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Nestell Guadalupian.Vp CONTENTS Volume 52 Number 1 2006 FAUNAL ANALYSIS 1 Galina P. Nestell and Merlynd K. Nestell Middle Permian (Late Guadalupian) foraminifers from Dark Canyon, Guadalupe Moun- tains, New Mexico SYSTEMATICS 51 Neil E. Tibert and Christopher P. Dewey Velatomorpha, a new healdioidean ostracode genus from the early Pennsylvanian Joggins Formation, Nova Scotia, Canada PALEOECOLOGY 67 Marci M. Robinson and Randolph A. McBride Benthic foraminifera from a relict flood tidal delta along the Virginia/North Carolina Outer Banks SHORT NOTES 81 Patrizia Maiorano Reticulofenestra circus var. lata n.var.: a large reticulofenestrid (Coccolithophoridae) from the Early Oligocene 87 Hossein Hashemi and Mahtab Fahimi Dictyotidium senticogremium sp. nov., a new prasinophyte (Chlorophyta) phycoma from the Upper Devonian of northern Iran ANNOUNCEMENT The Editors Internet access for subscribers Middle Permian (Late Guadalupian) foraminifers from Dark Canyon, Guadalupe Mountains, New Mexico Galina P. Nestell and Merlynd K. Nestell Department of Earth and Environmental Sciences, University of Texas at Arlington, Arlington, TX 76019, USA email: [email protected]; [email protected] ABSTRACT: Late Capitanian foraminifers are described from two cores taken by Amoco on the north side of the entrance of Dark Canyon. The Amoco #1 core (400ft) penetrates proximal forereef debris of the Tansill Formation and bottoms in massive Capitan reefal limestone. The Amoco #2 core (468ft) penetrates proximal backreef lagoonal deposits of the upper Yates Formation and the Tansill Formation. Foraminifers are very diverse and abundant in these cores. Based on the fusulinid distribution, the Codonofusiella extensa (upper part of the Yates Formation), Yabeina texana, Paradoxiella pratti and Reichelina lamarensis zones (lower and middle Tansill Forma- tion) are present in the Amoco #2 core. Among small foraminifers, the first appearance of the genera Sengoerina and Crescentia are re- corded at the base of the Y. texana Zone and they disappear at the top of the P. pratti Zone. The genus Baisalina appears at the base of the R. lamarensis Zone and disappears at the top of that zone. The Ocotillo Silt Member and upper dolomitized part of the Tansill For- mation do not contain foraminifers. The Ocotillo Silt Member is present in the Amoco #2 core at a depth of from 88 to 112ft and, be- cause of its regional dip of approximately six degrees, would possibly project below the base of the Amoco #1 core, thus correlating the Capitan and Tansill formations in the Amoco #1 core with the upper Tansill Formation of the Amoco #2 core. The lower part of the Amoco #1 core (from 110 to 370ft) contains fusulinacean species of the Paraboultonia splendens Zone. The assemblage of small foraminifers is dominated by species of nodosariids and hemigordiopsids. Four new genera and sixteen new species of foraminifers are described from both cores. INTRODUCTION AND STRATIGRAPHIC SETTING ent in the lower part of the Guadalupian, but not in the upper The Guadalupe Mountains in southeastern New Mexico and part. In the Guadalupe Mountains, the last appearance of the West Texas is a northeasterly tilted tectonic block of Mid- very large fusulinacean Polydiexodina appears to be in the mid- dle/Upper Permian strata exhumed in the Tertiary (Hayes 1964) dle part of the reefal Yates Formation (top of Yates B, middle and forms the classic shelf-to-basin exposures of the Middle Capitanian, Bebout and Kerans 1993) (text-fig. 1). The Permian Capitan reef depositional system. The International fusulinacean assemblage in the overlying upper part of the Commission on Stratigraphy (ICS) recently published an Inter- Yates Formation, Tansill (equivalent to basinal Lamar and national Standard for the Middle Permian Guadalupian Series “post-Lamar”) Formation is dominated by small fusulinaceans consisting of the Roadian, Wordian and Capitanian stages, the such as Codonofusiella, Yabeina, Reichelina, and Paraboult- stratotype sections of which were designated in the Guadalupe onia. Wilde (1990) proposed a Zone of Paraboultonia for the Mountains (Gradstein 2000, Gradstein et al. 2004, Wardlaw et last fusulinacean interval at the top of the Guadalupian. Re- al. 2004). The lower boundaries of these stages have been cently, Wilde et al. (1999) formally introduced the name Reef drawn based primarily on the first appearance of conodonts: for Trail Member for the “post-Lamar” strata at the top of the Bell the Roadian - Jinogondolella nankingensis;theWordian-J. Canyon Formation and illustrated fusulinaceans from four suc- aserrata; and the Capitanian - J. postserrata (Glenister et al. cessive fusulinacean zones: Yabeina, Paradoxiella, Reichelina, 1999, Wardlaw et al. 2004) (text-fig. 1). The boundary of the and Paraboultonia. Unfortunately, in this paper (Wilde et al. Guadalupian with the overlying Lopingian Series is based on 1999, fig. 3), the Guadalupian/Lopingian boundary was drawn the evolutionary transition of the conodont Clarkina postbitteri with question in the lower part of the Reef Trail Member with hongshuiensis into C. postbitteri postbitteri (Wardlaw et al. the implication that the Paraboultonia Zone was entirely of 2004). In a recent paper, Lambert et al. (2002) illustrated speci- Lopingian age. Although Wilde et al. (1999) discussed the mens of C. p. hongshuiensis from the western part of the fusulinacean succession in this interval as support for this con- Apache Mountains in West Texas that conclusively demon- clusion, it was premature for Wilde to suggest the placement of strated that the Guadalupian Series is essentially complete in this boundary in the Guadalupian stratotype because the cono- the type area. dont succession in the type section of the Reef Trail Member had not been completely documented. In a recent paper, Lam- Fusulinaceans are also present in abundance in Guadalupian bert et al. (2002) clearly documented the upper boundary of the strata of West Texas and have been used for many years in both Capitanian in an undisturbed section in the western part of the regional and international correlations of the Permian System Apache Mountains to the south of the Guadalupe Mountains. In (Dunbar and Skinner 1937, Ross 1964, Wilde 1955, 1975, this section (Lambert et al. 2002, fig. 3), the last appearance of 1990, Skinner and Wilde 1954, 1955, Wilde and Rudine 2000, Paraboultonia is in bed E1, about two meters below the first Yang and Yancey 2000). Large fusulinaceans of several genera, beds of the Castile Formation and in the same bed with the first such as Parafusulina, Skinnerina, and Polydiexodina, are pres- appearance of the conodont Clarkina postbitteri hongshuiensis micropaleontology, vol. 52, no. 1, pp. 1-50, text-figures 1-8, plates 1-11, appendices 1-2, 2006 1 Galina P. Nestell and Merlynd K. Nestell: Middle Permian foraminifers from Dark Canyon, Guadalupe Mountains, New Mexico Henderson, Mei and Wardlaw. The base of the Lopingian has Only a few genera and species of fusulinaceans have been de- since been defined at the first appearance of the conodont scribed from the upper Guadalupian (upper Yates/Tansill or the Clarkina postbitteri postbitteri (Wardlaw et al. 2004). Thus, the equivalent Lamar/Reef Trail) strata in West Texas. Wilde et al. Zone of Paraboultonia is entirely Late Guadalupian in age and (1999) presented a thorough discussion of the history of the dis- there are no known Lopingian fusulinaceans in the West Texas covery of many of these forms. Two interesting and controver- succession. No evidence was documented in this study of small sial occurrences are the Tethyan form Yabeina texana,first foraminiferal species in the Dark Canyon succession to dispute reported from the McKittrick Canyon area by Skinner and this conclusion. Wilde (1955), and Paraboultonia splendens, first described from Seven Heart Gap in the Apache Mountains (Skinner and A number of well known canyons, among them McKittrick, Wilde 1954). Yabeina texana is the only true verbeekinid found Big, Slaughter, Rattlesnake, and Walnut, cut the east facing in North America, except for specimens from exotic terranes in scarp of the Guadalupe Mountains and expose various facies of the Pacific Northwest. Since its first discovery in the basal the forereef, reef and backreef deposits of the Guadalupian. Lamar Limestone Member of the McKittrick Canyon area, Dark Canyon, the northernmost of these canyons, trends Yabeina texana has also been reported from the Tansill Forma- east-west, and is located about 10 miles (15km) southwest of tion at the top of McKittrick Canyon (Tyrrell, personal commu- Carlsbad, New Mexico in the SW ¼ of Sec. 24, T23E, R25E, nication, 2005) and near the entrance of Dark Canyon (Tyrrell Eddy County, New Mexico (text-fig. 2). 1962, 1969). The two Amoco research cores were taken on the north side of DISTRIBUTION OF FORAMINIFERS IN THE CORE Dark Canyon near the entrance (text-figs. 3, 4) in the shal- HOLES low-water backreef Capitanian facies where the upper Guadalupian (Capitanian) has been subdivided into three for- The Amoco #1 core has 220ft of Tansill, 70ft of alternating mations (in ascending order): the Seven Rivers, Yates and Tansill/Capitan lithologies, and 110ft of massive Capitan Tansill formations. In this paper, small foraminifers and (text-fig. 5). Small foraminifers in the cored section consist of fusulinaceans from the Yabeina, Paradoxiella, Reichelina and 38 species belonging to 25 genera. The following species ap- Paraboultonia zones are described from these two cores. The pear in the massive Capitan facies: Tuberitina variabilis n. sp., Amoco #1 core (no. 4492) was taken near the mouth of the can- Vachardella longiuscula n. gen., n. sp., Agathammina sp. 1, yon and penetrates 398ft (121.3m) of proximal forereef debris Nodosaria cf. N. novizkiana Sosnina, N.cf.N. partisana of the Tansill Formation and bottoms in massive Capitan reef Sosnina, Geinitzina sp. 1, G. sp. 4, Tauridia sp. 4, Ichthyolaria facies. The Amoco #2 core (no. 4518) was taken approximately sp. 1, Calvezina sp. 1, Globivalvulina sp. 1, Abadehella sp. 2, 4000ft (1220m) to the west and penetrates 468ft (142.6m) of and Deckerella sp.
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