BACTERIA

TAXONOMY Phyla, Classes, Orders, Families, and Genera

Bacteria I. : Genera and Species Bacteria II. Taxonomy: Reclassified Species Bacteria III. Taxonomy: Phyla, Classes, Orders, Families, and Genera

bacteria.martinklvana.com

BACTERIA

TAXONOMY Phyla, Classes, Orders, Families, and Genera Bacteria III. Taxonomy: Phyla, Classes, Orders, Families, and Genera 35 Phyla & 79 Classes & 183 Orders & 430 Families & 3,144 Genera Martin Klvana 2nd Edition | 2020·04·18

bacteria.martinklvana.com Om Try-Ambakam Yajaamahe Sugandhim Pushti-Vardhanam Urvaarukamiva Bandhanaan Mrityor-Mukshiya Maamrtaat

Rg Veda 7.59.12

Contents 1

1 Introduction 3

2 Phyla 5

3 A 7 8 F 71 13 P 104

4 B 37 9 G 99 14 R 177

5 C 57 10 K 100 15 S 179

6 D 66 11 L 101 16 T 182

7 E 70 12 N 103 17 V 187

1

1 Introduction

Bacteria (sing. bacterium: mod. L., ad. Gr. (this book) is an idea of "a list of all genera βακτηρι´ oν, dim. of βακτρ´ oν stick, staff) are belonging to each family, a list of all families living things belonging to the Bacteria.1 belonging to each order, a list of all orders Taxonomy (irreg. f. Gr. ταξιζ´ arrangement, order; belonging to each class, and a list of all classes -νoµ´ια distribution) is an orderly arrangement belonging to each phylum with references to of (living) things into vertical and transverse primary/secondary publications containing series of named units, the so-called taxa (sing. the description of the families, orders, classes, taxon; phyla, classes, orders, families, genera, and phyla"; for the generic references, see and species), each defined by a set of attributes Bacteria I. Taxonomy: Genera and Species. The shared by its members. The names of phyla, material cause of this book includes a computer classes, orders, families, and genera consist of (Windows, Cygwin, Mintty, VIM, Python, Mik- one word (with the first letter capitalized). TeX, Zotero, Xara Designer Pro, EverNote, Trello), The formal cause of Bacteria III. Taxonomy: table, chair, eyeglasses, coffee, peanuts, electricity, Phyla, Classes, Orders, Families, and Genera and the Internet. The efficient cause of this book

3 is a skillful use of the means. The final cause is a be announced in the BACTERIA project on harmony between this book and its formal cause. ResearchGate. The names of phyla (and their classes, orders, families, and genera) are listed in alphabetical References order; the type classes, orders, families, and [1] Woese, C. R.; Fox, G. E. Proc Natl Acad Sci U S genera are not indicated; each reference contains A 1977, 74, 5088–5090. three links (active, ; inactive, ): URL, DOI, and PMID.  This version (2020·04·18) of Bacteria III. Taxonomy: Phyla, Classes, Orders, Fami- lies, and Genera includes the names of 35 phyla, 79 classes, 183 orders, and 430 fam- ilies (published from 1917 to 2019), and 3,144 genera, The latest version of this book is available at bacteria.martinklvana.com, researchgate.net/publication/324528662, and patreon.com/martinklvana. New versions will

4 2 Phyla

Abditibacteriota 7 60 7 Deferribacteres 66 8 Deinococcus–Thermus 66 Aquificae 26 Dictyoglomi 67 27 70 37 71 Balneolaeota 50 71 Caldiserica 57 90 Calditrichaeota 57 99 57 Kiritimatiellaeota 100 Chlorobi 57 Lentisphaerae 101 Chloroflexi 58 103 Chrysiogenetes 60 104 Coprothermobacterota 60 105

5 Rhodothermaeota 177 179 180 Tenericutes 182 Thermodesulfobacteria 183 183 187

6 3 A

Phylum: Abditibacteriota1 Edaphobacter Class: Abditibacteria1 Granulicella Order: Abditibacteriales1 Occallatibacter Family: Abditibacteriaceae1 Silvibacterium Abditibacterium Telmatobacter Phylum: Acidobacteria2 Terracidiphilus Class: Acidobacteriia3 Terriglobus Order: Acidobacteriales3 Order: Bryobacterales4 Family: Acidobacteriaceae2 Family: Bryobacteraceae5 Acidicapsa Bryobacter Acidipila Paludibaculum Acidobacterium Class: Blastocatellia6 Bryocella Order: Blastocatellales6 Chloracidobacterium Family: Arenimicrobiaceae4

7 Arenimicrobium Order: Thermotomaculales4 Brevitalea Family: Thermotomaculaceae4 Family: Blastocatellaceae6 Thermotomaculum Aridibacter Class: Thermoanaerobaculia4 Blastocatella Order: Thermoanaerobaculales4 Stenotrophobacter Family: Thermoanaerobaculaceae4 Tellurimicrobium Thermoanaerobaculum Family: Pyrinomonadaceae7 Class: Vicinamibacteria4 Pyrinomonas Order: Vicinamibacterales4 Class: Holophagae8 Family: Vicinamibacteraceae4 Order: Acanthopleuribacterales8 Luteitalea Family: Acanthopleuribacteraceae8 Vicinamibacter Acanthopleuribacter Phylum: Actinobacteria3 Order: Holophagales8 Class: Acidimicrobiia9 Family: Holophagaceae8 Order: Acidimicrobiales10 Geothrix Family: Acidimicrobiaceae10 Holophaga Acidimicrobium

8 Ferrimicrobium Actinomyces Ferrithrix Actinotignum Family: Iamiaceae11 Arcanobacterium Aquihabitans Boudabousia Iamia Bowdeniella Family: Ilumatobacteraceae12 Buchananella Desertimonas Falcivibrio Ilumatobacter Flaviflexus Family: None Fudania Aciditerrimonas Gleimia Class: Actinobacteria10 Mobiluncus Order: Acidothermales13 Pauljensenia Family: Acidothermaceae10 Schaalia Acidothermus Trueperella Order: Actinomycetales14,15 Varibaculum Family: Actinomycetaceae15,16 Winkia Actinobaculum Order: Actinopolysporales17

9 Family: Actinopolysporaceae18 Scardovia Actinopolyspora Order: Catenulisporales19 Haloactinomyces Family: Actinospicaceae20 Halopolyspora Actinocrinis Mzabimyces Actinospica Order: Bifidobacteriales10 Family: Catenulisporaceae21 Family: Bifidobacteriaceae10 Catenulispora Aeriscardovia Order: Corynebacteriales22 Alloscardovia Family: Corynebacteriaceae15 Bifidobacterium Bacterionema Bombiscardovia Caseobacter Galliscardovia Corynebacterium Gardnerella Turicella Metascardovia Family: Dietziaceae10 Neoscardovia Dietzia Parascardovia Family: Gordoniaceae10 Pseudoscardovia Gordonia

10 Jongsikchunia Millisia Williamsia Nocardia Family: Lawsonellaceae23 Rhodococcus Lawsonella Skermania Family: Mycobacteriaceae15 Smaragdicoccus Bactoderma Tomitella Mycobacterium Family: Segniliparaceae25 Mycobacteroides Segniliparus Mycolicibacillus Family: Tsukamurellaceae10 Mycolicibacter Tsukamurella Mycolicibacterium Order: Cryptosporangiales23 Stibiobacter Family: Cryptosporangiaceae18 Family: Nocardiaceae15,24 Aldersonia Family: None Amycolicicoccus Fodinicola Hoyosella Order: Frankiales13 Micropolyspora Family: Frankiaceae15,26

11 Frankia Salilacibacter Family: Motilibacteraceae27 Salininema Motilibacter Stackebrandtia Order: Geodermatophilales13 Order: Jiangellales30 Family: Antricoccaceae23 Family: Jiangellaceae31 Antricoccus Haloactinopolyspora Family: Geodermatophilaceae28 Jiangella Blastococcus Phytoactinopolyspora Cumulibacter Order: Kineosporiales32 Geodermatophilus Family: Kineosporiaceae18 Klenkia Angustibacter Modestobacter Kineococcus Order: Glycomycetales29 Kineosporia Family: Glycomycetaceae10 Pseudokineococcus Glycomyces Quadrisphaera Haloglycomyces Thalassiella Paraglycomyces Order: Micrococcales15

12 Family: Beutenbergiaceae18 Pseudactinotalea Beutenbergia Sediminihabitans Litorihabitans Tropicihabitans Miniimonas Family: Demequinaceae35 Salana Demequina Serinibacter Lysinimicrobium Family: Bogoriellaceae33 Family: Dermabacteraceae10 Bogoriella Brachybacterium Georgenia Dermabacter Oceanitalea Devriesea Family: Brevibacteriaceae15 Helcobacillus Brevibacterium Family: Dermacoccaceae33 Sediminivirga Allobranchiibius Spelaeicoccus Barrientosiimonas Family: Cellulomonadaceae34 Branchiibius Actinotalea Calidifontibacter Cellulomonas Demetria

13 Dermacoccus Humibacillus Flexivirga Humihabitans Luteipulveratus Intrasporangium Rudaeicoccus Janibacter Tamlicoccus Knoellia Yimella Kribbia Family: Dermatophilaceae15 Lapillicoccus Arsenicicoccus Marihabitans Austwickia Monashia Dermatophilus Ornithinibacter Kineosphaera Ornithinicoccus Mobilicoccus Oryzihumus Piscicoccus Oryzobacter Tonsilliphilus Pedococcus Family: Intrasporangiaceae10 Phycicoccus Aquipuribacter Terrabacter Fodinibacter Terracoccus

14 Tetrasphaera Aurantimicrobium Family: Jonesiaceae10 Aureobacterium Flavimobilis Canibacter Jonesia Chryseoglobus Populibacterium Clavibacter Sanguibacter Cnuibacter Timonella Compostimonas Family: Kytococcaceae23 Conyzicola Kytococcus Cryobacterium Family: Microbacteriaceae36 Curtobacterium Agreia Diaminobutyricibacter Agrococcus Diaminobutyricimonas Agromyces Frigoribacterium Allohumibacter Frondicola Alpinimonas Frondihabitans Amnibacterium Galbitalea Arenivirga Glaciibacter

15 Glaciihabitans Microterricola Gryllotalpicola Mycetocola Gulosibacter Naasia Herbiconiux Okibacterium Homoserinibacter Parafrigoribacterium Homoserinimonas Phycicola Huakuichenia Planctomonas Humibacter Plantibacter Klugiella Pontimonas Labedella Protaetiibacter Leifsonia Pseudoclavibacter Leucobacter Pseudolysinimonas Lysinibacter Puzihella Lysinimonas Rathayibacter Marisediminicola Rhodoglobus Microbacterium Rhodoluna Microcella Rudaibacter

16 Salinibacterium Micrococcoides Schumannella Micrococcus Subtercola Neomicrococcus Yonghaparkia Nesterenkonia Zimmermannella Paenarthrobacter Family: Micrococcaceae15,37 Paeniglutamicibacter Acaricomes Pseudarthrobacter Arthrobacter Pseudoglutamicibacter Auritidibacter Renibacterium Citricoccus Rothia Enteractinococcus Sinomonas Falsarthrobacter Specibacter Galactobacter Stomatococcus Garicola Tersicoccus Glutamicibacter Zhihengliuella Haematomicrobium Family: Ornithinimicrobiaceae23 Kocuria Ornithinimicrobium

17 Serinicoccus Family: Ruaniaceae38 Family: Promicromonosporaceae10 Haloactinobacterium Antribacter Ruania Cellulosimicrobium Family: Tropherymataceae23 Isoptericola Tropheryma Koreibacter Family: Yaniellaceae39 Krasilnikoviella Yaniella Luteimicrobium Order: Micromonosporales40 Myceligenerans Family: Micromonosporaceae15 Oerskovia Actinaurispora Paraoerskovia Actinocatenispora Promicromonospora Actinoplanes Xylanibacterium Actinorhabdospora Xylanimicrobium Allocatelliglobosispora Xylanimonas Allorhizocola Family: Rarobacteraceae33 Amorphosporangium Rarobacter Ampullariella

18 Asanoa Planosporangium Catellatospora Plantactinospora Catelliglobosispora Polymorphospora Catenuloplanes Pseudosporangium Couchioplanes Rhizocola Dactylosporangium Rugosimonospora Hamadaea Salinispora Jishengella Spirilliplanes Krasilnikovia Verrucosispora Longispora Virgisporangium Luedemannella Xiangella Mangrovihabitans Order: Nakamurellales13 Micromonospora Family: Nakamurellaceae41 Phytohabitans Humicoccus Phytomonospora Microsphaera Pilimelia Nakamurella Planopolyspora Saxeibacter

19 Order: None Mumia Family: Jatrophihabitantaceae23 Nocardioides Jatrophihabitans Pimelobacter Family: None Family: Propionibacteriaceae15,44 Psychromicrobium Acidipropionibacterium Order: Propionibacteriales42 Aestuariimicrobium Family: Actinopolymorphaceae23 Arachnia Actinopolymorpha Auraticoccus Flindersiella Brevilactibacter Tenggerimyces Brooklawnia Thermasporomyces Cutibacterium Family: Kribbellaceae23 Desertihabitans Hongia Friedmanniella Kribbella Granulicoccus Family: Nocardioidaceae43 Luteococcus Aeromicrobium Mariniluteicoccus Marmoricola Microlunatus

20 Micropruina Actinocrispum Naumannella Actinokineospora Propionibacterium Actinomycetospora Propionicicella Actinophytocola Propioniciclava Actinorectispora Propionicimonas Actinosynnema Propioniferax Alloactinosynnema Propionimicrobium Allokutzneria Pseudopropionibacterium Amycolata Raineyella Amycolatopsis Tessaracoccus Crossiella Order: Pseudonocardiales45 Faenia Family: None Gandjariella Bounagaea Goodfellowiella Family: Pseudonocardiaceae46 Haloechinothrix Actinoalloteichus Herbihabitans Actinobispora Kibdelosporangium

21 Kutzneria Umezawaea Labedaea Yuhushiella Lechevalieria Order: Sporichthyales23 Lentzea Family: Sporichthyaceae10 Longimycelium Epidermidibacterium Prauserella Longivirga Pseudoamycolata Sporichthya Pseudonocardia Order: Streptomycetales3 Saccharomonospora Family: Streptomycetaceae15,47 Saccharopolyspora Actinopycnidium Saccharothrix Actinosporangium Salinifilum Allostreptomyces Sciscionella Chainia Streptoalloteichus Elytrosporangium Tamaricihabitans Embleya Thermocrispum Kitasatoa Thermotunica Kitasatospora

22 Microellobosporia Streptomonospora Streptacidiphilus Thermobifida Streptomyces Family: None Streptoverticillium Allonocardiopsis Yinghuangia Sinosporangium Order: Streptosporangiales48 Family: Streptosporangiaceae50 Family: Nocardiopsaceae49 Acrocarpospora Actinorugispora Bailinhaonella Allosalinactinospora Desertactinospora Haloactinospora Herbidospora Lipingzhangella Microbispora Marinactinospora Microtetraspora Marinitenerispora Nonomuraea Murinocardiopsis Planobispora Nocardiopsis Planomonospora Salinactinospora Planotetraspora Spinactinospora Sphaerimonospora

23 Sphaerisporangium Family: Atopobiaceae52 Spongiactinospora Atopobium Streptosporangium Fannyhessea Thermoactinospora Lancefieldella Thermobispora Libanicoccus Thermocatellispora Olsenella Thermopolyspora Parolsenella Thermostaphylospora Family: Coriobacteriaceae10 Family: Thermomonosporaceae10 Collinsella Actinoallomurus Coriobacterium Actinocorallia Enorma Actinomadura Senegalimassilia Excellospora Order: Eggerthellales52 Spirillospora Family: Eggerthellaceae52 Thermomonospora Adlercreutzia Class: Coriobacteriia51 Asaccharobacter Order: Coriobacteriales10 Cryptobacterium

24 Denitrobacterium Order: Euzebyales56 Eggerthella Family: Euzebyaceae56 Ellagibacter Euzebya Enterorhabdus Order: Nitriliruptorales57 Enteroscipio Family: Nitriliruptoraceae57 Gordonibacter Nitriliruptor Paraeggerthella Class: Rubrobacteria58 Parvibacter Order: Gaiellales59 Rubneribacter Family: Gaiellaceae59 Slackia Gaiella Class: Nitriliruptoria53 Order: Rubrobacterales10 Order: Egibacterales54 Family: Rubrobacteraceae10 Family: Egibacteraceae54 Rubrobacter Egibacter Class: Thermoleophilia60 Order: Egicoccales55 Order: Solirubrobacterales61 Family: Egicoccaceae55 Family: Baekduiaceae62 Egicoccus Baekduia

25 Family: Conexibacteraceae63 Hydrogenivirga Conexibacter Hydrogenobacter Family: Parviterribacteraceae64 Hydrogenobaculum Parviterribacter Thermocrinis Family: Patulibacteraceae65 Family: Hydrogenothermaceae70 Patulibacter Hydrogenothermus Family: Solirubrobacteraceae63 Persephonella Solirubrobacter Sulfurihydrogenibium Order: Thermoleophilales61 Venenivibrio Family: Thermoleophilaceae63 Family: None Thermoleophilum Thermosulfidibacter Phylum: Aquificae66 Order: Desulfurobacteriales71 Class: Aquificae67 Family: Desulfurobacteriaceae72 Order: Aquificales68 Balnearium Family: Aquificaceae69 Desulfurobacterium Aquifex Phorcysia Calderobacterium Thermovibrio

26 Phylum: Armatimonadetes73 References Class: Armatimonadia73 Order: Armatimonadales73 [1] Tahon, G.; Tytgat, B.; Lebbe, L.; Carlier, A.; Family: Armatimonadaceae73 Willems, A. Systematic and Applied Microbiol- Armatimonas ogy 2018, 41, 279–290. Order: Capsulimonadales74  Family: Capsulimonadaceae74 [2] Thrash, J. C.; Coates, J. D. Bergey’s Manual Capsulimonas of Systematic Bacteriology, 2nd ed.; Springer: Class: Chthonomonadetes75 New York, 2011; Vol. 4; p 725. Order: Chthonomonadales75  Family: Chthonomonadaceae75 Chthonomonas [3] Cavalier-Smith, T. Int J Syst Evol Microbiol 76 Class: Fimbriimonadia 2002, 52, 7–76. 76 Order: Fimbriimonadales  Family: Fimbriimonadaceae76 Fimbriimonas [4] Dedysh, S. N.; Yilmaz, P. Int J Syst Evol Mi- crobiol 2018, 68, 3796–3806.



27 [5] Dedysh, S. N.; Kulichevskaya, I. S.; Hu- crobiol 2008, 58, 2597–2601.

ber, K. J.; Overmann, J. Int J Syst Evol Mi-  crobiol 2017, 67, 498–501. [9] Norris, P. R. Bergey’s Manual of Systematic Bac-  teriology; Springer: New York, 2012; p 1968.

 [6] Pascual, J.; Wüst, P. K.; Geppert, A.; Foe- sel, B. U.; Huber, K. J.; Overmann, J. Syst [10] Stackebrandt, E.; Rainey, F. A.; Ward- Appl Microbiol 2015, 38, 534–544. Rainey, N. L. Int J Syst Evol Microbiol 1997,

 47, 479–491.

 [7] Wüst, P. K.; Foesel, B. U.; Geppert, A.; Hu- [11] Kurahashi, M.; Fukunaga, Y.; Sakiyama, Y.; ber, K. J.; Luckner, M.; Wanner, G.; Over- Harayama, S.; Yokota, A. Int J Syst Evol Mi- mann, J. Int J Syst Evol Microbiol 2016, 66, crobiol 2009, 59, 869–873. 3355–3366.   [12] Asem, M. D.; Shi, L.; Jiao, J.-Y.; Wang, D.; [8] Fukunaga, Y.; Kurahashi, M.; Yanagi, K.; Han, M.-X.; Dong, L.; Liu, F.; Salam, N.; Yokota, A.; Harayama, S. Int J Syst Evol Mi- Li, W.-J. Int J Syst Evol Microbiol 2018, 68,

28 3593–3599. 2012; Vol. 5; p 162.

 

[13] Sen, A.; Daubin, V.; Abrouk, D.; Gifford, I.; [18] Zhi, X.-Y.; Li, W.-J.; Stackebrandt, E. Int J Syst Berry, A. M.; Normand, P. Int J Syst Evol Mi- Evol Microbiol 2009, 59, 589–608. crobiol 2014, 64, 3821–3832. 

 [19] Donadio, S.; Cavaletti, L.; Monciardini, P. [14] Buchanan, R. E. J Bacteriol 1917, 2, 155–164. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, 2012; Vol. 5; p 225.   [15] Skerman, V. B. D.; McGowan, V.; Sneath, P. H. A. Int J Syst Bacteriol 1980, 30, 225–420. [20] Cavaletti, L.; Monciardini, P.; Schumann, P.; Rohde, M.; Bamonte, R.; Busti, E.; Sosio, M.;  Donadio, S. Int J Syst Evol Microbiol 2006, 56, [16] Buchanan, R. E. J Bacteriol 1918, 3, 403–406. 1747–1753.

 

[17] Goodfellow, M. Bergey’s Manual of Systematic [21] Busti, E.; Cavaletti, L.; Monciardini, P.; Schu- Bacteriology, 2nd ed.; Springer: New York, mann, P.; Rohde, M.; Sosio, M.; Donadio, S.

29 Int J Syst Evol Microbiol 2006, 56, 1741–1746. New York, 1919; p 1040.

 

[25] Butler, W. R.; Floyd, M. M.; Brown, J. M.; [22] Goodfellow, M.; Cavaletti, L.; Monciardini, P. Toney, S. R.; Daneshvar, M. I.; Cooksey, R. C.; Bergey’s Manual of Systematic Bacteriology, 2nd Carr, J.; Steigerwalt, A. G.; Charles, N. Int J ed.; Springer: New York, 2012; Vol. 5; pp 235– Syst Evol Microbiol 2005, 55, 1615–1624. 243.   [26] Becking, J. H. Int J Syst Bacteriol 1970, 20, 201– [23] Nouioui, I.; Carro, L.; García-López, M.; 220.

Meier-Kolthoff, J. P.; Woyke, T.; Kyrpi-  des, N. C.; Pukall, R.; Klenk, H.-P.; Good- [27] Lee, S. D. Int J Syst Evol Microbiol 2013, 63, fellow, M.; Göker, M. Front Microbiol 2018, 9, 3818–3822. 2007.   [28] Normand, P. Int J Syst Evol Microbiol 2006, 56, [24] Castellani, A.; Chalmers, A. J. Manual of trop- 2277–2278.

ical medicine, 3rd ed.; Williams Wood & Co.: 

30 [29] Labeda, D. P.; Cavaletti, L.; Monciardini, P. [33] Stackebrandt, E.; Schumann, P. Int J Syst Evol Bergey’s Manual of Systematic Bacteriology, 2nd Microbiol 2000, 50, 1279–1285.

ed.; Springer: New York, 2012; Vol. 5; p 546. 

 [34] Stackebrandt, E.; Prauser, H. Syst Appl Micro- [30] Tang, D. P.; Cavaletti, L.; Monciardini, P. biol 1991, 14, 261–265. Bergey’s Manual of Systematic Bacteriology, 2nd  ed.; Springer: New York, 2012; Vol. 5; p 546. [35] Ue, H.; Matsuo, Y.; Kasai, H.; Yokota, A. Int J  Syst Evol Microbiol 2011, 61, 1322–1329. [31] Tang, S.-K.; Zhi, X.-Y.; Wang, Y.; Shi, R.;  Lou, K.; Xu, L.-H.; Li, W.-J. Int J Syst Evol [36] Park, Y.-H.; Suzuki, K.-I.; Yim, D.-G.; Lee, K.- Microbiol 2011, 61, 194–200. C.; Kim, E.; Yoon, J.-s.; Kim, S.-j.; Kho, Y.-H.;  Goodfellow, M.; Komagata, K. Antonie van Leeuwenhoek 1993, 64, 307–313. [32] Kämpfer, P. Bergey’s Manual of Systematic Bac-  teriology, 2nd ed.; Springer: New York, 2012; Vol. 5; p 561. [37] Pribram, E. J Bacteriol 1929, 18, 361–394.

 

31 [38] Tang, S.-K.; Zhi, X.-Y.; Wang, Y.; Wu, J.-Y.; of Systematic Bacteriology, 2nd ed.; Springer: Lee, J.-C.; Kim, C.-J.; Lou, K.; Xu, L.-H.; New York, 2012; Vol. 5; p 1137.

Li, W.-J. Int J Syst Evol Microbiol 2010, 60,  2113–2119. [43] Nesterenko, O. A.; Kvasnikov, E. I.;  Nogina, T. M. Mikrobiol Zhurnal 1985, 47, [39] Li, W.-J.; Zhi, X.-Y.; Euzéby, J. P. Int J Syst Evol 3–12.

Microbiol 2008, 58, 525–527. 

 [44] Delwiche, E. A. Bergey’s Manual of Determi- [40] Genilloud, O. Bergey’s Manual of Systematic native Bacteriology, 7th ed.; The Williams & Bacteriology, 2nd ed.; Springer: New York, Wilkins Co.: Baltimore, 1957; p 569.

2012; Vol. 5; p 1035. 

 [45] Labeda, D. P.; Goodfellow, M. Bergey’s [41] Tao, T.-s.; Yue, Y.-y.; Chen, W.-x.; Chen, W.-f. Manual of Systematic Bacteriology, 2nd ed.; Int J Syst Evol Microbiol 2004, 54, 999–1000. Springer: New York, 2012; Vol. 5; p 1301.

 

[42] Patrick, S.; McDowell, A. Bergey’s Manual [46] Embley, M. T.; Smida, J.; Stackebrandt, E.

32 Syst Appl Microbiol 1988, 11, 44–52. 36.

 

[47] Waksman, S. A.; Henrici, A. T. J Bacteriol 1943, [51] König, H. Bergey’s Manual of Systematic Bac- 46, 337–341. teriology, 2nd ed.; Springer: New York, 2012;

 Vol. 5; p 1968.

 [48] Goodfellow, M. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, [52] Gupta, R. S.; Chen, W. J.; Adeolu, M.; Chai, Y. 2012; Vol. 5; p 1805. Int J Syst Evol Microbiol 2013, 63, 3379–3397.

 

[49] Rainey, F. A.; Ward-Rainey, N.; Kroppenst- [53] Ludwig, W.; Euzéby, J.; Whitman, W. B. edt, R. M.; Stackebrandt, E. Int J Syst Bacteriol Bergey’s Manual of Systematic Bacteriology, 2nd 1996, 46, 1088–1092. ed.; Springer: New York, 2012; Vol. 5; p 1968.

 

[50] Goodfellow, M.; Stanton, L. J.; Simpson, K. E.; [54] Zhang, Y.-G.; Wang, H.-F.; Yang, L.-L.; Minnikin, D. E. J Gen Microbiol 1990, 136, 19– Zhou, X.-K.; Zhi, X.-Y.; Duan, Y.-Q.; Xiao, M.;

33 Zhang, Y.-M.; Li, W.-J. Int J Syst Evol Micro- [58] Suzuki, K. I. Bergey’s Manual of Systematic biol 2016, 66, 283–289. Bacteriology, 2nd ed.; Springer: New York,

 2012; Vol. 5; p 1968.

 [55] Zhang, Y.-G.; Chen, J.-Y.; Wang, H.-F.; Xiao, M.; Yang, L.-L.; Guo, J.-W.; Zhou, E.-M.; [59] Albuquerque, L.; França, L.; Rainey, F. A.; Zhang, Y.-M.; Li, W.-J. Int J Syst Evol Micro- Schumann, P.; Nobre, M. F.; da Costa, M. S. biol 2016, 66, 530–535. Syst Appl Microbiol 2011, 34, 595–599.

 

[56] Kurahashi, M.; Fukunaga, Y.; Sakiyama, Y.; [60] Suzuki, K. I.; Whitman, W. B. Bergey’s Manual Harayama, S.; Yokota, A. Int J Syst Evol Mi- of Systematic Bacteriology, 2nd ed.; Springer: crobiol 2010, 60, 2314–2319. New York, 2012; Vol. 5; p 1968.   [61] Reddy, G. S. N.; Garcia-Pichel, F. Int J Syst [57] Sorokin, D. Y.; van Pelt, S.; Tourova, T. P.; Ev- Evol Microbiol 2009, 59, 87–94. tushenko, L. I. Int J Syst Evol Microbiol 2009,  59, 248–253.

 [62] An, D.-S.; Siddiqi, M. Z.; Kim, K.-H.; Yu, H.-

34 S.; Im, W.-T. J Microbiol 2018, 56, 24–29. York, 2001; Vol. 1; pp 359–367.

 

[63] Stackebrandt, E. Acta Microbiol Immunol [67] Reysenbach, A. L. Bergey’s Manual of Sys- Hung 2004, 51, 449–462. tematic Bacteriology, 2nd ed.; Springer: New

 York, 2001; Vol. 1; pp 359–367.

 [64] Foesel, B. U.; Geppert, A.; Rohde, M.; Over- mann, J. Int J Syst Evol Microbiol 2016, 66, [68] Reysenbach, A. L. Bergey’s Manual of Sys- 652–665. tematic Bacteriology, 2nd ed.; Springer: New

 York, 2001; Vol. 1; pp 359–367.

 [65] Takahashi, Y.; Matsumoto, A.; Morisaki, K.; Omura,¯ S. Int J Syst Evol Microbiol 2006, 56, [69] Reysenbach, A. L. Bergey’s Manual of Sys- 401–406. tematic Bacteriology, 2nd ed.; Springer: New

 York, 2001; Vol. 1; pp 360–367.

 [66] Reysenbach, A. L. Bergey’s Manual of Sys- tematic Bacteriology, 2nd ed.; Springer: New [70] Eder, W.; Huber, R. Extremophiles 2002, 6, 309–

35 318. [74] Li, J.; Kudo, C.; Tonouchi, A. Int J Syst Evol

 Microbiol 2019, 69, 220–226.

 [71] Gupta, R. S.; Lali, R. Antonie van Leeuwenhoek 2013, 104, 349–368. [75] Lee, K. C.-Y.; Dunfield, P. F.; Morgan, X. C.;

 Crowe, M. A.; Houghton, K. M.; Vyssot- ski, M.; Ryan, J. L. J.; Lagutin, K.; McDon- [72] L’Haridon, S.; Reysenbach, A.-L.; Tin- ald, I. R.; Stott, M. B. Int J Syst Evol Microbiol dall, B. J.; Schönheit, P.; Banta, A.; 2011, 61, 2482–2490. Johnsen, U.; Schumann, P.; Gambacorta, A.;  Stackebrandt, E.; Jeanthon, C. Int J Syst Evol Microbiol 2006, 56, 2843–2852. [76] Im, W.-T.; Hu, Z.-Y.; Kim, K.-H.; Rhee, S.-K.;

 Meng, H.; Lee, S.-T.; Quan, Z.-X. Antonie Van Leeuwenhoek 2012, 102, 307–317. [73] Tamaki, H.; Tanaka, Y.; Matsuzawa, H.;  Muramatsu, M.; Meng, X.-Y.; Hanada, S.; Mori, K.; Kamagata, Y. Int J Syst Evol Mi- crobiol 2011, 61, 1442–1447.



36 4 B

Phylum: Bacteroidetes1 Family: Bacteroidaceae3 Class: Bacteroidia1 Acetofilamentum Order: Bacteroidales1 Acetothermus Family: Barnesiellaceae2 Bacteroides Barnesiella Family: Balneicellaceae4 Coprobacter Balneicella Family: Dysgonomonadaceae2 Family: Muribaculaceae5 Dysgonomonas Duncaniella Proteiniphilum Muribaculum Family: Paludibacteraceae2 Paramuribaculum Paludibacter Family: None Family: Tannerellaceae2 Phocaeicola Parabacteroides Family: Porphyromonadaceae6 Tannerella Falsiporphyromonas

37 Fermentimonas Anaerocella Gabonibacter Mucinivorans Macellibacteroides Rikenella Microbacter Family: Williamwhitmaniaceae9 Oribaculum Williamwhitmania Petrimonas Order: Marinilabiliales10 Porphyromonas Family: Marinifilaceae11 Family: Prevotellaceae7 Ancylomarina Alloprevotella Butyricimonas Capsularis Labilibaculum Hallella Marinifilum Paraprevotella Odoribacter Prevotella Family: Marinilabiliaceae12 Xylanibacter Alkaliflexus Family: Rikenellaceae8 Alkalitalea Acetobacteroides Anaerophaga Alistipes Carboxylicivirga

38 Geofilum Family: Salinivirgaceae14 Labilibacter Salinivirga Mangroviflexus Class: Chitinophagia15 Marinilabilia Order: Chitinophagales16 Natronoflexus Family: Chitinophagaceae17 Saccharicrinis Arachidicoccus Thermophagus Arvibacter Family: Prolixibacteraceae13 Asinibacterium Draconibacterium Aurantisolimonas Mangrovibacterium Chitinophaga Maribellus Cnuella Mariniphaga Compostibacter Meniscus Crenotalea Prolixibacter Deminuibacter Roseimarinus Dinghuibacter Sunxiuqinia Edaphobaculum Tangfeifania Ferruginibacter

39 Filimonas Paracnuella Flaviaesturariibacter Parafilimonas Flavihumibacter Parasediminibacterium Flavipsychrobacter Parasegetibacter Flavisolibacter Pseudobacter Flavitalea Pseudoflavitalea Gynurincola Puia Haoranjiania Rurimicrobium Heliimonas Sediminibacterium Hydrobacter Segetibacter Hydrotalea Taibaiella Lacibacter Terrimonas Nemorincola Thermoflavifilum Niabella Vibrionimonas Niastella Class: Cytophagia18 Niveitalea Order: Cytophagales3,19 Panacibacter Family: Bernardetiaceae20

40 Bernardetia Mariniradius Garritya Mongoliibacter Hugenholtzia Mongoliicoccus Family: Catalimonadaceae21 Mongoliitalea Catalinimonas Negadavirga Family: Cyclobacteriaceae22 Nitritalea Algoriphagus Pleomorphovibrio Aquiflexum Rhodonellum Belliella Shivajiella Cecembia Family: Cytophagaceae3,23 Chimaereicella Allopseudarcicella Cyclobacterium Aquirufa Echinicola Arcicella Fontibacter Arcticibacterium Hongiella Arsenicibacter Indibacter Arundinibacter Lunatimonas Cytophaga

41 Dyadobacter Pseudarcicella Edaphorhabdus Ravibacter Ekhidna Rhabdobacter Emticicia Rhodocytophaga Fibrella Rudanella Fibrisoma Runella Flectobacillus Siphonobacter Flexibacter Spirosoma Fluviimonas Sporocytophaga Huanghella Taeseokella Jiulongibacter Telluribacter Lacihabitans Family: Flammeovirgaceae24 Larkinella Cesiribacter Leadbetterella Fabibacter Litoribacter Fabivirga Nibrella Flammeovirga Persicitalea Flexithrix

42 Fulvivirga Hymenobacter Imperialibacter Nibribacter Limibacter Pontibacter Marinicola Rufibacter Marinoscillum Family: Microscillaceae20 Marivirga Eisenibacter Nafulsella Microscilla Perexilibacter Family: Mooreiaceae21 Rapidithrix Mooreia Reichenbachiella Family: None Roseivirga Chryseolinea Sediminitomix Luteivirga Tunicatimonas Ohtaekwangia Family: Hymenobacteraceae15 Thermoflexibacter Adhaeribacter Family: Persicobacteraceae15 Botryobacter Aureibacter Effluviibacter Fulvitalea

43 Persicobacter Cryomorpha Family: Raineyaceae25 Luteibaculum Raineya Owenweeksia Family: Thermonemataceae15 Phaeocystidibacter Thermonema Salibacter Class: Flavobacteriia26 Vicingus Order: Flavobacteriales27 Family: Flavobacteriaceae30 Family: Blattabacteriaceae28 Actibacter Blattabacterium Aequorivita Family: Crocinitomicaceae15 Aestuariibaculum Brumimicrobium Aestuariicola Crocinitomix Aestuariimonas Fluviicola Aestuariivivens Lishizhenia Algibacter Salinirepens Algitalea Wandonia Algoriella Family: Cryomorphaceae29 Antarcticibacterium

44 Antarcticimonas Capnocytophaga Apibacter Cellulophaga Aquaticitalea Changchengzhania Aquibacter Chishuiella Aquimarina Chryseobacterium Arcticiflavibacter Citreitalea Arenibacter Cloacibacterium Arenitalea Coenonia Ascidiimonas Confluentibacter Aurantiacicella Corallibacter Aurantivirga Costertonia Aureicoccus Croceibacter Aureisphaera Croceitalea Aureitalea Croceivirga Aureivirga Cruoricaptor Bergeyella Daejeonia Bizionia Dokdonia

45 Donghaeana Gaetbulimicrobium Elizabethkingia Galbibacter Empedobacter Gangjinia Epilithonimonas Gelatiniphilus Eudoraea Gelidibacter Euzebyella Geojedonia Flagellimonas Gillisia Flavicella Gilvibacter Flavimarina Gramella Flaviramulus Hanstruepera Flavirhabdus Hoppeia Flavivirga Hwangdonia Flavobacterium Hyunsoonleella Formosa Ichthyenterobacterium Frondibacter Imtechella Fulvibacter Jejudonia Gaetbulibacter Jejuia

46 Joostella Maripseudobacter Kaistella Maritimimonas Kordia Marixanthomonas Kriegella Meridianimaribacter Krokinobacter Mesoflavibacter Lacinutrix Mesonia Leeuwenhoekiella Moheibacter Leptobacterium Muricauda Litoribaculum Muriicola Lutaonella Myroides Lutibacter Namhaeicola Lutimonas Neptunitalea Mangrovimonas Nonlabens Maribacter Oceanihabitans Mariniflexile Olleya Marinirhabdus Ornithobacterium Marinivirga Persicivirga

47 Pibocella Sabulilitoribacter Planktosalinus Salegentibacter Planobacterium Salinimicrobium Polaribacter Sandarakinotalea Pontirhabdus Saonia Postechiella Sediminibacter Pricia Sediminicola Pseudobizionia Sejongia Pseudofulvibacter Seonamhaeicola Pseudotenacibaculum Siansivirga Pseudozobellia Sinomicrobium Psychroflexus Snuella Psychroserpens Soonwooa Pustulibacterium Spongiibacterium Riemerella Spongiiferula Robertkochia Spongiimicrobium Robiginitalea Spongiimonas

48 Spongiivirga Zeaxanthinibacter Stanierella Zhouia Stenothermobacter Zobellia Subsaxibacter Zunongwangia Subsaximicrobium Family: Ichthyobacteriaceae31 Sungkyunkwania Ichthyobacterium Taeania Family: Schleiferiaceae32 Tamlana Schleiferia Tenacibaculum Class: None Ulvibacter Order: None Vitellibacter Family: Lentimicrobiaceae33 Wautersiella Lentimicrobium Weeksella Family: None Wenyingzhuangia Toxothrix Winogradskyella Class: Saprospiria34 Xanthomarina Order: Saprospirales20 Yeosuana Family: Haliscomenobacteraceae20

49 Haliscomenobacter Saprospira Phaeodactylibacter Family: Sphingobacteriaceae39 Portibacter Anseongella Family: Lewinellaceae20 Arcticibacter Lewinella Mucilaginibacter Family: Saprospiraceae35 Nubsella Aureispira Olivibacter Membranicola Parapedobacter Saprospira Pedobacter Class: Sphingobacteriia36 Pelobium Order: Sphingobacteriales37 Pseudopedobacter Family: Crenotrichaceae3 Pseudosphingobacterium Crenothrix Solitalea Family: Filobacteriaceae38 Sphingobacterium Filobacterium Phylum: Balneolaeota20 Family: Saprospiraceae35 Class: Balneolia15 Membranicola Order: Balneolales16

50 Family: Balneolaceae40 Dal’Molin, C. G. O.; Palfreyman, R. W.; Aliifodinibius Nielsen, L. K.; Cooper, M. A.; Morrison, M.; Balneola Hansbro, P. M.; Hugenholtz, P. Microbiome Fodinibius 2016, 4, 36.

Gracilimonas  Rhodohalobacter [3] Skerman, V. B. D.; McGowan, V.; Sneath, P. Family: Soortiaceae41 H. A. Int J Syst Bacteriol 1980, 30, 225–420. Soortia 

References [4] Fadhlaoui, K.; Ben Hania, W.; Postec, A.; [1] Krieg, N. R.; Ludwig, W.; Euzéby, J.; Whit- Hamdi, M.; Ollivier, B.; Fardeau, M.-L. Int J man, W. B. Bergey’s Manual of Systematic Bac- Syst Evol Microbiol 2016, 66, 4692–4696. teriology, 2nd ed.; Springer: New York, 2011;  Vol. 4; p 25. [5] Lagkouvardos, I.; Lesker, T. R.; Hitch, T.  C. A.; Gálvez, E. J. C.; Smit, N.; Neuhaus, K.; [2] Ormerod, K. L.; Wood, D. L. A.; Lachner, N.; Wang, J.; Baines, J. F.; Abt, B.; Stecher, B.; Gellatly, S. L.; Daly, J. N.; Parsons, J. D.; Overmann, J.; Strowig, T.; Clavel, T. Micro-

51 biome 2019, 7, 28. [9] Pikuta, E. V.; Lyu, Z.; Hoover, R. B.; Liu, Y.;

 Patel, N. B.; Busse, H. J.; Lawson, P. A. Int J Syst Evol Microbiol 2017, 67, 4132–4145. [6] Krieg, N. R. Bergey’s Manual of Systematic Bac-  teriology, 2nd ed.; Springer: New York, 2011; Vol. 4; p 61. [10] Wu, W.-J.; Zhao, J.-X.; Chen, G.-J.; Du, Z.-J.

 Int J Syst Evol Microbiol 2016, 66, 4243–4249.  [7] Krieg, N. R. Bergey’s Manual of Systematic Bac- teriology, 2nd ed.; Springer: New York, 2011; [11] Iino, T.; Mori, K.; Itoh, T.; Kudo, T.; Vol. 4; p 85. Suzuki, K.-i.; Ohkuma, M. 2014, 64, 3660– 3667.   [8] Krieg, N. R.; Staley, J. T.; Brown, D. R.; Hed- [12] Ludwig, W.; Euzéby, J.; Whitman, W. B. lund, B. P.; Paster, B. J.; Ward, N. L.; Lud- Bergey’s Manual of Systematic Bacteriology, 2nd wig, W.; Whitman, W. B. Bergey’s Manual ed.; Springer: New York, 2011; Vol. 4; p 49. of Systematic Bacteriology, 2nd ed.; Springer:  New York, 2011; Vol. 4; p 54.

 [13] Huang, X.-F.; Liu, Y. J.; Dong, J.-D.; Qu, L.-

52 Y.; Zhang, Y.-Y.; Wang, F.-Z.; Tian, X.-P.; Evol Microbiol 2011, 61, 518–523.

Zhang, S. Int J Syst Evol Microbiol 2014, 64,  875–881. [18] Nakagawa, Y. Bergey’s Manual of Systematic  Bacteriology, 2nd ed.; Springer: New York, [14] Hania, W. B.; Joseph, M.; Bunk, B.; Spröer, C.; 2011; Vol. 4; p 370. Klenk, H.-P.; Fardeau, M.-L.; Spring, S. Envi-  ron Microbiol 2017, 19, 1134–1148. [19] Leadbetter, E. R. Bergey’s Manual of Determi-  native Bacteriology, 8th ed.; The Williams & [15] Munoz, R.; Rosselló-Móra, R.; Amann, R. Wilkins Co.: Baltimore, 1974; p 99. Syst Appl Microbiol 2016, 39, 281–296. 

 [20] Hahnke, R. L.; Meier-Kolthoff, J. P.; García- [16] Munoz, R.; Rosselló-Móra, R.; Amann, R. López, M.; Mukherjee, S.; Huntemann, M.; Syst Appl Microbiol 2016, 39, 491–492. Ivanova, N. N.; Woyke, T.; Kyrpides, N. C.; Klenk, H.-P.; Göker, M. Front Microbiol 2016,  7, 2003.

[17] Kämpfer, P.; Lodders, N.; Falsen, E. Int J Syst 

53 [21] Choi, E. J.; Beatty, D. S.; Paul, L. A.; Feni- Cunha, A.; Egas, C.; da Costa, M. S. Int J Syst cal, W.; Jensen, P. R. Int J Syst Evol Microbiol Evol Microbiol 2018, 68, 982–989.

2013, 63, 1219–1228. 

 [26] Bernardet, J. F. Bergey’s Manual of Systematic [22] Nedashkovskaya, O. I.; Ludwig, W. Bergey’s Bacteriology, 2nd ed.; Springer: New York, Manual of Systematic Bacteriology, 2nd ed.; 2011; Vol. 4; p 105.

Springer: New York, 2011; Vol. 4; p 423. 

 [27] Bernardet, J. F. Bergey’s Manual of Systematic [23] Stanier, R. Y. J Bacteriol 1940, 40, 619–635. Bacteriology, 2nd ed.; Springer: New York,

 2011; Vol. 4; p 105.

 [24] Yoon, J.; Adachi, K.; Park, S.; Kasai, H.; Yokota, A. Int J Syst Evol Microbiol 2011, 61, [28] Kambhampati, S. Bergey’s Manual of System- 2342–2347. atic Bacteriology, 2nd ed.; Springer: New

 York, 2011; Vol. 4; p 315.

 [25] Albuquerque, L.; Polónia, A. R. M.; Bar- roso, C.; Froufe, H. J. C.; Lage, O.; Lobo-da [29] Bowman, J. P.; Nichols, C. M.; Gibson, J. A. E.

54 Int J Syst Evol Microbiol 2003, 53, 1343–1355. [33] Sun, L.; Toyonaga, M.; Ohashi, A.; Tour-

 lousse, D. M.; Matsuura, N.; Meng, X.-Y.; Tamaki, H.; Hanada, S.; Cruz, R.; Yam- [30] Reichenbach, H. Bergey’s Manual of Systematic aguchi, T.; Sekiguchi, Y. Int J Syst Evol Mi- Bacteriology, 1st ed.; The Williams & Wilkins crobiol 2016, 66, 2635–2642. Co.: Baltimore, 1989; Vol. 3; pp 2011–2013.  

[31] Takano, T.; Matsuyama, T.; Sakai, T.; Naka- [34] Hahnke, R. L.; Meier-Kolthoff, J. P.; García- mura, Y.; Kamaishi, T.; Nakayasu, C.; López, M.; Mukherjee, S.; Huntemann, M.; Kondo, H.; Hirono, I.; Fukuda, Y.; Sori- Ivanova, N. N.; Woyke, T.; Kyrpides, N. C.; machi, M.; Iida, T. Int J Syst Evol Microbiol Klenk, H.-P.; Göker, M. Front Microbiol 2018, 2016, 66, 580–586. 9, 304.  

[32] Albuquerque, L.; Rainey, F. A.; Nobre, M. F.; [35] Krieg, N. R.; Staley, J. T.; Brown, D. R.; Hed- da Costa, M. S. Int J Syst Evol Microbiol 2011, lund, B. P.; Paster, B. J.; Ward, N. L.; Lud- 61, 2450–2455. wig, W.; Whitman, W. B. Bergey’s Manual

 of Systematic Bacteriology, 2nd ed.; Springer:

55 New York, 2011; Vol. 4; p 358. Bacteriol 1998, 48, 165–177.

 

[36] Kämpfer, P. Bergey’s Manual of Systematic Bac- [40] Xia, J.; Ling, S.-K.; Wang, X.-Q.; Chen, G.- teriology, 2nd ed.; Springer: New York, 2011; J.; Du, Z.-J. Int J Syst Evol Microbiol 2016, 66, Vol. 4; p 330. 2225–2233.

 

[37] Kämpfer, P. Bergey’s Manual of Systematic Bac- [41] Amoozegar, M. A.; Khansha, J.; Mehr- teriology, 2nd ed.; Springer: New York, 2011; shad, M.; Shahzadeh Fazeli, S. A.; Vol. 4; p 330. Ramezani, M.; de la Haba, R. R.; Sánchez-  Porro, C.; Ventosa, A. Int J Syst Evol Microbiol [38] Ike, F.; Sakamoto, M.; Ohkuma, M.; Kajita, A.; 2017, 67, 113–120. Matsushita, S.; Kokubo, T. Int J Syst Evol Mi-  crobiol 2016, 66, 150–157.



[39] Steyn, P. L.; Segers, P.; Vancanneyt, M.; San- dra, P.; Kersters, K.; Joubert, J. J. Int J Syst

56 5 C

Phylum: Caldiserica1 Chlamydia Class: Caldisericia1 Chlamydophila Order: Caldisericales1 Family: Parachlamydiaceae8 Family: Caldisericaceae1 Neochlamydia Parachlamydia Phylum: Calditrichaeota2 Family: Simkaniaceae8 Class: None Simkania Order: None Family: Waddliaceae9 Family: None Waddlia Calorithrix Phylum: Chlorobi10 Phylum: Chlamydiae3 Class: Chlorobia11 Class: Chlamydiia4 Order: Chlorobiales5,12 Order: Chlamydiales5,6 Family: Chlorobiaceae5 Family: Chlamydiaceae5,7 Ancalochloris

57 Chlorobaculum Family: Anaerolineaceae15 Chlorobium Anaerolinea Chloroherpeton Bellilinea Pelodictyon Leptolinea Prosthecochloris Levilinea Class: Ignavibacteria10 Longilinea Order: Ignavibacteriales10 Ornatilinea Family: Ignavibacteriaceae10 Pelolinea Ignavibacterium Order: None Family: Melioribacteraceae13 Family: None Melioribacter Flexilinea Phylum: Chloroflexi14 Class: Ardenticatenia17 Class: Anaerolineae15 Order: Ardenticatenales17 Order: Aggregatilineales16 Family: Ardenticatenaceae17 Family: Aggregatilineaceae16 Ardenticatena Aggregatilinea Class: Caldilineae15 Order: Anaerolineales15 Order: Caldilineales15

58 Family: Caldilineaceae15 Family: Kallotenuaceae20 Caldilinea Kallotenue Litorilinea Class: Dehalococcoidia21 Class: Chloroflexia18 Order: Dehalococcoidales21 Order: Chloroflexales18 Family: Dehalococcoidaceae21 Family: Chloroflexaceae18 Dehalococcoides Chloroflexus Dehalogenimonas Family: Oscillochloridaceae19 Class: Ktedonobacteria22 Chloronema Order: Ktedonobacterales22 Oscillochloris Family: Dictyobacteraceae23 Family: Roseiflexaceae18 Dictyobacter Heliothrix Tengunoibacter Roseiflexus Family: Ktedonobacteraceae22 Order: Herpetosiphonales18 Ktedonobacter Family: Herpetosiphonaceae18 Family: Thermosporotrichaceae24 Herpetosiphon Thermosporothrix Order: Kallotenuales20 Order: Thermogemmatisporales25

59 Family: Thermogemmatisporaceae25 Order: Chrysiogenales33 Thermogemmatispora Family: Chrysiogenaceae34 Class: Thermoflexia26 Chrysiogenes Order: Thermoflexales26 Desulfurispira Family: Thermoflexaceae26 Desulfurispirillum Thermoflexus Phylum: Coprothermobacterota35 Class: Thermomicrobia27 Class: Coprothermobacteria35 Order: Sphaerobacterales28 Order: Coprothermobacterales35 Family: Sphaerobacteraceae28 Family: Coprothermobacteraceae35 Nitrolancea Coprothermobacter Sphaerobacter Phylum: Cyanobacteria11 Order: Thermomicrobiales29 Class: None Family: Thermomicrobiaceae30 Order: None Thermomicrobium Family: None Thermorudis Prochlorococcus Phylum: Chrysiogenetes31 Prochloron Class: Chrysiogenetes32 Prochlorothrix

60 Rubidibacter Vol. 4; p 844.

 References [5] Skerman, V. B. D.; McGowan, V.; Sneath, P. [1] Mori, K.; Yamaguchi, K.; Sakiyama, Y.; H. A. Int J Syst Bacteriol 1980, 30, 225–420. Urabe, T.; Suzuki, K.-i. Int J Syst Evol Micro-  biol 2009, 59, 2894–2898.

 [6] Storz, J.; Page, L. A. Int J Syst Bacteriol 1971, 21, 332–334. [2] Kublanov, I. V. et al. Front Microbiol 2017, 8, 195.   [7] Rake, G. W. Bergey’s Manual of Determinative [3] Garrity, G. M.; Holt, J. G. Bergey’s Manual Bacteriology, 7th ed.; The Williams & Wilkins of Systematic Bacteriology, 2nd ed.; Springer: Co.: Baltimore, 1957; p 957. New York, 2001; Vol. 1; pp 119–166. 

 [8] Everett, K. D. E.; Bush, R. M.; Andersen, A. A. [4] Horn, M. Bergey’s Manual of Systematic Bacte- Int J Syst Bacteriol 1999, 49, 415–440.

riology, 2nd ed.; Springer: New York, 2010; 

61 [9] Rurangirwa, F. R.; Dilbeck, P. M.; Craw- Merkel, A. Y.; Karnachuk, O. V.; Ravin, N. V.; ford, T. B.; McGuire, T. C.; McElwain, T. F. Bonch-Osmolovskaya, E. A.; Kublanov, I. V. Int J Syst Bacteriol 1999, 49, 577–581. Environ Microbiol 2012, 15, 1759–1771.

 

[10] Iino, T.; Mori, K.; Uchino, Y.; Nakagawa, T.; [14] Garrity, G. M.; Holt, J. G. Bergey’s Manual Harayama, S.; Suzuki, K.-i. Int J Syst Evol of Systematic Bacteriology, 2nd ed.; Springer: Microbiol 2010, 60, 1376–1382. New York, 2001; Vol. 1; pp 427–446.

 

[11] Cavalier-Smith, T. Int J Syst Evol Microbiol [15] Yamada, T.; Sekiguchi, Y.; Hanada, S.; 2002, 52, 7–76. Imachi, H.; Ohashi, A.; Harada, H.; Kam-  agata, Y. Int J Syst Evol Microbiol 2006, 56, [12] Gibbons, N. E.; Murray, R. G. E. Int J Syst 1331–1340. Bacteriol 1978, 28, 1–6. 

 [16] Nakahara, N.; Nobu, M. K.; Takaki, Y.; [13] Podosokorskaya, O. A.; Kadnikov, V. V.; Miyazaki, M.; Tasumi, E.; Sakai, S.; Gavrilov, S. N.; Mardanov, A. V.; Ogawara, M.; Yoshida, N.; Tamaki, H.; Ya-

62 manaka, Y.; Katayama, A.; Yamaguchi, T.; Palisoc, M. M.; Williams, A. J.; Takai, K.; Imachi, H. Int J Syst Evol Microbiol Dohnalkova, A. C.; Ming, H.; Yu, T. T.; 2019, 69, 1185–1194. Dodsworth, J. A.; Li, W.-J.; Hedlund, B. P. Int

 J Syst Evol Microbiol 2013, 63, 4675–4682.

 [17] Kawaichi, S.; Ito, N.; Kamikawa, R.; Sug- awara, T.; Yoshida, T.; Sako, Y. Int J Syst Evol [21] Löffler, F. E.; Yan, J.; Ritalahti, K. M.; Microbiol 2013, 63, 2992–3002. Adrian, L.; Edwards, E. A.; Konstantini-  dis, K. T.; Müller, J. A.; Fullerton, H.; Zin- [18] Gupta, R. S.; Chander, P.; George, S. Antonie der, S. H.; Spormann, A. M. Int J Syst Evol van Leeuwenhoek 2013, 103, 99–119. Microbiol 2013, 63, 625–635.

 

[19] Keppen, O. I.; Tourova, T. P.; Kuznetsov, B. B.; [22] Cavaletti, L.; Monciardini, P.; Bamonte, R.; Ivanovsky, R. N.; Gorlenko, V. M. Int J Syst Schumann, P.; Rohde, M.; Sosio, M.; Dona- Evol Microbiol 2000, 50, 1529–1537. dio, S. Appl Environ Microbiol 2006, 72, 4360–  4369.

[20] Cole, J. K.; Gieler, B. A.; Heisler, D. L.; 

63 [23] Wang, C.-m.; Zheng, Y.; Sakai, Y.; Toyoda, A.; 2119–2127.

Minakuchi, Y.; Abe, K.; Yokota, A.; Yabe, S.  Int J Syst Evol Microbiol 2019, 69, 1910–1918. [27] Garrity, G. M.; Holt, J. G. Bergey’s Manual  of Systematic Bacteriology, 2nd ed.; Springer: [24] Yabe, S.; Aiba, Y.; Sakai, Y.; Hazaka, M.; New York, 2001; Vol. 1; p 447. Yokota, A. Int J Syst Evol Microbiol 2010, 60,  1794–1801. [28] Stackebrandt, E.; Rainey, F. A.; Ward-  Rainey, N. L. Int J Syst Evol Microbiol 1997, [25] Yabe, S.; Aiba, Y.; Sakai, Y.; Hazaka, M.; 47, 479–491. Yokota, A. Int J Syst Evol Microbiol 2011, 61,  903–910. [29] Garrity, G. M.; Holt, J. G. Bergey’s Manual  of Systematic Bacteriology, 2nd ed.; Springer: New York, 2001; Vol. 1; p 447. [26] Dodsworth, J. A.; Gevorkian, J.; Despujos, F.;  Cole, J. K.; Murugapiran, S. K.; Ming, H.; Li, W.-J.; Zhang, G.; Dohnalkova, A.; Hed- [30] Garrity, G. M.; Holt, J. G. Bergey’s Manual lund, B. P. Int J Syst Evol Microbiol 2014, 64, of Systematic Bacteriology, 2nd ed.; Springer:

64 New York, 2001; Vol. 1; p 447. New York, 2001; Vol. 1; p 421.

 

[31] Garrity, G. M.; Holt, J. G. Bergey’s Manual [35] Pavan, M. E.; Pavan, E. E.; Glaeser, S. P.; of Systematic Bacteriology, 2nd ed.; Springer: Etchebehere, C.; Kämpfer, P.; Pettinari, M. J.; New York, 2001; Vol. 1; pp 421–425. López, N. I. Int J Syst Evol Microbiol 2018, 68,  1627–1632.

[32] Garrity, G. M.; Holt, J. G. Bergey’s Manual  of Systematic Bacteriology, 2nd ed.; Springer: New York, 2001; Vol. 1; p 421.



[33] Garrity, G. M.; Holt, J. G. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, 2001; Vol. 1; p 421.



[34] Garrity, G. M.; Holt, J. G. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; Springer:

65 6 D

Phylum: Deferribacteres1 Phylum: Deinococcus–Thermus5 Class: Deferribacteres2 Class: Deinococci6 Order: Deferribacterales3 Order: Deinococcales7 Family: Deferribacteraceae4 Family: Deinococcaceae8 Calditerrivibrio Deinobacter Deferribacter Deinobacterium Denitrovibrio Deinococcus Flexistipes Family: Trueperaceae9 Geovibrio Truepera Mucispirillum Order: Thermales10 Petrothermobacter Family: Thermaceae11 Seleniivibrio Calidithermus Family: None Marinithermus Caldithrix Meiothermus

66 Oceanithermus New York, 2001; Vol. 1; p 465.

Rhabdothermus  Thermus [3] Huber, H.; Stetter, K. O. Bergey’s Manual Vulcanithermus of Systematic Bacteriology, 2nd ed.; Springer: Phylum: Dictyoglomi12 New York, 2001; Vol. 1; p 465. Class: Dictyoglomia13  Order: Dictyoglomales14 Family: Dictyoglomaceae15 [4] Huber, H.; Stetter, K. O. Bergey’s Manual Dictyoglomus of Systematic Bacteriology, 2nd ed.; Springer: New York, 2001; Vol. 1; pp 465–466. References 

[1] Garrity, G. M.; Holt, J. G. Bergey’s Manual [5] Weisburg, W. G.; Giovannoni, S. J.; of Systematic Bacteriology, 2nd ed.; Springer: Woese, C. R. Syst Appl Microbiol 1989, New York, 2001; Vol. 1; pp 465–471. 11, 128–134.

 

[2] Huber, H.; Stetter, K. O. Bergey’s Manual [6] Garrity, G. M.; Holt, J. G. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; Springer: of Systematic Bacteriology, 2nd ed.; Springer:

67 New York, 2001; Vol. 1; p 395. New York, 2001; Vol. 1; p 403.

 

[7] Rainey, F. A.; Nobre, M. F.; Schumann, P.; [11] da Costa, M. S.; Rainey, F. A. Bergey’s Manual Stackebrandt, E.; da Costa, M. S. Int J Syst of Systematic Bacteriology, 2nd ed.; Springer: Bacteriol 1997, 47, 510–514. New York, 2001; Vol. 1; pp 403–404.

 

[8] Brooks, B. W.; Murray, R. G. E. Int J Syst Bac- [12] Patel, B. K. C. Bergey’s Manual of Systematic teriol 1981, 31, 353–360. Bacteriology, 2nd ed.; Springer: New York,

 2011; Vol. 4; p 775.

 [9] Albuquerque, L.; Simões, C.; Nobre, M. F.; Pino, N. M.; Battista, J. R.; Silva, M. T.; [13] Patel, B. K. C. Bergey’s Manual of Systematic Rainey, F. A.; da Costa, M. S. FEMS Microbiol Bacteriology, 2nd ed.; Springer: New York, Lett 2005, 247, 161–169. 2011; Vol. 4; p 776.

 

[10] Rainey, F. A.; da Costa, M. S. Bergey’s Manual [14] Patel, B. K. C. Bergey’s Manual of Systematic of Systematic Bacteriology, 2nd ed.; Springer: Bacteriology, 2nd ed.; Springer: New York,

68 2011; Vol. 4; p 776.



[15] Patel, B. K. C. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, 2011; Vol. 4; p 776.



69 7 E

1 Phylum: Elusimicrobia  Class: Elusimicrobia1 [2] Zheng, H.; Dietrich, C.; Radek, R.; Brune, A. Order: Elusimicrobiales1 Environ Microbiol 2016, 18, 191–204. Family: Elusimicrobiaceae1 Elusimicrobium  Class: Endomicrobia2 Order: Endomicrobiales2 Family: Endomicrobiaceae2 Endomicrobium

References

[1] Geissinger, O.; Herlemann, D. P. R.; Mörschel, E.; Maier, U. G.; Brune, A. Appl Environ Microbiol 2009, 75, 2831–2840.

70 8 F

Phylum: Fibrobacteres1 Class: Tissierellia9 Class: Chitinispirillia2 Order: Tissierellales9 Order: Chitinispirillales2 Family: Tissierellaceae9 Family: Chitinispirillaceae2 Tissierella Chitinispirillum Family: Gottschalkiaceae10 Class: Chitinivibrionia3 Andreesenia Order: Chitinivibrionales3 Gottschalkia Family: Chitinivibrionaceae3 Class: Bacilli11 Chitinivibrio Order: Bacillales7 Class: Fibrobacteria4 Family: Alicyclobacillaceae12 Order: Fibrobacterales5 Alicyclobacillus Family: Fibrobacteraceae6 Effusibacillus Fibrobacter Kyrpidia Phylum: Firmicutes7,8 Sulfobacillus

71 Tumebacillus Calditerricola Family: Bacillaceae7 Camelliibacillus Aeribacillus Cerasibacillus Aliibacillus Compostibacillus Alkalibacillus Desertibacillus Alkalilactibacillus Domibacillus Allobacillus Falsibacillus Alteribacillus Fermentibacillus Amphibacillus Fictibacillus Anaerobacillus Filobacillus Anoxybacillus Geobacillus Aquibacillus Gracilibacillus Aquisalibacillus Halalkalibacillus Aureibacillus Halobacillus Bacillus Halolactibacillus Caldalkalibacillus Hydrogenibacillus Caldibacillus Lentibacillus

72 Lysinibacillus Pontibacillus Marinococcus Pradoshia Melghiribacillus Pseudogracilibacillus Microaerobacter Psychrobacillus Natribacillus Pueribacillus Natronobacillus Saccharococcus Nosocomiicoccus Salibacillus Oceanobacillus Salibacterium Ornithinibacillus Salimicrobium Parageobacillus Salinibacillus Paraliobacillus Salipaludibacillus Paralkalibacillus Salirhabdus Paucisalibacillus Salisediminibacterium Pelagibacillus Saliterribacillus Pelagirhabdus Salsuginibacillus Piscibacillus Sediminibacillus Polygonibacillus Sinibacillus

73 Streptohalobacillus Indiicoccus Swionibacillus Jeotgalibacillus Tenuibacillus Kurthia Tepidibacillus Marinibacillus Terribacillus Paenisporosarcina Texcoconibacillus Planococcus Thalassobacillus Planomicrobium Thalassorhabdus Savagea Thermolongibacillus Sporosarcina Virgibacillus Ureibacillus Viridibacillus Family: Listeriaceae13 Vulcanibacillus Brochothrix Family: Caryophanaceae7 Listeria Bhargavaea Family: None Caryophanon Alkalicoccus Chryseomicrobium Chungangia Filibacter Desulfuribacillus

74 Exiguobacterium Longirhabdus Gemella Marinicrinis Geomicrobium Oxalophagus Rummeliibacillus Paenibacillus Sinobaca Paludirhabdus Solibacillus Saccharibacillus Terrilactibacillus Thermobacillus Thermicanus Xylanibacillus Family: Paenibacillaceae14 Family: Pasteuriaceae7 Ammoniibacillus Pasteuria Ammoniphilus Family: Sporolactobacillaceae15 Aneurinibacillus Caenibacillus Brevibacillus Pullulanibacillus Chengkuizengella Scopulibacillus Cohnella Sporolactobacillus Fontibacillus Tuberibacillus Gorillibacterium Family: Staphylococcaceae16

75 Abyssicoccus Marininema Aliicoccus Marinithermofilum Auricoccus Mechercharimyces Corticicoccus Melghirimyces Jeotgalicoccus Novibacillus Macrococcus Paenactinomyces Salinicoccus Paludifilum Staphylococcus Planifilum Family: Thermoactinomycetaceae17 Polycladomyces Baia Risungbinella Croceifilum Salinithrix Desmospora Seinonella Geothermomicrobium Shimazuella Hazenella Staphylospora Kroppenstedtia Thermoactinomyces Laceyella Thermoflavimicrobium Lihuaxuella Order: Lactobacillales18

76 Family: Aerococcaceae19 Desemzia Abiotrophia Dolosigranulum Aerococcus Granulicatella Dolosicoccus Isobaculum Eremococcus Jeotgalibaca Facklamia Lacticigenium Globicatella Lactosphaera Ignavigranum Marinilactibacillus Suicoccus Pisciglobus Family: Carnobacteriaceae20 Trichococcus Alkalibacterium Family: Enterococcaceae21 Allofustis Bavariicoccus Alloiococcus Catellicoccus Atopobacter Enterococcus Atopococcus Melissococcus Atopostipes Pilibacter Carnobacterium Tetragenococcus

77 Vagococcus Class: Clostridia25 Family: Lactobacillaceae7,22 Order: Eubacteriales7,26 Lactobacillus Family: Catabacteraceae27 Paralactobacillus Catabacter Pediococcus Family: Caldicoprobacteraceae28 Sharpea Caldicoprobacter Family: Leuconostocaceae23 Family: Christensenellaceae29 Convivina Christensenella Fructobacillus Family: Clostridiaceae7 Leuconostoc Alkaliphilus Oenococcus Anaerobacter Weissella Anaeromicrobium Family: Streptococcaceae7,24 Anaerosalibacter Floricoccus Anaerosolibacter Lactococcus Anaerosporobacter Lactovum Anaerovirgula Streptococcus Anoxynatronum

78 Anoxynatronum Hydrogenispora Asaccharospora Intestinibacter Brassicibacter Irregularibacter Caldisalinibacter Keratinibaculum Caloramator Lactonifactor Caloranaerobacter Lutispora Caminicella Natronincola Clostridiisalibacter Oceanirhabdus Clostridium Oxobacter Crassaminicella Paeniclostridium Fervidicella Paraclostridium Fonticella Proteiniborus Garciella Proteiniclasticum Geosporobacter Romboutsia Haloimpatiens Salimesophilobacter Hathewaya Sarcina Hungatella Serpentinicella

79 Sporosalibacterium Pseudoramibacter Tepidimicrobium Rhabdanaerobium Terrisporobacter Family: Gracilibacteraceae32 Thermobrachium Gracilibacter Thermohalobacter Family: Heliobacteriaceae33 Thermotalea Heliobacillus Tindallia Heliobacterium Youngiibacter Heliophilum Family: Defluviitaleaceae30 Heliorestis Defluviitalea Family: Lachnospiraceae34 Family: Eubacteriaceae31 Abyssivirga Acetobacterium Acetatifactor Alkalibacter Acetitomaculum Alkalibaculum Agathobacter Aminicella Anaerobium Anaerofustis Anaerobutyricum Eubacterium Anaerocolumna

80 Anaerosacchariphilus Falcatimonas Anaerostipes Frisingicoccus Anaerotaenia Fusicatenibacter Anaerotignum Herbinix Blautia Hespellia Bryantella Howardella Butyrivibrio Johnsonella Catenibacillus Kineothrix Catonella Lachnoanaerobaculum Cellulosilyticum Lachnobacterium Coprococcus Lachnospira Cuneatibacter Lachnotalea Dorea Marvinbryantia Eisenbergiella Mediterraneibacter Extibacter Merdimonas Faecalicatena Mobilisporobacter Faecalimonas Mobilitalea

81 Moryella Aminipila Muricomes Anaerobranca Murimonas Anaerovorax Natranaerovirga Carboxydocella Oribacterium Dethiosulfatibacter Parasporobacterium Fenollaria Pseudobutyrivibrio Fusibacter Robinsoniella Guggenheimella Roseburia Kallipyga Schaedlerella Marinisporobacter Sellimonas Mogibacterium Shuttleworthia Sedimentibacter Sporobacterium Soehngenia Stomatobaculum Sporanaerobacter Syntrophococcus Thermaerobacter Family: None Family: Oscillospiraceae7 Acidaminobacter Acetanaerobacterium

82 Acetivibrio Herbivorax Acutalibacter Hungateiclostridium Agathobaculum Hydrogenoanaerobacterium Anaerobacterium Intestinimonas Anaerofilum Lawsonibacter Anaerotruncus Mageeibacillus Butyricicoccus Massiliimalia Caproiciproducens Monoglobus Cellulosibacter Oscillibacter Dysosmobacter Oscillospira Ercella Paludicola Ethanoligenens Papillibacter Faecalibacterium Petroclostridium Fastidiosipila Pseudobacteroides Flavonifractor Pseudoflavonifractor Fournierella Ruminiclostridium Harryflintia Ruminococcus

83 Ruthenibacterium Desulfonispora Saccharofermentans Desulfosporosinus Sporobacter Desulfotomaculum Subdoligranulum Desulfurispora Tepidibaculum Pelotomaculum Thermoclostridium Peptococcus Family: Peptococcaceae7,35 Sporotomaculum Cryptanaerobacter Syntrophobotulus Dehalobacter Thermincola Desulfallas Family: Peptoniphilaceae36 Desulfitibacter Anaerococcus Desulfitispora Anaerosphaera Desulfitobacterium Citroniella Desulfocucumis Ezakiella Desulfofarcimen Finegoldia Desulfofundulus Gallicola Desulfohalotomaculum Helcococcus

84 Murdochiella Proteinivorax Parvimonas Family: Symbiobacteriaceae39 Peptoniphilus Symbiobacterium Family: Peptostreptococcaceae37 Family: Syntrophomonadaceae40 Acetoanaerobium Dethiobacter Clostridioides Pelospora Filifactor Syntrophomonas Maledivibacter Syntrophospora Paramaledivibacter Syntrophothermus Peptoanaerobacter Thermohydrogenium Peptoclostridium Thermosyntropha Peptostreptococcus Family: Vallitaleaceae41 Proteocatella Petrocella Sporacetigenium Vallitalea Tepidibacter Order: Halanaerobiales42 Wukongibacter Family: Halanaerobiaceae43 Family: Proteinivoraceae38 Halanaerobium

85 Halarsenatibacter Order: Natranaerobiales44 Halocella Family: Natranaerobiaceae44 Haloincola Natranaerobaculum Halothermothrix Natranaerobius Family: Halobacteroidaceae42 Natronovirga Acetohalobium Order: None Fuchsiella Family: None Halanaerobacter Syntrophaceticus Halanaerobaculum Order: Thermoanaerobacterales45 Halobacteroides Family: None Halonatronum Caldicellulosiruptor Natroniella Mahella Orenia Family: Thermoanaerobacteraceae46 Selenihalanaerobacter Acetogenium Sporohalobacter Ammonifex Family: None Brockia Anoxybacter Caldanaerobacter

86 Caldanaerobius Family: Thermodesulfobiaceae47 Calderihabitans Thermodesulfobium Caloribacterium Order: Thermosediminibacterales48 Carboxydibrachium Family: Tepidanaerobacteraceae48 Carboxydothermus Biomaibacter Desulfothermobacter Tepidanaerobacter Desulfovirgula Family: Thermosediminibacteraceae48 Gelria Caldanaerovirga Moorella Fervidicola Thermacetogenium Thermosediminibacter Thermanaeromonas Thermovenabulum Thermoanaerobacter Thermovorax Thermoanaerobacterium Class: Erysipelotrichia49 Thermoanaerobium Order: Erysipelotrichales50 Thermobacteroides Family: Erysipelotrichaceae51 Thermodesulfitimonas Allobaculum Thermoterrabacterium Anaerorhabdus

87 Breznakia Longibaculum Bulleidia Longicatena Catenibacterium Solobacterium Catenisphaera Turicibacter Coprobacillus Class: Limnochordia52 Dielma Order: Limnochordales52 Dubosiella Family: Limnochordaceae52 Eggerthia Limnochorda Erysipelothrix Class: Negativicutes53 Faecalibacillus Order: Acidaminococcales54 Faecalibaculum Family: Acidaminococcaceae53 Faecalicoccus Acidaminococcus Faecalitalea Anaerospora Holdemanella Phascolarctobacterium Holdemania Succiniclasticum Ileibacterium Succinispira Kandleria Order: Selenomonadales53

88 Family: Selenomonadaceae54 Lucifera Anaerovibrio Methylomusa Centipeda Pelosinus Megamonas Propionispora Mitsuokella Sporolituus Pectinatus Sporomusa Propionispira Sporotalea Schwartzia Thermosinus Selenomonas Order: Veillonellales54 Zymophilus Family: Veillonellaceae7,55 Family: Sporomusaceae54 Allisonella Acetonema Anaeroglobus Anaeroarcus Dialister Anaeromusa Megasphaera Anaerosinus Negativicoccus Anaerosporomusa Quinella Dendrosporobacter Veillonella

89 Class: None Psychrilyobacter Order: None Family: Leptotrichiaceae60 Family: None Caviibacter Caecibacterium Leptotrichia Class: Thermolithobacteria56 Oceanivirga Order: Thermolithobacterales56 Sebaldella Family: Thermolithobacteraceae56 Sneathia Thermolithobacter Streptobacillus Phylum: Fusobacteria1 Class: Fusobacteriia57 References Order: Fusobacteriales58 [1] Garrity, G. M.; Holt, J. G. Bergey’s Manual Family: Fusobacteriaceae59 of Systematic Bacteriology, 2nd ed.; Springer: Cetobacterium New York, 2001; Vol. 1; pp 119–166. Fusobacterium  Hypnocyclicus Ilyobacter [2] Sorokin, D. Y.; Rakitin, A. L.; Gumerov, V. M.; Propionigenium Beletsky, A. V.; Sinninghe Damsté, J. S.; Mar- danov, A. V.; Ravin, N. V. Front Microbiol

90 2016, 7, 407. York, 2011; Vol. 4; p 739.

 

[6] Spain, A. M.; Forsberg, C. W.; [3] Sorokin, D. Y.; Gumerov, V. M.; Rakitin, A. L.; Krumholz, L. R. Bergey’s Manual of Sys- Beletsky, A. V.; Damsté, J. S. S.; Muyzer, G.; tematic Bacteriology, 2nd ed.; Springer: New Mardanov, A. V.; Ravin, N. V. Environ Micro- York, 2011; Vol. 4; p 739. biol 2014, 16, 1549–1565.   [7] Skerman, V. B. D.; McGowan, V.; Sneath, P. [4] Spain, A. M.; Forsberg, C. W.; H. A. Int J Syst Bacteriol 1980, 30, 225–420.

Krumholz, L. R. Bergey’s Manual of Sys-  tematic Bacteriology, 2nd ed.; Springer: New [8] Gibbons, N. E.; Murray, R. G. E. Int J Syst York, 2011; Vol. 4; p 739. Bacteriol 1978, 28, 1–6.  

[5] Spain, A. M.; Forsberg, C. W.; [9] Alauzet, C.; Marchandin, H.; Courtin, P.; Krumholz, L. R. Bergey’s Manual of Sys- Mory, F.; Lemée, L.; Pons, J.-L.; Chapot- tematic Bacteriology, 2nd ed.; Springer: New Chartier, M.-P.; Lozniewski, A.; Jumas-

91 Bilak, E. Syst Appl Microbiol 2014, 37, 23–34. Bergey’s Manual of Systematic Bacteriology, 2nd

 ed.; Springer: New York, 2009; Vol. 3; p 244.

 [10] Poehlein, A.; Yutin, N.; Daniel, R.; Galperin, M. Y. Int J Syst Evol Micro- [14] De Vos, P.; Ludwig, W.; Schleifer, K. H.; Whit- biol 2017, 67, 2711–2719. man, W. B. Bergey’s Manual of Systematic Bac-  teriology, 2nd ed.; Springer: New York, 2009; [11] Ludwig, W.; Schleifer, K. H.; Whitman, W. B. Vol. 3; p 269. Bergey’s Manual of Systematic Bacteriology, 2nd  ed.; Springer: New York, 2009; Vol. 3; pp 19– [15] Ludwig, W.; Schleifer, K. H.; Whitman, W. B. 20. Bergey’s Manual of Systematic Bacteriology, 2nd  ed.; Springer: New York, 2009; Vol. 3; p 386. [12] da Costa, M. S.; Rainey, F. A. Bergey’s Manual  of Systematic Bacteriology, 2nd ed.; Springer: [16] Schleifer, K. H.; Bell, J. A. Bergey’s Manual New York, 2009; Vol. 3; p 229. of Systematic Bacteriology, 2nd ed.; Springer:  New York, 2009; Vol. 3; p 392.

[13] Ludwig, W.; Schleifer, K. H.; Whitman, W. B. 

92 [17] Matsuo, Y.; Katsuta, A.; Matsuda, S.; [21] Ludwig, W.; Schleifer, K. H.; Whitman, W. B. Shizuri, Y.; Yokota, A.; Kasai, H. Int J Syst Bergey’s Manual of Systematic Bacteriology, 2nd Evol Microbiol 2006, 56, 2837–2842. ed.; Springer: New York, 2009; Vol. 3; p 594.

 

[18] Ludwig, W.; Schleifer, K. H.; Whitman, W. B. [22] Winslow, C.-E. A.; Broadhurst, J.; Bergey’s Manual of Systematic Bacteriology, 2nd Buchanan, R. E.; Krumwiede, C.; ed.; Springer: New York, 2009; Vol. 3; p 464. Rogers, L. A.; Smith, G. H. J Bacteriol

 1917, 2, 505–566.

 [19] Ludwig, W.; Schleifer, K. H.; Whitman, W. B. Bergey’s Manual of Systematic Bacteriology, 2nd [23] Schleifer, K. H. Bergey’s Manual of Systematic ed.; Springer: New York, 2009; Vol. 3; p 533. Bacteriology, 2nd ed.; Springer: New York,

 2009; Vol. 3; p 624.

 [20] Ludwig, W.; Schleifer, K. H.; Whitman, W. B. Bergey’s Manual of Systematic Bacteriology, 2nd [24] Deibel, R. H.; Seeley, H. W., Jr. Bergey’s Man- ed.; Springer: New York, 2009; Vol. 3; p 549. ual of Determinative Bacteriology, 8th ed.; The

 Williams & Wilkins Co.: Baltimore, 1974; pp

93 490–515. [29] Morotomi, M.; Nagai, F.; Watanabe, Y. Int J

 Syst Evol Microbiol 2012, 62, 144–149.

 [25] Rainey, F. A. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, [30] Jabari, L.; Gannoun, H.; Cayol, J.-L.; 2001; Vol. 3; p 736. Hamdi, M.; Fauque, G.; Ollivier, B.;  Fardeau, M.-L. Int J Syst Evol Microbiol 2012, [26] Buchanan, R. E. J Bacteriol 1917, 2, 347–350. 62, 550–555.  

[27] Lau, S. K. P.; McNabb, A.; Woo, G. K. S.; [31] Ludwig, W.; Schleifer, K. H.; Whitman, W. B. Hoang, L.; Fung, A. M. Y.; Chung, L. M. W.; Bergey’s Manual of Systematic Bacteriology, 2nd Woo, P. C. Y.; Yuen, K.-Y. J Clin Microbiol 2007, ed.; Springer: New York, 2009; Vol. 3; p 865. 45, 395–401. 

 [32] Lee, Y. J.; Oneynwoke, R. U.; Wiegel, J. [28] Yokoyama, H.; Wagner, I. D.; Wiegel, J. Int J Bergey’s Manual of Systematic Bacteriology, 2nd Syst Evol Microbiol 2010, 60, 67–71. ed.; Springer: New York, 2009; Vol. 3; p 910.

 

94 [33] Madigan, M. T.; Euzéby, J. P.; Asao, M. Int J Vol. 3; p 1008.

Syst Evol Microbiol 2010, 60, 1709–1710. 

 [38] Kevbrin, V.; Boltyanskaya, Y.; Zhilina, T.; Kol- [34] Rainey, F. A. Bergey’s Manual of Systematic ganova, T.; Lavrentjeva, E.; Kuznetsov, B. Ex- Bacteriology, 2nd ed.; Springer: New York, tremophiles 2013, 17, 747–756.

2009; Vol. 3; p 921. 

 [39] Shiratori-Takano, H.; Akita, K.; Yamada, K.; [35] Rogosa, M. Int J Syst Bacteriol 1971, 21, 234– Itoh, T.; Sugihara, T.; Beppu, T.; Ueda, K. Int 237. J Syst Evol Microbiol 2014, 64, 3375–3383.

 

[36] Johnson, C. N.; Whitehead, T. R.; [40] Zhao, H.; Yang, D.; Woese, C. R.; Bryant, M. P. Cotta, M. A.; Rhoades, R. E.; Lawson, P. A. Int J Syst Bacteriol 1993, 43, 278–286.

Int J Syst Evol Microbiol 2014, 64, 3538–3545. 

 [41] Quéméneur, M.; Erauso, G.; Frouin, E.; [37] Ezaki, T. Bergey’s Manual of Systematic Bacte- Zeghal, E.; Vandecasteele, C.; Ollivier, B.; riology, 2nd ed.; Springer: New York, 2009; Tamburini, C.; Garel, M.; Ménez, B.;

95 Postec, A. Front Microbiol 2019, 10, 1497. Vol. 3; p 1224.

 

[42] Rainey, F. A.; Zhilina, T. N.; Bouly- [46] Wiegel, J. Bergey’s Manual of Systematic Bac- gina, E. S.; Stackebrandt, E.; Tourova, T. P.; teriology, 2nd ed.; Springer: New York, 2009; Zavarzin, G. A. Anaerobe 1995, 1, 185–199. Vol. 3; p 1225.   [47] Mori, K.; Kim, H.; Kakegawa, T.; Hanada, S. [43] Oren, A.; Paster, B. J.; Woese, C. R. Syst Appl Extremophiles 2003, 7, 283–290. Microbiol 1984, 5, 71–80.   [48] Zhang, X.; Tu, B.; Lv, X.-m.; Lawson, P. A.; [44] Mesbah, N. M.; Hedrick, D. B.; Pea- Yang, M.; Dai, L.-r.; Zhang, H.; Tang, Y.-Q.; cock, A. D.; Rohde, M.; Wiegel, J. Int J Syst Deng, Y.; Cheng, L. Int J Syst Evol Microbiol Evol Microbiol 2007, 57, 2507–2512. 2019, 69, 3891–3902.  

[45] Wiegel, J. Bergey’s Manual of Systematic Bac- [49] Ludwig, W.; Schleifer, K. H.; Whitman, W. B. teriology, 2nd ed.; Springer: New York, 2009; Bergey’s Manual of Systematic Bacteriology, 2nd

96 ed.; Springer: New York, 2009; Vol. 3; p 1298. P.; Jumas-Bilak, E. Int J Syst Evol Microbiol

 2010, 60, 1271–1279.

 [50] Ludwig, W.; Schleifer, K. H.; Whitman, W. B. Bergey’s Manual of Systematic Bacteriology, 2nd [54] Campbell, C.; Adeolu, M.; Gupta, R. S. Int J ed.; Springer: New York, 2009; Vol. 3; p 1298. Syst Evol Microbiol 2015, 65, 3203–3215.

 

[51] Verbarg, S.; Rheims, H.; Emus, S.; Früh- [55] Rogosa, M. Int J Syst Bacteriol 1971, 21, 231– ling, A.; Kroppenstedt, R. M.; Stacke- 233.

brandt, E.; Schumann, P. Int J Syst Evol Micro-  biol 2004, 54, 221–225. [56] Sokolova, T.; Hanel, J.; Onyenwoke, R. U.;  Reysenbach, A.-L.; Banta, A.; Geyer, R.; [52] Watanabe, M.; Kojima, H.; Fukui, M. Int J González, J. M.; Whitman, W. B.; Wiegel, J. Syst Evol Microbiol 2015, 65, 2378–2384. Extremophiles 2007, 11, 145–157.

 

[53] Marchandin, H.; Teyssier, C.; Campos, J.; [57] Staley, J. T.; Whitman, W. B. Bergey’s Manual Jean-Pierre, H.; Roger, F.; Gay, B.; Carlier, J.- of Systematic Bacteriology, 2nd ed.; Springer:

97 New York, 2011; Vol. 4; p 747.



[58] Staley, J. T.; Whitman, W. B. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, 2011; Vol. 4; p 747.



[59] Staley, J. T.; Whitman, W. B. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, 2011; Vol. 4; p 748.



[60] Staley, J. T.; Whitman, W. B. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, 2011; Vol. 4; p 766.



98 9 G

Phylum: Gemmatimonadetes1 Int J Syst Evol Microbiol 2003, 53, 1155–1163. 1 Class: Gemmatimonadetes  Order: Gemmatimonadales1 [2] Pascual, J.; García-López, M.; Bills, G. F.; Ge- Family: Gemmatimonadaceae1 nilloud, O. Int J Syst Evol Microbiol 2016, 66, Gemmatimonas 1976–1985. Roseisolibacter Class: Longimicrobia2  Order: Longimicrobiales2 Family: Longimicrobiaceae2 Longimicrobium

References

[1] Zhang, H.; Sekiguchi, Y.; Hanada, S.; Hugen- holtz, P.; Kim, H.; Kamagata, Y.; Nakamura, K.

99 10 K

Phylum: Kiritimatiellaeota1 Class: Kiritimatiellae1 Order: Kiritimatiellales1 Family: Kiritimatiellaceae1 Kiritimatiella

References

[1] Spring, S.; Bunk, B.; Spröer, C.; Schumann, P.; Rohde, M.; Tindall, B. J.; Klenk, H.-P. ISME J 2016, 10, 2801–2816.



100 11 L

Phylum: Lentisphaerae1 References 2 Class: Lentisphaeria [1] Cho, J.-C.; Vergin, K. L.; Morris, R. M.; Giovan- 1 Order: Lentisphaerales noni, S. J. Environ Microbiol 2004, 6, 611–621. Family: Lentisphaeraceae3  Lentisphaera Order: Victivallales1 [2] Cho, J. C.; Derrien, M.; Hedlund, B. P. Bergey’s Family: Victivallaceae4 Manual of Systematic Bacteriology, 2nd ed.; Victivallis Springer: New York, 2011; Vol. 4; p 787. Class: Oligosphaeria5  5 Order: Oligosphaerales [3] Cho, J. C.; Hedlund, B. P. Bergey’s Manual 5 Family: Oligosphaeraceae of Systematic Bacteriology, 2nd ed.; Springer: Oligosphaera New York, 2011; Vol. 4; p 788.



[4] Derrien, M.; Cho, J. C.; Hedlund, B. P. Bergey’s

101 Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, 2011; Vol. 4; p 791.



[5] Qiu, Y.-L.; Muramatsu, M.; Hanada, S.; Kam- agata, Y.; Guo, R.-B.; Sekiguchi, Y. Int J Syst Evol Microbiol 2013, 63, 533–539.



102 12 N

Phylum: Nitrospirae1 Class: Nitrospira1 Order: Nitrospirales1 Family: Nitrospiraceae1 Leptospirillum Nitrospira Thermodesulfovibrio

References

[1] Garrity, G. M.; Holt, J. G. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, 2001; Vol. 1; pp 451–464.



103 13 P

Phylum: Planctomycetes1 Tundrisphaera Class: Planctomycetia2 Family: Planctomycetaceae3 Order: Planctomycetales3 Blastopirellula Family: Gemmataceae4 Gimesia Fimbriiglobus Mariniblastus Gemmata Pirellula Telmatocola Planctomicrobium Zavarzinella Planctomyces Family: Isosphaeraceae5 Planctopirus Aquisphaera Pseudopedobacter Isosphaera Rhodopirellula Paludisphaera Roseimaritima Singulisphaera Rubinisphaera Tautonia Rubripirellula

104 Schlesneria Order: Mariprofundales10 Thermogutta Family: Mariprofundaceae11 Thermostilla Mariprofundus Tuwongella Class: Acidithiobacillia12 Class: Phycisphaerae6 Order: Acidithiobacillales13 Order: Phycisphaerales6 Family: Acidithiobacillaceae14 Family: Phycisphaeraceae6 Acidithiobacillus Algisphaera Family: Thermithiobacillaceae15 Phycisphaera Thermithiobacillus Order: Sedimentisphaerales7 Class: Alphaproteobacteria16 Family: Sedimentisphaeraceae7 Order: Caulobacterales17,18 Sedimentisphaera Family: Caulobacteraceae18 Order: Tepidisphaerales8 Aquidulcibacter Family: Tepidisphaeraceae8 Asticcacaulis Tepidisphaera Brevundimonas Phylum: Proteobacteria9 Caulobacter Class: Zetaproteobacteria10 Phenylobacterium

105 Family: Hyphomonadaceae19 Thalassorhabdomicrobium Algimonas Woodsholea Asprobacter Order: Emcibacterales20 Fretibacter Family: Emcibacteraceae20 Glycocaulis Emcibacter Hellea Order: Iodidimonadales20 Henriciella Family: Iodidimonadaceae20 Hirschia Iodidimonas Hyphobacterium Order: Kiloniellales21 Hyphomonas Family: Kiloniellaceae21 Litorimonas Kiloniella Maribaculum Order: Kordiimonadales22 Maricaulis Family: Kordiimonadaceae23 Marinicauda Eilatimonas Oceanicaulis Kordiimonas Ponticaulis Family: Temperatibacteraceae24 Robiginitomaculum Temperatibacter

106 Order: Magnetococcales25 Marinicaulis Family: Magnetococcaceae25 Parvularcula Magnetococcus Order: Rhizobiales28 Order: Micropepsales26 Family: Aestuariivirgaceae29 Family: Micropepsaceae26 Aestuariivirga Micropepsis Family: Alsobacteraceae30 Rhizomicrobium Alsobacter Order: Minwuiales27 Family: Ancalomicrobiaceae31 Family: Minwuiaceae27 Ancalomicrobium Minwuia Pinisolibacter Order: None Siculibacillus Family: None Family: Bartonellaceae17 Amphiplicatus Bartonella Aquisalinus Grahamella Enterovirga Rochalimaea Hartmannibacter Family: Beijerinckiaceae32 Hyphococcus Beijerinckia

107 Methylocapsa Tardiphaga Methylocella Variibacter Methyloferula Family: Brucellaceae17,34 Methylorosula Brucella Methylovirgula Daeguia Pseudochelatococcus Falsochrobactrum Qingshengfania Mycoplana Family: Bradyrhizobiaceae33 Ochrobactrum Afipia Paenochrobactrum Agromonas Pseudochrobactrum Blastobacter Family: Chelatococcaceae35 Bosea Camelimonas Bradyrhizobium Chelatococcus Nitrobacter Family: Cohaesibacteraceae36 Oligotropha Breoghania Rhodoblastus Cohaesibacter Rhodopseudomonas Family: Hyphomicrobiaceae17

108 Angulomicrobium Pelagibacterium Aquabacter Prosthecomicrobium Arsenicitalea Rhodomicrobium Blastochloris Rhodoplanes Cucumibacter Seliberia Devosia Vasilyevaea Dichotomicrobium Youhaiella Filomicrobium Zhangella Gemmiger Family: Lichenihabitantaceae37 Hyphomicrobium Lichenihabitans Limoniibacter Family: Mabikibacteraceae38 Maritalea Mabikibacter Methyloligella Family: Methylobacteriaceae39 Methylorhabdus Balneimonas Methyloterrigena Meganema Paradevosia Methylobacterium Pedomicrobium Methylorubrum

109 Microvirga Aureimonas Neomegalonema Bauldia Protomonas Consotaella Psychroglaciecola Flaviflagellibacter Family: Methylocystaceae40 Fulvimarina Albibacter Jiella Chenggangzhangella Mangrovicella Chthonobacter Martelella Hansschlegelia Methylobrevis Methylocystis Methyloceanibacter Methylopila Mongoliimonas Methylosinus Oharaeibacter Pleomorphomonas Phreatobacter Terasakiella Pseudorhodoplanes Family: None Pyruvatibacter Amorphus Tepidicaulis Aurantimonas Family: Notoacmeibacteraceae41

110 Notoacmeibacter Pseudohoeflea Family: Phyllobacteriaceae42 Roseitalea Aminobacter Salaquimonas Aquamicrobium Thalassocola Chelativorans Thermovum Chelatobacter Tianweitania Corticibacterium Zhengella Defluvibacter Family: Rhizobiaceae17,43 Hoeflea Agrobacterium Lentilitoribacter Allorhizobium Mesorhizobium Carbophilus Nitratireductor Ciceribacter Oceaniradius Crabtreella Oricola Ensifer Phyllobacterium Gellertiella Pseudahrensia Kaistia Pseudaminobacter Liberibacter

111 Neorhizobium Family: Roseiarcaceae45 Pararhizobium Roseiarcus Pseudorhizobium Family: Salinarimonadaceae46 Rhizobium Salinarimonas Shinella Saliniramus Sinorhizobium Family: Xanthobacteraceae19 Family: Rhodobiaceae44 Ancylobacter Afifella Azorhizobium Anderseniella Labrys Butyratibacter Microcyclus Lutibaculum Pseudolabrys Microbaculum Pseudoxanthobacter Parvibaculum Starkeya Rhodobium Xanthobacter Rhodoligotrophos Order: Rhodobacterales47 Roseospirillum Family: Rhodobacteraceae48 Tepidamorphus Acidimangrovimonas

112 Actibacterium Amylibacter Acuticoccus Antarctobacter Aestuariibius Aquicoccus Aestuariicoccus Aquimixticola Aestuariihabitans Arenibacillus Aestuariivita Ascidiaceihabitans Aestuarium Boseongicola Agaricicola Brevirhabdus Ahrensia Carideicomes Albidovulum Catellibacterium Albimonas Celeribacter Albirhodobacter Cereibacter Aliiroseovarius Chachezhania Aliishimia Citreicella Alkalilacustris Citreimonas Alkalimicrobium Cognatishimia Amaricoccus Cognatiyoonia

113 Confluentimicrobium Histidinibacterium Cribrihabitans Huaishuia Defluviimonas Hwanghaeicola Dinoroseobacter Jannaschia Donghicola Jhaorihella Epibacterium Kandeliimicrobium Falsirhodobacter Ketogulonicigenium Flavimaricola Labrenzia Fluviibacterium Lacimonas Frigidibacter Leisingera Gaetbulicola Lentibacter Gemmobacter Limibaculum Haematobacter Limimaricola Halocynthiibacter Litoreibacter Halodurantibacterium Litorimicrobium Halovulum Litorisediminicola Hasllibacter Litorisediminivivens

114 Litorivita Neptunicoccus Loktanella Nereida Lutimaribacter Nesiotobacter Maliponia Nioella Mameliella Oceanibium Mangrovicoccus Oceanibulbus Maribius Oceanicella Marimonas Oceanicola Marinibacterium Oceaniglobus Marinovum Octadecabacter Maritimibacter Pacificibacter Marivita Paenimaribius Marivivens Paenirhodobacter Meridianimarinicoccus Palleronia Methylarcula Pannonibacter Monaibacterium Paracoccus Nautella Paradonghicola

115 Paraphaeobacter Profundibacterium Pararhodobacter Pseudaestuariivita Pelagibaca Pseudodonghicola Pelagicola Pseudogemmobacter Pelagimonas Pseudohalocynthiibacter Pelagivirga Pseudomaribius Phaeobacter Pseudooceanicola Phycocomes Pseudooctadecabacter Planktomarina Pseudopelagicola Planktotalea Pseudophaeobacter Pleomorphobacterium Pseudopuniceibacterium Pontibaca Pseudorhodobacter Ponticoccus Pseudoroseicyclus Pontivivens Pseudoroseovarius Poseidonocella Pseudoruegeria Primorskyibacter Pseudoseohaeicola Profundibacter Pseudovibrio

116 Psychromarinibacter Roseovarius Puniceibacterium Rubellimicrobium Rhodobaca Rubribacterium Rhodobacter Rubricella Rhodobaculum Rubrimonas Rhodophyticola Ruegeria Rhodosalinus Sagittula Rhodovulum Salinihabitans Roseibaca Salinovum Roseibacterium Salipiger Roseibium Sedimentitalea Roseicitreum Sediminimonas Roseicyclus Seohaeicola Roseinatronobacter Shimia Roseisalinus Silicibacter Roseivivax Silicimonas Roseobacter Sinirhodobacter

117 Staleya Xinfangfangia Stappia Xuhuaishuia Sulfitobacter Yangia Tabrizicola Yoonia Tateyamaria Youngimonas Thalassobacter Order: Rhodospirillales17,49 Thalassobius Family: Acetobacteraceae50 Thalassococcus Acetobacter Thiobacimonas Acidibrevibacterium Thioclava Acidicaldus Thiosphaera Acidiphilium Tranquillimonas Acidisoma Tritonibacter Acidisphaera Tropicibacter Acidocella Tropicimonas Acidomonas Vadicella Ameyamaea Wenxinia Asaia

118 Belnapia Neokomagataea Bombella Nguyenibacter Caldovatus Paracraurococcus Commensalibacter Rhodopila Craurococcus Rhodovarius Crenalkalicoccus Rhodovastum Dankookia Roseicella Elioraea Roseococcus Endobacter Roseomonas Gluconacetobacter Rubritepida Gluconobacter Saccharibacter Granulibacter Siccirubricoccus Humitalea Stella Komagataeibacter Swaminathania Kozakia Swingsia Muricoccus Tanticharoenia Neoasaia Teichococcus

119 Zavarzinia Ferruginivarius Family: Geminicoccaceae51 Fodinicurvata Arboricoccus Haematospirillum Geminicoccus Hwanghaeella Family: Rhodospirillaceae17,49 Indioceanicola Aestuariispira Inquilinus Algihabitans Insolitispirillum Aliidongia Lacibacterium Azospirillum Limibacillus Caenispirillum Limimonas Conglomeromonas Magnetospira Constrictibacter Magnetospirillum Defluviicoccus Magnetovibrio Desertibacter Marinibaculum Dongia Marispirillum Elstera Marivibrio Ferrovibrio Nisaea

120 Nitrospirillum Taonella Niveispirillum Telmatospirillum Novispirillum Thalassobaculum Oceanibaculum Thalassospira Oleisolibacter Tistlia Pararhodospirillum Tistrella Pelagibius Varunaivibrio Phaeospirillum Order: Rhodothalassiales52 Phaeovibrio Family: Rhodothalassiaceae52 Reyranella Rhodothalassium Rhodocista Order: Rickettsiales17 Rhodospira Family: Anaplasmataceae17,53 Rhodospirillum Aegyptianella Rhodovibrio Anaplasma Roseospira Cowdria Skermanella Ehrlichia Tagaea Neorickettsia

121 Wolbachia Family: Erythrobacteraceae19 Family: Holosporaceae54 Altererythrobacter Caedibacter Croceibacterium Holospora Croceicoccus Lyticum Erythrobacter Pseudocaedibacter Erythromicrobium Symbiotes Porphyrobacter Tectibacter Qipengyuania Family: Rickettsiaceae17,55 Family: Sphingomonadaceae58 Occidentia Aquisediminimonas Orientia Blastomonas Rickettsia Chakrabartia Order: Sneathiellales56 Erythromonas Family: Sneathiellaceae56 Hephaestia Oceanibacterium Novosphingobium Sneathiella Pacificimonas Order: Sphingomonadales57 Parablastomonas

122 Parasphingopyxis Class: Betaproteobacteria59 Polymorphobacter Order: Burkholderiales60 Rhizomonas Family: Alcaligenaceae61 Rhizorhabdus Achromobacter Rhizorhapis Advenella Sandaracinobacter Alcaligenes Sandarakinorhabdus Algicoccus Sphingoaurantiacus Ampullimonas Sphingobium Azohydromonas Sphingomicrobium Basilea Sphingomonas Bordetella Sphingopyxis Brackiella Sphingorhabdus Caenimicrobium Sphingosinicella Candidimonas Sphingosinithalassobacter Castellaniella Stakelama Corticimicrobacter Zymomonas Derxia

123 Eoetvoesia Family: Burkholderiaceae62 Kerstersia Burkholderia Oligella Caballeronia Orrella Chitinimonas Paenalcaligenes Cupriavidus Paracandidimonas Ephemeroptericola Paralcaligenes Formosimonas Parapusillimonas Hydromonas Parvibium Lautropia Pelistega Limnobacter Pigmentiphaga Mycetohabitans Pusillimonas Mycoavidus Saccharedens Pandoraea Taylorella Paraburkholderia Tetrathiobacter Pararobbsia Verticia Paucimonas Verticiella Polynucleobacter

124 Quisquiliibacterium Corticibacter Ralstonia Curvibacter Robbsia Delftia Thermothrix Diaphorobacter Trinickia Extensimonas Wautersia Giesbergeria Family: Comamonadaceae63 Hydrogenophaga Acidovorax Hylemonella Albidiferax Ideonella Alicycliphilus Kinneretia Aquabacterium Lampropedia Brachymonas Leptothrix Caenibacterium Limnohabitans Caenimonas Macromonas Caldimonas Malikia Calidifontimicrobium Melaminivora Comamonas Mitsuaria

125 Oryzisolibacter Thiomonas Ottowia Tibeticola Paucibacter Variovorax Pelomonas Verminephrobacter Polaromonas Xenophilus Pseudacidovorax Xylophilus Pseudorhodoferax Zhizhongheella Pulveribacter Family: None Ramlibacter Aquincola Rhodoferax Inhella Roseateles Methylibium Rubrivivax Piscinibacter Schlegelella Rhizobacter Simplicispira Rivibacter Sphaerotilus Thiobacter Tepidicella Family: Oxalobacteraceae64 Tepidimonas Collimonas

126 Duganella Parasutterella Glaciimonas Sutterella Herbaspirillum Turicimonas Herminiimonas Order: Neisseriales66 Janthinobacterium Family: Chromobacteriaceae67 Massilia Andreprevotia Naxibacter Aquaspirillum Noviherbaspirillum Aquitalea Oxalicibacterium Chitinibacter Oxalobacter Chitinilyticum Paraherbaspirillum Chitiniphilus Pseudoduganella Chromobacterium Solimicrobium Deefgea Telluria Formivibrio Undibacterium Gulbenkiania Family: Sutterellaceae65 Iodobacter Mesosutterella Jeongeupia

127 Laribacter Prolinoborus Leeia Rivicola Microvirgula Simonsiella Paludibacterium Snodgrassella Pseudogulbenkiania Stenoxybacter Silvimonas Uruburuella Vogesella Vitreoscilla Family: Neisseriaceae17 Order: Nitrosomonadales68 Alysiella Family: Gallionellaceae17,18 Amantichitinum Ferrigenium Bergeriella Ferriphaselus Conchiformibius Gallionella Crenobacter Sulfuricella Eikenella Sulfuriferula Kingella Sulfurirhabdus Morococcus Family: Methylophilaceae69 Neisseria Methylobacillus

128 Methylophilus Sulfurisoma Methylotenera Sulfuritalea Methylovorus Family: Thiobacillaceae71 Novimethylophilus Annwoodia Pseudomethylobacillus Thiobacillus Family: Nitrosomonadaceae70 Order: None Nitrosolobus Family: None Nitrosomonas Chitinivorax Nitrosospira Sulfuritortus Nitrosospira Order: Rhodocyclales72 Family: Spirillaceae17 Family: Azonexaceae71 Spirillum Azonexus Family: Sterolibacteriaceae71 Dechloromonas Denitratisoma Ferribacterium Georgfuchsia Quatrionicoccus Methyloversatilis Family: Rhodocyclaceae73 Sterolibacterium Azospira

129 Azovibrio Bradymonas Dechlorosoma Lujinxingia Niveibacterium Order: Desulfarculales76 Oryzomicrobium Family: Desulfarculaceae77 Propionibacter Desulfarculus Propionivibrio Desulfocarbo Rhodocyclus Dethiosulfatarculus Rugosibacter Order: Desulfobacterales78 Family: Zoogloeaceae71 Family: Desulfobacteraceae79 Aromatoleum Desulfatibacillum Azoarcus Desulfatiferula Thauera Desulfatiglans Uliginosibacterium Desulfatirhabdium Zoogloea Desulfatitalea Class: Deltaproteobacteria74 Desulfobacter Order: Bradymonadales75 Desulfobacterium Family: Bradymonadaceae75 Desulfobacula

130 Desulfobotulus Desulfocapsa Desulfocella Desulfofustis Desulfococcus Desulfopila Desulfoconvexum Desulfoprunum Desulfofaba Desulforhopalus Desulfofrigus Desulfotalea Desulfoluna Desulfurivibrio Desulfomusa Family: Nitrospinaceae81 Desulfonatronobacter Nitrospina Desulfonema Order: Desulfovibrionales82 Desulforegula Family: Desulfohalobiaceae83 Desulfosalsimonas Desulfohalobium Desulfosarcina Desulfonatronospira Desulfospira Desulfonatronovibrio Desulfotignum Desulfonauticus Family: Desulfobulbaceae80 Desulfothermus Desulfobulbus Desulfovermiculus

131 Family: Desulfomicrobiaceae84 Family: Desulfurellaceae88 Desulfomicrobium Desulfurella Desulfoplanes Hippea Family: Desulfonatronaceae85 Order: Desulfuromonadales89 Desulfonatronum Family: Desulfuromonadaceae90 Family: Desulfovibrionaceae86 Desulfuromonas Bilophila Desulfuromusa Desulfobaculum Malonomonas Desulfocurvibacter Pelobacter Desulfocurvus Family: Geobacteraceae91 Desulfohalovibrio Geoalkalibacter Desulfomonas Geobacter Desulfovibrio Geopsychrobacter Halodesulfovibrio Geothermobacter Lawsonia Trichlorobacter Pseudodesulfovibrio Order: Myxococcales17 Order: Desulfurellales87 Family: Anaeromyxobacteraceae92

132 Anaeromyxobacter Myxococcus Family: Archangiaceae17 Pyxidicoccus Angiococcus Simulacricoccus Archangium Family: Nannocystaceae94 Cystobacter Enhygromyxa Hyalangium Nannocystis Melittangium Plesiocystis Stigmatella Pseudenhygromyxa Vitiosangium Family: None Family: Kofleriaceae93 Minicystis Haliangium Family: Phaselicystidaceae95 Kofleria Phaselicystis Family: Labilitrichaceae92 Family: Polyangiaceae17 Labilithrix Aetherobacter Family: Myxococcaceae17 Byssovorax Aggregicoccus Chondromyces Corallococcus Jahnella

133 Polyangium Desulfobacca Racemicystis Desulfomonile Sorangium Smithella Family: Sandaracinaceae96 Syntrophus Sandaracinus Family: Syntrophobacteraceae100 Family: Vulgatibacteraceae92 Desulfacinum Vulgatibacter Desulfoglaeba Order: None Desulforhabdus Family: None Desulfosoma Deferrisoma Desulfovirga Dissulfuribacter Syntrophobacter Dissulfurimicrobium Thermodesulforhabdus Dissulfurirhabdus Class: Epsilonproteobacteria101 Family: Syntrophorhabdaceae97 Order: Campylobacterales102 Syntrophorhabdus Family: Campylobacteraceae103 Order: Syntrophobacterales98 Aliarcobacter Family: Syntrophaceae99 Arcobacter

134 Campylobacter Family: Nautiliaceae105 Dehalospirillum Caminibacter Halarcobacter Cetia Malaciobacter Lebetimonas Poseidonibacter Nautilia Pseudarcobacter Nitratifractor Sulfurospirillum Nitratiruptor Family: Helicobacteraceae104 Thioreductor Helicobacter Class: Gammaproteobacteria106 Sulfuricurvum Order: Vibrionales1 Sulfurimonas Family: Vibrionaceae17 Sulfurovum Aliivibrio Thiovulum Allomonas Wolinella Beneckea Family: None Catenococcus Hydrogenimonas Echinimonas Order: Nautiliales105 Enterovibrio

135 Grimontia Oceanimonas Listonella Oceanisphaera Lucibacterium Pseudaeromonas Paraphotobacterium Tolumonas Photobacterium Zobellella Salinivibrio Family: Succinivibrionaceae110 Thaumasiovibrio Anaerobiospirillum Vibrio Ruminobacter Order: Acidiferrobacterales107 Succinatimonas Family: Acidiferrobacteraceae107 Succinimonas Acidiferrobacter Succinivibrio Sulfuricaulis Order: Alteromonadales111 Sulfurifustis Family: Alteromonadaceae112 Order: Aeromonadales108 Aestuariibacter Family: Aeromonadaceae109 Agaribacter Aeromonas Agarivorans Dongshaea Alginatibacterium

136 Aliagarivorans Salinimonas Aliiglaciecola Tamilnaduibacter Alishewanella Family: Celerinatantimonadaceae113 Alkalimarinus Celerinatantimonas Alteromonas Family: Colwelliaceae114 Bowmanella Colwellia Catenovulum Litorilituus Glaciecola Piscinibacterium Hydrocarboniclastica Thalassomonas Lacimicrobium Thalassotalea Mangrovitalea Family: Ferrimonadaceae114 Marinobacter Ferrimonas Marinobacterium Paraferrimonas Neptunicella Family: Idiomarinaceae114 Paraglaciecola Aliidiomarina Planctobacterium Idiomarina Pseudobowmanella Pseudidiomarina

137 Family: Moritellaceae114 Psychrobium Moritella Shewanella Paramoritella Order: Arenicellales115 Family: None Family: Arenicellaceae115 Aestuariicella Arenicella Motilimonas Perspicuibacter Neiella Order: Cardiobacteriales116 Family: Pseudoalteromonadaceae114 Family: Cardiobacteriaceae117 Algicola Cardiobacterium Pseudoalteromonas Dichelobacter Psychrosphaera Suttonella Salsuginimonas Order: Cellvibrionales118 Family: Psychromonadaceae114 Family: Cellvibrionaceae118 Corallincola Agaribacterium Psychromonas Agarilytica Family: Shewanellaceae114 Cellvibrio Parashewanella Eionea

138 Gilvimarinus Marimicrobium Halioxenophilus Parahaliea Maricurvus Pseudohaliea Marinimicrobium Family: Microbulbiferaceae118 Pseudomaricurvus Microbulbifer Pseudoteredinibacter Family: Porticoccaceae118 Saccharophagus Porticoccus Simiduia Family: Spongiibacteraceae118 Teredinibacter Dasania Umboniibacter Marortus Family: Halieaceae118 Melitea Chromatocurvus Sinobacterium Congregibacter Spongiibacter Haliea Zhongshania Halioglobus Order: Chromatiales119 Kineobactrum Family: Chromatiaceae17 Luminiphilus Alishewanella

139 Allochromatium Thiobaca Amoebobacter Thiocapsa Chromatium Thiococcus Halochromatium Thiocystis Imhoffiella Thiodictyon Isochromatium Thioflavicoccus Lamprobacter Thiohalocapsa Lamprocystis Thiolamprovum Marichromatium Thiopedia Nitrosococcus Thiophaeococcus Pfennigia Thiorhodococcus Phaeobacterium Thiorhodovibrio Phaeochromatium Thiospirillum Rhabdochromatium Family: Ectothiorhodospiraceae120 Rheinheimera Acidihalobacter Thermochromatium Alkalilimnicola Thioalkalicoccus Alkalispirillum

140 Aquisalimonas Thiogranum Arhodomonas Thiohalomonas Ectothiorhodosinus Thiohalospira Ectothiorhodospira Thiorhodospira Halofilum Family: Granulosicoccaceae121 Halopeptonella Granulosicoccus Halorhodospira Sulfuriflexus Inmirania Family: Halothiobacillaceae122 Methylohalomonas Halothiobacillus Methylonatrum Thiofaba Natronocella Thiovirga Natronospira Family: Thioalkalibacteraceae123 Nitrococcus Guyparkeria Oceanococcus Thioalkalibacter Spiribacter Family: Thioalkalispiraceae124 Thioalbus Endothiovibrio Thioalkalivibrio Thioalkalispira

141 Thiohalophilus Cedecea Family: Thioprofundaceae125 Citrobacter Sulfurivermis Cronobacter Thioprofundum Enterobacillus Family: Wenzhouxiangellaceae126 Enterobacter Wenzhouxiangella Escherichia Family: Woeseiaceae127 Franconibacter Woeseia Gibbsiella Order: Enterobacterales128 Izhakiella Family: Budviciaceae128 Klebsiella Budvicia Kluyvera Leminorella Kosakonia Pragia Koserella Family: Enterobacteriaceae17 Leclercia Biostraticola Lelliottia Buttiauxella Levinea Calymmatobacterium Limnobaculum

142 Mangrovibacter Erwinia Metakosakonia Kalamiella Phytobacter Mixta Pluralibacter Pantoea Pseudescherichia Phaseolibacter Pseudocitrobacter Tatumella Raoultella Wigglesworthia Rosenbergiella Family: Hafniaceae128 Saccharobacter Edwardsiella Salmonella Hafnia Shigella Obesumbacterium Shimwellia Family: Morganellaceae128 Siccibacter Arsenophonus Trabulsiella Cosenzaea Yokenella Moellerella Family: Erwiniaceae128 Morganella Buchnera Photorhabdus

143 Proteus Serratia Providencia Yersinia Xenorhabdus Order: Immundisolibacterales129 Family: None Family: Immundisolibacteraceae129 Plesiomonas Immundisolibacter Family: Pectobacteriaceae128 Order: Legionellales130 Brenneria Family: Coxiellaceae131 Dickeya Aquicella Lonsdalea Coxiella Pectobacterium Diplorickettsia Sodalis Rickettsiella Family: Yersiniaceae128 Family: Legionellaceae17,132 Chania Fluoribacter Ewingella Legionella Rahnella Sarcobium Rouxiella Tatlockia Samsonia Order: Lysobacterales17,133

144 Family: Lysobacteraceae17,133 Alkanibacter Arenimonas Povalibacter Aspromonas Singularimonas Chujaibacter Steroidobacter Coralloluteibacterium Family: Rhodanobacteraceae134 Luteimonas Ahniella Lysobacter Aquimonas Metallibacterium Chiayiivirga Mizugakiibacter Dokdonella Pseudoxanthomonas Dyella Silanimonas Frateuria Stenotrophomonas Fulvimonas Thermomonas Luteibacter Vulcaniibacterium Oleiagrimonas Xanthomonas Pseudofulvimonas Xylella Rehaibacterium Family: None Rhodanobacter

145 Rudaea Methyloparacoccus Tahibacter Methyloprofundus Order: Methylococcales135 Methylosarcina Family: Crenotrichaceae17 Methylosoma Crenothrix Methylosphaera Family: Methylococcaceae136 Methyloterricola Methylobacter Methylovulum Methylocaldum Family: Methylothermaceae137 Methylococcus Methylomarinovum Methylocucumis Methylothermus Methylogaea Family: None Methyloglobulus Methylotuvimicrobium Methylohalobius Order: Nevskiales134 Methylomagnum Family: Algiphilaceae138 Methylomarinum Algiphilus Methylomicrobium Family: Nevskiaceae17,18 Methylomonas Alkanibacter

146 Fontimonas Family: None Hydrocarboniphaga Acidibacter Nevskia Alkalimonas Panacagrimonas Arsukibacterium Singularimonas Flintibacter Sinimarinibacterium Gallaecimonas Sinobacter Ignatzschineria Solimonas Litorivivens Stagnimonas Marinagarivorans Stenotrophobium Marinibactrum Family: Salinisphaeraceae134 Marinicella Abyssibacter Plasticicumulans Salinisphaera Pseudohongiella Family: Steroidobacteraceae139 Salicola Povalibacter Sedimenticola Steroidobacter Thiohalobacter Order: None Thiohalorhabdus

147 Thiolapillus Endozoicomonas Thiosocius Kistimonas Wohlfahrtiimonas Parendozoicomonas Family: Thorselliaceae140 Family: Hahellaceae145 Coetzeea Allohahella Thorsellia Hahella Order: Oceanospirillales141 Halospina Family: Alcanivoracaceae142 Sansalvadorimonas Alcanivorax Zooshikella Fundibacter Family: Halomonadaceae146 Ketobacter Aidingimonas Pelagibaculum Carnimonas Pleionea Chromohalobacter Family: Balneatrichaceae143 Cobetia Balneatrix Deleya Pokkaliibacter Halomonas Family: Endozoicomonadaceae144 Halotalea

148 Halovibrio Amphritea Kushneria Bacterioplanes Larsenimonas Bacterioplanoides Modicisalibacter Bermanella Pistricoccus Corallomonas Salinicola Litoribacillus Terasakiispira Litoribrevibacter Volcaniella Marinomonas Zymobacter Marinospirillum Family: Kangiellaceae147 Motiliproteus Aliikangiella Neptuniibacter Kangiella Neptunomonas Family: Litoricolaceae148 Nitrincola Litoricola Oceaniserpentilla Family: None Oceanobacter Spongiispira Oceanospirillum Family: Oceanospirillaceae149 Oleibacter

149 Oleispira Basfia Pontibacterium Bibersteinia Pseudospirillum Bisgaardia Reinekea Caviibacterium Thalassolituus Chelonobacter Family: Oleiphilaceae150 Conservatibacter Oleiphilus Cricetibacter Order: Orbales151 Frederiksenia Family: Orbaceae151 Gallibacterium Frischella Haemophilus Gilliamella Histophilus Orbus Lonepinella Order: Pasteurellales152 Mannheimia Family: Pasteurellaceae153 Mesocricetibacter Actinobacillus Muribacter Aggregatibacter Necropsobacter Avibacterium Nicoletella

150 Otariodibacter Cavicella Pasteurella Enhydrobacter Phocoenobacter Faucicola Rodentibacter Fluviicoccus Seminibacterium Moraxella Testudinibacter Paraperlucidibaca Ursidibacter Perlucidibaca Vespertiliibacter Psychrobacter Volucribacter Family: Pseudomonadaceae17,157 Family: Psittacicellaceae154 Azomonas Psittacicella Azomonotrichon Order: Pseudomonadales17,155 Azorhizophilus Family: Moraxellaceae156 Azotobacter Acinetobacter Chryseomonas Agitococcus Entomomonas Alkanindiges Flavimonas Branhamella Mesophilobacter

151 Oblitimonas Cysteiniphilum Permianibacter Francisella Pseudomonas Pseudofrancisella Rugamonas Family: None Serpens Fangia Thiopseudomonas Family: Piscirickettsiaceae162 Family: Ventosimonadaceae158 Cycloclasticus Ventosimonas Galenea Order: Thiotrichales159 Hydrogenovibrio Family: Beggiatoaceae17 Methylophaga Beggiatoa Piscirickettsia Thioflexithrix Sulfurivirga Family: Fastidiosibacteraceae160 Thioalkalimicrobium Facilibium Thiomicrorhabdus Fastidiosibacter Thiomicrospira Family: Francisellaceae161 Family: Thiofilaceae163 Allofrancisella Thiofilum

152 Family: Thiolineaceae163 Class: Oligoflexia167 Thiolinea Order: Bacteriovoracales168 Family: Thiotrichaceae164 Family: Bacteriovoracaceae169 Achromatium Bacteriolyticum Cocleimonas Bacteriovorax Leucothrix Family: Halobacteriovoraceae170 Thiobacterium Halobacteriovorax Thiomargarita Family: Peredibacteraceae171 Thioploca Peredibacter Thiospira Order: Bdellovibrionales172 Thiothrix Family: Bdellovibrionaceae173 Class: Hydrogenophilalia71 Bdellovibrio Order: Hydrogenophilales165 Micavibrio Family: Hydrogenophilaceae166 Vampirovibrio Hydrogenophilus Order: Oligoflexales167 Petrobacter Family: Oligoflexaceae167 Tepidiphilus Oligoflexus

153 Family: Pseudobacteriovoracaceae174 [3] Schlesner, H.; Stackebrandt, E. Syst Appl Pseudobacteriovorax Microbiol 1986, 8, 174–176. 168 Order: Silvanigrellales  Family: Silvanigrellaceae168 Silvanigrella [4] Kulichevskaya, I. S.; Ivanova, A. A.; Baulina, O. I.; Rijpstra, W. I. C.; Sin- References ninghe Damsté, J. S.; Dedysh, S. N. Int J Syst Evol Microbiol 2017, 67, 218–224.

[1] Garrity, G. M.; Holt, J. G. Bergey’s Manual  of Systematic Bacteriology, 2nd ed.; Springer: New York, 2001; Vol. 1; pp 119–166. [5] Kulichevskaya, I. S.; Ivanova, A. A.;

 Suzina, N. E.; Rijpstra, W. I. C.; Sin- ninghe Damsté, J. S.; Dedysh, S. N. Int J [2] Ludwig, W.; Euzéby, J.; Whitman, W. B. Syst Evol Microbiol 2016, 66, 837–844.

Bergey’s Manual of Systematic Bacteriology,  2nd ed.; Springer: New York, 2010; Vol. 4; pp 21–24. [6] Fukunaga, Y.; Kurahashi, M.; Sakiyama, Y.;

 Ohuchi, M.; Yokota, A.; Harayama, S. J Gen

154 Appl Microbiol 2009, 55, 267–275. [10] Makita, H.; Tanaka, E.; Mitsunobu, S.;

 Miyazaki, M.; Nunoura, T.; Uematsu, K.; Takaki, Y.; Nishi, S.; Shimamura, S.; Takai, K. [7] Spring, S.; Bunk, B.; Spröer, C.; Rohde, M.; Arch Microbiol 2017, 199, 335–346. Klenk, H.-P. Environ Microbiol 2018, 20,  2438–2455.

 [11] Emerson, D.; Rentz, J. A.; Lilburn, T. G.; Davis, R. E.; Aldrich, H.; Chan, C.; [8] Kovaleva, O. L.; Merkel, A. Y.; Moyer, C. L. PLoS One 2007, 2, e667. Novikov, A. A.; Baslerov, R. V.;  Toshchakov, S. V.; Bonch- Osmolovskaya, E. A. Int J Syst Evol [12] Williams, K. P.; Kelly, D. P. Int J Syst Evol Microbiol 2015, 65, 549–555. Microbiol 2013, 63, 2901–2906.

 

[9] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s [13] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, 2005; Vol. 2; p 1. Springer: New York, 2005; Vol. 2; p 60.

 

155 [14] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s 29, 3–4.

Manual of Systematic Bacteriology, 2nd ed.;  Springer: New York, 2005; Vol. 2; p 60. [19] Lee, K.-B.; Liu, C.-T.; Anzai, Y.; Kim, H.;  Aono, T.; Oyaizu, H. Int J Syst Evol Microbiol [15] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s 2005, 55, 1907–1919.

Manual of Systematic Bacteriology, 2nd ed.;  Springer: New York, 2005; Vol. 2; p 62. [20] Iino, T.; Ohkuma, M.; Kamagata, Y.;  Amachi, S. Int J Syst Evol Microbiol 2016, 66, [16] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s 5016–5022.

Manual of Systematic Bacteriology, 2nd ed.;  Springer: New York, 2005; Vol. 2; p 1. [21] Wiese, J.; Thiel, V.; Gärtner, A.; Schmaljo-  hann, R.; Imhoff, J. F. Int J Syst Evol Micro- [17] Skerman, V. B. D.; McGowan, V.; Sneath, P. biol 2009, 59, 350–356.

H. A. Int J Syst Bacteriol 1980, 30, 225–420. 

 [22] Kwon, K. K.; Lee, H.-S.; Yang, S. H.; Kim, S.- [18] Henrici, A. T.; Johnson, D. J Bacteriol 1935, J. Int J Syst Evol Microbiol 2005, 55, 2033–

156 2037. 67, 839–844.

 

[23] Xu, X.-W.; Wu, M.; Oren, A. The ; [27] Sun, C.; Xu, L.; Yu, X.-Y.; Zhao, Z.; Wu, Y.- Springer: Berlin/Heidelberg, 2014; pp 307– H.; Oren, A.; Wang, C.-S.; Xu, X.-W. Int J 312. Syst Evol Microbiol 2018, 68, 3856–3862.

 

[24] Teramoto, M.; Nishijima, M. Int J Syst Evol [28] Kuykendall, L. D.; Bell, J. A.; Lilburn, T. Microbiol 2014, 64, 3075–3080. Bergey’s Manual of Systematic Bacteriology,  2nd ed.; Springer: New York, 2005; Vol. 2; p [25] Bazylinski, D. A.; Williams, T. J.; 324. Lefèvre, C. T.; Berg, R. J.; Zhang, C. L.;  Bowser, S. S.; Dean, A. J.; Beveridge, T. J. [29] Li, X.; Salam, N.; Li, J.-L.; Chen, Y.-M.; Int J Syst Evol Microbiol 2013, 63, 801–808. Yang, Z.-W.; Han, M.-X.; Mou, X.; Xiao, M.;  Li, W.-J. Int J Syst Evol Microbiol 2019, 69, [26] Harbison, A. B.; Price, L. E.; Flythe, M. D.; 299–306.

Bräuer, S. L. Int J Syst Evol Microbiol 2017, 

157 [30] Sun, L.; Liu, H.; Chen, W.; Huang, K.; Bergey’s Manual of Determinative Bacteriology, Lyu, W.; Gao, X. Int J Syst Evol Microbiol 7th ed.; The Williams & Wilkins Co.: Balti- 2018, 68, 3902–3907. more, 1957; p 394.

 

[31] Dahal, R. H.; Chaudhary, D. K.; Kim, J. Int J [35] Dedysh, S. N.; Haupt, E. S.; Dunfield, P. F. Syst Evol Microbiol 2018, 68, 1955–1962. Int J Syst Evol Microbiol 2016, 66, 3177–3182.   [32] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s [36] Hwang, C. Y.; Cho, B. C. Int J Syst Evol Mi- Manual of Systematic Bacteriology, 2nd ed.; crobiol 2008, 58, 267–277. Springer: New York, 2005; Vol. 2; p 422.  

[33] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s [37] Noh, H.-J.; Baek, K.; Hwang, C. Y.; Manual of Systematic Bacteriology, 2nd ed.; Shin, S. C.; Hong, S. G.; Lee, Y. M. Int J Syst Springer: New York, 2005; Vol. 2; p 438. Evol Microbiol 2019, 69, 3837–3842.  

[34] Breed, R. S.; Murray, E. G. D.; Smith, N. R. [38] Choi, G.; Gao, M.; Lee, D.-S.; Choi, H. Int J

158 Syst Evol Microbiol 2017, 67, 3375–3380. New York, 2005; Vol. 2; p 393.

 

[39] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s [43] Conn, H. J. J Bacteriol 1938, 36, 320–321. Manual of Systematic Bacteriology, 2nd ed.;  Springer: New York, 2005; Vol. 2; p 567. [44] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s  Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, 2005; Vol. 2; p 571. [40] Bowman, J. P. Bergey’s Manual of Systematic  Bacteriology, 2nd ed.; Springer: New York, 2005; Vol. 2; pp 411–413. [45] Kulichevskaya, I. S.; Danilova, O. V.;

 Tereshina, V. M.; Kevbrin, V. V.; Dedysh, S. N. Int J Syst Evol Microbiol [41] Huang, Z.; Guo, F.; Lai, Q.; Shao, Z. Int J 2014, 64, 2558–2565. Syst Evol Microbiol 2017, 67, 2527–2531.   [46] Cole, J. K.; Morton, B. R.; Cardamone, H. C.; [42] Mergaert, J.; Swings, J. Bergey’s Manual of Lake, H. R. R.; Dohnalkova, A. C.; Kim, Y.- Systematic Bacteriology, 2nd ed.; Springer: M.; Kyle, J. E.; Maezato, Y.; Dana, K. L.;

159 Metz, T. O.; Romine, M. F.; Nelson, W. C.; 30, 7–27.

Lindemann, S. R. Int J Syst Evol Microbiol  2018, 68, 1591–1598.

 [51] Proença, D. N.; Whitman, W. B.; Vargh- ese, N.; Shapiro, N.; Woyke, T.; Kyrpi- [47] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s des, N. C.; Morais, P. V. Syst Appl Microbiol Manual of Systematic Bacteriology, 2nd ed.; 2018, 41, 94–100. Springer: New York, 2005; Vol. 2; p 161.  

[48] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s [52] Venkata Ramana, V.; Manual of Systematic Bacteriology, 2nd ed.; Kalyana Chakravarthy, S.; Ramaprasad, E. Springer: New York, 2005; Vol. 2; p 161. V. V.; Thiel, V.; Imhoff, J. F.; Sasikala, C.;

 Ramana, C. V. Syst Appl Microbiol 2013, 36, 28–32. [49] Pfennig, N.; Trüper, H. G. Int J Syst Bacteriol  1971, 21, 17–18.

 [53] Philip, C. B. Bergey’s Manual of Determina- [50] Gillis, M.; De Ley, J. Int J Syst Bacteriol 1980, tive Bacteriology, 7th ed.; The Williams &

160 Wilkins Co.: Baltimore, 1957; pp 980–984. [58] Kosako, Y.; Yabuuchi, E.; Naka, T.; Fuji-

 wara, N.; Kobayashi, K. Microbiol Immunol 2000, 44, 563–575. [54] Görtz, H. D.; Schmidt, J. A. Bergey’s Manual  of Systematic Bacteriology, 2nd ed.; Springer: New York, 2005; Vol. 2; pp 146–149. [59] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s  Manual of Systematic Bacteriology, 2nd ed.; [55] Pinkerton, H. Parasitology 1936, 28, 172–189. Springer: New York, 2005; Vol. 2; p 575.  

[56] Kurahashi, M.; Fukunaga, Y.; Harayama, S.; [60] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s Yokota, A. Int J Syst Evol Microbiol 2008, 58, Manual of Systematic Bacteriology, 2nd ed.; 548–552. Springer: New York, 2005; Vol. 2; p 575.

 

[57] Yabuuchi, E.; Kosako, Y. Bergey’s Manual of [61] De Ley, J.; Segers, P.; Kersters, K.; Systematic Bacteriology, 2nd ed.; Springer: Mannheim, W.; Lievens, A. Int J Syst Bacte- New York, 2005; Vol. 2; pp 230–233. riol 1986, 36, 405–414.

 

161 [62] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s Vol. 2; p 774.

Manual of Systematic Bacteriology, 2nd ed.;  Springer: New York, 2005; Vol. 2; p 575. [67] Adeolu, M.; Gupta, R. S. Antonie van  Leeuwenhoek 2013, 104, 1–24. [63] Willems, A.; De Ley, J.; Gillis, M.; Ker-  sters, K. Int J Syst Bacteriol 1991, 41, 445–450.

 [68] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; [64] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s Springer: New York, 2005; Vol. 2; p 863. Manual of Systematic Bacteriology, 2nd ed.;  Springer: New York, 2005; Vol. 2; p 623.  [69] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s [65] Morotomi, M.; Nagai, F.; Watanabe, Y. Int J Manual of Systematic Bacteriology, 2nd ed.; Syst Evol Microbiol 2011, 61, 637–643. Springer: New York, 2005; Vol. 2; p 770.  

[66] Tønjum, T. Bergey’s Manual of Systematic Bac- [70] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s teriology, 2nd ed.; Springer: New York, 2005; Manual of Systematic Bacteriology, 2nd ed.;

162 Springer: New York, 2005; Vol. 2; p 864. Springer: New York, 2005; Vol. 2; p 922.

 

[71] Boden, R.; Hutt, L. P.; Rae, A. W. Int J Syst [75] Wang, Z.-J.; Liu, Q.-Q.; Zhao, L.-H.; Du, Z.- Evol Microbiol 2017, 67, 1191–1205. J.; Chen, G.-J. Int J Syst Evol Microbiol 2015, 65, 1542–1549.   [72] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s [76] Kuever, J.; Rainey, F. A.; Widdel, F. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, 2005; Vol. 2; p 887. Springer: New York, 2005; Vol. 2; p 1003.  

[73] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s [77] Kuever, J.; Rainey, F. A.; Widdel, F. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, 2005; Vol. 2; p 887. Springer: New York, 2005; Vol. 2; p 1003.  

[74] Kuever, J.; Rainey, F. A.; Widdel, F. Bergey’s [78] Kuever, J.; Rainey, F. A.; Widdel, F. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; Manual of Systematic Bacteriology, 2nd ed.;

163 Springer: New York, 2005; Vol. 2; p 959. Manual of Systematic Bacteriology, 2nd ed.;

 Springer: New York, 2005; Vol. 2; pp 925– 926. [79] Kuever, J.; Rainey, F. A.; Widdel, F. Bergey’s  Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, 2005; Vol. 2; pp 959– [83] Kuever, J.; Rainey, F. A.; Widdel, F. Bergey’s 960. Manual of Systematic Bacteriology, 2nd ed.;  Springer: New York, 2005; Vol. 2; pp 948– [80] Kuever, J.; Rainey, F. A.; Widdel, F. Bergey’s 949.

Manual of Systematic Bacteriology, 2nd ed.;  Springer: New York, 2005; Vol. 2; p 988.

 [84] Kuever, J.; Rainey, F. A.; Widdel, F. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; [81] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s Springer: New York, 2005; Vol. 2; p 944. Manual of Systematic Bacteriology, 2nd ed.;  Springer: New York, 2005; Vol. 2; p 999.

 [85] Kuever, J.; Rainey, F. A.; Widdel, F. Bergey’s [82] Kuever, J.; Rainey, F. A.; Widdel, F. Bergey’s Manual of Systematic Bacteriology, 2nd ed.;

164 Springer: New York, 2005; Vol. 2; p 956. Springer: New York, 2005; Vol. 2; pp 1005–

 1006.

 [86] Kuever, J.; Rainey, F. A.; Widdel, F. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; [90] Kuever, J.; Rainey, F. A.; Widdel, F. Bergey’s Springer: New York, 2005; Vol. 2; p 926. Manual of Systematic Bacteriology, 2nd ed.;  Springer: New York, 2005; Vol. 2; p 1006. [87] Kuever, J.; Rainey, F. A.; Widdel, F. Bergey’s  Manual of Systematic Bacteriology, 2nd ed.; [91] Holmes, D. E.; Nevin, K. P.; Lovley, D. R. Int Springer: New York, 2005; Vol. 2; p 922. J Syst Evol Microbiol 2004, 54, 1591–1599.   [88] Kuever, J.; Rainey, F. A.; Widdel, F. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; [92] Yamamoto, E.; Muramatsu, H.; Nagai, K. Springer: New York, 2005; Vol. 2; p 923. Int J Syst Evol Microbiol 2014, 64, 3360–3368.  

[89] Kuever, J.; Rainey, F. A.; Widdel, F. Bergey’s [93] Reichenbach, H. Bergey’s Manual of System- Manual of Systematic Bacteriology, 2nd ed.; atic Bacteriology, 2nd ed.; Springer: New

165 York, 2005; Vol. 2; p 1143. Appl Environ Microbiol 2008, 74, 2051–2058.

 

[94] Reichenbach, H. Bergey’s Manual of System- [98] Kuever, J.; Rainey, F. A.; Widdel, F. Bergey’s atic Bacteriology, 2nd ed.; Springer: New Manual of Systematic Bacteriology, 2nd ed.; York, 2005; Vol. 2; pp 1136–1137. Springer: New York, 2005; Vol. 2; p 1021.

 

[95] Garcia, R. O.; Reichenbach, H.; Ring, M. W.; [99] Kuever, J.; Rainey, F. A.; Widdel, F. Bergey’s Müller, R. Int J Syst Evol Microbiol 2009, 59, Manual of Systematic Bacteriology, 2nd ed.; 1524–1530. Springer: New York, 2005; Vol. 2; p 1033.

 

[96] Mohr, K. I.; Garcia, R. O.; Gerth, K.; [100] Kuever, J.; Rainey, F. A.; Widdel, F. Bergey’s Irschik, H.; Müller, R. Int J Syst Evol Mi- Manual of Systematic Bacteriology, 2nd ed.; crobiol 2012, 62, 1191–1198. Springer: New York, 2005; Vol. 2; p 1021.

 

[97] Qiu, Y.-L.; Hanada, S.; Ohashi, A.; [101] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s Harada, H.; Kamagata, Y.; Sekiguchi, Y. Manual of Systematic Bacteriology, 2nd ed.;

166 Springer: New York, 2005; Vol. 2; p 1145. Stackebrandt, E. Int J Syst Evol Microbiol

 2004, 54, 41–45.

 [102] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; [106] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s Springer: New York, 2005; Vol. 2; p 1145. Manual of Systematic Bacteriology, 2nd ed.;  Springer: New York, 2005; Vol. 2; p 1. [103] Vandamme, P.; De Ley, J. Int J Syst Bacteriol  1991, 41, 451–455. [107] Kojima, H.; Shinohara, A.; Fukui, M. Int J  Syst Evol Microbiol 2015, 65, 3709–3713.

[104] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s  Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, 2005; Vol. 2; p 1168. [108] Martin-Carnahan, A.; Joseph, S. W. Bergey’s Manual of Systematic Bacteriology, 2nd ed.;  Springer: New York, 2005; Vol. 2; p 556. [105] Miroshnichenko, M. L.; L’Haridon, S.;  Schumann, P.; Spring, S.; Bonch- Osmolovskaya, E. A.; Jeanthon, C.; [109] Colwell, R. R.; Macdonell, M. T.; De Ley, J.

167 Int J Syst Bacteriol 1986, 36, 473–477. Syst Evol Microbiol 2011, 61, 1053–1060.

 

[110] Hippe, H.; Hagelstein, A.; Kramer, I.; [114] Ivanova, E. P.; Flavier, S.; Christen, R. Int J Swiderski, J.; Stackebrandt, E. Int J Syst Bac- Syst Evol Microbiol 2004, 54, 1773–1788. teriol 1999, 49, 779–782.   [115] Teramoto, M.; Yagyu, K.-i.; Nishijima, M. Int J Syst Evol Microbiol 2015, 65, 353–358. [111] Bowman, J. P.; McMeekin, T. A. Bergey’s  Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, 2005; Vol. 2; p 443. [116] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s

 Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, 2005; Vol. 2; p 123. [112] Ivanova, E. P.; Mikhailov, V. V. Microbiology  2001, 70, 10–17. [117] Dewhirst, F. E.; Paster, B. J.; La Fontaine, S.;  Rood, J. I. Int J Syst Bacteriol 1990, 40, 426– [113] Cramer, M. J.; Haghshenas, N.; Bag- 433.

well, C. E.; Matsui, G. Y.; Lovell, C. R. Int J 

168 [118] Spring, S.; Scheuner, C.; Göker, M.; New York, 2005; Vol. 2; p 58.

Klenk, H.-P. Front Microbiol 2015, 6, 281. 

 [123] Boden, R. Int J Syst Evol Microbiol 2017, 67, [119] Imhoff, J. F. Bergey’s Manual of Systematic 3919–3928. Bacteriology, 2nd ed.; Springer: New York,  2005; Vol. 2; pp 1–3.

 [124] Mori, K.; Suzuki, K.-i.; Urabe, T.; Sug- ihara, M.; Tanaka, K.; Hamada, M.; [120] Imhoff, J. F. Int J Syst Bacteriol 1984, 34, 338– Hanada, S. Int J Syst Evol Microbiol 2011, 339. 61, 2412–2418.   [121] Lee, K.; Lee, H. K.; Choi, T.-H.; Kim, K.- M.; Cho, J.-C. J Microbiol Biotechnol 2007, 17, [125] Kojima, H.; Watanabe, M.; Fukui, M. Int J 1483–1490. Syst Evol Microbiol 2017, 67, 3458–3461.  

[122] Kelly, D. P.; Wood, A. P. Bergey’s Manual of [126] Wang, G.; Tang, M.; Li, T.; Dai, S.; Wu, H.; Systematic Bacteriology, 2nd ed.; Springer: Chen, C.; He, H.; Fan, J.; Xiang, W.; Li, X.

169 Antonie van Leeuwenhoek 2015, 107, 1625– Springer: New York, 2005; Vol. 2; p 210.

1632. 

 [131] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s [127] Du, Z.-J.; Wang, Z.-J.; Zhao, J.-X.; Chen, G.-J. Manual of Systematic Bacteriology, 2nd ed.; Int J Syst Evol Microbiol 2016, 66, 107–112. Springer: New York, 2005; Vol. 2; p 237.

 

[128] Adeolu, M.; Alnajar, S.; Naushad, S.; [132] Brenner, D. J.; Steigerwalt, A. G.; Mc- S. Gupta, R. Int J Syst Evol Microbiol 2016, Dade, J. E. Ann Intern Med 1979, 90, 656.

66, 5575–5599. 

 [133] Christensen, P.; Cook, F. D. Int J Syst Bacte- [129] Corteselli, E. M.; Aitken, M. D.; Single- riol 1978, 28, 367–393.

ton, D. R. Int J Syst Evol Microbiol 2017, 67,  925–931. [134] Naushad, S.; Adeolu, M.; Wong, S.; So-  hail, M.; Schellhorn, H. E.; Gupta, R. S. An- [130] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s tonie van Leeuwenhoek 2015, 107, 467–485.

Manual of Systematic Bacteriology, 2nd ed.; 

170 [135] Bowman, J. P. Bergey’s Manual of Systematic Int J Syst Evol Microbiol 2012, 62, 2743–2749.

Bacteriology, 2nd ed.; Springer: New York,  2005; Vol. 2; pp 248–252. [139] Liu, Q.; Liu, H.-C.; Zhou, Y.-G.; Xin, Y.-H.  Int J Syst Evol Microbiol 2019, 69, 1404–1410. [136] Whittenbury, R.; Krieg, N. R. Bergey’s Man-  ual of Systematic Bacteriology, 1st ed.; The [140] Kämpfer, P.; Glaeser, S. P.; Nilsson, L. Williams & Wilkins Co.: Baltimore, 1984; K. J.; Eberhard, T.; Håkansson, S.; Guy, L.; Vol. 1; pp 256–261. Roos, S.; Busse, H.-J.; Terenius, O. Int J Syst  Evol Microbiol 2015, 65, 444–451. [137] Hirayama, H.; Abe, M.; Miyazaki, M.;  Nunoura, T.; Furushima, Y.; Yamamoto, H.; [141] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s Takai, K. Int J Syst Evol Microbiol 2014, 64, Manual of Systematic Bacteriology, 2nd ed.; 989–999. Springer: New York, 2005; Vol. 2; p 270.

 

[138] Gutierrez, T.; Green, D. H.; Whitman, W. B.; [142] Golyshin, P. N.; Harayama, S.; Tim- Nichols, P. D.; Semple, K. T.; Aitken, M. D. mis, K. N.; Yakimov, M. M. Bergey’s Manual

171 of Systematic Bacteriology, 2nd ed.; Springer: Springer: New York, 2005; Vol. 2; p 299.

New York, 2005; Vol. 2; p 295. 

 [146] Franzmann, P. D.; Wehmeyer, U.; Stacke- [143] Krishnan, R.; Lang, E.; Midha, S.; Patil, P. B.; brandt, E. Syst Appl Microbiol 1988, 11, 16– Rameshkumar, N. Syst Appl Microbiol 2018, 19. 41, 570–580. 

 [147] Wang, G.; Tang, M.; Wu, H.; Dai, S.; Li, T.; Chen, C.; He, H.; Fan, J.; Xiang, W.; Li, X. [144] Bartz, J.-O.; Blom, J.; Busse, H.-J.; Mvie, J. B.; Int J Syst Evol Microbiol 2015, 65, 4488–4494. Hardt, M.; Schubert, P.; Wilke, T.; Goess-  mann, A.; Wilharm, G.; Bender, J.; Kämpfer, P.; Glaeser, S. P. Syst Appl Micro- [148] Kim, H.; Choo, Y.-J.; Cho, J.-C. Int J Syst Evol biol 2018, 41, 73–84. Microbiol 2007, 57, 1793–1798.

 

[145] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s [149] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; Manual of Systematic Bacteriology, 2nd ed.;

172 Springer: New York, 2005; Vol. 2; p 271. 1979.

 

[150] Golyshin, P. N.; Chernikova, T. N.; Abra- [154] Xin, D.; Bisgaard, M.; Busse, H.-J.; ham, W.-R.; Lünsdorf, H.; Timmis, K. N.; Olsen, R. H.; Hess, C.; Aalbæk, B.; Yakimov, M. M. Int J Syst Evol Microbiol Olsen, J. E.; Christensen, H. Int J Syst Evol 2002, 52, 901–911. Microbiol 2019, 69, 350–355.

 

[151] Kwong, W. K.; Moran, N. A. 2013, 63, 2008– [155] Orla-Jensen, S. J Bacteriol 1921, 6, 263–273.

2018. 

 [156] Rossau, R.; Van Landschoot, A.; Gillis, M.; [152] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s De Ley, J. Int J Syst Bacteriol 1991, 41, 310– Manual of Systematic Bacteriology, 2nd ed.; 319.

Springer: New York, 2005; Vol. 2; p 850. 

 [157] Winslow, C.-E. A.; Broadhurst, J.; [153] Pohl, S. Inaug. Diss., Phillips-Universität Buchanan, R. E.; Krumwiede, C.; Marburg/Lahn. Mauersberger, Marburg, Rogers, L. A.; Smith, G. H. J Bacteriol

173 1917, 2, 505–566. 2005; Vol. 2; pp 199–200.

 

[158] Lin, J. Y.; Hobson, W. J.; Wertz, J. T. Int J Syst [162] Fryer, J. L.; Lannan, C. N. Bergey’s Manual Evol Microbiol 2016, 66, 2869–2875. of Systematic Bacteriology, 2nd ed.; Springer:  New York, 2005; Vol. 2; p 180. [159] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s  Manual of Systematic Bacteriology, 2nd ed.; [163] Boden, R.; Scott, K. M. Int J Syst Evol Micro- Springer: New York, 2005; Vol. 2; p 131. biol 2018, 68, 2226–2239.   [160] Xiao, M.; Salam, N.; Liu, L.; Jiao, J.-Y.; Zheng, M.-L.; Wang, J.; Li, S.; Chen, C.; [164] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s Li, W.-J.; Qu, P.-H. Int J Syst Evol Microbiol Manual of Systematic Bacteriology, 2nd ed.; 2018, 68, 347–352. Springer: New York, 2005; Vol. 2; p 131.  

[161] Sjöstedt, A. B. Bergey’s Manual of Systematic [165] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s Bacteriology, 2nd ed.; Springer: New York, Manual of Systematic Bacteriology, 2nd ed.;

174 Springer: New York, 2005; Vol. 2; p 763. [169] Davidov, Y.; Jurkevitch, E. Int J Syst Evol

 Microbiol 2004, 54, 1439–1452.

 [166] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; [170] Koval, S. F.; Williams, H. N.; Stine, O. C. Int Springer: New York, 2005; Vol. 2; p 763. J Syst Evol Microbiol 2015, 65, 593–597.

 

[167] Nakai, R.; Nishijima, M.; Tazato, N.; [171] Piñeiro, S. A.; Williams, H. N.; Stine, O. C. Handa, Y.; Karray, F.; Sayadi, S.; Isoda, H.; Int J Syst Evol Microbiol 2008, 58, 1203–1209.

Naganuma, T. Int J Syst Evol Microbiol 2014,  64, 3353–3359.

 [172] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; [168] Hahn, M. W.; Schmidt, J.; Koll, U.; Ro- Springer: New York, 2005; Vol. 2; p 1040. hde, M.; Verbarg, S.; Pitt, A.; Nakai, R.; Na-  ganuma, T.; Lang, E. Int J Syst Evol Microbiol 2017, 67, 2555–2568. [173] Garrity, G. M.; Bell, J. A.; Lilburn, T. Bergey’s

 Manual of Systematic Bacteriology, 2nd ed.;

175 Springer: New York, 2005; Vol. 2; pp 1040– 1041.



[174] McCauley, E. P.; Haltli, B.; Kerr, R. G. Int J Syst Evol Microbiol 2015, 65, 522–530.



176 14 R

Phylum: Rhodothermaeota1 Family: Salisaetaceae4 Class: Rhodothermia1 Roseithermus Order: Rhodothermales2 Salisaeta Family: Rhodothermaceae3 Longibacter References Longimonas [1] Munoz, R.; Rosselló-Móra, R.; Amann, R. Syst Natronotalea Appl Microbiol 2016, 39, 281–296. Rhodothermus Rubricoccus  Rubrivirga [2] Munoz, R.; Rosselló-Móra, R.; Amann, R. Syst Family: Rubricoccaceae1 Appl Microbiol 2016, 39, 491–492. 1 Family: Salinibacteraceae  Salinibacter [3] Ludwig, W.; Euzéby, J.; Whitman, W. B. Salinivenus Bergey’s Manual of Systematic Bacteriology, 2nd

177 ed.; Springer: New York, 2011; Vol. 4; p 457.



[4] Park, M.-J.; Oh, J. H.; Yang, S.-H.; Kwon, K. K. Int J Syst Evol Microbiol 2019, 69, 1213–1219.



178 15 S

Phylum: Spirochaetes1 Order: Spirochaetales7,9 Class: Spirochaetes2 Family: Borreliaceae3,4 Order: Brachyspirales3,4 Borrelia Family: Brachyspiraceae5 Borreliella Brachyspira Cristispira Serpulina Family: None Order: Brevinematales3,4 Exilispira Family: Brevinemataceae6 Family: Spirochaetaceae7 Brevinema Alkalispirochaeta Order: Leptospirales3,4 Clevelandina Family: Leptospiraceae7,8 Diplocalyx Leptonema Hollandina Leptospira Marispirochaeta Turneriella Oceanispirochaeta

179 Pillotina Aminiphilus Pleomorphochaeta Aminivibrio Rectinema Aminobacterium Salinispira Aminomonas Sediminispirochaeta Anaerobaculum Sphaerochaeta Cloacibacillus Spirochaeta Dethiosulfovibrio Treponema Fretibacterium Phylum: Synergistetes10 Jonquetella Class: None Lactivibrio Order: None Pyramidobacter Family: None Synergistes Rarimicrobium Thermanaerovibrio Class: Synergistia10 Thermovirga Order: Synergistales10 Family: Synergistaceae10 References Acetomicrobium [1] Garrity, G. M.; Holt, J. G. Bergey’s Manual

180 of Systematic Bacteriology, 2nd ed.; Springer: [6] Paster, B. J. Bergey’s Manual of Systematic Bac- New York, 2001; Vol. 1; pp 119–166. teriology, 2nd ed.; Springer: New York, 2011;

 Vol. 4; p 545.

 [2] Cavalier-Smith, T. Int J Syst Evol Microbiol 2002, 52, 7–76. [7] Skerman, V. B. D.; McGowan, V.; Sneath, P.

 H. A. Int J Syst Bacteriol 1980, 30, 225–420.

 [3] Gupta, R. S.; Mahmood, S.; Adeolu, M. Front Microbiol 2013, 4, 217. [8] Hovind-Hougen, K. Int J Syst Bacteriol 1979,

 29, 245–251.

 [4] Gupta, R. S.; Mahmood, S.; Adeolu, M. Front Microbiol 2013, 4, 322. [9] Buchanan, R. E. J Bacteriol 1917, 2, 155–164.

 

[5] Paster, B. J. Bergey’s Manual of Systematic Bac- [10] Jumas-Bilak, E.; Roudière, L.; teriology, 2nd ed.; Springer: New York, 2010; Marchandin, H. Int J Syst Evol Micro- Vol. 4; p 531. biol 2009, 59, 1028–1035.

 

181 16 T

Phylum: Tenericutes1 Spiroplasma Class: Mollicutes2,3 Order: Haloplasmatales9 Order: Acholeplasmatales4 Family: Haloplasmataceae9 Family: Acholeplasmataceae2,5 Haloplasma Acholeplasma Order: Mycoplasmatales2,10 Order: Anaeroplasmatales6 Family: Mycoplasmataceae2,10 Family: Anaeroplasmataceae6 Eperythrozoon Anaeroplasma Haemobartonella Asteroleplasma Mycoplasma Order: Entomoplasmatales7 Ureaplasma Family: Entomoplasmataceae7 Williamsoniiplasma Entomoplasma Order: Mycoplasmoidales11 Mesoplasma Family: Metamycoplasmataceae11 Family: Spiroplasmataceae8 Mesomycoplasma

182 Metamycoplasma Phylum: Thermotogae16 Mycoplasmopsis Class: Thermotogae17 Family: Mycoplasmoidaceae11 Order: Kosmotogales18 Malacoplasma Family: Kosmotogaceae18 Mycoplasmoides Kosmotoga Phylum: Thermodesulfobacteria12 Mesotoga Class: None Thermococcoides Order: None Order: Mesoaciditogales19 Family: None Family: Mesoaciditogaceae19 Thermosulfuriphilus Athalassotoga Class: Thermodesulfobacteria13 Mesoaciditoga Order: Thermodesulfobacteriales14 Order: Petrotogales18 Family: Thermodesulfobacteriaceae15 Family: Petrotogaceae18 Caldimicrobium Defluviitoga Thermodesulfatator Geotoga Thermodesulfobacterium Marinitoga Thermosulfurimonas Oceanotoga

183 Petrotoga [3] Edward, D. G. F.; Freundt, E. A. Int J Syst Order: Thermotogales20 Bacteriol 1967, 17, 267–268. 18 Family: Fervidobacteriaceae  Fervidobacterium Thermosipho [4] Freundt, E. A.; Whitcomb, R. F.; Barile, M. F.; Family: Thermotogaceae21 Razin, S.; Tully, J. G. Int J Syst Bacteriol 1984, Pseudothermotoga 34, 346–349. Thermotoga 

[5] Edward, D. G. F.; Freundt, E. A. J Gen Micro- References biol 1970, 62, 1–2.

[1] Murray, R. G. E. Bergey’s Manual of Systematic  Bacteriology, 1st ed.; The Williams & Wilkins Co.: Baltimore, 1989; Vol. 3; pp 31–34. [6] Robinson, I. M.; Freundt, E. A. Int J Syst Bac- teriol 1987, 37, 78–81.   [2] Skerman, V. B. D.; McGowan, V.; Sneath, P. H. A. Int J Syst Bacteriol 1980, 30, 225–420. [7] Tully, J. G.; Bov’e, J. M.; Laigret, F.; Whit-

 comb, R. F. Int J Syst Bacteriol 1993, 43, 378–

184 385. 111, 1583–1630.

 

[8] Skripal’, I. G. Int J Syst Bacteriol 1983, 33, 408– [12] Garrity, G. M.; Holt, J. G. Bergey’s Manual 408. of Systematic Bacteriology, 2nd ed.; Springer: New York, 2001; Vol. 1; pp 389–393.   [9] Antunes, A.; Rainey, F. A.; Wanner, G.; [13] Hatchikian, E. C.; Ollivier, B.; Garcia, J. L. Taborda, M.; Pätzold, J.; Nobre, M. F.; Bergey’s Manual of Systematic Bacteriology, 2nd da Costa, M. S.; Huber, R. J Bacteriol 2008, ed.; Springer: New York, 2001; Vol. 1; p 389. 190, 3580–3587.   [14] Hatchikian, E. C.; Ollivier, B.; Garcia, J. L. [10] Freundt, E. A. Int Bull Bact Nom Tax 1955, 5, Bergey’s Manual of Systematic Bacteriology, 2nd 67–78. ed.; Springer: New York, 2001; Vol. 1; pp 389–  390.

 [11] Gupta, R. S.; Sawnani, S.; Adeolu, M.; Alna- jar, S.; Oren, A. Antonie van Leeuwenhoek 2018, [15] Hatchikian, E. C.; Ollivier, B.; Garcia, J. L.

185 Bergey’s Manual of Systematic Bacteriology, 2nd Ohkuma, M. Int J Syst Evol Microbiol 2016, ed.; Springer: New York, 2001; Vol. 1; p 390. 66, 1045–1051.

 

[16] Reysenbach, A. L. Bergey’s Manual of Sys- [20] Reysenbach, A. L. Bergey’s Manual of Sys- tematic Bacteriology, 2nd ed.; Springer: New tematic Bacteriology, 2nd ed.; Springer: New York, 2001; Vol. 1; pp 369–387. York, 2001; Vol. 1; pp 369–370.

 

[17] Reysenbach, A. L. Bergey’s Manual of Sys- [21] Reysenbach, A. L. Bergey’s Manual of Sys- tematic Bacteriology, 2nd ed.; Springer: New tematic Bacteriology, 2nd ed.; Springer: New York, 2001; Vol. 1; p 369. York, 2001; Vol. 1; p 370.

  [18] Bhandari, V.; Gupta, R. S. Antonie van Leeuwenhoek 2014, 105, 143–168.



[19] Itoh, T.; Onishi, M.; Kato, S.; Iino, T.; Sakamoto, M.; Kudo, T.; Takashina, T.;

186 17 V

Phylum: Verrucomicrobia1 Ereboglobus Class: Spartobacteria2 Geminisphaera Order: None Nibricoccus Family: None Oleiharenicola Terrimicrobium Opitutus Class: None Order: Puniceicoccales3 Order: None Family: Puniceicoccaceae3 Family: None Cerasicoccus Limisphaera Coraliomargarita Class: Opitutae3 Pelagicoccus Order: Opitutales3 Puniceicoccus Family: Opitutaceae3 Ruficoccus Alterococcus Class: Verrucomicrobiae4 Cephaloticoccus Order: Verrucomicrobiales5

187 Family: Akkermansiaceae6 Bacteriology, 2nd ed.; Springer: New York, Akkermansia 2011; Vol. 4; p 795. 7 Family: Rubritaleaceae  Rubritalea Family: Verrucomicrobiaceae5 [2] Sangwan, P.; Chen, X.; Hugenholtz, P.; Brevifollis Janssen, P. H. Appl Environ Microbiol 2004, 70, Haloferula 5875–5881. Luteolibacter  Persicirhabdus [3] Choo, Y.-J.; Lee, K.; Song, J.; Cho, J.-C. Int J Phragmitibacter Syst Evol Microbiol 2007, 57, 532–537. Prosthecobacter  Roseibacillus Roseimicrobium [4] Hedlund, B. P.; Gosink, J. J.; Staley, J. T. Antonie Verrucomicrobium Van Leeuwenhoek 1997, 72, 29–38.

 References [5] Ward-Rainey, N.; Rainey, F. A.; Schlesner, H.; [1] Hedlund, B. P. Bergey’s Manual of Systematic Stackebrandt, E. Microbiology 1995, 141, 3247–

188 3250.



[6] Hedlund, B. P.; Derrien, M. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, 2011; Vol. 4; p 809.



[7] Hedlund, B. P. Bergey’s Manual of Systematic Bacteriology, 2nd ed.; Springer: New York, 2011; Vol. 4; p 812.



189

BACTERIA

TAXONOMY Phyla, Classes, Orders, Families, and Genera