy'g961lgdy SI'uno:(HJl{gd) '186 '8nV 'alprsftooJ 'r 'uonIBJ 7919taqua 1t1y t ZZ Jo t&NN tu{ tOI Jo lds tu)tZI :(HJUAd) uloN llV'996I eunlp'ur.ruflun1q o1pEor uo urperctl JoN solrul9:(HIUAd) 09LI tuaqBDX t9 '81,6I '8urqqoundloo11o 's1lg '8L6I ,(sI I 9I g1r1 urqyc8unq:(EJuad) pg91tuaq3oS1 t 9 AEINZ 'urTrlorla/t\Jo N'e^reseg11t113utdooc:BppltlJ:(IXAd) tgEg.raddotI OS'886Idlnlt'g,II .rII S,ZI oZ€'uolloojgJoANu{ gZ'{Bad(urqutsn0),{tunoJJo sedots IAN:(Hll{gd) g7p9nddo11 qg '986I 'n, 'pue[oH rdcAg tO "6I l S,ZI o€€ l] I Jo,^dN :(HJUSd) EZZ; laddoll Q'S'98618nY LZ 'a,gt.gII 'pu 'c s.z?.zt lstraqErcpad tuo{ p5 q1no5urre31e1 3uol3 s tu)ttl :(HJxgd) z0tt '4,9S 'o^r0sel{ernl?N nddo11q g'996leunltI .LI I S,,0t,L0 oz€'urluqcgJo rld tu{ 6'8 s,uosualoS :(HJUgd)96€t./r./doH O S'986Ioun I EI'A,LV.Ltt S,50.Zt'qqog lhuo idru{ t:(HIUAd ZZII 'ld3s 'a,9t 'IOou '1coa nddoll o S'BL6l 9 .8I I S.8Eo0€ cluuoglo gsg txl 6€ Sutuourqe,{p16 '696I '^oN 'ultuuel '696I 'ururtu?J :(HIXAd) a^ar.t9 ['g gZ :(fffXAa) a^atr1 [ g ,(8]l 0Z 'utqsx?3 'V') :(HJXAJ 60LI uaat7 A t' L96I 3nV I t Jo A s3[Itu€ :(HJXgd) Zt06I )auprD) '996I 'oetrng '616I '3ut11nqel1nS lro 8 t cry1 :(HJdAd) I LZ6nup.tDt V J fulq 1g :(HIUgd) 'llrH '3ut11nqe11ng puattl V l'886I ounl I unouu) JoN tu{ g :(HJUgd)?, DrI If t'8261 8ny '9.89 ':r8uoo1rl N :GJdgd',\\S N) (.aUaqDJ|g E ILtBtuz^oC )1'9t6t tdc5at .8I S,0Z.1g Jo 'Bluolscld '()troU :(HJ)iAd'SNVJ) Z6 alDpuaddulJW t'L96I qcrnl^I9 JogN sc[tu , ploJuos '(tr.r) =) s{JoU.ur{ur{ro^ :(HIxSd'SNVJ) 68 apPaaddryJ'W'J'L96I qrrcr{ 9 utqlrr?JJo A scuurI'S:(HJUAd'gNV)) SgalppuaddtLlCW g' L96Il:jutL[g'ulpclowJo,^dSsclttu 9't:(HI)Igd ',4dSN'1AN 'SNVJ 'OV) '9,8[.tII 80L6.tntoo.tgIf IW'LB6l Atnt9I S,80oZ€'Sulporlon} ',^(SN"ISW 'SNVC 'qV) Jo S ru{ It :(HJUSd t0L6.t too.tg'lIf W'L86I,{lnf gI'Sulp?lmno ',^[SN'-rAhI 'SNVJ) Jo S ur)tZ :(HJUSd 08E6l loo.tg'lf|1l'986I ,(tnlZ'g,gI .rII S.09"08 'psor - ',(SN "ISN'SNYJ) crucdJoord lrqquU uo puolsproo) xnopBJJo S tuL't:(HIdAd '986I 'uut 'Sutudurl SgL6rDloo.tg H I'W 9Z'9,9, "9II 5,71"19 Jo l(\ tu{ I€ :GJUgd',,\\SN '1AW'SNVC) '986I WI6t lootg'If fW uulgZA,90.rtI S.ZI.It'Lucqr1c1lr:{161o16ur4 1g 'gNVJ) 'H 'plror :(HJdAd"\\sN 8ttu -/27arr8 1't41'186Iuvt 9Z'g,rI . I S,Z9oOt llc{Bcrguo ' 'qag 'frnpunoq peor8uru,{ar1.;o 16 tu{ t'0 :(HJUAd'ASN'SNVJ)916r./Dlootg lf l W E861 tI ',^\SN'SNVJ) '286I g'c^resad crnlBNSurdoocrrpprq3 :(HJUAd SZ9, talootg tl In ldrs I 'a,gs.LII 'rurl7{nN 'r s.6 t oI t Jort[ url 9z :(HJugd "/y\sN'sNvJ) 029L nloo.tg lf f hl'286I 'ldcs 'A,SZ 'pll SI .8I I 9,97.19 urru8unNuo ulpollowJo N ruI S 6 :GJUAd',t\SN'SNYJ) ,1,t9 'qrulJllng .tattoo.tgH f W'6L6I 8nV?Z'g.tl .6II S,r0 .I € sprt,lnolssorJ uraqlnos Jo ldN url t SI 'SNVJ) 'S,00 'srpm8loo:)Jo :(HIUAd'./tSN I Ltg ntlootg flf 141'61613nV €Z .0ZI S.9I .I g ld Lrr){EZI :(HJdAd',,\d.SN'gNVC) 6889 ra{oots IW'6L6I 8nV0Z'4,20.0ZIS.9I oIg'ullppog 'gNVJ) 'tucqlel Jotd urt 9'ZZ:(HJ){gd I69Z irloo.tg 11[ W'0161,{lnf 9I spro./!\olulrreqroltex '1aN'SNYJ) '016I selru gI :(HJuad 6ZlZ t22lou8',HI n u?I 9I'(cls) {ro}I proJpu€s Jo N s 'n :(HJ)IAd) u'ssrrrog l,tl' wp s'RurutzlJ'Iueqlo^BIA urrud el?puexo :G{JUgd) t}S rtldv A t '696I 'urcqc1e1p.{16 'pautwDxa aunl gg Jo N satlu Z :YITy11JS1Y NIIAJS:I11 saawaadg

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'r3ddoH 'O aparyufN - ueppw ,rrrr adstt"T sa]dassniKtDrn? uaqdals T ra{oorg HIW Nuytsiavol. 8, No. I (1991)

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Figure 17. Holotype off. capillosaBrooker &Hopper subsp.cdpr'ttosa. '986I '$lrol{ eunl tI'S.gf .8II S,ZZoZ€ roue^ rsJo,4[s url 8:GJXad) EIl, raddoH'o's '986I 'A,6t.8II 'slcol{ ounltI S,,0€.6Iozg reuo^Irrs JoN 'x:l t'0:(HJxad) zlrr.nddoH os 'S86I '9.9S "IIII^I'SNVJ) eunl tI "8I I S,,0g,IZ.Z€'sdurnH eqJJo ids tu)Ig:(HJ{Ad'/\&SN €S96 'qlruspqlJog'peo11qAnoroqesogJo r loot{'Il IW'LB61 .{e116y g uDIg'0:(-1!trr,1',41SN'HIXAd 'SNYJ'qV) '986I 'p?or - IBV6r looig'H'lW TcOrc'A,r0 .6II S,rt oZ€ IroU ltoH 8uu€8uld 'nd.sN'HIUAd '[INY3) puu {cor uee^ueq's^lasauIJol{ uo8€l( :(14 I ZIZ6 r loorg'H'I'W '986I 'ereqlrd 'r '11&sN"E 'gNVc) qrrul^t gI Jo s tu)ts'0 :(HJusd ^I t9I6 tapoq H I 14 '986I uelgZ'A.90"LII S,ZI oI€'uaqclewdidJo i txlIe :(HHgd'l1[sN"IahI'sNv))7816 're(I "I! Dloo.tg H I'/I'S86I 8'A,02 "LII S,LZ.ZE'ululpuoXJoANN:(HJESd'A\SN ^l'SNYJ) '^oN "IahI 9906Dtootg H I r{'s86I t':I.0t.9I I s,9Zo0g'€rsqlldJo s uq I :(Idldgd'/\[sN ' 'a,82 €rNy))gTgsnloorg'H lW'r86l EnV6 .61I S,€ZoZ€'{c?Jl rlsou{Eooqs uo {3PJl uep^H 'uapfg uro{ uDtt'9 :(gNyJ) tZEgrr{oorg'H I W'6161'8ny g1 3oA uI g gI :GIUAd) 9t9Z t loorg H'I w'0L61 .{1n171'uepdgspruaol{3ol{lloHJo li\N sellurg:(Hldgd"I!MI'SNVJ) z99z 'uop,{g3o raYoorgH I W'0L61f1n171 g selur 9 8 :VIIYUISnV NU!IJSE,^&p dulaDxa suawlrdds 'sdelpl puorq3o ssdols ',{pues 'llq?q ,(11e,re:E sardnrJopu?'lpl tx 9 ol eettBuoqt ,{q sorJodsqns pclddl eqluro{ sreJJIplI '(99I 'uo!tDusnll! :066I) Slulalx d' ror{oolg rnopJ

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Figure 18. Hololype of E. capillosa Brooker &Hopper subsp.polyclada Brooker & Hopper. 'proqnJ 'u,^aoJq ,{lerBrol Isouoqdsqns 'peuesxe 'rrJ 'JBIndnJ 'etnral 'paxogur tqEIIpeas lou se^p^ S'0x ,'0 ol lrnrC uooslou sJe,trol{ Ip 'lrore 'urt 'urroJrsnJ ,{1a1e1durocro,{1ged seuo reuur sueuelsJolno ouos t'0 x 6 I ol ro proluBII?spng 'epl,r 'pere1(ou-I ruc g'0 ol seljunped I ol saJuecserouulueerS-,te:? ,(yemr :o ,ter8dlsnonJrdsuoJ 'llnp 'ul' 'el?loecuBl'Suqsuroll?'eloloued 'f:Ieq 'uee:3'rur 'ale^o S'I x 0I ol so^?o11ynpy g't x I ol 'SuqeulellB usql's{ed9-?roJolrsoddo Sulupue:8u1pees eqtJo se^Be'I e8u?ro lqEuqre^o,{a$rysg 'snocnq8 'relnpuB13 'suels 1ouEalqJu?rg slelqcuaqgo qld qloous qlr,r oo4 II€tusJo oolp] I

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WrrlIiIl1*l SOUTHWESI .i i- r' \:rti I I lir i iii l-.1

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Figtrrc 19. Eucalfpus L ,idd distribution, buds, fruits, adult leavesand silhouetleofa fifth node seedlingleaf(scale bar I cm).

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Specimensexamined. WESTERN AUSTRALIA: N of Bullabulling,25 May 1964,J.S. Beard3336 (PERTH);44km SEof Coolgardie,3l' 15' S 121'28' E,9 Nov.l98l,M.l.H. Brooter 7055 (CANB, NSW,PERTH);8.9kmW ofCoolgardie Norsemanroad onHyden track,32' 0l'S 121' 35'E'7Nov. 1983,M.I.H. Brooker 8351(CANB, NSW, PERTH):74 km W of highwayon Lake King Road, 32'42'5120" 48'F,,l2Feb1985,M.I.H.Brooker8845(CANB,MEL,NSW,PERTH)iNWofNorth Ironcap,E of Hyden,32" 20'S ll9" 37'E,24 A:ug.1988,M.l.H. Brooker10054 (AD, CANB' MEL, NSW, PERTH);North Ironcapsummit,32o 21'S 119'40'E,M.I.H Brooker10060 (AD, CANB' MEL, NSW, PERTH); on Hyden track betweencross roads and Marvel Loch' 8 Dec. 1966, S. Chambers184 (PERTH);46 km from Coolgardiealong Eyre Highway towa-rdsNorseman, 31" 15'S,121" 29'E, l Feb.1979,M D.Crisp 5624 (CBG,NSW,PERTH);Hyden-NorsemanRoad' 135 milesW of EyreHighway, 6 Feb,1963,A.S. George 4334 (PERTI{); 21 km NW ofHoltRock P.O.on tlack to Hy den,32"32'S 119" 16'E,5 Oct, 1976,L. Haegi l20g (AD, PERTTD;96.6 km W of Norseman-CoolgardieRoad on trackto Hyden,3 km w of McDermidRock, 32' 01'S 120' 46'E' 7 Nov. 1983,K. Hill 615andL.Johnson,D.Blaxell,l.Brooker mdS. Llopper (CANB, NSW' PERTH): 115.7km E oflake King,32' 41'S120' 42'E, U Sept.1976,R. Hnatiukl60778 (PERTH);' Kumarl Road,l6kmWNWof Kumarlsiding,24.5 km NE ofPeakCharles,32" 44'S 121' 20'E,6 Sept.1982' S.D.IIopper250l QERTH);9.6 km SEalong mainroad from DiggersRock toHatters Hill,32' 45'S 119"36'E,30 Aug.1986,5.D. Hopper 5305 (PERTH); Hatters Hill,32' 51'S 119'59'8,30 Aug. 1986,SD. I'lopper5308 (PERTH); 15.9 km S of Mt Hollandon LakeCronin-Southern Cross road, 32" 18'S 119'45'E,2 Sept.1986, S.D.llopper 5402 (PERTH); Top slopesof NorthIroncap, 33' 22'S 119' 40'E, 3 Sept.1986, Sr. tlopper 5430(PERTH); 13.5 km NE ofHattersHill onLake Hopetrack, 32o 44'00'5 120"04'00'E, 28 Sopt.1988, S, . Hopper6868 (PEF.TH); 5.6 km NE of HattersHill on LakeHope track, 32'47'30'5 120'01'00'E,28 Sept.1988, SD Hopper687 | (PERTH);Hatters Hill, 3 Sept. 19?0,K.R. Newbeys.n. (PERTH);North Ironcap,7 J:ulyL919, K. Newbey5213 (PERTH);29 km SW of McDermidRock, 114km W ofNorseman,15 July 1979, K.Newbey5304 (CANB, PERTH); 0.3 km SEofHatters Hill,40 km NE oflake King,8Aug 1979' K. Newbey5449 (PERTH): 3 km NW of LakeCronin, c. 83km E of Hyden,17 Jttly 1981,KNewbey 8305 (PERTH); Gibrallar,c. 24 km W of Coolgardie,11 April 1966,P.G Wilson 4102 utd S.G.M.Carr (PER'|H); near Gibraltar, c.22 km WSWof Coolgardie,25Aug. L968,P.G.Wilson182 PERTH).

DistributionandhaDilcl. Ccntral and southern goldfields, particularly from Coolgardie to Norseman, and southto thc Kumarl - Pcak Charlesarea and west to the North Ironcap- HattersHill area (Figure19). E. livida growsin a pure tall malleeformalion, usually confined to small elevated breakawaysof decorrposinggranite or rarely bandedironstone. Detailcd ecologicalnotes on associatesarrd habi tats of E. livida arc givenin Newbeyand Hnatiuk (1988: 24, 27,99' 111)'where the speciesis refen'edto asE. aff. wandooor E. aff. redwtca'

Conservalio,tr/a/r.t. Locally abundantbut in widely disjunctpopulations on vacantCrown land. Poorlyrepresented on naturereserves,

Floweringperiod. Unknown.

Etymology.Theepithet refers lo the canopycolour (-atin /rvidas,blue or leadcncolour).

Notes.E.lividq,which maybe a treeor mallee,is theeastern and south-eastem representative of 'tr0proqnJ le^ urueeqdsqnsturueg uorlsn8uuorqluedfq eJn ado rlcqoq,crpur^-oprlled,ululoJJustu1lnpc [lloC 'auq?t8 'oxry str1uEId'opr^€Ulcdns oosu8 o:dsuerorregur eJnJoJ .snlp u g pe..oollBuJ,,xeln.rC

,:eddog (92'92'tZ sorn8rC),iou'ds 4 ralroorgDtlottpqaq sntdtptng.S 'opun{u8ru'g pueosolltdoc A uoo.rloqsopurS:e1ur aq ,(eurpuu llnq cql 'sorzuow IJ€lq {rcl ((HJr[Id) Z€06InaUDg trJ Jols.^r p uB ccJB eoJreg ery1cq1 ur'3 0)lsem eq1 o1 suoqplndod'l:osc6elJolcr4 leotg eqluor,J soruoc utrol lccrd,{l eql.,{Jrcq sSurtpoes eql puD snoJntrla or€slelqJueJq oqJ nsolltdot E pre oopuDtftg u€qllln{ puespnq,so^ecl rellutus qlr^\ pu€Ircq qBno:psrq lrup q1r,udllcnsn ocr1 B sl lI oopuc,rUcsop porsaocsrp ,{11ue:sr c s r oputrlu7tu.V salop

'(uonoq 'rrpr,y' pu€IJulq'r.rtu uge-I) nq )tlBpoqt o1sjegc: lcqlrdo crl lneds a\L.tBoloweg

'u,roulull pouad Suuauo111

'a^Jaser sJnlDuc uo psploJclloN puq u,\\olc luesc^uo ',{a,\lns suouplndodlJUn fsrp,^roJ uto{ u,rou),I JoqlJnIJopccu ur lnq,oJnl,{lqrssod.sn:rts uouDMasuoJ

'suruld pur sodolsluecrfpe pue elruu:8 Sursoduro:cp jo s,{u,1\eltcJq ol pcuUuoJpu0lpoo,^\ ,rol ? sruJoJlJoroq,{1.'(27 ornBrg) gcscq nrrolcl^ luclg cql uorl .{1uou,nrouy rrlrqDq puD uo!rnEl$!O

'(HJUqd) 'uosnu g puowq ll'IL6I ^oN r r{lt-I3o5 ,{:1unotxcJrurds ,4puns '.Iaw 'gNV)) ,lrosco ur uorra^?'I:(Hluad'A\sN 1898n\oo.tg t w'ng ,(onl euolrr^ 'pautwoxa 1uu9'dr:1srrego 16N ru:l SIZ'uoscg oID'I Jo A\:Vl.lVUJSnV NUAJSA16 saawnadg '(ur t'0 x S0 oD lln{ pup(urJ Z 0 x I ot) spnq'(urrS I x I I ol) sc^cclllnpp rclleus '{Joq 'q3 '1csuq aqtpuB sSurlpscs lucJsoqnd oql Icnlq no: cql^q flo]0lB dopilrlt g ruo{ sJaJJr(I

'(1.91 'uottD.I$ry11 :066.l)SruIct>t d' ro{oorg .tttoloJ

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'pajyp snquor,rrtldenbsnq4cn.r3 sulsuqele'snlnpe sr!loJlc snqrluof,scqndsln1ue1d'rseq orElu oleJrlJocep-uou oc4roc ,41e1e19 oop uDtr oldf,ponA qV

'raddo11 (gZ'ZZsorn8l{) ,rou'ds 4r n4ootg npunluSrusnfitprng', 'punoJE ,(uolspal?^olo uo ,{uelnJnJed,{IunoJ ,4Jpur FrluotodIBJnllnJruoq '(IZ seqlI emArC)uoqelo8e^ Jorllo o^oqu luaE.roruc ,tlpnsn lueuruop snonJrdsuoJ c sr Dpt^ll fl

',funq 'salJunpod eres8uqpees eqJ lrn{ pm spnq lsnqorelou prrusolaol 'slelqcuaq '3' ,(e:3,(13ur1rgs snoonulS-uo u xqtu!DSollldoJntl]o{sJeJJrptI ,,oop?r,n,, serceds:edns

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Figure 21. 8. livida showing mallee habit (NE of North lroncap)' 'sllnlJ 'spnq = rsq apcs) Pel Surpeesapou rlIJ e Jo atanows pu?se^?al Unp? .uoonqulstp ,lzlrfr Stusnitp"ng .ZZeJnR$

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-;;-1i ";::.:'.1:4:.-1 .i.""-- i\t t tt r' :.::.

Figurc 23. Holotype of t. ieriJunda Brcoker &Hoppel

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equalsthe hypanthiumat thejoin, E. abditaappers notto be closelyrelated to E. phaenophyllaor E. arschnqea.Itsrelationships require further invcstigation.

7. Eucalyptussparsiroma Brooker &Hopper, sp.nov. (Figures29, 30, 31)

Frutex("mallee") ad 5 m altuscaulibus exilibus et canopiisparvis terminalibus. Cortex peculiads, superlatera occidentale in taeniisnon dccorticantibus tenentibus, orientale laevis vel omnino corticae aspero.Medulla ramulorum glandulifera. Folia adulta anguste lanceolata, ad 7 x 0.9 cm, nitentia, viridia. Inflorescentiaeaxillares, ad 11-florae; pedunculi complanati. Alabastra pedicellata, ad 1x 0.3 cm, operculodistincte hypanthio angustiore. Fructus pedicellati, cupulati, ad 0.8 x 0.5 cm. Seminasubsphaerica vel cuboidea.

Typas:0.8 km E ofChinocuproad, Western Austmlia, 11 Janruy 1988,M.1.H.Brooker9850 (holo: PERTH;iso: AD, CANB, MEL, NSW).

A malleeto 5 m tallwith slcndcr slcms and small lerminal crowns of smallnarrow leaves. Bark sometimesunusual wilh thewestem sides ofthe slems covered with non-decorticated ribbons ofrough barkwhile the easternsides are smooth pinkish grey and light greyiother mallees with roughbark allround the funk. Pith ofbranchlets glrmdular. Leaves ofthe seedling remaining opposite for 2 pairs, thenaltemating, elliptical to ovate,to 7 x 2.5cm, blue-green. Adult leaves nanowly lanceolate, to 7 x 0.9cm, concolorous,glossy, grcen. Inflorescencesaxillary, unbranched, to 1l-flowered. Pedunclesflattened, to I cm long.Buds pedicellate, fusiform, to 1 x 0.3cm, with operculum distinctlynarrower lhan hypanlhium.Some outer stamens orect, inner onespartly or completely inflexed,all fertile.Flowers not seen. Fruit pcdicellate, cupular, to 0.8 x 0.5cm; valves not cxserted. Seedlight grey-brown, subspherical to cuboid.

Specimensexaminerl. WESTERNAUSTRALIA: type locality, 11 January l988,M.l.l1Brooker9849 (AD,CANB, MEL, NSW,PERTH); N of Nyabing- Pingruproad on Chinocup road, 8 March1988, M,II L Brooker9902,9903& S.D.I lopper (AD,CANB, MEL, NSW, PERTH)10.9 km Eof Tincurrin Nth roadon StockRoute 3, 32"56'S 117o 48'E, 12Decembcr !988, M.l.H Brooker 10144 (AD, CANB,MEL, NSW,PERTH); Pingrup,22Feb. 1952, C.A. Gardner 10322 (PERTH).

Distributionand hablrat.Known only from nearTincunin andlhe southernside of LakeChinocup (Figure29), where it growsin whiteorpale brown sand or sandyloam on flat to gentlysloping terain. AtLakeChinocup,it formsan emergentshrub mallee formationw\1.}]lE. scyp hocalyx, E.aff. flocktoniae, E. aff. occidentalisalnd, E. phaenophyllastbsp. phaenophyl/a over hcath or low scrubof uncinata,sometrmes over a BoryasphaerocephalaR. Br. herbfield.

ConservationJ/a/rd. Requiresfurther survey.Appffenlly geographicallyrestricted but locally abundantand represcntcd on a naturcreserve.

F lower in g p eriod. December-March.

Etymology.The specificepilhet refers to thecharacteristic crown (Latin sparsus, sparse and cama, crownof leaves). .(TUJ I '.uorFqutsrp .62 = sreq ale.s) leal 3u!lp3es epou qUIJ E Jo atFnoqtls pu? Fel llnpe'sltruJ'spnq ,rzorrs./Dds srudqort4 ern?lf

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Notes.E.sparsicomahasglossyleavesandtheopelculumnalTowelthanthehypanthiumlite n. phaenophylla,butdiffers in its terminalcrown, rough balk hcldin looseribbons, smaller buds and its smallererect leaYcs.

8.E ucallptusaruchtt d€a Brooker & Hopper,nom. et stat. nov ' E. r edunc a Schaterv ar ' melanophloia Benth.":Fl. Aushal ." 3:253 (1867).Type:Chceanga thicket, Murchison River, Westem Ausfralia' Oldfields.n. (holo: K; iso:NSW)

A malleeto 7 m tall or a trecto 10m. BaIk loughand tightly heldon lowel partof stcmsat least, darkgrey ro grey-black.Pith of branchletsglmdular. Leavcs of theseedling lemairing opposite fol 3 or + pairs,inenalternating, deltoid to broadlylanceolate, to 12 x 5 cm, bluc-greento glaucous' slightly or very hairyparticularly on margins'Adult leaveslancoolate, to 9 x 1'5 cm, vcry glossy gre-en.inflorestencesio 13-flowered; pedunclcs to 2 cm long.Buds shortly pedicella(e, fusiform, to i. g ^ 0.3.r, op"."ulumnirnower than hypanthium at the jo in,horn-shaped, usually recurved at thc tip.Flowcrswhitc.Fruit shortlypcdicellatc, shorlly cylin&ical (iustlongefthar bfoad), to 0.7 x 0.5cm. Sccdlighf brown, more or lessspherical (Figures 5a, 32,33 '34,35).

Etymology.A ncw specificepil.hct is rcquirodfor thistaxon because the varictal ]name melanophloia uscdby Benthamis alrcadyin useat lhc specicslevel, i.e. Silvcr-leavcd ltonbar* E. melanoPhloitt spidclyclustors F.Mucll. of Queenslandand northern New South Walcs. The ncw namc lcfels to the of budsseen particularly in lhc frcshstatc (Lalin a| achnaeus,sp\dery)' "phaenophylla", Notes.D. arachnaeabclongs to thc supcrspccics I gloup ol'spccicslhrt is characlcriscdbythc opcrculum bcing clcarly narrowcr than thc hypanthium alld by thcglossy grccn lcuvcs.E. arachnaeais widcsprcad allhough sporadic as it chalactcrislicallyoccupies low slonyliscs or latcitic brcakawaysas in thc MorscbyRangc. It ditfcrs from relatcdspccics (E. phoenophyllq, E. tumirla,E. histophylla)by therough bark in maturcspccimens, vcrrucosc seedlings and morc dclicatebuds which arc oltcn scer in spidcrymasscs wilhin thc crown'

Thcrearc two subspecics.

8a.Eucalyptus araclzaea Brookcr& Hoppcrsubsp. aruchnaea, subsp' tlov '

Calourillustation. Brooker & Kleinig(1990: 181).

A malleeto 7 m tall (Figurcs32, 33).

Specimensexamined' wESTERN AUSTRALIA: 3 miless of Cz.Inamah,22May|962,.1..E.11. Ap|in 1480(PERTH); Berween Dongara and Norlhampton, Sept I932,W.E Blackall (PERTH);15 milcs from Northamptonon Lynton road, April 1952,G.E. Broclcway (PFRTH); 5 milesE of Brookton' 22 hly 1969,M.LH. Brool17.y :(HJNAd)u's rr,lolJ 9'996I ldcs8Z'eo8Dq eI I:(HJ-Ilg,d.)nupntT aurDJ.VZ6Ildog,s11y11 us8uo^\:(gNVJ)8 6g7aqptq.ng 1 p' Lt6I ldosZ,uolptueoJo N salluZZ :GJUAd.,^[SN,.IAhi 'SNVJ'(v)08001 n2lootg Lff W'886I ldrs0Z.9.91.SII S.00 ernloNsqqseug '{ro.rl oot.e^rosou qlnos s-N uo(ld-o oruoJJoN u)t I :(HIUAd'leSN'TgI I'SNVJ'qV) E g6atoo.tg,H.l.N '886I usl II'iI.IT, "LU 5,61.gg'ur8e6goA\NrxlI r:(HJdAd.,4 ,SN,.IqN,qNVC,CV),596 ' '(pso{ ,su.ro( nlootg H I W Lg6Ieunt ? rpr8qloolJo S) fi g0I uolleJo-I?Holcl^ prru,rauooJ '{ropped ',$SN 'HJl{Ad 'SNVC) ,986I dlcor :(.IU.{ VLt6 ralootg'H I t{ ounf LZ,S.,g0"LII 'a^rosaU 'l1[SN'HId'Ed'gNy)) S.€Zozg ornl?Nureoid:(TAhl 9LZ6r gIW,9B6I AeINg 'uoldu8qtroNJo.l1['peog.(uourr11rq3 loo.tg ruo:;puog qog 6uop ur{ 6 €I :GJUAd'l[SN,TAhI.SNVJ) 'tI'I 's86I 'a, ,ouurv ,nsN ..IaL/{ IE06 r loolg w cunl g Lz o9I I S.LZ06Z Jo A\s,/\\:(Hluad 'gNvf,) ?I88.ra?oorS ffI ht'986t qodt'A,90.9II S.t0 06Z,reuroHll^lJo AIUH:(HJUAd,.4d.SN 'gNy))EZLSDllootg'gIW'Vg6I'1xOIE'A,6E "r g.gg.99,1 .{u.rq8rq uo uoldruuq.uoNJo N url 'E86I 9'02:GIUAd'nd,SN'SNVJ) 98I8 ratootg H I W ounl6Z.g.It oglI S,I9 o0€,€el?sttrH ',(?^rlrer 'SNVJ) uuEuol61r1 pu?esnoq s,de,rruo3 ure,rleq :(HJUAd'/$,SN gEIg.tDtoo.tgltf lltt '€86I 'prror ,SNYJ) ,iuhl ,Z uo^rpurrJ uo a8puqre,rrg uosrqrrntr{Jo N ru{ 9 I I :(HJ){Sd'/tSN 'a.9e .uoldueqtroN 6118t lootg [I'fW'e86I ,{ul{ SZ "tII S.tg o8Z pueuolptcrog uoo,rleq 'duC 'SNV)) 'A,€€ ?Jreqle,roH:(HJUAd',t[SN 9g6Lntoo.tg H'f W'Eg6I'gog,9l .9I I 's11r11uuEuo16 5.69 .0€ spru,r,rolur8ulp3;ogp ur{ g'91:(HIUAd'A\SN'SNVJ) gt6r rrl oorg l! J I4l.Eg6I 'uBI '9.0I 'pBoU .l\SN ,gNVJ) 9Z .SI I S,S0 .62 sturng uo djJeq,rBrsJo tdN ru{ €Z :(HJUAd 'Eg6I'ueI .uotdur?rtuoN 6E6Lnloorg'H'I'W SZ'A,ZZ"V\ S./l .gZ puera^rU llnH uoe^\l6g :(HJuad'./dsN'sNvJ)EEgL'zEgLt tootg H I w'286r'ldos 'a,6r 'nd,sN'qNvJ) sI .Lrr s.8tor€'prrrrol. uoulrxrx€J uroq ur{ 0t :(Hluad I}gLr tootgH I w,286I'arryg(,,a,87,.gu S,t9 o0g'8urue^\BrdSurtt:(H11{Ad',ld.SN'SNVC) 869, npotg'H'lI I'Zg6I'6nV92,A.8€ .9II 'reAot 's1151 'id,SN 'gNvJ) S,ZSoot orp?r ol p?or ueEuol6:GJUAd g68st nlootg H'tt[ '286I '8nV 'g,It 'ruBr€J ,ldSN 9Z "9II 5,69 o0€ l? Surssorclpr Jo g ur)t 9 ZI :GJUAd 'gNVJ) I8Sr'089,nlootg'H'I W'ZB6I 8nV9Z'A.99"9II S,69o0g.tu8rsJJo 9.,{e^\.HUrequoN l?3rcJog ur{ 8z:GIJrEd'tdsN'sNvJ) \Lg Lt loorg'I.t I7tt1'7961.8ny61,eue8ur3peglo 1r1,p1 '286I 'rpurg rpreqlooJ:GIIUAd',^NSN'flNV3) IESLDlootg H'l W .{sI I IZ ryurgJo fl ur{ 9.ZI :C UAd',^d.SN'SNv))IZS Lmlootg H I W'286I IEINIZ'pII p€oqueer9-^\orooOpuu pU e8pnw

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31 Aug. 1980,G.,r. Ke ighery 3334 QERTITI; c. 15.4km S ofNorthamptonalong Nw CoastalHwy' 28' 31'S 114"37'E ,21 At:g.1983,C.M Ilnclr 66(PERTH);HowathanaHillResorye,2l road miles N of Geraldton,25 Oct.1974,D. & N. McFarlandNMlOg?(CANB' NFLD' PERTH);ibid,9Feb' 1975,D. &. N. McFarland NM1087 (CANB, PERTH): SE from Po Gregory,26 May 1952' Road, D.H. Penny@FjR"]:F'); Watheroo,4 Nov. I954,R.D Royce4941@ERTII); 15 miles Lvnton 1 March 1966,E.M. Scrymgeourn3,274 & S.G.M Carr (PERTH);370 miles NW Coastal Highway,3 March L966,D.M.Scrymgeour 3L3 & S.GM. Car (PERTI{);

Distibution andhabitdt. Westeln palt of northernand central wheatbelt, to nolthof theMulchison River in the north-west,and to wagin in the south(Figure 32). Occursin tal1mallee often with E. pluricaulis,E.erythronema,E. ancep s,E.flocktoniae ,E. pyriformis,E' astringens andE gardneri, on gravellylateritic breakaways and slopes.

Consenationstatus.Widespreacl ancllocally abundant in disjunctpopulalionsthatarewellrepresented on conservationrescrves.

F lower i ng p eriod. APril.

Notes.A malleesubspecies wcst and nolth-wcst of subsp.artecta. Alot'tgwilh D. subangusta,lhis subspeciesis a commonmembcr of tho seriesthroughout the northcmwhcatbclt'

8b.Eucallptus arachnaeaBrooker&:llropper subsp. dff€claBIookcI& Hoppcr, subsp. nov. (Figurcs 5a,32a,34,35)

A subspecietypica habitu arboreo altiorc (ad 10 m) et foliis adultisnitentioribus'

Typas:west of Morawa,29' 08'S115" 44'E, Wcstern Australia,3 Feb. 1988'M'lII Brooker9879 &. C. Sounness(holo: PERTH; iso: AD, CANB, MEL' NSW).

It differsfrom the typicalsubspecies by thetree habit to 10 m tall andglossiel adult leaves.

Specimensemmined. WESTERN AUSTRALIA: typclocality, 9 Sopt'1987 ,M I 11 Brooker9757, 9?60(AD, CANB, MEL, NSW,PERTH).

Distibution and habirar.Known only from thc typelocalily westof Molawa(Figule 32) wheleit formsa low openforc stw\thq. subangustastbsp. pusilla. Thesite is a shallowcreeklinc on terrain high in thelandscape. Broombush (Me laleuca uncinata) is promincntin the undelstolcy.The soil is gravellyloam.

Conservationstatus. ln urgentnced of lufiher survey. The type locality hasbecn left uncleared deliberatelyby the presentowner. This is thc raresttaxon known in the senes'

Flowering period. Unknown.

I I

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ate.su,{W - ueprcn ,rnrads^z7 sauessntdqDrng laddoH O ueqdersd, re{oor8 .H.l.ll Nuyrsiavol. 8, No. I (1991)

Fig]ure35. E. arachnaca subsp.arr.".rd trec habit at type locality, and hrnk and bad<. '(ruc I =req atert lEatSunpres 'sllruJ'spnq'uorlnqulsrp apouWJrJ s Joaltanoqln pu? sa^pal llnpp ,rr{fdou2rqd dsqnselltqdouatqd sndqDrng gt atnFlJ

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'reddoH ae'srsu,q{ - uaprBt{ ,r'1.!aclsr,r"l seues sxldtlo2l1A C uaqdals T raloorg H I }\l NuylsiaVol. 8, No. I (199r)

atrR00iyFr0noF fo36r {r!rraa}i ti,

FiSure. 37 tlolotype of E. phaenophtlla Brcoket & Hopwr svbsp. ptaenophltta I

'4req pue srualspuv'(decuorl quoN Jo iAtD trq?qaat]FtlJD (qdoua td dsqns,/tqdo "rfd t'8[arnBIC

eeoca}rtrN- urpr?t^ ,r!z rddst^4 sauasstlldtl'rrg l.ddoH Orcqdais Treloorg HIW NuyrsiaVol. 8, No. I (1991)

Etymology.From theLatin arrect s, upright,alluding to the hee habit.

Nores.This subspeciesoccurs to theeast of therange ofE. arachnaeasubsp. arachneea,fromythich itis completelyisolated (Figue 32). Itis a strikinglree to 10 m tall growingon very stonysites with avarietyof associatedspecies includingE. stowardiiMaiden,E. oldfieldiiF -Muell, E. subangusta subsp.pusilla, E. obtusilloraDCurd E. ftoristesJohnson & Hill, all ofwhich notablygrow on sandy soilselsewhere. E. a rachnae a stbsp.areoa iseasilyrecognisedamong thesebecause ofits treeform, heightand very glossyleaves, smaller than those of the equallyshiny leaves of E. stowardii.

9. Eucaltptusphaenophylla Brooker & Hopper,sp. nov. @igures36,37 ,38,39, 40)

Eucalyptoarachnaeae Brooker & Hopper affinis, caulibuscortice laevi, plartulae minus Dubescentibus.et floribus flavidibus differt.

Typas:N of Nyabing-Pingruproad on ChinocupRd, 33'34'S, 118' 23'8, WesternAustralia, 8 March 1988,M"LH Brooker9901 (holo: PERTH; iso: AD, CANB, MEL, NSW).

A malleeto 4 m tall with greyover coppery smooth stoms, occasionally wilh partlydecorticatcd ribbonsor flakesof barknear base. Leaves of thesecdling remaining oppositc for 3 or 4 pairs,thcn alternating,ovate, to 9 x 6 cm,bluish green, glabrous. Adult leaveslanceolate to broadlylanceolate, to 10x 1.5cm, glossy green. Inflorescences to 13-flowered;pedunclesto2 cmlong. Buds pedicollato, fusiform,to 1.8x 0.4 cm. Flowerspale yellow. Fruit pedicellate,obconical, cylindrical or barrcl- shaped,to 0.7 x 0.5 cm. Seedlight grey-brown,more or lessspherical.

Etymology.The specificepithet refcrs to theshiny leaves (Grcek phaeno, shrny and phyllon,leaf).

Notes.E. phaenophyl/a may be seen as a southernsister spcciestoE. arachnaea,from which it diffcrs in the smoothbark (which may often be blackish)and non-venucoseseedlings. The massed inflorescenceclusters ggnerally lack thespidery effect of E. arachnaea.The narrowedoperculum seenbest in E. arachnqea is still clearlyevident in E. phaenophyllastbsp. p haenop hyll a blutis lcss pronounced.This charactermay be seenparticularly in theyoung buds o f othertaxa, yiz., E. tumida andE. histophylla.It is a variablechara cter in E.p haeno p hyllas|bsp. interjacens which we consider to be a form linking E. phaenophyllasubsp. phaenophylla and E. rumida.

Thereare two subspecics.

9a.Eucallptus phdenophyllnBrooket & Hoppersubsp. pl, aenophylla @i.grrres 36, 37, 38)

Colourillustation Brooker& Kleinig (1990:182).

Stemsmore orless straight. Buds to 2.5mm diam;operculum always conspicuously narrower than hypant}liumat join.

-. I I

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'raddoH .Earystr,at - uapIBI^I ,rkzads^"7 seu.s sntdqDrnq ( ueq&ls ? ra)toorg H I I^t Nuytsia Vol. 8, No. I (1991) oft),27 Sept.1975,J.W. Green4478 (PERTH); 27.2 rniles W of Ravensthorpc(5.4 miles W of West River), 28 Sept. 1975,J.W. Green4527 (PERTH);7.6 km NE of Dumblcyung,29 Oct. I975, J.W. Green4544(PERTH); 34km Wof LakeKing,29Oct. 1975,,/.W.Greer4561 (PERTH); 13 km W ofLate King,29 Oct.1975,J.W.Green4568,4569 (PERTH); 13 km E ofFitzgeraldtownship onRavensthorpe to Jcrramunguproad, 3 | Oct.1975, J.W. Green 4617 (PEP.'|H); Soulhern Ironcap, SEslopes,llNov. 1978,J.W. Green4898 (PERTH); l0kmNof DunnRockonOldNewdegateRoad, 18Nov. 1986,"r.W. Greea 5043 (PERTH); 54 km E ofNewdegateonOld NewdegateRoad, 18 Nov. 1986,1.W.Green 5051(PERTH); 26 km W ofRavensthorpeon Old OngcrupRoad,22 Oct. 1987, J.W. Green5572(PER'II{); 5.5 krn NW of Ongerupon property of K. Newbey,33' 55'S 118' 27'8, 23Qct.1983,K. ill338,L. Johnson& D. Blaxell(NSW,PERTH);0.7 km S alongNormanRd frorn CowalellupRd, SE of Ongerup,34' 11'S 118'43'8, 23 Oct. 1983,K. IIiU 344,L. Johnson& D. Blarell(NSW, PERTH); 9 kmNE ofKondinin,ncartrig point,3z" 27'S 118" 21'E,8 Nov. 1983, K.IIill &7 ,L. Johnson,D.Blcuell,I. Brooker& S.tlopper (CANB,NSW, PERTH): Cargonocking Hill, 6.5km SW of Kulin on roadto Harrismith,32" 42'S I18'06'E, 8 Nov. 1983,K. Hill 655, L. Johnson,D.Blatell,I.Brooker& S.llo2pei-, (CANB, NSW,PERTH); 16 km SEofKulin,32' 43'S 118' 17'E,8 July1977 ,I?.J. Hnatiuk1 70154 (PERTH); 5 km WNWof Ongcrup,4.5km N of Foster Rd fromOngerup Rd,33" 55'S 118' 27'E,31July1982, S.D. Ilopper 2404 (PERTH); 16 km SEof CowalellupRock, NE comcro[ CorackcrupNalure Reserve, 34" 08'S 118"42'8,31 July 1982, S.D,Hopper 2405 (PE'F.'IH); Stirling Rangc Nalioral Park,1 km S of SaltRiver Rd on internalEW fircbreak, 34" 20'S Il7" 42'E, 9 Oct. 1982, S.D. Ilopper 2648 (PERTH); c. 14 km SW of Ravensthorpc,1.7 km W of Moir Rd on thctrack to PhillipsRivcr, 30 Scpt. 1981, S.D. tlopper 616I (PERTH);l2milesEof LakcGracc,Fcb. 1958,PR.lef ies580207 (PERTH); 1km onRollands Rd from CarmodyRd (bcforeClarc Rd), 33'23'S 120"52'E, 20 Jan.1981,, C.l. Keighery3717 (PERTH);Rcscrvc No. 20U6,25 km NE of Nyabirrg,19 Jan.1978, .LM. Koch N30 (PERTH); Pingrup,2lkm S at 110 milepcg,2TFcb . L97 5,O.W.Loneraga, L231 (PERTH);Nyabing, Dcc. 1957, V.McDoudle083,084 (PERTH);6.4 km from Jcnamungup towards Ongcrup,5 Jan.1978,8. Mul Lins 371 (CANB); 14 milcsE of Ongcrup,29 April 1962,K. Newbeys.n. (PERTH);4 milcs E of Dumbleyungand 2 mileson road S, 30 Oct. 1968, M.E. P hillips s.n. (CANB, CBG 021982); Tarin Hill, nearLakeGrace,Junc 1924,llalph & Stanford(PERTH); 18 miles E ofLakc King, May 1969, B.A.Rockel,460 (CANB); 10 milesE of Broomchill,13 lat\.1954,R.D.I?oyce 4788 (PERTH); 25.5km (16milcs) E of LakeKing townsile, 9 Aug.1968, R.A. Sffiey 405(PERTH); about 30 krn N of BremerBay in lhc FitzgcraldRivcr Rescrvc,7 Oct. 1970.P .G. Wilson10203 (PERTH).

Distributionqnd habitat. Southernwheatbelt from Wickcpinlo thc StirlingRange, and to eastof Jenamungup(Figure 36). A commoncomponent of malleecommurilios ort plains and undulating terrain.

Consemation statu.s.Widespread and abundant.Wcll rcprcscntcd on conservationreserves.

F lowering period. Ianujry.

Notes. E. phaenophylla subsp.phaenopilll.r is the most aburdant taxon in thc scrics. In the past it has been called E. r edunca.It differs from subsp.interjacens in its more upright straight-stemmcd habit, and its more slcndcr buds with the operculum noticeably naffower than lhe hypanlhium at the join.

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Figure 40. Holotype of E. phaenophylla Brooker & Hopper subsp.intarl.er.!_ '9761'eout:edsgruru:(HJUf,d '3uruuu1q4;o Z08t a)toY O'U'Vg6I\teI71 g ru0Z:GJUAd) tglt '191p'gg1p 'u€I 'tlqeuoorgJo actoy O A'196I tI A setrul0I :(]fl'J-].dgd)gVL.tVL$nDJ.S.l L6I 'unoladoH tdos EI pu€etuoqtsue^?U uoo./rUoq :(VSU'HJXAd'.I'SNVJ) 16llnDd'7'S'0L6I trBlgI 'odroqtsuo^el{Jo td solfiuI € :(HJUEd'qy) 699yp.totqttg flV'g96l TrO 0Z lelul so)tolslets?or 'r'988 're^rl{ oqtJo ldNN tu{ IZ uortsro'J Suno^rp.ou :(HJUgd'qNvJ) gtgz ?Vsrl,Vu.S,IL6I 'Jc( 0Z'psod dn8unrsrof-eJuerodsg'Surssorcrelrg Sunol :(HJdEd) s,11227111'g961 1de5.feg 'rdeS'.(sg .I reurerg;o1q'p11drus.rS :(HJUAd) sd4zry'1'r96I reuorg:(HJUgd) t6z19plapult A '91,6I 'pU tdes ZZ'A,gt "OU S,g€.tg edroqlsuc.trguo unolodoHrxo{ ru{ tZ :(HJUAd) '286I .ldos 'g,rS 're,nr11 .dnur16un14 6997nddo11 O S 0I .0Zl S.8t "€t dnpu::o;3o,t| uq I.g 'tro 'pBoU JogS uq 9'EI:(HDl!Id) Llg,uaatg l1't'9L6I I € dn8unura.rclo1 edroqlsua,rell uo 'A'l'g ,odroqlsue^eu drqsumol:c,rra plurc8zlrglo g tu1 g1 :(Hldgd t69t 1/,,tg L6I tco 0E Jo S 'edroqtsue^?UJo r!rr{9I :GJUad Z6SVuaalt A I'SL6I loo 0€ S ru{ SI :(HJUAd) LBSVuaatg ',At'9L6Itto0t'odroqlsuc^sdJoSrr{€Z:(HJUAd)gBfiuaarg,llt'L96Irl.mbl,El .dnje8uo 'a.Oto0zl Joasclrrx09:(HJuad) SILsI a3n29 s'v'6L6I cunl I 5,7g.gg,4aer3dnullpq '9961'3ny 're,rr6 rscu'200I uortsro-IpteUpto :(HIUAd ,9I9I nup.tD] V J g1 plauplo:(SNYJ) 'Lg6I ''lsm 'qNvC '(V) 'ft1 EEEZllDlDll V tlo 6 dnSunuruJJelrrrou :(HJUAd ,rggg7nypry '896I 'ro^rUptcUplOJo 'f€ tro IZ qlnouJoNur{ 0t . I I uolluro-J:(HJdgd'Oy)69661ngctn [p '896I'ldos9Z'lclulsc{otsluls?o3Jo\Iur1 '1co:3 '006 91 c dnlurstro1 Surpeel dllnE ur uorleJo-I 'L96I 'unolcdog;o l(tJIUSd'SNVJ)tlzalDpuaddltlJ W r qor?W9I 1.1salrur6.ZI :(HJUAd) t69t 'pg6l 'too 'g. 'pr?ol{ uowSng y 1,t1 St .0ZI S,9Z.t t'19 LI t c^rosol lurodlsol\ uopU uosurrly\?X Jo N ur{ € g :GJuad'SNVJ) arNry,lis T I90I umuStngy 1,tJ'€g6IqsrBl^l 9Z,A,LE "OZI 'pl{lulodlsold S.8Z.gg uo'{rol{trcqo,rur{onwJogSg ru{ Zt :(HJ){Ad',1\SN'.IghI'SNYJ) €688 'p0od Dloo.tg'lIlW'S86l qlray{IZ'A.20 o0ZI S,0€ og € retcold'psotssacr? oSueg sd:oqlsue,reg "Mht'sNvc) '986I 'a,9€.8II :(HJdgd ",\\sN tr88 r too.lg lrf w qcr?II t s.€I otg'p€or dnrolrproJ uopuor,{rgrcucrq-dn:cSuOJoN ur 00Z'r:(HJUAd'A\.SN'.If .4l'gNVJ) Egggnlootg 'H 'qad0Z'9,60 'peog 'A IW'S86I "0ZI S.tg .gg edroqtsua^Bgrro uorug l] J otJJo-unl uonJunf 'ANVJ) IJoN rx{ 8'9 :(HJ,dgd',\\SN 6L0BDtoolg IIf W'E861IIrdV0I'g,ZI .0ZI S,St.€€ 'pDor 'IlIW.VL6I .rdv unotcdoHrro JJo-urnl uouo^pla lo S Iu)I t' :(SNVC) 99tt Dloorg L 'roru 'o^roscd 1yr11dnlupueg :(HJdAd'SNVJ) 0tl, Dlootg'I[IW'rL6I'tdy g pt?re84rgo't S 're,,rrg JJo-urnlspeorssoJJ pleraSzlrg ruo.4 selrur 9 S:(HJtEd) fDtltzlJo.tggg'W61 ue1 dnul11u4 'l'O 'gL6I ,pEor :(HJl{Ad 689IUaxDM aunlZZ'9,00 oIZI g, St "€t etuoqlsue^?)Iuo rruEredsa '\r's '9961 'a8ueu 'SNYJ) lo td rlx{06 :(HJdAd.) tpuuag W 3' 1de5 edroqtsuc^?d:(HJUAd 6gSZ 'ldcs 'o8uuu 'p tJauaagW fl'896I Z cdrorllsuo^€)I:VI.IVdJSnv N!IIJSA,4[ aunuoxasuawnadg

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D.L. Serventy@ERTH); 33.5 km N of Hopetounalong road to Ravensthorpe,33o 40'S 120"06'8, 4 Sept.1986, P.S. S hort 2698,M. Amerena& B.A.Fuhrer (MEL, PERTH);6 km from Munglingup alongroad to Ravensthorye,33" 41'S 120"48'E, 5 Nov. 1982,A. StidzLL59 (PERTH);l1 milesE of Newdegateonthe highway to Lake King,20March 1970, M.D.Tindale222 &B -R.Maslin (NSW, PERTH);20kmWof BremerBay township, l Oct. 1966,P.G.Wilson4344(CANB,PERTH);20 km Nof Hopetoun,5Oct. 1966,P.G.Wikon 5515(CANB, PERTH);Block 1136,Oldfield district, 33" 30'S120'45'8, 28 Sept.1968, P.G. ffi/son 8063 (CANB, K,PERTH);7 miles Nof Hopetoun, LL Oct. 1967, D. Yoirrg304 @ERTH).

Distributionand habitat.Jerramungup and Boxwood Hills eastto RavensthorpeRange, possibly extendingwestwards to Katanningand Broomehill(Figure 39). Commonin species-richmallee communitieson plainsand undulating terlain.

ConsemationstatuJ. Abundant and well representedin FitzgeraldRiver NationalPark and adjacent Ieserves.

F lower ing period. January-March.

Et moloBy.The subspecific ep i thet(Latin interj acens, coming between) indicates the morphological and geographicalsituafion of lhis subspeciesbetween E. phaenophyllasvbsp. phdenophylld and E. tumida.

Notes. This subspeciesappears to link E. phaenophyllasrbsp, phaenophylla nd E. tumida. The narowedoperculum maybe seen in theimmature buds ofsubsp. interjacens urdboth buds and fruits aresomewhat robust resembling those of E. tumidato theeast which has larger leaves, buds and fruit, The budsof subsp.interjacens ue Dsrallyfusiform while thoseofE. tumidaare allannid.

L0.Eucallptus luteala Brooker & Hopper,sp. nov. (Figures41,42)

"mallee" Frutex ad 4 m altus cortice emortuoferenti laxo ad basin. Medulla ramulorum glandulifera.Folia adultaleviter nifentia, viridia vel olivacea.Alabastra robusta fusiformia, ad 2 x 0.3 cm, operculoaequanti hypanthium diametro. Flores luteoli. Semina subsphaerica laevia.

Typus:2.5 km SW of 90-mile Tank, NE of Lake King, Westem Australia, 12 Feb. 1985, M.I.[L Brooker8847 (holo: PERTH; iso: AD, CANB, MEL, NSW).

A malleeto 4 m tall with looserough bark to 1 m, smoothabove. Pith of branchletsglandular. Leavesof theseedling remaining opposite for 2 or 3 pairs,then altrernating, ovate to lanceolate,to 7 x 3 cm, glabrous.Adult leavesnarrowly lanceolate, to 11 x 1.2cm, slightlyglossy, green to olive- green.Inflorescences to 1l-flowered;peduncles to 1.5 cm long, conspicuouslyflattened. Buds distinctlypedicellate, fusiform, to 2 x 0.3 cm; operculumequal in width to hypanthiumat thejoin, to 1.4cm long.Flowers pale yellow. Fruit pedicellate,barrel-shaped to obconical,to 0.7 x 0.6 cm. Seedpale grey-brown, subspherical .slrruJ .spnq ,uolnqulslp = sr€q elecs) Jeet Swlpees epou qug B Jo atenoqlrs puB Jsel tlnp? ,/or,trl tfird{prn| .b afiAilc.

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Colourillustration. Brooker & Kleinig (1990:183).

It differs from -8.phaenophylla in the largcr adultleaves (up to 11 x 2 cm), the more robust allantoidbuds (to 2 x 0.3 cm) andfruit (to 0.8x 0.7 cm)and the operculum on maturcbuds equalling lhewidth of lheh)?anthium al thejoin.

Specimensemmined. WESTERN AUSTRALIA: slopeof Mt Ragged,19 Oct. 1970'T E.I7.Aplin 434?@ERTII); 35 milesE of Espcrance,4 Nov. 1962,J.S. Beard 2317 (PERTH); 25 milesNE of Conclingup,16 Sept. 19?0, "r.S. Beard 6351 (PERTH); Scadden, IsFeb.1970,M I.H Brookr2503 (CANB, MEL, PERTH);Tower Hill, Mt Ragged,10 April 1974,M.I H. Brooker4530 (CANB' PERTH);1 km NELanesRd on CoolinupRd, NE ofEsperance,33' 4I'S 122' 2TE 12 Artg 1982, M.I.H.Brooker7 545 (CANB,NSW,PERTII);2.7 kmW ofStylesRoadon NorwoodRoad,33o 27'S 12L"52'E,9 April1983,M.LH. Brooker 8068 (CANB, MEL, NSW,PERTH); Ned's Corncr Road, 3km Nof Cascadestownsitc,33" 28'S 121' 06'E,9 Aprtl'1983,M.I.II.Br-ortdr 8075 (CANB' NSW' PERTH);11.4 km from IsracliteBay trackon Mt Raggedtrack, 8 Aptil1985, M I.LI Brooker 891'7 (CANB,MEL, NSW,PERTH); 10 km dueSSW ot Mt Burdett,0.7 km NE of ScrddcnRd otr wittenoomRoad, 33' 33'36"5122' 07'21"E,2 Aug.1983, M.A. Burgn.an MAB 1593& S.McNe e (PERTH);520.6 mile peg on Norsemar-SalmonGums Rd, 29 March 1968,S.G M Catr 625 (PERTH)lGrasspatch,49 mls N of Esperancc,31 March1968, S.G.M. Caff 628,631(PERTH); 1.7miles S of GibsonsSoakon Norseman-Espcrancc Rd,3l March1968, S.G.M. Carr 642 (PER'[H)1 39.3milcs W of Esperancc,15 March !967, G.M.Chipendale 192(CANB, PERTH); 32.6 milcs E ofEsperance,25 March 1968,G.M.Chippendale 409 (CANB, PERTH); c. 75 km NE o[Espcrance' 1 Arg. 1980,M.A. Clementsl8L5 (CBG 8005162,PERTH); Mt RaggedRangc, 2.5 km S of Towcr Peat,33' 28'S 123"28'E,6 Jan.1979, M.D. Crisp4845 (CBG, NSW, PERTH); c. 40 km NE of Condingup,junction of BeaumontRoad and Parmango Road, 33" 30'S 122' 49'8,26 Nov. 1985' D.B.Forernan 1253 (MEL, PERTH): 2.3 miles N of SalmonGums, 15Dcc 1940,C.A. Gardner s-n. (PERTH);N of Scadden,Dcc. 1940,C.A. Gordner s.n. (PERTH)l 5 milessoulhward lrom Salmon Gums,7 Nov. 1953,C.A. Gardner 11171' (PERTH); 46 km N of IsraelircBay, 12 Fcb.1966, C.A.Gardner 16381 (PERTH): Mt Ragged,7 Dcc. 1960,A.S. George2109 (PERTH); 8 km S ol' Ravensthorpc,30Oct. 19?5,J.W. Green4603 (PERTH);Capc Arid NationalPark,33'45'S 123"00'E, 19 Sept.1976, R. Llnatiuk761072 (PERTH); along Point Malcolm Road, 33" 47'S 123"45'E,20 Scpt. 197 6,R.1'lnatiukl61139 (PERTH);5 km WSWof GoraHill,8 4 kmNEof Cape Arid N.P.W boundary,Mt Raggedtrack, 33'33'S I23" l9'8,7 Sept.1982, S.D. Hopper 2522 (PERTH);MtRagged,8km Eof Mt Symmons,Cape Arid NationalPark,33' 27'S 123 " 29'E,8Sept. 1982,S D. Hopper2533,2534eERTfD; SpringdalcRoad,5.5kmEofHopetoun-Ravensthorpcroad, 33" 55'S 120' 13'E,29 Sept.1987, SD. Ilopper6"144 (PERTH); 1 km S of junctionbctwccn CoomalbidgupandGriffithsRoads,33' 30'S 121' 20'E,21 Jan.1981,G.1. Keighety 3706 (PERTH); GriggsRoad, 10 km E ofLort River,21 Jan.198I, G.l. Keighery3755 (PERTH); 15 milcs SW of MountRagged, June 1973, O. Loneragdn 34 (PERTH);Kau RockReseryc,0.8 km from Coolittup Rd alongKau Rock Road,33" 33'S 122'E,22 Sept.1985.L.LNu.nn258 (PERTH);ncar YoungRiver, 20Oct. 1968, A. Orchard1669 (CANB); 10 m iles S of RedLake siding, l8 April 1953,R.D. Royce 4061(PERTH);W face ofMountRaggcd, 3 Oct.1970,R.A. Sdjf'ey 1322 (CANB, PERTH);Sheoaks Hill,29 kmE of MtRaggedand 19 km W of IsracliteBay,3Oct. 1970,R.A. Saffi'ey 1370 (CANB, PERTH);5 milesN of Scaddcn,10 April1966,E.M. Scrymgeolr 458 (PERTH); 9'8 km E ofScadden on ScaddenRd,2lAug- 1982,P.van der Moezel 163 (PERTH); 6 km E of Scaddcnon ScaddenRd, 'lBcreqdsqns'u.,'{orq-dor6 IqAIIpees ul3S 0 x 6 0 ol'lBJgpull^t 'peduqs 'el?lleJlpedllnJd'ueoslou sre,{lolg'ulofoqllB unlqluedfq ueql:a.'iro:reu,{lsnonotdsttoc 1ou 'ruroJrsnJ'olElte3lpad 'uooJ8 -u:oq runlnoredo:ruJ g 0 x z ol spng'8uol tttJ8I ol "{ssol8Fnq8lls 'uJJ 'elslooJusl 'se^eol ?uunlsur I'I x II ol lJoJo ploq ol oleloeouel,(l,a\olJuu 11npy snorqelE 'urJ 'clploeJuul 'Eu[uuJolle 'sjlud i x II ol ol alu^o uew t-Z loJ ollsoddoSulull]tuoJ Eullpoos oql 'qsppld ',ter8 ',(aB 'qloouls gose,reel elud-ro u.ro:q 1{3ll 1q311 lrup UDg ll€l uI t ol oallBulV

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Fig\\e 45. Eucalyptur hirtopl)//d distribution, and buds, fruits, adult leavesand silhouetleofa fifth node seedlingleaf (scale bar = r cm). 'raddoH 't T ra{oorg ,trlC{dortrrl Jo ad,{roloH gt ernBIJ

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Specimensexamined. WESTERN AUSTRALIA: 131mile peg on Norsemanroad,IT lttly 1952, P.H.Barrett s.n. (PERTH);c. 26 km NW of Balladoniaon Norsemanroad, 32' 16'5 123"24'E' 22 Au,g.1979,M.LH.Brooke r 6447 (CANB, NSW,PERTH);30 km Wof Balladonia,28April 1982; M.I.H. Brooker 7485 (CANB, NSW, PERTH);37.8 km NW of Balladoniatowards Norseman, 32" l0'S 123" 15'E,10 Feb. 1985,M.I.H.Brooker8835 (CANB, MEL, NSW,PERTH);40km NW ofBalladonia,32' 10'S 123" l2'E,91an.1986,M.LH.Brooker9147 (CANB, MEL, NSW,PERTH); 33 km W of BalladoniaRoadhouse on Hwy 1, 32" 12'S 123' 19'8, 14 Nov. 1983,K. Hill705 & D. ,B/arell(NSW, PERTA; DundasNature Reserve, 4 km E of BoinganingRock, 21 Aug. 1989, S.D.IIopper7359 (PERTID; Dundas Nature Reserve, Boingaring Rock,21 Au e.1989,S D. Hopper 7362(PERTH); Mt Coobaninya,22Au g. !989,5D. Hopper7408(PERTH);Mt Buraminya,23 Aug. L989,S D. Hopper7446(PERTH);48 km fromNorseman on Esperanceroad,3 Jan.1978,E. Mullins 33i (CANB); DundasNature Reserve, Boingaring Rocks, 40 km WSW of BalladoniaHotel, 32'i3'05"S 124'21' 50"8, 30 Sept1987 , K. Newbey 11778 (PERTH).

Distributionand habr'rat. Between FraserRange and Balladonia and southwards on graniterocks to Mt Buraminya(Figure 45). E. histophyllais knownfrom sitesnoflh-west and west-south-wcst of Balladorriawhere it growsin tall matleewith E. eremophila(Dicls) Maiden,E. faseri (Brooker) Brooker,E. sp. nov. aff. transcontinentalisMaiden, E. leptophyllaand' E. indurataBrooker and Hopperin€d. However, it is commonthough sporadically distributed around granite outcrops further south,where it is oftenthe only eucalyptpresent.

Conservqtionstatus. Poorly surveyed but knownon DundasNature Reserve.

F lower i n g p eri o d. U nknown.

Erymology.Thespecific epithet refers to theerect leaves (Greek listos, uprightattd phyllon,leaf).

Notes.E. htstophylld,as the epithet implies, is characterisedby theerect leaves which make it easy to recognisein the field. The leavesare narrower than those of E. tumida,and only becomeglossy within the crownby thesecond year. E. histophyllahasnot been seen in flowerso comparisonsfor this characterwith thebeautiful, yellow-flowered E. flavida havenot beenpossible.

Surveysby S.D.Hopper in 1989establishedlhat E. histophyl/aoccurs predomirlantly on granite outcrops,an unusualhabitat for taxain theseries.

13.Eucallptus clivicolaBrooker & Hopper,sp. nov. (Figurcs2,47 , 48,49)

Arborparva ("mallet") vel frutex("mallee") ad 7 m altacaulibusexilibus etcortice laevi cinereo extuscupdno intus. Folia adultanitentia viridia. Inflorescentiaead 11-flores.Alabastra operculo aequantihypanthium diametro, redunco. Fructus doliiformes, ad 0.9 x 0.6cm. Semina subsphaerica vel rarocuboidea.

Typus:9.6km southof theEldverton tum-off on the Hopetounroad, Western Australia, 10 April L983,M.1.H.Brooker 8078 (holo: PERTH; iso: CANB, NSW).

Colourillustration. Brooker& Kleinig (1990:186).

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Distributionand habitat.North andeast of Kalgoorlie(Figure 50). E. flavida growsin openlow woodlandsandmalleeusuallyon gently undulating terrain. Soils are red loam with qualtzoI calclete. Associatedeucalypts are E.aff .longicornis,E.grffithsiiMaiderl E. atf. tt'a scontinentalisMaiden, E. clelandii(Maiden) Maiden and E. rigidula Maiden'

Conseflationst.,tus. Locally abundantin disjunctpopulations, a few of whichare on reservessuch asQueen Victoria SpdngNature Reserye.

F lowerin g p eriod.November-Decembcr.

Etymology.Ane\N specific epithet is neededfor thistaxon because Maiden's varietal nal:l.e oxymitra is alreadyin useat the specieslcvel for Sharp-cappedMallee, E. oxymita Blxkely, of the central Australianregion. The new specific epithet refers to thecolour ofthe flowers (Latin/avidrs, yellow)'

Notes.The glossy grcen l ceolatejuvenileleaves ofthis species are unique in theseries. Its long slenderuncinatebuds, ycllow stamens and long barrel-shaped fruits suggest affi Irit\es wilh E rcdunca, E.pluricaulis, E. varia,E. densaand E. gardneri,butlheconsistent presencc of pilhglands and the "redunca" juvenileleaves in E./avidd isolatesthis species somewhat from the superspec ies . The copiousflower production and inlanddistribulion of E. flavidu indicatesuilability as a hardy ornamental.Birds such as Yellow-thrcated Miners and Red Wattlebirds have been observed taking nectarfrom flowersin the wild (Hopperunpubl.).

15.Eucallptus crirpal4 Brooker & Hopper,sp. nov. (FiguLes5c, 52, 53, 54)

Frutex("mallee") ad 5 m altuscortice inferiore decorlicanti crispxto superiore laevi cineraceo, foliis adultishebetibus vel nitentibus,et alabastrisplerurnque brevioribus crassiodbusque quam affirribus.Semina cuboidca vel plano-ovoidea.

7lpr.r: nea.rYandarrooka, Wcstem Australia, 13 March 1986,M.l.tl. Brooker9201 &S.D llopper (holo:PERTH; iso: CANB, MEL, NSW).

Colourillusnation. Brooker & Kleinig(1990: 180).

A malleeto 5 m tall, erector spreading.Bark towardsbase of stemsheld ill partlydecoflicated curls,smooth grey above. Pith ofbranchlets glandular. Leaves ofthe seedling remaining opposite for about2 pairs,then alternating, ovate to dcltoid,to 10x 9 cm,glabrous, slightly glossy, blue-grcen. Adult leavespetiolate, altemating, lanceolale to falcate,to 9 x 1.5cm, dull or glossy,green. Inflorescencesto 13-flowered. Peduncles to 1.6cm long. Buds to 1x 0.4cm, shorterand broader than mostrelated specics, operculum cylindrical to conical.Fruit sessileto shortlypedicellate, obconical to cupular,to 0.5x 0.5cm. Seed light grey-brown, cuboid to compressed-ovoid.

Specimensexamined. WESTERN AUSTRALIA: Type locality, 13 Much 1986,M.I.fl. Brooker 9205,9208,9209& Sr. Hrpp?r(CANB, MEL, NSW,PERTH); NE ofEneabba,21 Nov. 1986, M.I.H.Brookr9555(CANB, MEL, NSW,PERTH); l kmN ofEW fenceonN-S track, west of South EneabbaNature Reserve, 30' 00'S 115' i6'E, 20 Sept.1988, M.I.H. Brool

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Distributionand habitat.Knownflom lateriticbleakaways south of Mingenewwhere it occursin severalsmall pure stands,and a singleoccunence near Eneabba (Figure 52)

Conseryationstatus. Rare and vulnerable. Currently declaled asEndangered Flora undertlreWildlife conservationAct (Hopperet a\.1990).Population sizes ale smalland will needto be monitoledto ensureagainst accidental destruction.

Floweringperiod. ? APil.

Erymology.The specificepithet refers to the curled,pafily detachedbasal bark (Latin crispatus, curled).

Notes.E. crispatais a raremallee known only from two localities. The crisped brsrl brrk is unique in theseries. ihe budsare relatively short and fat thoughvariable while the seedling leaves vary from ovateto cordate(sim ilar to E. accedensand E.leprophloiaBrooker & Hopperined. although not seon to segregatewithin family lorsof fourpalents tested). The seedare somewhat ovoid While of fairly unifo-rm-appearancein the field, the speciesmay be of recognisablyleccnt hybrid origin with the parentageof E. accedens and E. arachnaea,bolhof whichco-occu with E crispata'

16.Eucalyptus subdt gastd (Blakely) Brooker & Hopper,comb. et sfat.nov'; E reclunca Schater ,,Key var.subangusta Blakely, to theEucalypts." p. 111(1934). Leclotlpe(here designated): Cunderdin,Western Australia, W.Z F ttzgeralds.n. (Figures 3, 5f, 12b,55,56,57' 58' 59' 60' 6l ' 62)

A malleeor malletwith smooth,grey over pink or copperybark. Pith of blanchletsglandular. Leavesof theseedling remaining opposite for 4-6 pairs,then altenatillg, lnnccolale to ovate,to 9 x 4 cm,dull, green, blue-grey or glaucous,glabrous. Adult lcavcs petiolate, alternalinS, narrowly lanceolatetolanceolate, to 1I x 1 cm,dull or rarcly glossy (in subsp. virescen s),light green, maturing to blue-green.Inflorescences to l7-floweredipeduncles to I cm long. Buds shoflly pediccllafc, fusiform,to I x 0.3cm, operculum oblong, obtuse or acutcat tip; few strmens elect, flowers while. Fruitshortly ped icellate, cupulxr (i.e. about as long as broad), to 0.5 x 0.5cm (as brord as long)' Secd light grey-browu,subspherical to cuboid.

Typificarion:lntheprotologuefor var.r ubangusta,Bl|lkely presumably refers to Maiden'stlcatment of i. rrdurto ru. irSl,JlfolraBenth. that alludes fo twoDrummond specimens, 67 and 187, which arethe syntypes of vir. angustifutia(now E. xanthonenn).There are lhree furlher specimens which aremosisimply refened to by theirlocatities, 1,i2. cunderdin, Knutsford and Goldcn valley. Blakely thencited ore oflhese, cunderdin, in theillustmtions (Plrte 140, number6) which becomes the type of E. redwtcavar. subangusta.

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7602,(CANB,NSW,PERTH);c. 1.5kmNof Dowerinon CadouxRd,3l. 13'S 117.03,8, 14Sept. 1982,M.LH. Brooker76M,7605 (CANB, NSW, PERTH);4.3 km N of Dowerinon CadouxRd, 31' 11'S 117'03'E,14 Sept.1982, M.I.H. Broolcer 7608 (CANB, NSW, PERTH)l 21 km NE of Doodardingtowards Koorda, 30" 5L'5 ll7" 22'E 14 Sept. 1982,M.LH. Brooker 7612 (CANB, NSW, PERTII); 6 km SW of Koorda towardsDoodarding, 30.52'S It7" 27'8,14 Sept. 1982, M.I.H. BrookerT6L4(CANB, NSW, PERTH); Comer Scotsman'sRd and Clcary-paynesFindRoad, 30" 22'S tt1" 34'E, 11 Jan. 1983,M.I.H. Brooker 7914(CANB, NSW, PERTH);0.5 km W of WandanaRdat N end,N or Yuna,28' 03'S1 15. 03'E, 25 Ian. t983, M.I.H. B rooler 7938 (CANB. NSW,PERTH); c.4kmWof Kalannie,30. 20'S 11?"04'E,8Feb.1983, M.1.H. Brooker 7953 (CANB,NSW, PERTH); 6.1 km N of MerindoN. Roadon Kulja CentralRoad, 30" l8'S 117. 18'8, 9 Feb.1983, M.LH. Brooker T96I (CANB,NSW,PERTH);101 km Wof Cootgardic-Norsemanroad on Hydentrack, 32" 04'S I20' 42'8,1 Nov. 1983,M.I.H. B rooker 8355 (CANB, NSW,pERTH); Mason'sroad E of Watheroo,2Sept. L984, M.I.H. Brooker8687 (CANB, MEL, NSW, PERTH); 0.5km Sof Tammin,31'39'S 117' 29'E, 15 Nov. 1985, M.1.[L Brooker 9104 (CANB,MEL,NSW, PERTH);17.7 km N of MurchisonRiver on NW CoastaLHwy,2T.40'S 114. 42,E,6 May 1986, M.I.H.Brooker9266(CANB,MEL,NSW,PERTH);NboundaryofKalbzrriNarionalpark,2T.24,s lL4" 26'E,9 Oct.1986,M.1.H.Brooker 9473 (CANB, MEL, NSW,PERTH); 3.5 km W of Dowerin, 31' 13'S1160 59'E,23 tnly 1987,M.I.H. Brooker 9726 (AD, CANB, MEL, NSW,pERTH); S of Pintharukaroad, 9.6 km from Mingenew-Morawaroad via YandanookaRoad, W of Morawa, 29o06'S 115'38'E, 9 Sept.1.987, M.LLI. Brooker 9754 (AD, CANB, MEL, NSW, pERTH); 0.5 milesS of Wubin,21Oct.1966,G.M.C htppentlale 59 (CANB,PERTH); 3.6 miles S of Kalannic, 2I Ocl t966, G.M. Chippenrlale65 (CANB,PERTH); 22.8 miles SW of Merrcdin,6 March 1967. G.M.C hippendale 88 (CANB,PERTH); 1.4 miles W of Karatce,7 March 1967, G.M.C hippendale 104(CANB,PERTH); 9.1 milcs E ofPithara,9Aug. 1967, G.M.Chippendate250 (CANB, pERTH); 9.4milesNEofBruceRock, 10 Aug. 19 67,G.M. Chippendale 256 (CANB, pERTH);Cadoux.3 Nov. t981, II. Demarz9126 (PERTH); Mogumbcr, March 1901,Dr Diels & prirzei (pERTH);6 km E of Koorda,12 Nov. 1974,Forests DepartmentLlSz (PEP'TH); Dowerin, 19 Aug.1920, C.A. Gardner 657(PERTI{); 2 milesS of Dalwallinugare, Rabbit-proof Fence, 14 Jan.1947, C A. Gardner8523 (PERTH);Bendcring, 19Oct. t949, CA. Gardner9478(PERTH); 3 milesN of yerecoin,21Aug. 1957, J.W. Green1486 (PERTH); Tammin - CundcrdinRd, 26 JDne1959, B.J. cleve (PERTH);; Tammin,Oct.1959 ,B.J.Grieve (PEP.TH);40 mites S ofMullewa,3 Aprit 1967,8.11olrn s.n. (CANB); KalbarriNation al Park, c.2km NEofJungaDam,l3 Sepr. 1979, S.D. Hopper1301 (pERTH);5.2 km Nof Yerecoinsiding,4 km S ofPiawaningsiding, 30. 53'S 1t6.24'E,26 Alg.l9B2,SD.Hopper 2483(PERTH);5 kmWNWof ScrivenerRocks,32.19'30'5 118. 46,8,1.4luteJ.985,5 D.Ilopper 4415(PERTH); Dowcrin, west, 11 Nov. 19?4,O.W. Loneragan L233 (pERTH);6 milesout of WonganHills,3Oct. 1962,F. Lullfitz LI659 (PERTH);Wongan Hills,3 Oct 1903,A.Morrison (PERTH);9 milesfrom WonganHills towardsPiawaning, 3 Oct. 1962,M.E. phillips s.n.(CANB, CBG024965); Walebing, Feb. 1,950, Popplewell Bras. (PERTH); 7 milesN of BruceRock. 14 Anril 1953,RD. Royce3983 (PERTH); Water Reserve, Ceron, 10 miles S ofperenjori.1f Aprit 1962, R.D.Royce 6813 @ERTH); Watheroo Nationalpark, W. of Watheroo,8Oct. 1971,R.D. RoycegT 62 (PERTH);14.6 miles Lynton Road, 1 March 1966,E.M.Scrymgeour 272 & S.G.M.Car (pERTI{)l 390.8miles NW CoastalHighway, 3 March, 1966,E.M. Scrymgeour322,323 &, S.G.M.Carr (PERTH);212.6 miles Morawa-Perth Road, 5 Mnch t966,E.M. Scrymgeour393 & S.G.M.Car (PERTID;W of Manmanningtownsite, next to AvonLoc. 19405,30.51,S 1l?.05'E,24 March 1,987,B.H. Smith840 (AD, CHR,FRI, HO, MEL, NSW.pERTm. '(uc I =req stsrs) Fal Sunpaas '99 '|q!ruJ 'sPnq pu! 'uoqnqlrrslP ', tt1311oqns'dxqns Dtsk?uDqns snilDznfl ernev epou qlJu s Jo qlanoqlls puB sa^sel 1FPB

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Distribution and habitat. Centraland northernwheatbelt, north to Kalbarri and south-eastto Norseman@igure 55). Occursin low to tall mallee,usually on plainsof yellow or red sand.

Conservationstatus. Widespread and abundant. Well representedon naturereserves.

Floweringperiod. January- ?

Nores.This is oneofthe mostcommon mallees in thenorthernwheatbelt. It difforsfrom subsp.pnsllla in its largerbudsandfruits, its consistentmallee habit, and its preferencefor light sandysoil. Subsp. subangustalackstheglaucous branchlets and consistent acuminate operculum ofsubsp. cerina lthas dull leavesunlike the glossyleaves of subsp.vtrescels. l6b. Eucalyptussubangusta (Blakdry) Brooker & Hoppersubsp. pasil/a Brooker & Hopper,subsp. nov.(Figures 56, 57, 58, 59)

A subspecietypicahabitu variabili frutice ("mallee") vel arboro ("mallet") alabaslris ftuctibusque parvioribusdiffert.

Iyprs:15.8 km eastof Mullewatowards Pindar, Western Australia, 12 Jute 1985'M I.LI Brooker 9039(holo: PERTHi iso: CANB, NSW,MEL).

Colourillustration Brooker& Kleinig (1990:169).

It differsfrom the typical subspecies in the variable mallee or mallethabit, smallcr buds (to 0.7 x 0.2cm) and fruit (to 0.3 x 0.3cm).

Specimensexamined. WESTERN AUSTRALIA: YunaEast Reserve,24 Oct' 1974,1.5 Beard'7155 (PERTH);53 milesE of Dalwallinu,2 Oct. 1976,1.5.Beard7977 (PERTH); Between Kulja and Koorda,13 Oct.1937,W.E.Blackall35L9 (PERTH); 59 km NW of Wubintowards Paynes Find, 20lan.1976, M. Blackwel/ 34 (CANB);6.5 km S of MtGibson turn-offon GtNlhn Highway,20 Oct. I978,J. Briggs222 (CANB); 32.4 km NE Carnamahtowards Percnjori, 19 Oct 1978,J. Bri8gs314 (CANB);40milesNEof Wubintowards Paynes Find, 11Aug. 1969,M.1J'|. Btooker 1972,1973 (PERTH);15.3 km NE of Calingiritowards Wongan Hills, 30'59'S 116"33'E, 16 Feb. 1983, M.I.H.Brooker 7966 (CANB, NSW, PERTH)l 15.4 km E of Mullewarowards Pindar, 28'22'S ll5' 41'E,24lan.1984,M.l.H. Brooker 8414 (CANB, MEL, NSW,PERTH); 84 km SW of Paynes Find.29' 40'S 117o08'E, 18 Oct.1984, M.l.H. Brooker 8718 (CANB, NSW, PERTH); 15.8 km E of Mullewatowards Pindar,28" 3l'S 115'40'E, 12June 1985,M.l.ILBrooker 9039 (CANB, MEL, NSW,PERTH); 39 km NE of Wubin,29'53'S 116'53'8, 15April 1986,M.l.H. Brooker 9226 (CANB, MEL, NSW, PERTH);82.5 km S of PaynesFittd, 18 April 1986,M.IJ'|. Brookr 9262 (CANB,PERTH, NSW, MEL); Stoke'sfarm, Franco Road, W of Morawa,29" 08'S 115'44'E, 3 Feb.1988, M.l.H. Brooker9880 (AD, CANB, MEL, NSW, PERTH);69 milepeg from Geraldton onMagnet-Geraldton Rd, 19 March 1968,5.G.M. Carr 455 (PERTH);4 miles SSE of Kulja,24June 1978,S. Charltons.n. (CANB, PERTH); 0.4 milesE of Pithara,E of railwayline, 21 Jan.1966, G.M. Chippendale63 (CANB, PERTH);Tardun, 8 Aug. 1967,G.M. Chippendale246 (CANB, 'eln aoo 'olrrl,r IeJIuot flluolsrsuoJpue slalqJusrq,{xu,1,1 aql ,{q solJedsqnslerrdfl eq1uo{ sloJJrplI

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Figure 56. DistribDtionof Eucalypt|s subangustasubsp.plrttla (o), subsp.c?rid (v) and subsp.viercrrd (E), andbuds, fruils' aaluftleaves and sillrouclteofa fiflh node seedlingleaf Gcale bar= I C]..n)of Eucaryptussubars6td sobsp p,r//ld. 'raddoH t raloorg ,llrrd dsqnsreddol{ ? reloorg (,{talel8) ,tsttSrrqrs g Jo ad,{lotoH /9 arnSI{

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Figorc 58. t. rrlrargrBld subsp.prBilld mnllcl babit nearMt 6ibsorl. and !lxnk and b:r*.

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t 'ueer8 'sa,ree111npe,(sso1E dpq311s lq8rl eql ur sarradsqnspctdfl sq uoq sroJJIq

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'DuDlqvaqs'7 ptrcoqalqdoxol JJD ? sc qJnssslJeds qttar' stuuol deloper uo ,(ypnsn'pwlqnJqs Jo oollpur 'puplpoo,4\ /(ol ul s,nor8naua: 'dsqnsasnSuDqns g '(gS 'llaql?eq.A{ ',tr,tlqDq emSrd)ssorJ uJaqlnos Jo ls?e ol Iuoqlol uro{ I?ruecuolsEll puDuoynqlr$!O 'HJUAd) 'W't'S 'OZIV '9 'ssor3 'c 'eluolsei[ ttDJ T tZL uostt6 A'996I pdy 71 ueqtnos Jo rt[ uq 0t 'lt '8L6I 'p?or puE urq"r?J uee,rleg :(I{Jl{gd) 9691tuaq81ay Aetn8'a,zv "ozl s.s0 oz€ '1coy "IfU 'SNVJ) ueuresroN-uap,{H deuy go16 uq ZE:(Hlffgd'.^eSN I 8IW )addoH Q'S'8861 'A,rE ,{InI9 "8I I S.tg o0€'!IlI Euruunr1ce4 Suop peog uoslrrol IJo SuDl g'I pw drpunoq A\Jog 'e1'JeseU 'O'S '986I '9.9t uDI€'0 arnleg EurdoocrrpprqJ:GIUgd) SIt? raddoH aunl tI o8I I S 'sIrol{ 'I68I ']3o "Y'.^d.'proJslnu) ,,0€.6Iozg roue^lrrsJo i[Ilrt[ uDts :O[sN) ,tu1aHu 6 rEeu 'ro(I6'!I.20 :GIXgd)$r1aH U'I68I "6I I S,€9.0€'(LtrugtlngJo N) delJe^ueplogmeN:(HJugd 'glltf J)06appuaddtltJ W't'L96Iq)nr tr9'urq8rreJJoAselrur I'9:(HJlIgd'ASN'TaI I'SNVC 'qV) 'p8ou LZL6r loorg'H I n'Lg6I flr{eZ'ruoqlelJo gS seurBg',.eppuexo,,:(HJXAd'ir\SN 'Ttr 'qY) 'txo 'qJuUIIng I'SNV3 V6V6'86t6Dloolg'H'l W'9861 Z7'9.,89"811 S,Z0 0Ig Jo 'ulllepog 1I\SII :Gruad'A,sN'SNVJ) 9908rapo.tg'H I W',t86l IIrdV''!I.0t .8II S.ZZoI€ ',{A.SN'SNVC) 'qeg. 'A.Zt.8II Jo ifid'unt tZ :GIUAd VL6L'o\ootg'H'I'W'88 rl S.I9.0g 'pue lsee'peor,1\-g'e,T rase11emp1q Surdooirupplq3:(HJUEd'nd.SN'gNVJ) tZ9 L)qooq H'l'W '286I 'ldes 'lurelnN 'r 9I'A,99.Ltt 5,6I "Ig Jo td ur{ SZ :(HJXAd I6LI ntootg'H'I'W '696I 'esmo3 'pautuDx] eunl t JIoc ulperal Jo g selluiz :vITruJSnv Nur[s:L/y\ suawlrdds

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Figure 61. Ilolotype of E. rllbd,grsrd (BIakely) BrcokeL& Hopper rubsp. vlrercerr Brooker & LIoppcr. 'dsgns rrq?q aalEru 3ur^.,oqs .eatsan^ DtsnSuoqrs t Z9 arn8il

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Nuy(siaVol. 8, No. I (1991) ll8

Narembeen'17 J ttly !970'l 'S' Beard Specimensexamined. WESTERN AUSTRALIA: 14milesE of (CANB);25'7km Sof Koorda l'sio (CANB);20km NW of Hvden,4o ct.rg'75,D'BldxellwT5l75 I'H'Brooker7616(CANB' Nsw' ,"*.0. wyufi"rchem,31"00;S til" zl'e' 14 Sept'1982, M (CANB' MEL' NSW' penffD, frlo.on'. RoadE of Watheroo,2S;pr 19-34,M ILI Brooker8687 pithara, April 1986,M.I.H. Brooker9225 i,enrnii betweenMiling and 10. zi's rte. so'e, 15 Roadalong Jones Road' rtael-,NSw, PEi'TtI); w of Manmanning,2'8 km E of Hourigan ic-em, tvpelocalitv' :o' zo'! t tS" S:'e ,23oct.1986,MLH.nrooker9497 ICANP,MEL'NSw'PERTH); (AD' CANB' MEL' NSw' PERTII)' ii i"e.l98S,u.t.H. Brook"r 10046,1004? & C't' Ranford

it isknownfrom thetype locality Distributionandhabirol.Eastern part ofthe cental wheatbeltwhere 56)' Soilsoccupied range andat a few scatteredsites between Manmanning and Watheroo Figue kochiiandE' erythronema)to white clay ir"* V"U"* t""a t withE. flocktoniae,E.sheathiana,E (with E. yilgarnenstsandE. erythronema)'

roadverges in largelycleared Conseryationstatus.poorlyknown. All fourrecordedpopulationsarcon fuflher su ey' agriculturalland. The subspecies,therefore, is in urgentreed of

F Iow e r in g p eri od. Unknown.

of the adult leaves'which Etymology.Thesubspecific epiihet refers to the colour andglossiness (Latill vi idis' greenand -esceas' contrast"tothe dull blue-greenleaves of all other subspecies becoming).

Notes.Thissubspeciesistheleast.knowninthespecies.Itisnotablyvariableinitssitelequirements' differingfrom theother with soilsranging from sandto whiteclay. Morphologically it is consistent' manifestin theolder leaves threesubspeclsln its gre€nglossy adulileaves' These features become withinthe crown.

17. Eucalptus subtirtSBrooker & Hopper,sp' llov (Figures63' 64)

AbEucalyptosubangtlJ'a(Blakely)BIooker&Hoppelfoliisjuvenilibusadultisqueangustioribus, nervissecund;iis infirmis,et operculissemper attenuatis differt'

36'E' Westen Australia' Typus:0.7 km southof rail crossing,south of Norseman'32'-32-'S 121' ttrea. lj/ts, ul.a. Brooker8841(holo: PERTH; iso: CANB, MEL' NSw)'

Colourillustration. Brooker & Kleinig(1990: 171)

glandular'Leaves A malleeto 3 m tall.Bark smooth, dark grey and pinkish grey' Pith ofbranchlets lanceolateto lanceolate' oftheseedling remaining opposite for 3 ora pairs,then alternating' nanowly green;secondaly veins to 8 x 2 cm. Adult leaveslinear, to 8 x 0.6 cm, dull to slightlyglossy,light weakwithindistinctconnectionstothemidribandintmmalginalvein.Inflorescencestollflowered; Flowerswhite' to o.6.tn long.Buds short, to 0.7x 0 3 cm,opercula conical' attenuate f"Jonlt",'riuii cuboid' se.tite, .upotar,tJO.4 x 0.4 cm. Seecllight grey-brown'subspherical to '(ur. I = req 'sl!ruJ'spnq luolrrq!{slp '€9 elscs)Jeal Sqtpeesepou rtlJlJ s Jo errenoqF pue se^eelllnps pw ttllqrs nt, dgornt aJnS!8

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.. !rriuor/Bror\orunr rN:rr/R ! |

Figlrrc64.llolotype of f..v/rrilr.rllrookcr & llofper. 'ploqnJ ol lucusqdsqns'u,tro:q-,{ar31q311pee5'urc 9 0 x 9 0 ol'JBtndnJ ',\\ollo.( 'e1u11ecypecldp:oqs lrnJC apd lo ellq.^\,{ursolJ slo,tolC uloI lu unlqlucd'(quuql Je',r\orrEu 'tu.ro3rsn;'a1ellecrpcd puupeo:q 'etenuaiieurntn edo:uc t 0x I ol ,(tuoqsspn g pouelsg'{lEuorts ,(3uol urJI>) UOqssolJunpad.lJsrs pleq'u,roJJ eprsul dsso[6 Suunluur'11np ls:r; lu'uear3-el1lo 'ru3 'elBtoecuul '3ir1s 'snolqe18'uae:8-qs1n1q'u3 x ol'otcloocu?l 6'0x 8 ol ,41,tor.reu sa^ealllnpv € 0I rypuorqoieleno'aulleuiollB oql's{ed 9lo tJoJellsoddo auturcurerBwlpaes eqlJo sa^ge-I'JulnpwlS 'osuop qtl4\ ur ol eoll€I slslqJmq Jo qlld u,^oJJ sulolsqloours delS qsllutd :e,'roderS IIBI z tr '(ldsN ''Iahl 'sNvJ '9.92 'qv :osr:HJdad :oloq)tI86.raloo.{ s H lw'L86l loquc^oNtz'ell0llsnY urelsei& 06II ;pooU - p€od plo Suol? wI.-6'0Vrndq S, Z "gi a,rI)I 6{e'I olcBsp,^{aNuolJ edroqlsue^8u .lroJJIp ooc€lll6ruoocerourr olos ul uuoJsoJOlo'a.roqsnSuu olqturdi(q olnJrado'sllEuEIdtuoJ oplg^ stlnJunpod 'snquuollurall^et srtlnp0slltoJ 'erollsJtrl urnlels enb e r.;et?ocawsn|uDqns oldtplng setpeds'

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L - in qel'PotJad?ulntt ol{

'o^Josaruoll8^JesuoS 3 uo 'sntDls peprorerloN puEtu,^aol3 ltrcol,',\ uo stlouqndod lJUnfslp fqclud ul puo;dsap116 uo!lD^JasuoJ '(99 'u1e1dpuas ut sllospalnlxol auu uo puulqnJqsro ee11nu:'tro1 u! '{pucurtuopoJd orn8l{) puDuollnqltlslo uBuasroNJo qlnos?eJe llf,un)-soljcr]J {Dedeqt ol Iu0J a[ I i]6urN'JDJlqoq 'HJUSd) 66?Z url '{uBJ elll^l raddoH O'S'286I'ldos9'g,z9 "0?.1S,zl"Zt'e?uqlreuerg'oss8lgltr\IJogss S'8I tuBuosroNJo sellu :(HJtAd) ,itourNJoA u{ SI :(HJl{gd 09II I raup tDg'V 3'896I ^oN , S ,Z 'W't '896I 'pr uo Eedeltur g'rrt :GIUgd Zl9 Lto) S tlrrBN 6Z trutxosroN-stun9uoul8s II9 rrDJ rlt t's'8g61qr$tr tr0z'pr ucuesloN-sunc uoups uo 6edollulrtt:(HJugd'}qJ)uosTrot 'v '1 '9869 'q'7t1'6161ldes '4,6I 'salrBr.lJ 01psor T roltDJ l,869 dslq 0Z "IZI S,9V"ZE {Ded auqe fen,q?1qocucrodsg-ueuasloNJo i[S uDt8Z :(HJuad) ' aNrw'S T 6V6IuDu3)ng V W'886I '3nV 'g,,Zt.8Z gfl I I .0ZI S,,I g,ZI .gg'lIH dnulfnoJJo tDI€Z :(SNVJ)ZI naSptqrng J'N' Lr6I '1cle5 settul9Z:(HD{gd'leSN'gNY3)0I SLr lootg H I W'286I^uWZ'srlrEq:) 96'uetuesropJoS '^oN 'sunc psod:Gtuaa rapoq H IW'1861 0I uotulBspu? {eod ol "{[sN'gNv3) IL}L 'ottrsrodsg ue*".roN u"i^teg:(HIUAd',4ASN'SNya) y6yTn1ootg H f W'0L6I'qe1.gl sprs',*tol 'I eunl uauesroNJoSseltu t'92:(HJ){Ad',{[SN'gNVJ) ZrgI r9l amq'l 0'816I IZ'g,St .IZ.I S.Itozt'peoraouEr€dsAuoucluasroNJoSuDtgt:YIIYUJSOVNUAJS:^.'pzulwDrasuawtraos

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Figlurc65. Ducalyplusniclor.reda disLribulion,and buds, fru;ts, adult lcavesand sjlhoucrteofA fifth node secdlinplcaf (scale bar = I cm), 'raddoH 6, rqoor8 ,uarpro./rrtu g Jo 3d,{toloH 99 arn8rc

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NuytsiaVol. 8, No. I (1991)

FiBDrc6'1. E. icroscheaa showing compact mallee habit (SE of Newdegarc). 'lrnJJJelndn3 AllualslsuoJ 'Jo eq1pue urnlnuado 1?3ruoJ I qlt^\ spnq elellecrpad dpcurlsrp u?ql leqlB.I poJod4 oqseql urJoJJrp 'g q1qNotsn?uoqnsgJo suuoJ po^eel-r'ro:ruu pue silrJqr,r qll,t'posnJuoJ oq ^3u lI soleolre,rro:l?u pu?{.r"qqloours'ornlelsre,rolsllu!srlolpaw'x]J.l.o]gsrellplI'llruJrulndnsolpaduqs-laueqIpus puespnq e1eulJunJepuals'ueunJ" snonctdsuoc e tgt,n so,leal,tro:reu dlpnsn 11np'lIqeq eelPu oql,{q pesuelJsnqssllI 'BeJsdnSunuErel - eSuegSuq:r15eq13o oolplu uotutxoc ? aqol poslu8otol1r\ou sl DwauotltuDxfl'se\ Jods polJelloJ f,lerur pue enJsqo us se(g16l) eppueddlqJ.{q pepreSeg saro,11 'proqnc 'u,uorq-,(uE 'u.lc o1lucuaqds-qns 1q311pae5 S'0 x S'0 ',t1o113,4 'elnJa o1'rulndnc ol padeqsloueq'olqlacrpod ,{1uoqs llnrg oIEddJe^ ol ellqm sle,tolC 'IsJluoJ 'uJ 'ttuoJlsnJ'alulloJlpod,{nroqs unln ado g'0x ,'I ol spnq'3uo1ur3 ,'0 ol salJunped 'uoorE '11np'ulc 'uotunJu :pore.rou-II ol dnsacueJsoJouul 1q?11 9'0 x 6'0ol snonJrdsuoJuJppns B 'snorqu18'uoo:8 ',{sso13 'uc qlr,r 'urql rJaeurt01 eleloecuel ,(1,t'ouuu se^sal llnpv dpqBtls a 1 x g1 '8un?uJal[€ 'sr?d ol'el8loJsuel ot ele^o ueql 9-z JoJollsoddo Sululuursr 3u11pees eq1 ;o sa,reol 'relnpuelE 'u^\oro slelqcuarqJo qlrd osuoprotlleJ pu€ surals doJE qloous qlt,,trtu t ol oell€urV '69 '99 (91 sarn8rg)18I ou'uorlcelloJ 'Q98:D '1d,, rltspuounanlo puz DwauolluDy g toJse :satltl E97:t ,.'lslsnv qluagoyolysnSuo 're^ '>I 'C '-I{ 'A 'gOJ 'r^lq 'ou 'uollrolloJ rensqrs orunpat A :(TEL{ :osl) 19 pr' puowwnlq '€lpuelloH?^oN "XX '3sol^l'cos'leN'llnq'zolnl :ad,(Jl(LV8i EgI:I ,t rauoqluox snldtpxng'61

's!l!lqnsg 'sllos Joasoql uur{l lepsoJq oJ? se^eal sll .(?lcallq^\ f^eoq JoJ ecueJoJsld sll pu?slnclodo po,l\ouuu 'o3B 'llquq eq1'selcunped paualluu ,{tEuors qlla dssolSouoJeq l€q'l so^?ol lJare JJIIS Jell?tussll u! sreJJIpll q3!q^\ur ot1'otsnSuoqns g ol pollp sl{lqBqold satcadspere,r.ocslp .{[uetol slqJ'saroN 'oollBru 'uro3 'DuatlJs 'sotclu 'Ioorc slql Jo orrtlglsllsurs eql ol Surpnlp plle lleJJls oql li.J.oJd't3oloutJa

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Figure 68. Distribution of E. xanthonena nbsp. xantlonet a (a, andD.xa tho ena srbsp.upposita (o), and buds,fruirs, adult feavesand silbouelteof a iifih node seedlingleal of E. xanthonenn subsp.rdnrlrner?a (scatebar = I cm). '(lgIN) 'z.tlll .69 oa?uotltuo{dsqns ow?ltoqt|lox t lo adlpsl arn8u

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Theepithet given by Turczaninowsuggests yellow flowers, butan inspection of scveralisotypes andother collections indicates that the stamens arc white to verypale yellow when fresh and darken in colouron drying.

Thereare two subspecies.

19a.EucalJptus xanthonemaT\rcz srbsp. xsnth.onema (Figures 68, 69)

Colourillustration Brooker& Kleinrg Q99A:l12).

Adult leavesgreen, thin, to 0.7 cm wide.

Specimensexamined. WESTERN AUSTRALIA: 10- 15 km Nof Dumbleyung,on MtPlcasant Naturc Reserveno. 15f97,1985, D. Backshall108 (PERTH); Stirling Rangc Nat. Park, 25 Sept.1975, J.S.BeardT 468(PERTH);4.2 km WofNcedilup-Boxwood Hill Rd,onroad to Ongcrup,6 Oct. 1982, M.1.H.BrookerT 086 (CANB, NSW,PERTH);roadside NE of Newbey'slarm, Ongcrup,6 Oct. 1982, M.I.H. Brooker7681 (CANB, NSW PERTH);Stirling Rangc, 2.5 km alongN boundaryfirc trail from Bluff Knoll Road,7 Oct. 1"982,M.I.H. Brooker 7696 (CANB, NSW, PERTH);0.4 km SE of ChesterPass Road on Bluff Knoll Road,Slirling Rangc, 34'21'S 118' 13'E,21 March 1983, M.l.IL Brooker8029(CANB, NSW,PERTH);Bcaufo Rivcr crossing, Albaly Highway,33' 50'S 117'04'E,21 Nov. 1983,M.l.ll. Brooker 8367 (CANB, NSW, PERTH); 8.4 kn W of Jcramungup- Albanyroadon StockRoad,34' 0l'S 118"52'8,23 Nov. !983,M.1.11. Brooker 8377 (CANB,NSW, PERTH);1.7 km S of Jerramungup,33'57'S 118'55'E,3 March 1,985,M.lll. Brooker8873 (CANB,MEL, NSW,PERTH);E of Amelup,34"16'5 118' 15'E,13April 1985,M.I.ll. Brooker 8951(CANB, MEL, NSW,PERTH);6.6kmSofJcrramungup on Albany road,33' 58'S 118'55'E, 27Nov. 1985,M.1.ll. Brooker 9I2I (CANB,MEL, NSW,PERTH); 2.9 km N of PicsscvilleRoad, 33'09'S 117'03'8,9Dcc. 1985,M.I.l1. Brooke r 9145(CANB, MEL, NSW,PERTH); 37 km SE ofPingrup-Bordcnroad on RPF road,33" 50'S 118" 45'E,27 Jan.1987,M.lJI.Brooker9567 (AD, CANB,MEL, NSW,PERTH);Nccdilup, i mostoppositeloadingshed,33o57'S118.47'E, 13Jan. 1988,M.I.II.Brooker 9866 (AD, CANB, MEL, NSW,PERTH); 110 km NE of Albany,8 Nov. 1978, R.J.Cranfield LlLl (PERTH);lTkmSof Jcrramungup,2lNov. 1979,ri'. DemarzD7844 (CANB, PERTH);Ravcnsthorpe Dislrict, Nov. 1944,C.A. Gardner s.n. (CANB, PERTH); Fitzgerald Rivcr, 11Nov. 1935,CA. Gardner& AJ. Milesi(PERTH); 19 milcs E of Dumbleyung,21 Fcb.1966, A.S.George7560&S.G.M. Cnrr (PERTH); 31 miles E o[ Cranbrook,12 March 195'7,J.W.Green 1168A(PERTH); 34milesEof Cranbrook, 12 March 1957,J.W. Green 1170 (PER-fH)l 2kmEof Jerramungup,3lOct.1975,J.W. Greel4622 (PERTH);5 km NW of Ongcrup,33'55'S ll8" 27'8, 23Oct.1983, K. Hill332,L.lohnson &D. B/axe1l(CANB,NSW, PERTH);Nsideof Stirling Rangc; CamelLatc NatureReserve, N of StirlingRange, SW corneron SaltRiver Road,22 March 1982, S.D.Ilopper 2109 (PERTH);5 km WNW of Ongcrup,4.5 km N ofFoster Rd from OngerupRd, 33" 57'S 118'27'8, 31 July 1982,SD. Ilopper2403 (PERTH); Stirling Range N.P., adjacent to caravanpark, N boundaryof park,34'20'S 118"12'E, 23 Nov. 1983,S.D. Llopper 3591,3592 (PERTH);4.4 km E of QuissRoad on N boundaryof FitzgeraldRiver NationalPark, 33.58'S ll9" l4'8, I Ocl 1987, SD. Hopper6168 (PERTH); Gnowcllen Rd, 6 km N Ellen'sPeak, Amelup to CapeRiche, 1l May 1982,G.J. Keighery4837 (PERTH); Phillips Ranges,n.d., Maxwell3gS (MEL); 3 milesNW of Ongerup,17 Jute 1952,K. Newbey244 (PERTH);5 km NW of Ongerup, 14Sept. 1969, K Newbey2877 (PERTH); 10km Eof Broomehill,13 Jan. 1954, R.D. I?oyce 4789 I puuq1r,n .{1rur;ye slr ol Eulpnlp'eprs fq opls'lsup8e paJe1d'sursoddo 'upelerll ntotg't8o1ow{lg

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NuyrsiaVol.8, No. I (199r) 134

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118"43'E, 23 Oct. lg83,K. Htlt 341(CANB, NSW, PERTH); 11.6 km SEof CowalellupRock, NE cornerof CorackerupNat. Res.,34' 08'S 118' 42'E, 31 July 1982,S D. Llopper2406 (PERTH); 1.5kmSSEofChillinup on StockwellRd,34'21'30'S 118'38'30'E, 1Feb. 1988, A Napier& A. Taylor230 (CAM, PERTH).

Distrlbutionand habitat.Pallinup River ald CorackerupNaturo Reserve arcas, usually on stony breakaways(Figure 74). Growsin low closedoropenforest wilhE. redactaBfooker&Hopperized., E. platypusandE. dnnulata.

Conservationstatus. Locally abundantin disjunctpopulations across a narrowgeographical range. Known on onenature reserve.

Floweringp eriod.F ebruary.

Etymology-Frcmthe Grcck, melano, dark, black and2/ritros' bole, llurrk, alluding to theblack butt'

Nores.This is a recentlydiscovered spccies drawn to theatlenlion of SDH irt 1982by the late KenNewbey, who studied in detailthe vegelation of thccorackerup Nalurc Rescrve as part of his M.Phil.degree at MurdochUniversity.

we considerthe affinities ol E. melanoprir.dto be wilh E. xanthonema,[rcm whichit diffels in the mallethabit, rough basal bark, broader leaves, and occurrclccon brcakaways'and with E. medialis,whichisamallee, not ofbreakaways, with larger lcaves and only loose rough basal bark'

22.Euc{tlyptus prcteterraissaBrcoker & Hopper, sp. nov. (Figures77,78' 79)

fubor ("mallet")ad l0 m alfacorlice pro parle maxima llcvi cremeo,cilrcraceo vel subroseo, saepefragmen(is nondecorticantibus adbasin. Medulla rarnulor um glandulifera. Folio cdulta viridia, hebeta.Inflorescenliae axillaros, ad 15-llorae;pedunculi valde complanati. Alcbastra pcdicellata, fusiformiavel levitcrcurvata, ad 1.5x 0.4cm; opefculum decresccns in apicernlcnuem. Fructus pedicellati,obconici, doliiformes vel cupulati, ad 0.7 x 0.5crn. Semina cuboidea vel lcviter plano- ovoidca.

Typrd:Beaufort Inlet, north side,Western Australia, 29 Nov. 1984,M.l l!. Broaker 8744 (holo: PERTH:iso: CANB, NSW).

A malletto 10m tallwilh mostly smooth, cream, grey or pinkish brlk butolfen with loosely held non-decorticateddead fragments held on the lower 1 m.Black horizontal i'rsect scar s oltenpresent. Pithofbranchlets glandular. Leaves of theseedling remain iD g oppositefor abouf 4 or 5 pain,then altemating,ovate, to 7 x 2.5cm, green to blue-green,glabrous. Adult leaves petiolate, alternating' lanceolate,to l0 x 2 cm, concolorous,dull, green.Inflorescerces axillary, unbranched, to 15- flowered;peduncles strongly flattened, to 1.8cm long. Buds pedicellate, fusiform or slighlly curved, to 1.5 x 0.4cm; operculum tapedng to a finepoint Someoutcr stamens ercct, irner ones partly or completelyinflexed, all fertile.Flowers creamy white to paleyellow. Fruit pediccllate,obconical, cupularor barrel-shaped,to 0.? x 0.5 cmi valvesnot exserted.Sced light blownishgrey, cuboid to slightlycompressed-ovoid.

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attenuate,to 2.6 x 0.5 cm,opercula recurved at the tip; stamens sometimes reduced to a singlewhorl, outererect, inner when present parfly or completelyinflexed, all fertile.Flowers pale yellow. Fruit pedicellate,barel-shaped, to 1.1x 0.7 cm. Seedlight grey-brown,more or lessspherical.

Notes-E. gardneri(blue mallet) is typicallya malletoflateritic breakaways. The trunk ofthe typical subspecieshas imperfecfly decorticated flakes of bark and the crownis easilyrecognised from a distanceby the dull bluishleaves. In thepast several other taxa have been incltdedin E. gardneri. We considerthem to be thedistinct species E. pluricaulis,E. densaand E. rednca whichdiffer in habit,habitat, crown formation and leaf characters (see notes under each specie sbelow). E. gardneri is usedfor roadsideplanting and is a sourceof nectarfor honeyproducers in winter(Smith 1969).

Thereare two subspecies.

23a.Eucalyptus gardneriMaiden stbsp. gardneri (Figures 5d, 10b,80, 81, 82)

ColouriLlusn'aion Brooker & Kleinig(1990: 175)

Budsnarrowly fusiform, matule operculum more than l5 mm long.

Specimensexaminerl. WESTERN AUSTRALIA: Hyden, T Sept.1966,M. Barrru 70 (PERTH)i13.3 kmWSWof Wickepintowards Conigin,32'49'S lITo 22'8,4 May 1983,M.l.LI. Brooker 8094 (CANB,NSW, PERTH); 2.7 ktn E of WedinNth roadon Narrogin-Hanismith road, 32. 55'S 117"43'E,4May 1983,M.l.L!. Brooker 8100 (CANB, NSW, PERTH)I 6.1 km W of Jitarningrail crossingtowards Wickepin, 32' 46'5 l17o57'E, 5 May 1983,M.l L-l .Brooke r. 8107 (CANB, NSW, PERTH);c. 9 km NE of Koldininon Trig Road, 32' 27'S I 18' l7'E, 8 Nov. 1983,M.Ltt. Brooker 8363(CANB, NSW,PERTH);NW of Hyden,32' 18'S I l8' 43'E,9Alg.1984,M.1 .LI .Brookzr8632 (CANB,MEL, NSW,PERTH); NW of Burngup,33o 00'S I l8' 39'8,9 Dec.1985, M .l.LLBrooker 9138(CANB, MEL, NSW,PERTH); 8.3 km N ofNyabingon Kukerinroad,33.26'5 118o 08'8, 9Dec.1985,M.l .Brooker9l43(CANB, MEL, NSW, PERTH);Corrrer ofPiesseville rord and road to north,33' 10'S 117' 03'8, 9 Dcc. 1985,M.l.lt. Brooker.9146 (CANB, MEL, NSW,PERTH): Toorackie,S of Williams,33' l0'S 116'-59'E, L9 Fcb. 1986, M.LH. Brooker 9184 (CANB, MEL, NSW,PERTH); 5 km E of Cadoux,2llly 1986,M.l.H..Broater (CANB, PERTH, NSW, MEL): 26.6rnilcs W of Lakecrace on rd 10Dumbleyung, 4 ApLil 1968, S.C.M. Calr. 683 (PERTH); 9 km NE of Kondinin,ncar lrig point,32' 27'S1 l8' 21'E,27 Jan.1979, M.D. Crisp 5520 (CBG, NSW, PERTH);1 km N of Nurogin-Waginroad, 16 Sept.1984,D. Forematr 729 (CANB);Wagin, 13 June 1920,C.A. Gardner 509 (PERTH); Lyndhurst, nearWagin, 15 June 1920,C.A. Gardnerg (pERTH); Bendering,6Feb.1922,C A. GardnerI230 (PERTH)lLyndhursr, WagiD, T Feb.1923,C A.Gardner 1408(PERTH)I Wagin, 7 Feb. 1923,C.A. Gardner 1908 (PERTH); 4 milesN of Wagin,nr Jaloran "Lyndhurst", Road, C.A. Gardner 1996 (PERTH)I Wagin, 2 Sepr. 1923, C.A. Gardner 1496 (PERTH);HalTi sm i th,6Much 1924,C A. Gardner1615,2115 (PERTH); 22 mitesE of Dumbleyung, 21Feb. 1966,A.S. George 7 561& S.G.M.Can'(PERTH); 5km SEof Kukerin turnoff,39.9 km NE of Dumbleyung,33' 10'SI 18' 05'E,5 Alg. 1982,S.D. Hopper 2456 (PERTH); Bcndering Nature Reserve,18 km ENE ofBendedng,32' 21'30'S118'29'30' E, 14June 1985, S.D, Ilopper 4421 (PERTII);3.25 miles NE ofManmanning,on N sideofAyon Loc. 24909,30. 50'S 117" 09'E,6 July

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(CANB,NSW, PERTH); Upper Dalyup River, 12Nov. 1981,M.LH. Brooker 7118 (CANB, NSW, PERTH);roadside NE of Newbey'sfarm,6 Oct 1982,M.I.H. Broolcer 7680 (CANB); 19.6km N of Rayensthorpe-Jeramunguproad towardsLakc King, NW of Ravensthorpe.33" 25'S 119"56'E. 7 !\ne 1983,M.I.H.Brooker8172(CANB, NSWPERTIT); SheoakRock track,Eof Hyden,32' 24'S I19" 27'E,9 Atg.1984,M.I.H.Brooker8625 (CANB, MEL, NSW,PERTH);13.8kmNof Needilup just S of RPFroad,30Nov. 1984,M.tH.Brooker 8747(CANB, NSW,MEL, PERTH);1 km E of TarcoRoad,33"14'S 119' 23'8,15 July 1987, M.l.H.Brooker9114,9715 (AD, CANB,MEL, NSW, PERTH);5.6km fromMillsteedRd onLong CreekRd, S oflakeKing,15 July1987,M.1.11 Brooker 9717(AD, CANB, MEL, NSW,PERTID; 43 km N of Needilup,33' 33'SIl9" 17'8,16July 1987, M.I.II.Brooker9725(AD, CANB,MEL, NSW,PERTH);24miles NofKojonup, l2December 1951, N.T.Burbidge 3674 (CANB); c.61 krnEof Hydcn,13 Aug. 1965,C-4. Gardner16114 (PEP.TIi)1 Forrestaniacomer,4 km S thcnI km E fromNorseman-Hyden ffack, 32" 27'S 119'41'8,7 Nov. 1983,K. Hill638 &L.lohnson,D. Blaxell,I.Brooker, S. Hopper (CANB, NSW, PERTH);37 km NNE of Holt Rock, 73 km E of Hyden, 32" 26'5 lI9'39'E, 16 Aug. 1978,S.D. Ilopper 1062 (PERT[I);2 kmSW of Middlc Ironcap,32 km ENE of HoltRock,32' 35'S 119' 4 4'E,17 Aug.1978, SD. Ilopper 1093@ERTT{); 9 krn WNW of Ongcrup,6 km ENE of OngcrupRock, 33'54'S 118"2?'E, 31 July 1982,S D. Ilopper2410 (PERTH); 8 km NNW of BanddupHill, N boundiryof West Point Farm, 33'24'S lZ0' 22'E,4 AL;.g.1982, S.D. Ilopper 2439(PERTH); 25 km NE of BanclalupHill, 33" 29'S 120'33'E, 4 Aug. 1982,S.D. llopper 2442,2443 (PERTH); 23kmWof LakeKing settlement, 30.5 km E ofNewdegatc,33' 06'S119" 27'E,5 Aug. 1982, S.D .I Iopper2454 (PERTH);4.2km E ofLakcKing general storc, 9 km WSWofBurkctt Rocks,33' 05'S 119'45'E, 5 Scpt.1982, S D. Llopper2492 (PEF.Ti].); S otJilbadgiNature Resefvc, whcrc tho road turns duc N, 34' 13'S119o 50'E, 2 Scpt.1986, S.D.,l lopper 5403 (PERTH); Frzrnk Hann National Park: 17.7 km E ofRabbitProofFence on LakcKing - Norscmanroad (at major bcnd NE), 33" 00'S 120' 15'E, 27 Sept.1988, S-D. Ilopper 6853(PERTH); Frank Hann National Park: 27 km SW of Nincty Milo Tank on Lake King - Norscmanroad,32'51'S 120028'E, 27 Scpt.1988, S.D. Ilopper 6854 GERTH);W cndofLake Hope;2.3 km SEofHatters Hill - Lakc Hopetrack along lrack to Ninety MileTank,32' 32'30"S 120' 19'8,28S ept.1988,5.D.l lopper 6860 (PERTH):W cnd ofl-akcHopc; 11.4km SW of Ninety Milc Tank trackalong Hattcrs Hill - Lakc Hope track,32'36'00"5 120' 12'30"E,28S cpt.1988,S.D.IIopper 6865 (PERTH); 15 km NE ofHattcrs Hill ak)ngLakcHopc lrack,32o 43'30"S 120' 05'00'E, 28 Sept.1988, S.D. i/opper 6ft67 (PERTH); 4 km NE of Halters Hill alongLake Hope track,32' 48'S 120' 00'30"E,28 Sept.1988,SD.llopper 6874 (PERTH); 20 kmESEofMtGibbs,FrarkHannNational Park, c.40km ENEoflakc King, I 1Au g.1979,K.Newbey 5504(PERTH); 15 km E o[ DunnSwamp, c. 90 km ENE ofRavensthorye,15 Nov. 1980,K. Newbey 8135(PERTH); Ravensthorpe,26 Junc 1924,I?alph & Stanfurd €ERTH);6 milesNof Ongcrup, May 1969,B. RockelA23 (PERTH); 15 milcs N ofRavensthorpe,May 1969,8.A.IlockelA52 (CANB); SWof gateinRabbithoofFence, E oflake Kirrg,T Aug.1968,R.A. Saffi'ey 336 (CANB, E, L, NSW, PERTH).

Distributionand habirar. Southem whcatbclt ftom Ongerupeast to theRavcnsthorpe area and north- eastto theHyden- Lale Hopcdistrict (Figure 83). Forms pure slands in closedor openlow forcstwith sparseunderstorey, usually on heavysoils in broadvallcy floors.Occasionally it is foundon higher groundin gentlyundulating tenain '(ur.I =reqapcs) 'strruJrsprq 'uolnq$s!p Jsat Supees apouqUtJ s Jo etanoqtls pw se^ealllnpe puB ,rurp .dsqns,f,I, ap ntdtlorng.tB etnelc

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Figure 84. Holotype of g. dersa Brookcr & Hopper subsp./e,ra. ') Dqpuv ltlru] puulrquq t.lF$,szrl dsqns,srrp A 98 ,JnBU

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Conservationstatus. Locally abundantin scattered disjunctpopulations, some on naturerese esand manyon undisturbedvacant Crown land east of theRabbit Proof Fence.

Floweringperiod.Newbey & Newbey(1987) recorded peak flowering at Ongerup bctween May and August.

Notes.This subspeciesis frequentlyseen in the southernwheatbelt as a dense-crownedmallet occupyingeither Iowlying sitesor higherground. With its compactstature, blue leaves and yellow flowersit hasconsiderable ho iculturalmerit.

Newbey& Newbey(1987) noted that numbers of Yellow-throatedMiners and Red Wattlebirds rosesharply and peaked during the flowering ofE. denscsubsp. densa (referred to asE. gardneriby theseauthors) in 1978near Ongerup. The subspeciesappears to be an importantnectar resource for birdselsewhere as well (Hopper,unpubl.).

24b. Eucalyptusdensd Brooker & Hopper slbsp. improceraBrooker & Hopper,subsp. nov. (Figures86, 87)

A subspecietypica statura inferiore, habitu fruticoso differt.

Typus:7.2kmW alongAerodrome Road from Lake King - Ravenslhorperoad, Western Australia, 19 May 1987,M.I.H.Brooker 9646 & P. Grayling(holo: PERTH; iso: CANB, NSW, MEL, AD).

Colourillusn'adon Brooker& Kleinig (1990:177).

MaUeeto 3 m tall wilh slenderstems and grey overcoppery smooth bark.Pith ofbranchlets usually withoutglands. Adult leaves to 9 x 0.8cm, dull, bluish green. Buds to 2 x 0.4cm. Flowers yellow. Fruit barrel-shapedto truncate-pyriform,to 1 x 0.8 cm.

Specimensexamined. WESTERN AUSTRALIA: Hopetoun Phins, 8 Nov. 1952,P.H . Barreu 15,16 (PERTH)IRavensthorpe Range, May 1.979,E.M. Bennett (PERTH); 15.1 km from EastMt Barren carpark on HamersleyDrive, 33' 55'S 119"55'E, 10Apdl 1983,M.l.H . Brooker8080 (CANB, NSw, PERTH);20.1 km S of Jenamungupon AlbanyRoad, 34'09'S 118"56'E, 10 April 1983, M.I.H.Brooker808l(CANB, NSW,PERTH);Between BeaufortlnletandJeramungup,3 June1983, M.I.H.Brooker 8158 (CANB, NSW, PERTH); type locality, 19 May 1987, M.I.H.Brooker 9645 &. P. Grayling(AD,CANB, MEL, NSW,PERTII); 11.8km W of Ravensthorpe- Lake King roadon AerodromeRoad, L6lu'ly 1987,M.1. . Brookr 9722 & S.D.Hopper (AD, CANB, MEL, NSW, PERTH)ic. 12 km N of Gairdnertum-off on highway,34"08'S118'57'E, 12 Jan. 1988, M.I.LI Brooker9851 (AD, CANB,MEL, NSW,PERTH);29 km NNE ot CoujinupHill, 33o05'S 120'30'E,l?May 1983,MA. Burgman1335&5. McNee(PERTH)I 1.4 miles from Jerramungup towardsAlbany, 1 Nov. 1968,E.M. Canning6945 (CBG, PERTH);between Ravensthorpe & Ongerup,3April 1968,S.G.M.Curr670 (PERTH); nearNoTreeHill, SofRavensthorpe,3 Dec. 1960, A.S.George 1979 (PERTH); 8 milesS of Ravensthorpe,14 March1957,1.W. Greer 1218(PERTH); 12.5km W of Ravensthorpe,31 Oct. 1975, J.W. Greez4608 (PERTH); 2.5 krn S of Ravensthorpe (Hwy1) along MoLRoad,30 Sept. 1987 ,5.D. Hopper6159 (PERTH);c. 14 km SWof Ravensthorpe, '!.r'upto8 E oeq peleleJeql ol lselluoJ '{SotowtJg rq '(rue1s's.rlnn: puu'f,uew'unld ur1z1)lrqsq ealpu oWol sre;er1eq11de cglcads e\L

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Notes.E. pluricauhs hasbeen referred to asa malleeform of E. gardrcri for manyyea-rs (e.g. Elliott andJones 1986). It differs from E. gardnerirn the malleehabit and smoolh bark, retaining these featureswhen growing on lateritic breakawayswilhin the geographicaltutge of E. gardneri. E. pluricaulisresembles E. reduncarnhabit but is clearlydistinguished by theblue-green leaves and, to a lesserextent, the longer more attenuate buds. It differsfrom E. vqria in its smallerstature, its lessdense canopy, its largerjuvenile and adult leaves and its purplishnew growth.

Thereare two subspecies.

25a.Eucalyptus plurrcaalrs Brooker & Hoppersubsp. p/arlcaalls (Figures 88, 89)

Colourtllustralion Brooker & Kleinig(1990: 178).

An erectmallce with blue-green adult leaves, and wilh inllorescencesup to I l-flowered.

Specimensexaminerl. WESTERNAUSTRALIA:41 milesEof Brookton,22 July 1969,M.l.l L Brooker 1868(PERTH);9 milcs S of Wickepin,3 Nov.1969,M.1.H. Brooker 2254 (CANB, NSW, PERTH); 16miles E of Quairadingtowards Corrigin, 12 July 1970, M.l.Ll . Brooker2637 (CANB, K, MEL, NSw, PERTH);road from The Humpsto Mt Walker,32" 14'S 118' 57'E, 3l Dac. 1979, M.I.l1.Brooker 6745 (CANB, NSW, PERTH);Williams' farm (Hi VaUee)Tootbardi rd, N of Badgingara,30'08'S 115"22'E, 19 Aug.1982, M.l.LL Brooker 7565 (CANB, NSW, PERTH); ToolbardiRd, N of Badgingara,19 Aug. \982,M.lll, Brooker'1572(CANB, NSW, PERTH)i WonganHills, c. 100m S of radiotowcr, 30' 52'SI 1603 8'E, 26 Aug.1982, M.l.l l . Brookerl599 (CANB,NSW, PERTH):c.4 km N of MtMichaudon EW track,30"07'S I l50 12'E,l March1983, M.III. Brooker7989 (CANB, NSW, PERTH); Hill SEol Ml BcDifl,30o15'S 115" 15'8,3 March !983,M.l J L Brooker'801 I (CANB,NSW, PERTH); 15.3 km S of Wickcpintowards Taalblin Lake, 32"53'S117"34'E,4May1983,M.l.LLBrooker8097(CANB,NSW,PERTH);c.l0kmESEof Wickepin,32' 40'S i 17' 35'E,5May 1983,M,I J-l. Brooker'8103 (CANB, NSW,PERTH);4.2 km Nof T junction,SEof MtAdams,29" 27'S 115' 19'F.27May 1983,M.l.Il.Brooker8140(CANB, NSW,PERTH);Newman Block, Chomley road, 19 June 1986.M.lJ'l.Brookcr9358 (CANB, NSW, PERTH);Newman Block, Chomley road, NW of Wagin,33'09'S 117' 10'E, 19 Juue1986, M.II I. Brooker936l(CANB,PERTH, NSW, MEL); Dryandr{t StateForest, Dryandm Block,27 June 1986,M.LH. Brooker 9367 (CANB, PERTH,NSW, MEL); Gura Road,Dlyandla StateForest, 27 June1986, M.l.H. Brooker 9372 (CANB, PERTH, NSW, MEL); E of Badgingarra,30' 34'S, 115'38'E,21 July 1986,M.l.II. Brooker9393 (CANB, PERTH, NSW, MEL); NorthBungulla NatureReserve,23 l:uly 1987, M.LIt. Brooker 9730 (CANB, PERTH, NSW, MEL): c. 16km E of YerramullaRoad, 30' 17'S,115" 27'E,14 4u9.1987 , M.l.L! .Brookef 9738 (CANB, PERTH, NSW, MEL); YandanHill, 30'45'S ll5'36'E, l8 Aug. 1987,M.l.ll. Brooker9739 (CANB, PERTH, NSW,MEL); 29 km NNEof CoujinupHill, 33o05'S 120' 30'E, l?May 1983,M.A. Btu gman1335 & S.McNee (PERTH); Coorow, 21 May 1961,n.S. Colenan(PERTH); Mudge Road, 2-3 km N of Coorow-GreenHeadRd,30'0l'S 115'44'E,6 Sepl. 1984,D .8. Forenan 573 (AD, CBG, HO, MEL, NSW,PERTH); Goomallirtg, 17 Aug. 1920,C.A. Gardner 130 (PERTH); Durnbleyung, 12 Nov. 1931,Gardner & Blackall1340 (PEP.TH): SE slopes of MtLesueur,neir the summit,25 Aug. 1948, CA. Gardner9088 (PERTH); 4 km NW of WongarHills (re:u railwaylinc), 30 Aug. 1976, A.M. George93(PERTH); NW ofBadgirrgana, 13Aug. 196 5,A.5. G eorge 6165 (?ERTH); Northam, Oct.19001ILGregory (PERTH); SEslopeofMl Lesucur, NEofJurien,30' 11'S 115o l2'E,17 Jriy ur{ 8'rS:GJUEd'/ASN'SNVJ) 6691.nloolg H J I/!'286I tcO/'o8u?U SulFuSJoN'dntoulvJo S 'I3olq ssauunos:Gllugd'nasN'gNvJ) LSgLraryoq H I w'2861'1log'peol dn:e8uguo peo11 'l3o ueurroNJo iA l]r)lr'?:(HJxad "4[sN'gNY)) gLgLDloors Hf w'786I g'o8u?u SurTrrlsot 'gNy)) 'o^r6sau p?oruouaprogJoS uDt ? :(HJUgd 9t , nlootg H I W'VL6I|rfV 9 pFro8zlrJ '016I '8nV spre,trol6urssor: ppraEzllggo S solur I'9I :(HJUAd) IILZnloo"tg H IW €'o eseu ppreS4rg'pue1g1y,l1Jo N'speorssortJo N selu t:vI.JVUlSOy NllgJ sglypr rj woxasuawnadg

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SofJerramungup on Albany road,34' l9'S 118'46'E, 10 Apil1983, M.l.H.Brooter 8083 (CANB' NSW,PERTH);1.2 km S along HillsRoadfrom TarinRockRoad,33" 06'S 118' 10'E'16 Dec. 1987' M.l.H.Brooker9835(AD, CANB,MEL, NSW,PERTH);Corackerup Reserve, lFeb 1977'T Evans (CANB, K, MEL, PERTH); 12 milesW of Ongerup,13 March 1957,J.W. Greer 1174(CANB' PERTH);6km NNW of Bluff Knoll: Nboundary(Stirling Range National Park),4.8 km Eof Chester PassRcl, 34' 20'S 118' 15'E,24 March 1982,S D. Llopper2140 (PERTH); 9.8 km SWof Boxwood Hill,3.8km NW ofToompup-Bremer Bay Rcl, 34' 13'S1 18' 43'E,30 July 1982, S D. Hopper2395 (PERTH);2 km SW of TarinRock, Tarin Rock Nature Reserye, 33' 02'S 118' 12'E' 5 Aug. 1982' SD. Hopper2455 (PERTH)i 5.5 km SW of Ward Hill, 21 km E of Wagin on rd to Dumbleyung, 33'17'S11?'35'E,5Sept. 1982,SD.Hoppet2489 (PERTH); 5kmNof MonginingHill' 6.5 km S of CoomelberrupRock,33' 31'S 117'48'E, 1l Sept.1982, S.D. Llopper2565 (PERTH);8.8 km W ofWoodanilling,3.8 kmE of AlbanyHwyon WoodanillingRd,33' 33'S 117' 08'E 24July 1984' SD. Hopper3829 QERTH); Gnowellen Rd, 6 km N EllcnPeak, nr Amelupto CapeRiche, 11 May 1982,G.1.Keighery4838 (CANB, PERTH); Gnowangerup, nolth east,27 Feb. 197 5,O.W. Loneragan L234(PERTH); 11mEof Broomehill,13Jan. \954,R.D Royce4791(PERTH).

Distributionand habirar. Tarin Rock south to stirlingRangeand Fitzgerald N:tlionalPark (Figure 88). Growsin low to tallmallee on highground on fine-tcxluredgravelly loams.

Conservationstdt s. Widespreadin smalldisjunct populations, a few of whichoccul on nature rescrves.

F lower ing p eriod. March-June.

Etymology.Thesubspecific epithet refers to thepredominant colour of thecrown (Lalrn porPhyt eus, purple).

Notes.This subspecies has potential as an orDamental wilh ils purylelcaves and ycllow flowers. It isillustrated in Elliott and Jones ( 1986, p.100) underthe namc D.3arrlaeri. NoIth-western populations nearWooclanillingand Tarin Rock (e.9. B rooket 9835, LIoppet2455,3829) arc more upright and may beintergrades with the typical subspecies, but are still striking in theirpurplc-coloured new growth.

26.Eucalyptus varid Brooker& Hopper,sp. llov. (Figure91, 92, 93)

"mallee" Frutex habituvario, foliis juvenilibus et adultis immaturis glaucis vel flavo-viridibus, nonporphyreis non cinereis. Alabastra pedicellata, fusiformia, ad 2.5 x 0.4cm, operculis attenuatis, reduncis.Fructus pedicellati, doliiformes, ad 1 x 0.6cm. Seminaplus minusve sphaerica.

Iypzs:90km Eof EsperanceP.O.,Western Australia,33' 45'S I22" 47'E,7Feb1989,MI H Brooker 10168,5.D.Hopper & D.Vincent(holo : PERTH;iso: AD, CANB,MEL, NSW). 'raddoH ? Je{oorg ,a.rtr{dlod dsqns raddoH T, ra\oor[ trlnorurlld glo ad,{roloH 06 dnStg

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Nuytsia Vol. 8, No. 1 (1991)

A malleeofvariable habit, low andeffuse or erectto 7 m tall, with roughor smoothbark. Leaves of theseedling remaining opposite for 3-5pairs, then altemating, oyate, to 8 x 4 cm,glabrous, blue- greenor yellow-green.Pith ofbranchletsusually without glands. Adultleaves petiolato, alternating, lanceolate,to 8 x 1.3cm, dull, light bluishgreen, in thefirst 1-2years, older leaves inside crown and rarelyseen, glossy, green. Inflorescences to L1-flowcred;peduncles to 1.2 cm long. Budspedicellate, fusiform,to 2.5 x 0.4 cm, attenuate,recurved at lip. Flowersnot seen. Fruit pedicellate,barrel- shaped,to 1 x 0.6 cm. Seedlight grey-brown,more or lessspherical.

Etymology.From theLatin varirs, varying,refering to thc variablcstature, bark, adult leaf width andhabitat.

Notes.E. yaria rcsemblesE. pluricaulisbut it differsparticula.rly in tholcaf colourandwidth. Tho newgrowLh is bluc-green or yellowgreen, nevcrpurplish, while the adult leavcs arc light bluishgreen and nevergrey. The adult andjuvenile leavesare generallynarrower and thinnerthan thosoof E. pluricaulis.E. varlais pcrhapsmore closely related to E. redunc a, whichhas similar colouration on the ncw growth,but r. reduncadiffcrs in ils broadergreener adult lcavcs,lowcr slatureand consistentlysmooth bark.

Thcrcarc two subspecies.

26a.Eucalyptus voria Brookor& Hoppcrsubsp. r'aria

A mallcewith smool.hbark, varying in habitfrom a dwarfpl:url I m tall to an erectmallcc 7 m tall,not growing in salincsituations (Figures 91, 92).

Specimensexamtned. WESTERN AUSTRALIA: 17.4 miles N of Esperance,18 Scpt. 1971, K.M.AUan163 (AD, CANB, PERTH); 16.3 milcs S of Scaddcn,14 Nov. 1970,.1Baker 77 (CANB): c. 15 km E ofEspcranccon roadto Capelt: Grand,33' 47'S 122"00'E,22 Jttnc1978, D.F. Blaxell 1678(CANB, NSW, PERTH); 18 km EDalyup,33" 48'S 121' 44'E,28Dcc.1979, M.t.H. Brookr 6699 (CANB, NSW, PERTH); I km S of FisheriesRoad on Tyncl's Road,33' 43'S 122"09'E, 9 April 1983,M.I.II. Brooker 8066 (CANB, NSW, PERTH); 19.4km N of FisheriesRoad on DempsterRoad, 33" 36'5 122' 00'E, 9 Aprtl 1983,M.l.H. Brooker 8067 (CANB, NSW, PERTH); corncrDalyup North and Speddingup West Roads, 7 Fcb.1989, M.l .LI. Brooker10166 (AD, CANB, NSW,MEL, PERTH)I45.1 milcs E of Espcrance, 25 March 1968,G.M. Chippendale 408 (CANB, PERTH);Buyi BillanakHomestead, c. 12km SEof CondingupPeak, 19 Sept.1968, N.N. Dozn er 2665 (AD, PERTH)rhcadwalcrs of ThomasRiver, c. 8 km SW of BoyatupHill, 5 Oct. 1968, N.N.Donner 29ll (AD, CANB,PERTH);Loc. 900 in gully lcadingto Yerritupcreek, 26 sept.1968, Hj. Eichler 19989(CANB); 10 miles N of Esperance,20 Oct. 193I, Gardner& Blackall 1093 (PERTH);67miles E ofEsperance,11Dec.1960,A.5. George 2187 (PERTH); 82km EofEsperance on Mcrrivalc Road,between Alexander Road and Daniels Road, 21 Nov. 1986,"r.iV. Green 5170 (PERTH);Maidavale Road, 33'48'S 122'20'E,19 Scpt.1976, R.J. Hnatiuk 761004 (PERTH); Escarymentof tributaryofThomas River, 9.5 krn SSWofBoyatup Hill, 3 km NNW ofBclinup Hill, 33" 49'S 123"02'E,9 Sept.1982, S-D. Hopper255l (PERTH);4km ENE of MtHawes,15 km SW of CondingupPcak ,33' 49'S122' 25'8,9 Sept.1982,SD. Hopper2552 (PERTH);2km from mouth ofThomasRiver(Cape Arid NationalPark),12 Oct,. 1985, G .J. Keighery1833 pBF.'l}l);near Howick Hill, 3 km E of woolshedsof Mt Howick Station, Localion 259, 18 Sepl.1968, A.E. Orchard1066 ' u moulun' Po ua d 3u t n,to1 g

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Distibution endharitdl. Coastalarcas from CapeRiche east to FitzgeraldRiver NationalPark and the RavensthorpeRange (Figure 94). Usually in low or tall malleecommunities on plains and undulatingterrain in sand.On windsweptcoastal slopes it is reducedto an almostprostrate shrub in denselow heath.At the type localityit occurson the lower slopesof a lateriticrise andon nearby surroundingplain withE. goniantha,E. redacta,E.tetragona, E. angulosaatld Banksiamedia.Inthe RavensthorpeRange it is an upright mallee occurringwith E. uncinata,E. aff. occidentalis, E. conglobata,E. eremophilaand E. desmondensis,

Conserlationstalus. Occurs in smalldisjunct populations, most ofwhich areon reserves,including theFitzgerald River NationalPark.

F lowerin g p eriod. Not known.

Nole^r.As discussedearlier, the nameE. reduncahas been mis-applied for manyyears. It is now recognisedto beofres{ricted south coastal dislribulion and charactefised by lhe mallee habit, finally glossyadult leaves, long uncinate buds and pale yellow flowers. These latter characters indicate ils affinity with E. varia,E. gardnerl,E. densaattd E. pluricatLlis.

A notablefoature of E. redunc a is thallh e canopyis almostalwrys predominantly dull andgreen. Most leavosappear to retainthe dull wax of new growthfor a long period.The glossyadult leaves arerare and usually confined to olderwood inside lhe canopy. Also, thenew growrh of E. reclunca is yellowgreen, a featureshared with E, vdria,and contrastirtg with the purplish-blue new growlh of E. gardneri,E. pluricaulisand E. densa.

Thepreviously broad application of thenamc E. rcdw1caIo malleesexlcnding well inlandand to thenorlh of typicalpopulations on thesouth coast parallels tho taxonomic hislory of manyother south-westAustralian groups (e.g. Borya nitida (Liliaceae) and its alliesrecently described by Churchill(1987)). Many generaand speciesgroups were first collecl"edby botanistssuch as Labillardidreand Robert Brown who travelledwilh sea-goingexplorers along the south coast. Subsequenttreatments ofthe flora used the south-coastal taxa as a bascline,and often placed within themsimilarlooking taxa collected inland and further norlh. We anl.icipalcthrt [uturedetailed studies of south-wcsterntaxa will bringto lightother examples of thistrend.

SubseriesDesmondenses

Eucalrptusser. Leeispennae subseries Desn0rdensrs Brooker & Hopper,subser. nov.

A subserietlpicacaulibus gracilibus, canopio macilentopenduloque, alabaslds brevibus crassisque, filamentisstaminum omnium inflexis differt.

E. desmondensisMaiden & Blakelv

Malleewith smoothorbasally rough bark. Stems slender, crown thin, finally drooping. Branchlets shinyred with wax overlay;pith glandsabsenf. Peduncles stout, flaltened, widening distally. Buds shortand thick. Operculum t equalin lengthto hypanthium.All sfaminalfilamenls inflexed. Flowers vellow. Seedsoherical. efuoqlsue^Q{Jols€6-tluou-ls?aol puB'looddnllnJ?uuJpuep?orrrotr^lJolsa,r ol eSuvgad:oqlsua,reg 'sJod) .g eWuJ peeJdsepr/t\ pue luepunq? sr sorcods eq11eq1 pegodar oq,r CuuroJ lleuuoq Iq ^a^ns ',.srqlJo 'a8ueg luocere ,(qpatrroddns sr srqJ lsuoeql ol eJmlslpouros pue ed:oqlsua^?Ueql Jo eo.Ie 6ErBI.{lrsJ?ra^opqnqlqsrp,,sltl tu4 pel?ls(096I) roupreC'peor unotedoH-edloqlsue^egoql Suop uees,{lrs?elsour q lnq uorlnqrlsrypolJlsser!osr slsuapuotusap sntd(.pJng'ptlqDq puo uo\ q!.Os!O

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'(Z9I 'uollDtsn :066I) Arurol>I? reloorg ! rnop)

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'o8ueg edroqlsue,relleql uo pueol luacBlpefrlunoc ,t1IIq ^\ol serdnJJolreJoq^r lclqsry edjoqlsue^E{ eW ol peulJuocsr sauasqns sql ur sepodsel8uls oqJ

- .H.I.I/tl 9Lt oeoceu,{l l u.pt?l^l ?t!r./adsr&I sauas shtd|pnq :raddoH ( uaqdals ? ra)loolg r76 Nulrlis vol.8. No. I (I991) on bothsides of theCarlinup road, pa.rticularly on granitic sands (Figure 98). It grcwsas an emergent from heathwith associatessuch as A//ocqsuqrina campestt'is.

Conseryationstatus. Vulnerable. No populationsare known [o be on conscrvationreseryes. All are on Crownland covered by miningleases or onprivate property. The species is locallyabundant, and recoyerswell from thelignotuber after bushfires.

F loweri n g p erio d. May to Novernber.

Notes.E. desmonder.fisis the only species in subseriesDes mondenses.Itdiltcrs from all otherspecies in thescries by thepeculiar slender straggly habit somewhat reminiscent of lhe quiteunrelated E. sepulcralisF.Muell. and E. lansdov,neanaLF,.Brown. The peduncles are stout, the buds are fat andrelativcly shofi, and all staminal filaments arc inflexed. All inl'loresceDceslructures are glaucous. Theflowers are yellow. This combination of chalaclcrsmakes il clcrrlydivclgcnt fiom subseries Levispermae,several species of whichgrow in closeproximity, viz. D. recluttca,E. phaenophylla stbsp.interjacens, E. clivicola,E. gardneri stbsp.ravet$thorpetlsis and E. (lansa.

Nuturalhybrids

Systematicsludics ofnatural hybridisation ineucalypts have bccn much neglccled, although some datahave accumulated (Griftin e r a/. 1988).In thecase of lhe le,rsy.re,'r?d?, atfcmpls to reviewthe importanceof naturalhybridisation would have been futile before lhe currcnt revision in viewof the unusuallylarge nurnber of undescribedlaxa we have documenlcd.

In anyevent, Griffin et a/. ( 1988)noted thal no records ol intelspecil'ichybrids within the series existcdin Chippendalo and Wolf's ( 1984) EUCALIST hcrbiuium datl blse, nor in thelilerature. Two 'B pufativehybrids with membc$of olherseries of sectiot'r isactorfu werelisled, onc erch with a memberof seriesCor utae(i.e. E. gomphocephalax E. u,atuloo- whichwe havebeen unable to relocateand considcr a doubtfuldctcnnination bccause the putaliveprrcnts are allopatric rnd tuonomicallyisolated) and serics LaJo phleboe (E. lotophleba x E. u,artdoo- well documented, see below).No natural hybrids bctween Zevr'sperrrrze specics and thosc ofother sections within subgenus 'Symphyomyr tus' weLcknowlr.

Griftrnet al. (1988)supporlcd Pryor and Johnson's (1981) conclusion thrt membcrsof section 'B 'Symp iseclaria'had a" lowpropensity to hybrid ise". lt wass uggestcd lh0t, wilh in hyonyrtus',"the largesection -Bi.rcctaria (113 species) scems to hrvcthe most well developed barriers at thespecies level,since inter amd even inlraseries combinations are rare." Howcver, Griffin e/ ai. (1988)notcd thatsuch indicalioDs could be due to inadequatesrmpling or taxonomicinteryretation, rather than reflectingreal varialion in ralesof naturalhybridisation.

In thecase o[ lheLeyispennoe, we htve establishedthrough fi01d, gl0sshouse and hcrbarium studiesthe occurrcnce of naturalhybrids derived from many species combinittions. Specimens are citedbelow for six illterseclionalcrosses. seven inlersedes crosses and 12 crosses wilhin the series Levispermae.Becausemost of thesehybrids are easily overlooked (we collected specinrens from all putativehybrids encounlercd during the study, and these were few), if secmsprobable that additional combinationswill bediscovered in thefuturc. -689t0

'(urr I = req al?cs)JselSullpees tpou qUlJ pJo alanoqlls puv sa^eol lFp?'slrruJ'spnq puB'uo]rnqllslp trtt?rotusap sni\ornl 86arfi$

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'reddoH eyaal1til4 - uapIeINamuradslAt| saues wdqDznt C ueqders 4 rrloorg HlI^l NuytsiAVol. 8, No. I (1991)

FigDre99. Isotype of E. derlro,rdeartr Maiden & Bl.*ely.

I I

'puoursoc 1I^lrPru llqrq eellvtrlns?/aProraraP g 0o I ci nnI:I

'rddoH a€3rE'u,&{- uaprurI arrzads^4 saqass'.t]dq@ry C mqdals ?ra)loorg IlII^l Nuytsia Vol. 8, No. I (1991)

Theevidence for hybridisationin individualsdepends on morphologicalintermediacy between putatiyeparents in habit, bark, canopy,leaves, buds, fruits and seeds,combined with the rare occurrencein thc field in mixedpopulations ofboth parents.Where the parental taxa diffcr in their cotyledonsand seedling atlibutes, wc havcalso grown progeny of theputative hybrids to testfor segregatron.

We list belowspecimens that mcot these criteda. Rather than name these hybrids with theirown epithet,we haveused the hybrid formula ofthe putative parents in alphabeticalorder. In oneinstancc (E.crispata)wehavenamedplants ofpossiblc hybridorigin butin thiscase therearemanyindividuals inknownpopulalions,andlhese arerelativelymorphologicallyuniform. Theydo notvary continuously from the form of oneparent to thatof the otheras is seenin a hybridswarm.

Justasprevious taxonomic ffeatlnents ofthc scrics havc underestimated thenumberofrecognisablc t:xa,past heatments ofnatural hybridisation (e.9. Griffin era/. 1988)zrc hcrc shown to bcinadequate. Untilrevisionaryand extensivefield studies ofmany othcr g rotps 1n Eucalyptus havebeen completed, weurge caution in attemptsto derivcsignificartconclusions aboutmtes ofhybridisation from thcdata basesthat were reviewed by Griflil et al. (1988).

Intersectionalhybrids

Eucalyptusarachnaea Brookcr & Hoppcrslbsp. arachnaea x D. obtusifloraDC.

Specimensemmined. WESTERN AUSTRALIA: brcakawaySSE of Mt Horncr,north of Pinchcr's Road,29' 10'S115' 08'E, 4 Feb.1985, M.LI L Brool

Eucd,lyptusconglobata (R.Br. cx Bcnth.)Maiden x E. wandooBlxkely slbsp.wandoo

Specimenexamined. WESTERN AUSTRALIA: 3.7 km from highwayon road to Woodanilling, 33'33'S 117'07'E,8 March1988, M.I.II. Brooker 9897 (AD, CANB, MEL, NSW,PERTH).

Eucalyptusdesmondensrs Maiden & Blakelyx /J. incrassataLabill.

Specimensexamined. WESTERN AUSTRALIA: c. 10 km S of Ravensthorpeon Hopctounroad, 33' 36'S120' 08'E,23 Jlune1978,D.F. B/arell 1696 (NSW, PERTH); hack toZ . benneuiae,c.50m off roadN of Mine, 33'38'S 120'08'E, 12 April 1985,M.LII. Brooker8943 (CANB, NSW, PERTH).

Eucalyptus incrassatdLabill. x E. nedialir Brooker & Hoppcr

Specimensexamined. WESTERN AUSTRALIA: 0.9 km S of Bluff Knoll road on firetrail ncar ChcsterPass road, Sftuling Range, 10 Oct. 1982,M.I-IL Brooker7735 (CANB, NSW, PERTH); 6.5km WNW of Bluff Knoll, 0.9km SW of Bluff Knoll Rd, 34" t9'S 118"13'8, 5 Mav 1982, T

'(HIUAd) gggYnddoP q'g'9961 cun167'puo113urpe:run|-4ro1Suoppeog qgoSl ur8ueq;o A ull8'Z:(HJUS A S6EVnddoq QS '986I 'S.9I 'pN eunl EI .LII S,,08,I0"Zt {lo^ Jo eprss uo SuIIIeqInqol JJo-urnlJo a ru 09 'Surps.nBno "IAhl'gNVC 'CV) Jo td ur{ 9I :(HJUAd'nd.SN 9896.ralootgH I n'L86I l\rdY tz 'a.Ig nloorg "9II S.tt oIg'aulHtelJ :GJuad'asN'1r ^I'gNVl'oY) S896'V896 H I W '1,86I 'pU :(HIUAd IIrdV EZ'A,rt.9II S.tE.I€ uolrequnquo IIeH u,4tolqsrrlulo{ uIrI9'9 '^\sN 'gNV '9,9t 'p11ut8uro3 "IaI^I J'qY) LLg6Dloo.ts H f w' L86rqrr8hl I € .rI1 5.p7 .76 'ulale{3lld '60IZl 'paurulDra -urde1cr16 Jo crlqs e^rosc{ :VITV)IJSOV N1I1IISEAA. sudtarrdds

orprrrd dsqnsf,lo{Elg o1puna'fl x'\luegDqalqdoxol sntd(plng

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sntdqDrns laddoH uatl&ls J3:loorg I8I ae.rsufw uaprgl "rw6dst^qsauas O ?8 HIl a

Nuytsia vol. 8, No. I (r991)

Eucalyptus phaenophylla Brooker & Hopper s\bsp- phaenophllls x E. spathulata Hook var' spalhulata

Specimenexamined. WESTERN AUSTRALIA: 0.3 km N of BeaufortRiver, Albany Highway' 37 Ocr.197 5. J.W. Green4625 (PERTH).

Eucaljptusphdenopftylta Brooker & Hoppersttbsp. phaenophylla x E ' uncinalaTrjfcz' specimenexamined.. WESTERN AUSTRALIA: 13kmENEof Amelup,intelsection of sandalwood Roadand Salisbury Road, 5 Oct. 1981, SD. Hopper6185 (PERTID.

Eucallptus recondila Brooker & Hopper ined.x E. xanthonema Turcz. subsp- appositaBrcoker& Hopper specimenexamined.]wF/S IERN AUSTRALIA: StirlingRange Nationzll PaIk, adjacent to caravan (PERTH)' Eark,Nboundary ofNP,34" 20'S 118" l2'E,23 Nov. 1983,5.D Llopper3588' 3589,3590

Eucallplus spatlla,lntaKook. ''lar.spathulata x E. wandooBlakely srtbsp wandoo specimenexamtned. WESTERN AUSTRALIA: 3.7 km from highwayon road to woodanilling, 33'33'S 117'07'E, 8 March1988, M.I.tl. Brooker 9898 (AD, CANB' MEL, NSW'PERTH)'

E. ser.Levi\perma€ hybrids

EucalyptusabditaBrooker & Hopperx E. ardchnaeaBrooker& Hoppersubsp. arachnaea

Specimenexamined. WESTERN AUSTRALIA: MtMisery, 18August 1988, M.1 lI. Brooker10031 (CANB).

E ucalyptusaraclmaea Brooker & Hopper subsp-aracLnaea x E . capillosaBrooker& Hoppersub sp' capillosa

Specimensexamined. WESTERN AUSTRALIA:4 km W of MtBcbb,32'05'S117" 47'E, 13 June 1985,S D. [Iopper 4398(PERTH); Sorenson's Nature Reservc, 8.7 km W of Babakin'32" 07'30"5 tIl' 55'E, 13 Jvrc 1985,SD. Hopper4403 (PERTH).

Eucalyptusarachna€o Brooker & Hopper s\bsp.arachnaea x E. wandooBl:akely subsp. ),at'd"

Specimensemmined. WESTERN AUSTRALIA: 12.6km E of rail crossingat Carani,30'59'S 116' 31'E,26 Aug.1982,M.I.H. Brooler 7589 (AD, CANB,NSW, PERTH).

Eucsllptus capillosaBrooket & Hoppersubsp. capillosd x E. subsngusta(Blakely) Brooker & Hoppersubsp. celina Brooker& Hopper '(HJrigd) '19tr7nddog 1Jg'7861 3r1Y9Z'A,0V .9Il 'sl?H 'drqsu.4\ol s,Ig.0g {3otuotrt Jo gN ru)t 6 sllrH uBSuoldJo .&\N ru{ 8 :GJuad'nd,sN 'gNy)) 'qod9I'fl,89.9II Lg6Ln4ootg HI W'886I S,69o0g'stllfl u?8uolA spre.'rol uISulFJ 'ld.SN'S ',H '2861'EnY '9,0t JoiIN rul €'9I :(HJUSd NYJ) 969L tzlool| f I I 9Z "9II S.ZS.09 'd€3 'J 'dolo8pu ol p?orJo d\ urll I SIIIHuu8uol& :VI'MIS1V NdS.lSE,4&p autwoxa suawnadg

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'reddoH aEary'u&{ - uaptel l ?lz Ddst^"7 sauas snrltlDcng O uaqdelS ?9 r3{oorB H'I t{ Nuylsia vol. 8, No. I (1991)

Eucaljptuswandoa Blakely subsp.wandoo x E. xanthonemaTnrcz.s\bsp. xanthonema

Specimensexamined. WESTERN AUSTRALIA: Berufort River Crossing,Albany Highway' 33'30'S 117'04'E, 21 Nov. 1983,M.I.H. Brooker 8368 (CANB, NSW, PERTH); 3.7 km E of AlbanyHwy on WoodanillingRd, 33' 33'S 117"08'E,26 Nov. 1987, M.LLI.Brooker 9828 (AD, CANB, MEL, NSW,PERTH).

Eucalyptuswandoo Blakely subsp.wandoo x E. xantllonemeTurcz. subsp. ttpposita Brcoker & Hopper

Specimensemmined. WESTERN AUSTRALIA:2.2 km alongN boundaryfiretrail from Bluff Knoll Rd, Stirling Rango,7 Oct. 1982,M.I.H. Brooker1694 (CANB, NSW, PERTH);2.5 km alongN boundaryfire trail (Stirling RangeNationalPark) from BluffKnollRoad, T Oct.1982,M.1.[LBrooker 7697(CANB, NSW, PERTH); 0.4 krn SEof ChesterPass road on Bluff Knoll road,Stirling Range, 34' 21'S118' 13'E,21March 1983,M.LLI.Brooker8030 (CANB, NSW,PERTH); SaltRiverRoad, Nof StirlingRange N.P.,22 Feb. 1985, M.LLI. Brooker 8868 (CANB, MEL, NSW, PERTH): Stirling RangeNalional Park, 100 km S of Bluff Knoll Rangcr'sresidence, 34' 19'30'S118'11'15'8, 2l Mry 1989,A. Ro.se1063 (PERTH).

Intergradesbetween species of E, ser.Levispermae

The following caseis an itrstanccof discrcte,morphologically uniform populaliorsthat are intermediatebctwccn wheatbclt wandoo and dcsert wandoo.

Eucalyptuscapillosa Brookcr & Hopper subsp.capillosa - E. nigrifunda Brookcr & Hopper intergrades

Specimensexamined. WESTERN AUSTRALIA: 2 km E of Dio HardyRangc road on Dicmals- Menziesroad, 29'43'S 119' 34'8, 16 Oct. 1984,M.LLI. Brooker 8694 (CANB, MEL, NSW, PERTH):23.8 km SE of Perril Vale on Menziesroad,29" 14'S 120'14'E,23 June1.987, M.1.H.Brooker 9662, 9662a (AD, CANB, MEL, NSW,PERTH); 30 km SE of PeronVale Slation, 29'13'S 120"16'E, 13 June1988, n./. Cranfiald6999 (PERTH); Lakc Barlee,18 Oct. 1966, C.A.Gardner 19032 (PERTH).

Acknowledgements

We are gmteful to B. Rockel, C. Sounnessand C. Ranford for growing specimensof all taxa in the glasshouseat CSIRO, Pe h, to SusanPafick and JohnRainbird for line drawings, to JanRayner and Raelene Hick for word processing, to Andrew Brown for cartoSraphy and other technical assistance,and to our colleagesL.A.S. Johnson,K. Hill andD.F. Blaxell for discussions.The late K.R. Newbey alerted SDH to the existenceof several undcscribed taxa of the Levispermae in lhe Ongerup and Jerramungupdisfficts on a field trip in 1982. ('elunoqI3I,.I 'SpIaDI 'ra{oorg 'I 'srd,{pcngol eptnc pI3Jc, (ts6l) pu€ H I I :ssard elr{sJ) .. ?IlBrlsnv (IlelsBe-r{1nos 1o1 v c 'eI-9 :, "lslfnN sllB4snvualsarlA ol rlurepuasep:ds .IF9H..I 'sa,ruaSry (686I) q 'mddoH 'H 'rc{oorg lAauaerql Butsgitruoc(3BacBu4,,,0 srld&a:rg 1o salras,,aau v S PUB I'W '9I€-Itg r'uo{ seDe'lspuPldn ^\eu 33lut Jo uold9csap :9 ?tsr,{nN?llv4snv ura$ad\ lsa^\-qFos 'ra{oor8 3ql uo soloN (986I) ( 'IeddoH pus ll l I"{ aqr pue (""ereu,{I D r,41i1oyg i ,,"";f.i1n*lrrlr{,, tlnor8 leulroJul s '8ZI-€II rt els nN (286I) 'raddoHpue 5e{oorB .lltwsnv uaFa/AJo (aeaceu.{},,1)s?t4r, A Ps?, lsa',. sni[lDtflg'r! sd\cedsqns/'reN G s I{ I I^I '08€-€lt:9?tsl^nN BrlulsnV urslse,^AJo uolSar BlBqlld (986I) ot{turo{ salceds^\au o^\l'(aBocsu,(I/{) trtrrrrq?td'Apw blorl\te!sndq'eng AiA'eqtrloce8papueH I I''{'leloorg '06€-18e:9rlsl,{nN Pllelsnv uralsaiAJo uolsr^rc feuaqu4x el{ru! 'Datznz '(g86:D 'ra4ooJg salceds,{au eettll '(aBacerriy} o\l&!dotol\. g pw t4rsa(lrlr'E s\dtlD'ng J ]'euoc Pu? H l l'{1 prrB 'IEZ-OZZfZBrslAnN ?llerrsnv urelsarrd uro'J sntdqocng lo sepedst aua^lJ (8/6I) J C'll€x"lg H I I^l'rrloorg ptre sl^eDPu nzao! sni&Dc'14 (886I) H I I I ral9ors 'tgg-gzgi 9 "tsr,(nN(aea3?u,{ l) salcods"heu Papler xls Pall 'Itr€-r9€ f9 ersdnN e?acsu,!,{- ,{lalelgaqpuv serrassniKlotng) aql salcadsqnspu? selcads ^\eN (986T) HlI^l'raIooIB uosuqol? ro,&A,,rr?o SocKlDC seltes stltilKlocrg,, Ieuuorq Jo '92-I r' sluoutug .uosgqol4'lo/fida,u,r,l,4oselJasqnsaqluopesgqs'4d(1,,,?snueSeqlJoJaJ,l,^o.saFasr'^6uv.(I86I)'H.I.W..lo{ooJg

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raloorg 'H e7a.e$lw - tr;{wl taoturads!^4 selraswiltl\rng raddoH C ueqdals?t I I^I 186 Nuytsia Vol. 8, No. 1 (1991)

"Field " (Inkata Brookef, M.I.H. and KleiniS, D.A. (r990). Guide to Eucalypts,vol. 2, south-westemand souhem Australia Press:Melboume.)

Burbidge,N.T.(1952).ThesignificanceofthemalleehabitinS1]6dlPl..t.ProcRoySocQueensland62:73-78' cam, D.J. and carr, s.G.M. (1969). Oil glandsand ducrsin Eucdrypr"r L'H6rit. I. The pt oem and the pith. Austral. J. Bot. 1'7t471-513. .|he carr' D.J. and carr, s.G 'M. (|9s0,. lzhna,n,aade,' a natural Sroup of westem Australian Eucalypts. Austral' J. Bot' 28:523-550.

r carr, D.J. Can, S.G.M. and Lcnz, J'R. (1986). Leat venationi\ Ducalrplrr and olher Seneraof : implications fo systemsof classification ofvenation. Austral. J. Bot.34:53 62.

"Eucalyprs Chippendal€,c.M. (1973). of rhe Wcstem Ausrralian Goldficlds." (Australjan Govemmeni Publishing Scrvice: Canberra.) ''Flora chippendale, G.M. (198s). Eucat,p|||s, A|lSopho|a (Myrtaceae). of Austr,llia'' vol. 19, (Australial Govcnrmenl hblishing Service:Canbena )

Chippendale,c'M. andwolf, L'I' (1984).EUCALISTi con)puteriseddnll rctricval syslcm fo r 'lcd 4,/,'IlJ(Mydaceac)' Austral' Forst Res.14: 147-152.

"Flora (Australian Govcmmeni PublishiDg Sclvice: Churchi , D.M. (198't). Borla (I-iliaccae). of Auslralia" 45i 268-2?9. Canbena.)

Davis,P.H. and lleywood, v.H. (1963)..,PrinciplcsofAngiosperm ." (olivef and Boyd: Edinburgh ^nd London.) "Encyclopaedia Elliot, W.R. snd Jones, D.L. (1986). of Australj^n Plants Sui(ablc for Cultivation." Vol 4. (Lothian: Mclboume.)

"The Erickson, R. (1969). Drummonds of Ilawthomdcn." (Larnb Patcrson:Osbomc Park' Pc(h )

c Ardner,c.A. (r 945). Taxonomy and thc spccic! conccpr w irh spccial rcIcrcnc} to Eltcarypnrs. Au stral.For. 9 | ?- I I .

cardncr,C.A.(1952-1966).,.TrcesofWesrcmAusrralia"scrics.J. Agric. WcstcmAustralia (Govcmment Printcrr Pcrth ) "Eucalypts Garclner,C.A. (19?9, 198?). of Westcm ADstralia." (Covcmment Printerr Pcrth.)

"Australia " Giles. E. (r889). Twice Travcrscd (Sampson,Low, Marslon, Scarle& Rivington I London )

" " Grant, V. (1981). Specialion 2nd edn. (Columbia Univers;lv Press:New York') grddl)pl,rr criffin, A.R., BurSess,I.P. and Wolf, L. (1988). Pattems of natural and manipulated hybridisation in lhc gcnus L'H6rit. - a rcview. Austral. J. Bot 36:41-66.

tlickey, L.J. (19?3). Classilicalion ofthe architcciureof dicolyledonousleaves Amer. J Bo1 60: I7-33

Hill, K.D. (1989).Malleeeucalyprcommunitjcs:theirclassificaiionandbiogeograplry..l'rJ.C.Noble&RA.Bradstock(edt .,Mcdirerran€anLandscapes in Ausrralia: Mallee Ecosysremsand rheir Manasemcnl", pp 93-108. (Cornmonwealth and Scientific Induslrial RescarchOrganisationr East Melboume )

Hill, K.D. and Johnson,L.A.S. (199r). Sysrcmaricsrudies in the eucalypts- 4 New taxa in Srcdl)rttur (Mynaceac). Telopea4: 321-349.

"A " Ilolliday, I. and Watton, G. (r980). Field Guide to Eucalypts (Rigbv: Svdney.)

Hopper, S.D. (19?9). Biogeographicalaspccls of speciationin the southwcsiAustralian flora. Annual Review ofEcology and Systematicsl0: 399 422.

Hopper, S.D. (19S2). An excorsioninto southemWestem Australian eucalypls.Swans l2 (1): ll-17

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Indexto new speciesand subspecies of Eucalyptusser. Levispermae C)agenumber for maindescription in bold)

E. abdita L,',7,29,34, 59, 60, 67, 62, tgl t,3, 13,t7 , t 9, 28,62, 6E,76, 104,109 E. arachn4eas[bsp, arachtaea l,'7,31, 61, 66, 67 , 6E,',76, 160,179,181 E. arachtwea s\bsp. arrecta t,'7, t5,29,66,70,'.7 1,72,76 E. copillosa 1,7, 16, 1',7,18, 23, 28, 37, 4t, 5r E. capillosa snbsp.capillosa 1,7, 13,18, 29, 41, 42, 43, 44, 45, 47, 105, 181, r82, 183 E. capillosa subsp.poficlada t,7 , t3, 34,42, 45 , 46,47 , 55, t05 E. clivicola 1,7, 10,16, t7 ,2t, 29, 92,94 ,95 , 96,99, 145 , t'l5 E. citpata t,7,15, 16,|',t,21,29.3r, 33,100, 101, 102, 103, 104, 105 E.de$a 1, 7, 16,19, 23, 29, 100,144, 149, |'.13, l'15 E. dtns.t $nbsp.densa 1,7, 13,30, 149, 15t, 152,153, 154, 155 E. densas[bsp, improcera t,'7, t3, 32,L54, 156 E. desmondensis r, 5,6,'t , 17, 19,2r,22, 28,29 ,31, 149, 1',73,l7 4, t75 , tI6, r11, 118,179 E.flarida r,3,'7, 13,16, r7 , 19,20,21,22,23,29,3r,92,97,98, 99, 100 E, gadneri 1,3,5,6,7,8,13, 15, 16, 11,t9,20,21,23,29,70, 100, 143, 144, 149, 154, 155, r59,16r,r't3, 175, 182 E. Sadneri snbsp.Sardneri r,1 ,30, t44, 145, 146,t47 , 148, t49 E. gardnerisnbsp, rdwks lhorpensis r,7,30,145,146, r49 E. heberifolia t,1 ,29,34,5L, 55 , 56,57, 58, 61, 182 E, histophrlla 1,7,t'7 ,23,2a,32,68,',7 6,89, 90, 9r, 92,r80 1,1,17,19,28,30,34,47, 48, 49, s0, 51, 52, 105 E.luleoh 1,1,2r, 23, 32,E2, 83 , 84, 85 E. nedialis 1,'t ,28,34, 125,t3t, 132,133 , 134, r35, 136,143 , 1'19 1,7,13,29,30, 135, 136, 137, 138, 139, 143 t, 1, 11, 19,21 ,22,32, tos, l2l, t22,r23 , 124 E. nisrifundg r,'7, 13,L't ,23, 28,29, 41, 51, 53, 54, 105, 183 E. phaenophylla r,3,1, fl, 19,23,55,62,68,76, 85, 88,89,99 E.phaenophrlla snbsp. interjacens r,7 ,3t,32,55,76,78,79, 80, 81, 82, 175 E. p haenophy I le sttbsp.p haenop hy I la r,'7,28,3r,55,62,13,'14, 75, 76, 78, 82, 180, r 8t, r82 E.plwbaulis 1,7,13,23,28,29,61,10,100, 144, 149, 155, r59, 163, 173 E. pllricaulis s'nbsp.plwicaulb 1,7,23,35,157, 158,159, 180, 182 E. pluricaulissnbsp. poryhyrea r,'7, 14,r't ,23,34, rs't ,160, r62, 180 E.praeternissa r, 1, 16,30ta6 , 140, r4t, t42, t43 E.redurca t,3,4,5,6,1,8,19,24,25,26,29,34,'t8, r$, n5, 135,r43, r44, t59, 163, 164, 768, 169,t7 0, r71 , t72, r't3, t'15 E. reduncavar. anqustifolia 4,5,104,125 E. reduncavzr, elata 4,5,6,35 E . redwDa var melanophloia r,4,5,6,68 E. reduncav{, otjt itra l,6,99,100 E. reduncavfi. redwrca 5 009-t6zt-6160

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